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<document id="CB06ED26B99AB9693A76D3E28503A893" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFE6E51A531A86DB618DFB15" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) bettotanus Cameron 1930" docType="treatment" docVersion="13" lastPageNumber="157" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="156" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="816DC772129ECDD5A0BDF596C78DFF1D">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<heading id="D0F981C0FFE6E515531A86DB67E9FE5A" box="[831,1250,467,494]" centered="true" fontSize="12" level="2" pageId="39" pageNumber="156" reason="2">
<taxonomicName id="4C0E4D2FFFE6E515531A86DB67E9FE5A" ID-CoL="7GHYK" authority="CAMERON, 1930" authorityName="Cameron" authorityYear="1930" box="[831,1250,467,494]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="bettotanus" subGenus="Zyras">
<emphasis id="B97AEABEFFE6E515531A86DB607EFE5A" box="[831,885,467,494]" italics="true" pageId="39" pageNumber="156">Zyras</emphasis>
(
<emphasis id="B97AEABEFFE6E51553A686DB60B2FE5A" box="[899,953,467,494]" italics="true" pageId="39" pageNumber="156">Zyras</emphasis>
)
<emphasis id="B97AEABEFFE6E51553E286DB673DFE5A" box="[967,1078,467,494]" italics="true" pageId="39" pageNumber="156">bettotanus</emphasis>
<bibRefCitation id="EF9F4B5DFFE6E515541886DB67E9FE5A" author="CAMERON, M." box="[1085,1250,467,494]" pageId="39" pageNumber="156" pagination="160 - 168" refId="ref57450" refString="CAMERON, M. 1930: Staphylinidae from British North Borneo, with descriptions of new species. - Journal of the Federated Malay States Museums 16: 160 - 168." type="journal article" year="1930">CAMERON, 1930</bibRefCitation>
</taxonomicName>
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<paragraph id="8BB136ACFFE6E515531A86F36077FDBB" blockId="39.[831,1250,467,527]" box="[831,892,507,527]" pageId="39" pageNumber="156">
(
<figureCitation id="13352A29FFE6E515536186F3607CFDBB" box="[836,887,507,527]" captionStart="Map 9" captionStartId="69.[151,196,1235,1256]" captionTargetBox="[151,1398,232,1226]" captionTargetId="figure-70@69.[151,1398,232,1226]" captionTargetPageId="69" captionText="Map 9: Revised distributions of Zyras bettotanus (black circles), Z. nilgiriensis (star), and Z. longilobatus (white circles) in the" figureDoi="http://doi.org/10.5281/zenodo.5742419" httpUri="https://zenodo.org/record/5742419/files/figure.png" pageId="39" pageNumber="156">Map 9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE6E515533F8524661CFD34" pageId="39" pageNumber="156" type="reference_group">
<paragraph id="8BB136ACFFE6E515533F85246781FDF5" blockId="39.[794,1303,555,641]" box="[794,1162,555,577]" pageId="39" pageNumber="156">
<treatmentCitationGroup id="AB1E1182FFE6E515533F85246781FDF5" box="[794,1162,555,577]" pageId="39" pageNumber="156">
<treatmentCitation id="0AAF10BDFFE6E515533F8524678DFDF5" author="CAMERON, M." box="[794,1158,555,577]" page="168" pageId="39" pageNumber="156" year="1930">
<taxonomicName id="4C0E4D2FFFE6E515533F8524678DFDF5" ID-CoL="7GHYK" authority="CAMERON, 1930: 168" authorityName="Cameron" authorityPageNumber="168" authorityYear="1930" box="[794,1158,555,577]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="bettotanus">
<emphasis id="B97AEABEFFE6E515533F852460BEFDF4" box="[794,949,555,577]" italics="true" pageId="39" pageNumber="156">Zyras bettotanus</emphasis>
<bibRefCitation id="EF9F4B5DFFE6E515539E8523678DFDF5" author="CAMERON, M." box="[955,1158,555,577]" pageId="39" pageNumber="156" pagination="160 - 168" refId="ref57450" refString="CAMERON, M. 1930: Staphylinidae from British North Borneo, with descriptions of new species. - Journal of the Federated Malay States Museums 16: 160 - 168." type="journal article" year="1930">CAMERON, 1930: 168</bibRefCitation>
</taxonomicName>
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.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB136ACFFE6E515533F85446601FDD4" blockId="39.[794,1303,555,641]" box="[794,1290,587,609]" pageId="39" pageNumber="156">
<treatmentCitationGroup id="AB1E1182FFE6E515533F85446601FDD4" box="[794,1290,587,609]" pageId="39" pageNumber="156">
<treatmentCitation id="0AAF10BDFFE6E515533F854467CAFDD4" author="CAMERON, M." box="[794,1217,587,609]" page="17" pageId="39" pageNumber="156" year="1939">
<taxonomicName id="4C0E4D2FFFE6E515533F854467CAFDD4" authority="CAMERON, 1939 b: 17" authorityName="Cameron" authorityPageNumber="17" authorityYear="1939" box="[794,1217,587,609]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="drescheri" subGenus="Zyras">
<emphasis id="B97AEABEFFE6E515533F85446045FDD5" box="[794,846,588,609]" italics="true" pageId="39" pageNumber="156">Zyras</emphasis>
(
<emphasis id="B97AEABEFFE6E51553798544609BFDD5" box="[860,912,588,609]" italics="true" pageId="39" pageNumber="156">Zyras</emphasis>
)
<emphasis id="B97AEABEFFE6E51553B8854360FAFDD4" box="[925,1009,587,608]" italics="true" pageId="39" pageNumber="156">drescheri</emphasis>
<bibRefCitation id="EF9F4B5DFFE6E51553D2854367CAFDD4" author="CAMERON, M." box="[1015,1217,587,609]" pageId="39" pageNumber="156" pagination="1 - 29" refId="ref57528" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">CAMERON, 1939b: 17</bibRefCitation>
</taxonomicName>
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;
<emphasis id="B97AEABEFFE6E51554EE854367E6FDD4" bold="true" box="[1227,1261,587,608]" pageId="39" pageNumber="156">syn</emphasis>
.
<emphasis id="B97AEABEFFE6E51554DD8543660EFDD4" bold="true" box="[1272,1285,587,608]" pageId="39" pageNumber="156">n</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB136ACFFE6E515533F8564661CFD34" blockId="39.[794,1303,555,641]" box="[794,1303,619,641]" pageId="39" pageNumber="156">
<treatmentCitationGroup id="AB1E1182FFE6E515533F8564661CFD34" box="[794,1303,619,641]" pageId="39" pageNumber="156">
<treatmentCitation id="0AAF10BDFFE6E515533F856467B7FD34" author="ASSING, V." box="[794,1212,619,641]" page="140" pageId="39" pageNumber="156" year="2016">
<taxonomicName id="4C0E4D2FFFE6E515533F856467B7FD34" authority="ASSING, 2016 a: 140" authorityName="ASSING" authorityPageNumber="140" authorityYear="2016" box="[794,1212,619,641]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="atrapicalis" subGenus="Zyras">
<emphasis id="B97AEABEFFE6E515533F85646045FD35" box="[794,846,620,641]" italics="true" pageId="39" pageNumber="156">Zyras</emphasis>
(
<emphasis id="B97AEABEFFE6E51553798564609BFD35" box="[860,912,620,641]" italics="true" pageId="39" pageNumber="156">Zyras</emphasis>
)
<emphasis id="B97AEABEFFE6E51553B8856360F5FD34" box="[925,1022,619,640]" italics="true" pageId="39" pageNumber="156">atrapicalis</emphasis>
<bibRefCitation id="EF9F4B5DFFE6E5155421856367B7FD34" author="ASSING, V." box="[1028,1212,619,641]" pageId="39" pageNumber="156" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">
ASSING, 2016a:
<specimenCount id="9D08FD25FFE6E51554BC856367B7FD34" box="[1177,1212,619,640]" pageId="39" pageNumber="156" type="generic">140</specimenCount>
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f.;
<emphasis id="B97AEABEFFE6E51554FD856367F1FD34" bold="true" box="[1240,1274,619,640]" pageId="39" pageNumber="156">syn</emphasis>
.
<emphasis id="B97AEABEFFE6E515552085636619FD34" bold="true" box="[1285,1298,619,640]" pageId="39" pageNumber="156">n</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE6E515533F85A16734FB95" pageId="39" pageNumber="156" type="materials_examined">
<paragraph id="8BB136ACFFE6E515533F85A167C0FCF5" blockId="39.[794,1399,681,1057]" pageId="39" pageNumber="156">
<emphasis id="B97AEABEFFE6E515533F85A1673FFD75" bold="true" box="[794,1076,681,705]" pageId="39" pageNumber="156">Type material examined</emphasis>
:
<taxonomicName id="4C0E4D2FFFE6E515546F85A267D8FD75" authorityName="Cameron" authorityYear="1930" box="[1098,1235,681,705]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="bettotanus">
<emphasis id="B97AEABEFFE6E515546F85A267D8FD75" box="[1098,1235,681,705]" italics="true" pageId="39" pageNumber="156">Z. bettotanus</emphasis>
</taxonomicName>
:
<materialsCitation id="3B663CF1FFE6E51554CC85A167C0FCF5" ID-GBIF-Occurrence="3450162481" collectingDate="1927-08-23" collectionCode="BMNH" collectorName="M. Cameron. Bequest. &amp; V. Assing" location="N. Borneo" pageId="39" pageNumber="156" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B5880EFFE6E51554CC85A16647FD75" box="[1257,1356,681,705]" pageId="39" pageNumber="156" type="holotype">Holotype</typeStatus>
<specimenCount id="9D08FD25FFE6E515557985A2667DFD75" box="[1372,1398,682,705]" pageId="39" pageNumber="156">♂:</specimenCount>
<location id="8ED16077FFE6E515530185C1609EFD55" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE6E515530185C1609EFD55" box="[804,917,713,737]" name="N. Borneo" pageId="39" pageNumber="156">N. Borneo</location>
.
<location id="8ED16077FFE6E515538685C2670BFD55" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE6E515538685C2670BFD55" box="[931,1024,714,737]" name="Bettotan" pageId="39" pageNumber="156">Bettotan</location>
, /
<location id="8ED16077FFE6E515540185C267B0FD55" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE6E515540185C267B0FD55" box="[1060,1211,713,737]" name="Nr. Sandakan" pageId="39" pageNumber="156">Nr. Sandakan</location>
.
<date id="FFB0106CFFE6E51554EF85C1667DFD55" box="[1226,1398,713,737]" pageId="39" pageNumber="156" value="1927-08-23">
<collectingDate id="EFF4E984FFE6E51554EF85C1667DFD55" box="[1226,1398,713,737]" pageId="39" pageNumber="156" value="1927-08-23">Aug. 23rd 1927</collectingDate>
</date>
/
<taxonomicName id="4C0E4D2FFFE6E515530885E26728FCB5" authority="Cam." authorityName="Cameron" authorityYear="1930" box="[813,1059,745,769]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="bettotanus">
<emphasis id="B97AEABEFFE6E515530885E260D5FCB5" box="[813,990,745,769]" italics="true" pageId="39" pageNumber="156">Zyras bettotanus</emphasis>
Cam.
</taxonomicName>
Type /
<collectorName id="26FB537AFFE6E515545A85E2667CFCB5" box="[1151,1399,745,769]" pageId="39" pageNumber="156">M. Cameron. Bequest.</collectorName>
B.M. 1955-147. / Holotype / Holotypus
<taxonomicName id="4C0E4D2FFFE6E51554ED8402608BFCF5" authority="Cameron" authorityName="Cameron" authorityYear="1930" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="bettotanus">
<emphasis id="B97AEABEFFE6E51554ED8402667DFC95" box="[1224,1398,777,801]" italics="true" pageId="39" pageNumber="156">Zyras bettotanus</emphasis>
Cameron
</taxonomicName>
, rev.
<collectorName id="26FB537AFFE6E51553908422671CFCF5" box="[949,1047,809,833]" pageId="39" pageNumber="156">V. Assing</collectorName>
2017” (
<collectionCode id="ED1FAE69FFE6E515544C842167B4FCF5" box="[1129,1215,809,833]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="39" pageNumber="156">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB136ACFFE6E515533F84426734FB95" blockId="39.[794,1399,681,1057]" pageId="39" pageNumber="156">
<materialsCitation id="3B663CF1FFE6E515533F84426766FBB5" ID-GBIF-Occurrence="3455147308" collectingDate="1926-08-07" collectingDateMax="1926-08-09" collectingDateMin="1926-08-07" collectionCode="BMNH" determinerName="V. Assing" location="Java" pageId="39" pageNumber="156" specimenCount="1" stateProvince="Java" typeStatus="lectotype">
<taxonomicName id="4C0E4D2FFFE6E515533F84426098FCD5" authorityName="Cameron" authorityYear="1939" box="[794,915,841,865]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="drescheri">
<emphasis id="B97AEABEFFE6E515533F84426098FCD5" box="[794,915,841,865]" italics="true" pageId="39" pageNumber="156">
<originalDetermination id="A8108E50FFE6E515533F84426098FCD5" box="[794,915,841,865]" pageId="39" pageNumber="156">Z. drescheri</originalDetermination>
</emphasis>
</taxonomicName>
:
<typeStatus id="54B5880EFFE6E515538684426707FCD6" box="[931,1036,842,866]" pageId="39" pageNumber="156" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D08FD25FFE6E515543084426721FCD5" box="[1045,1066,842,865]" pageId="39" pageNumber="156"></specimenCount>
, present designation: “Batoerraden,
<collectorName id="26FB537AFFE6E515534F846160D4FC35" box="[874,991,873,897]" pageId="39" pageNumber="156">G. Slamat.</collectorName>
<collectingRegion id="49CAF84EFFE6E51553CE84626712FC35" box="[1003,1049,874,897]" country="Indonesia" name="Jawa Timur" pageId="39" pageNumber="156">Java</collectingRegion>
,
<collectorName id="26FB537AFFE6E515540C846167CEFC35" box="[1065,1221,873,897]" pageId="39" pageNumber="156">F.C. Drescher.</collectorName>
<date id="FFB0106CFFE6E51554F58461666AFC35" box="[1232,1377,873,897]" pageId="39" pageNumber="156" value="1926-08-07" valueMax="1926-08-09" valueMin="1926-08-07">
<collectingDate id="EFF4E984FFE6E51554F58461666AFC35" box="[1232,1377,873,897]" pageId="39" pageNumber="156" value="1926-08-07" valueMax="1926-08-09" valueMin="1926-08-07">7-9.VIII.1926</collectingDate>
</date>
/
<taxonomicName id="4C0E4D2FFFE6E515533F8482672CFC15" authority="Cam Cotype" authorityName="Cam Cotype" box="[794,1063,905,929]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="drescheri">
<emphasis id="B97AEABEFFE6E515533F84826098FC15" box="[794,915,905,929]" italics="true" pageId="39" pageNumber="156">Z. drescheri</emphasis>
Cam Cotype
</taxonomicName>
/ M. Cameron. Bequest. B.M. 1955-147. / Syntype / Lectotypus ♂
<taxonomicName id="4C0E4D2FFFE6E51554AB84A2604AFC55" authority="Cameron" authorityName="Cameron" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="drescheri">
<emphasis id="B97AEABEFFE6E51554AB84A26621FC75" box="[1166,1322,937,961]" italics="true" pageId="39" pageNumber="156">Zyras drescheri</emphasis>
Cameron
</taxonomicName>
, desig.
<collectorName id="26FB537AFFE6E51553B384C260F3FC55" box="[918,1016,969,993]" pageId="39" pageNumber="156">V. Assing</collectorName>
2017 /
<taxonomicName id="4C0E4D2FFFE6E515547784C26679FC55" authority="Cameron" authorityName="Cameron" authorityYear="1930" box="[1106,1394,969,993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="bettotanus">
<emphasis id="B97AEABEFFE6E515547784C26609FC55" box="[1106,1282,969,993]" italics="true" pageId="39" pageNumber="156">Zyras bettotanus</emphasis>
Cameron
</taxonomicName>
, det.
<determinerName id="7E6DDA68FFE6E515536B84E260BBFBB5" box="[846,944,1001,1025]" pageId="39" pageNumber="156">V. Assing</determinerName>
2017” (
<collectionCode id="ED1FAE69FFE6E515542A84E1676EFBB5" box="[1039,1125,1001,1025]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="39" pageNumber="156">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B663CF1FFE6E515545984E16730FB95" ID-GBIF-Occurrence="3450162363" collectingDate="1926-08-07" collectingDateMax="1926-08-09" collectingDateMin="1926-08-07" collectionCode="BMNH" determinerName="V. Assing" location="Java" pageId="39" pageNumber="156" specimenCount="2" stateProvince="Jawa Timur" typeStatus="paralectotype">
<typeStatus id="54B5880EFFE6E515545984E16606FBB5" box="[1148,1293,1001,1025]" pageId="39" pageNumber="156" type="paralectotype">Paralectotype</typeStatus>
♀: same data as
<typeStatus id="54B5880EFFE6E515534A830160DBFB95" box="[879,976,1033,1057]" pageId="39" pageNumber="156" type="lectotype">lectotype</typeStatus>
(
<collectionCode id="ED1FAE69FFE6E51553F883016738FB95" box="[989,1075,1033,1057]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="39" pageNumber="156">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE6E51A533F8341627FFC15" lastPageId="40" lastPageNumber="157" pageId="39" pageNumber="156" type="discussion">
<paragraph id="8BB136ACFFE6E515533F83416619F9F5" blockId="39.[794,1399,1097,1761]" pageId="39" pageNumber="156">
<emphasis id="B97AEABEFFE6E515533F8341608CFBD5" bold="true" box="[794,903,1097,1121]" pageId="39" pageNumber="156">Comment</emphasis>
: The original description of
<taxonomicName id="4C0E4D2FFFE6E51554F683426652FBD5" authorityName="Cameron" authorityYear="1930" box="[1235,1369,1097,1121]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="bettotanus">
<emphasis id="B97AEABEFFE6E51554F683426652FBD5" box="[1235,1369,1097,1121]" italics="true" pageId="39" pageNumber="156">Z. bettotanus</emphasis>
</taxonomicName>
is based on a unique
<typeStatus id="54B5880EFFE6E51553C283616748FB35" box="[999,1091,1129,1153]" pageId="39" pageNumber="156" type="holotype">holotype</typeStatus>
from “
<collectingCountry id="F319763CFFE6E51554B483616621FB35" box="[1169,1322,1129,1153]" name="Malaysia" pageId="39" pageNumber="156">North Borneo</collectingCountry>
; Bettotan” (
<bibRefCitation id="EF9F4B5DFFE6E5155377838160FEFB15" author="CAMERON, M." box="[850,1013,1161,1185]" pageId="39" pageNumber="156" pagination="160 - 168" refId="ref57450" refString="CAMERON, M. 1930: Staphylinidae from British North Borneo, with descriptions of new species. - Journal of the Federated Malay States Museums 16: 160 - 168." type="journal article" year="1930">CAMERON 1930</bibRefCitation>
).
<bibRefCitation id="EF9F4B5DFFE6E515542C838267D6FB15" author="HLAVAC, P. &amp; NEWTON, A. F. &amp; MARUYAMA, M." box="[1033,1245,1161,1189]" pageId="39" pageNumber="156" pagination="1 - 151" refId="ref57922" refString="HLAVAC, P.; NEWTON, A. F. &amp; MARUYAMA, M. 2011: World catalogue of the species of the tribe Lomechusini (Staphylinidae: Aleocharinae). - Zootaxa 3075: 1 - 151." type="journal article" year="2011">HLAVÁČ et al. (2011)</bibRefCitation>
list the species as incertae sedis, but
<bibRefCitation id="EF9F4B5DFFE6E51553D483A16764FB75" author="PACE, R." box="[1009,1135,1193,1217]" pageId="39" pageNumber="156" pagination="107 - 156" refId="ref58804" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE (2008)</bibRefCitation>
assigned it to
<taxonomicName id="4C0E4D2FFFE6E51554DA83A2667CFB75" authority="sensu" authorityName="SenSU" box="[1279,1399,1194,1217]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEABEFFE6E51554DA83A26633FB75" box="[1279,1336,1194,1217]" italics="true" pageId="39" pageNumber="156">Zyras</emphasis>
sensu
</taxonomicName>
strictu in a key to the
<taxonomicName id="4C0E4D2FFFE6E515542183C26786FB55" box="[1028,1165,1225,1249]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97AEABEFFE6E515542183C26736FB55" box="[1028,1085,1226,1249]" italics="true" pageId="39" pageNumber="156">Zyras</emphasis>
species
</taxonomicName>
of Borneo.
<bibRefCitation id="EF9F4B5DFFE6E515552E83C16060FAB5" author="CAMERON, M." pageId="39" pageNumber="156" pagination="1 - 29" refId="ref57528" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">CAMERON (1939b)</bibRefCitation>
described
<taxonomicName id="4C0E4D2FFFE6E51553D783E26763FAB5" authorityName="Cameron" authorityYear="1939" box="[1010,1128,1257,1281]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="drescheri">
<emphasis id="B97AEABEFFE6E51553D783E26763FAB5" box="[1010,1128,1257,1281]" italics="true" pageId="39" pageNumber="156">Z. drescheri</emphasis>
</taxonomicName>
based on an unspecified number of
<typeStatus id="54B5880EFFE6E51553EC82026728FA96" box="[969,1059,1290,1314]" pageId="39" pageNumber="156" type="syntype">syntypes</typeStatus>
from “Batoerraden: G. Slamat”, “Bandoeng Dago”, and “G. Tangkoeban Prahoe”. Two of the
<typeStatus id="54B5880EFFE6E5155347824260B5FAD6" box="[866,958,1354,1378]" pageId="39" pageNumber="156" type="syntype">syntypes</typeStatus>
, a male and a female, were found in the Cameron collection at the BMNH. The male is designated as the
<typeStatus id="54B5880EFFE6E5155346828160CFFA15" box="[867,964,1417,1441]" pageId="39" pageNumber="156" type="lectotype">lectotype</typeStatus>
.
<taxonomicName id="4C0E4D2FFFE6E51553F582826777FA15" authorityName="ASSING" authorityYear="2016" box="[976,1148,1417,1441]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="atrapicalis">
<emphasis id="B97AEABEFFE6E51553F582826777FA15" box="[976,1148,1417,1441]" italics="true" pageId="39" pageNumber="156">Zyras atrapicalis</emphasis>
</taxonomicName>
was recently described based on material from the Chinese province Yunnan. An exmination of the type material of all three names revealed that it is conspecific, so that
<taxonomicName id="4C0E4D2FFFE6E515546382E267B6F9B5" authorityName="Cameron" authorityYear="1939" box="[1094,1213,1513,1537]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="drescheri">
<emphasis id="B97AEABEFFE6E515546382E267B6F9B5" box="[1094,1213,1513,1537]" italics="true" pageId="39" pageNumber="156">Z. drescheri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E4D2FFFE6E51554D782E2667DF9B5" authorityName="ASSING" authorityYear="2016" box="[1266,1398,1513,1537]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="atrapicalis">
<emphasis id="B97AEABEFFE6E51554D782E2667DF9B5" box="[1266,1398,1513,1537]" italics="true" pageId="39" pageNumber="156">Z. atrapicalis</emphasis>
</taxonomicName>
are placed in synonymy with
<taxonomicName id="4C0E4D2FFFE6E5155477810267D3F995" authorityName="Cameron" authorityYear="1930" box="[1106,1240,1545,1569]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="bettotanus">
<emphasis id="B97AEABEFFE6E5155477810267D3F995" box="[1106,1240,1545,1569]" italics="true" pageId="39" pageNumber="156">Z. bettotanus</emphasis>
</taxonomicName>
. For a detailed description and illustrations see
<bibRefCitation id="EF9F4B5DFFE6E515544C81216605F9F5" author="ASSING, V." box="[1129,1294,1577,1601]" pageId="39" pageNumber="156" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a)</bibRefCitation>
.
</paragraph>
<paragraph id="8BB136ACFFE6E515533F81416725F955" blockId="39.[794,1399,1097,1761]" pageId="39" pageNumber="156">
The coloration of this species is remarkably variable. While the head and pronotum are usually blackish in material from most regions, some specimens from
<collectingRegion id="49CAF84EFFE6E515556E8182667DF915" box="[1355,1398,1674,1697]" country="Indonesia" name="Jawa Timur" pageId="39" pageNumber="156">Java</collectingRegion>
have the head and pronotum reddish, or the head reddish and the pronotum brown.
</paragraph>
<paragraph id="8BB136ACFFE6E51A533F8001627FFC15" blockId="39.[794,1400,1801,2081]" lastBlockId="40.[189,795,233,929]" lastPageId="40" lastPageNumber="157" pageId="39" pageNumber="156">
<emphasis id="B97AEABEFFE6E515533F8001677AF895" bold="true" box="[794,1137,1801,1825]" pageId="39" pageNumber="156">Additional material examined</emphasis>
:
<materialsCitation id="3B663CF1FFE6E51554AD80016679F835" ID-GBIF-Occurrence="3450162431" collectingDate="1988-04-18" collectingDateMax="1988-04-27" collectingDateMin="1988-04-18" collectionCode="BMNH" collectorName="Brendell" country="Thailand" elevation="300" latitude="15.466666" location="Umphang district" longLatPrecision="1285" longitude="98.8" pageId="39" pageNumber="156" specimenCount="3" stateProvince="Tak province">
<emphasis id="B97AEABEFFE6E51554AD800167E1F895" bold="true" box="[1160,1258,1801,1825]" pageId="39" pageNumber="156">
<collectingCountry id="F319763CFFE6E51554AD800167E1F895" box="[1160,1258,1801,1825]" name="Thailand" pageId="39" pageNumber="156">Thailand</collectingCountry>
</emphasis>
:
<specimenCount id="9D08FD25FFE6E515552580016637F895" box="[1280,1340,1801,1825]" pageId="39" pageNumber="156" type="generic">3 exs.</specimenCount>
,
<collectingRegion id="49CAF84EFFE6E51555758001607CF8F5" country="Thailand" name="Tak" pageId="39" pageNumber="156">Tak province</collectingRegion>
,
<location id="8ED16077FFE6E51553AD80216740F8F5" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE6E51553AD80216740F8F5" box="[904,1099,1833,1857]" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Umphang district" pageId="39" pageNumber="156" stateProvince="Tak province">Umphang district</location>
,
<taxonomicName id="4C0E4D2FFFE6E515547880216718F8D5" authority="Bae Stream, Thung Yai Wildlife Sanctuary" authorityName="Bae Stream, Thung Yai Wildlife Sanctuary" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="156" phylum="Arthropoda" rank="species" species="song">
<location id="8ED16077FFE6E515547880216614F8F5" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE6E515547880216614F8F5" box="[1117,1311,1833,1857]" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Song Bae Stream" pageId="39" pageNumber="156" stateProvince="Tak province">Song Bae Stream</location>
,
<location id="8ED16077FFE6E515551580216718F8D5" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE6E515551580216718F8D5" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Thung Yai Wildlife Sanctuary" pageId="39" pageNumber="156" stateProvince="Tak province">Thung Yai Wildlife Sanctuary</location>
</taxonomicName>
,
<geoCoordinate id="EE3A506BFFE6E515540180416772F8D5" box="[1060,1145,1865,1889]" degrees="15" direction="north" minutes="28" orientation="latitude" pageId="39" pageNumber="156" precision="925" value="15.466666">15°28'N</geoCoordinate>
,
<geoCoordinate id="EE3A506BFFE6E51554AE804167D0F8D5" box="[1163,1243,1865,1889]" degrees="98" direction="east" minutes="48" orientation="longitude" pageId="39" pageNumber="156" precision="925" value="98.8">98°48'E</geoCoordinate>
,
<quantity id="4CF69B49FFE6E51554C880416624F8D5" box="[1261,1327,1865,1889]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="156" unit="m" value="300.0">
<elevation id="0023D19FFFE6E51554C880416624F8D5" box="[1261,1327,1865,1889]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="156" unit="m" value="300.0">300 m</elevation>
</quantity>
, evergreen rain forest,
<date id="FFB0106CFFE6E51553F980616778F835" box="[988,1139,1897,1921]" pageId="39" pageNumber="156" value="1988-04-18" valueMax="1988-04-27" valueMin="1988-04-18">
<collectingDate id="EFF4E984FFE6E51553F980616778F835" box="[988,1139,1897,1921]" pageId="39" pageNumber="156" value="1988-04-18" valueMax="1988-04-27" valueMin="1988-04-18">1827.IV.1988</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE6E515548B8061660DF835" box="[1198,1286,1897,1921]" pageId="39" pageNumber="156">Brendell</collectorName>
(
<collectionCode id="ED1FAE69FFE6E515553280616660F835" box="[1303,1387,1897,1921]" pageId="39" pageNumber="156">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFE6E515533F8081672BF7B5" ID-GBIF-Occurrence="3450162343" collectingDate="1988-04-27" collectingDateMax="1988-05-06" collectingDateMin="1988-04-27" collectionCode="BMNH" collectorName="Brendell" country="Thailand" county="Umphang district" elevation="300" latitude="15.5" location="Thung Yai Wildlife Sanctuary" longLatPrecision="1284" longitude="98.8" municipality="Mae Chan" pageId="39" pageNumber="156" specimenCount="11" stateProvince="Tak province">
<specimenCount id="9D08FD25FFE6E515533F80816069F815" box="[794,866,1929,1953]" pageId="39" pageNumber="156" type="generic">11 exs.</specimenCount>
,
<collectingRegion id="49CAF84EFFE6E515534E808160FEF815" box="[875,1013,1929,1953]" country="Thailand" name="Tak" pageId="39" pageNumber="156">Tak province</collectingRegion>
,
<collectingCounty id="62D04E20FFE6E51553DB808167B2F815" box="[1022,1209,1929,1953]" pageId="39" pageNumber="156">Umphang district</collectingCounty>
,
<collectingMunicipality id="6BD5ACD6FFE6E51554E780826624F815" box="[1218,1327,1929,1953]" pageId="39" pageNumber="156">Mae Chan</collectingMunicipality>
Mae Klong confluence,,
<location id="8ED16077FFE6E51553C780A1661EF875" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE6E51553C780A1661EF875" box="[994,1301,1961,1985]" country="Thailand" county="Umphang district" latitude="15.5" longLatPrecision="1284" longitude="98.8" municipality="Mae Chan" name="Thung Yai Wildlife Sanctuary" pageId="39" pageNumber="156" stateProvince="Tak province">Thung Yai Wildlife Sanctuary</location>
,
<geoCoordinate id="EE3A506BFFE6E515553B80A16678F875" box="[1310,1395,1961,1985]" degrees="15" direction="north" minutes="30" orientation="latitude" pageId="39" pageNumber="156" precision="925" value="15.5">15°30'N</geoCoordinate>
,
<geoCoordinate id="EE3A506BFFE6E515533F80C16061F855" box="[794,874,1993,2017]" degrees="98" direction="east" minutes="48" orientation="longitude" pageId="39" pageNumber="156" precision="925" value="98.8">98°48'E</geoCoordinate>
,
<quantity id="4CF69B49FFE6E515535180C160BDF855" box="[884,950,1993,2017]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="156" unit="m" value="300.0">
<elevation id="0023D19FFFE6E515535180C160BDF855" box="[884,950,1993,2017]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="156" unit="m" value="300.0">300 m</elevation>
</quantity>
, evergreen rain forest,
<date id="FFB0106CFFE6E515548180C16642F855" box="[1188,1353,1993,2017]" pageId="39" pageNumber="156" value="1988-04-27" valueMax="1988-05-06" valueMin="1988-04-27">
<collectingDate id="EFF4E984FFE6E515548180C16642F855" box="[1188,1353,1993,2017]" pageId="39" pageNumber="156" value="1988-04-27" valueMax="1988-05-06" valueMin="1988-04-27">27.IV.6.V.1988</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE6E515533F80E16079F7B5" box="[794,882,2025,2049]" pageId="39" pageNumber="156">Brendell</collectorName>
(
<collectionCode id="ED1FAE69FFE6E51553A180E160DDF7B5" box="[900,982,2025,2049]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="39" pageNumber="156">BMNH</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFE6E51A541580E161ACFE95" ID-GBIF-Occurrence="3450162373" collectingDate="1988-04-27" collectingDateMax="1988-05-06" collectingDateMin="1988-04-27" collectorName="Brendell" country="Thailand" county="Umphang district" elevation="300" lastPageId="40" lastPageNumber="157" latitude="15.5" location="Thung Yai Wildlife Sanctuary" longLatPrecision="1284" longitude="98.8" municipality="Mae Chan" pageId="39" pageNumber="156" specimenCount="1" stateProvince="Tak province">
<specimenCount id="9D08FD25FFE6E515541580E1676AF7B5" box="[1072,1121,2025,2049]" pageId="39" pageNumber="156" type="generic">1 ex.</specimenCount>
,
<collectingRegion id="49CAF84EFFE6E515545580E167F4F7B5" box="[1136,1279,2025,2049]" country="Thailand" name="Tak" pageId="39" pageNumber="156">Tak province</collectingRegion>
,
<collectingCounty id="62D04E20FFE6E515552B80E1606DF795" pageId="39" pageNumber="156">Umphang district</collectingCounty>
,
<collectingMunicipality id="6BD5ACD6FFE6E51553568F0260E8F795" box="[883,995,2057,2081]" pageId="39" pageNumber="156">Mae Chan</collectingMunicipality>
Mae Klong confluence,
<location id="8ED16077FFE6E51A55228F016274FEB5" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE6E51A55228F016274FEB5" country="Thailand" county="Umphang district" lastPageId="40" lastPageNumber="157" latitude="15.5" longLatPrecision="1284" longitude="98.8" municipality="Mae Chan" name="Thung Yai Wildlife Sanctuary" pageId="39" pageNumber="156" stateProvince="Tak province">Thung Yai Wildlife Sanctuary</location>
,
<geoCoordinate id="EE3A506BFFE9E51A51AD87E162D7FEB5" box="[392,476,233,257]" degrees="15" direction="north" minutes="30" orientation="latitude" pageId="40" pageNumber="157" precision="925" value="15.5">15°30'N</geoCoordinate>
,
<geoCoordinate id="EE3A506BFFE9E51A51C087E1613DFEB5" box="[485,566,233,257]" degrees="98" direction="east" minutes="48" orientation="longitude" pageId="40" pageNumber="157" precision="925" value="98.8">98°48'E</geoCoordinate>
,
<quantity id="4CF69B49FFE9E51A521A87E1618AFEB5" box="[575,641,233,257]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="40" pageNumber="157" unit="m" value="300.0">
<elevation id="0023D19FFFE9E51A521A87E1618AFEB5" box="[575,641,233,257]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="40" pageNumber="157" unit="m" value="300.0">300 m</elevation>
</quantity>
, edge of
<location id="8ED16077FFE9E51A52FF87E26011FEB5" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE9E51A52FF87E26011FEB5" box="[730,794,234,257]" country="Thailand" county="Umphang district" latitude="15.5" longLatPrecision="1284" longitude="98.8" municipality="Mae Chan" name="Karen" pageId="40" pageNumber="157" stateProvince="Tak province">Karen</location>
clearing,
<date id="FFB0106CFFE9E51A5105860162CEFE95" box="[288,453,265,289]" pageId="40" pageNumber="157" value="1988-04-27" valueMax="1988-05-06" valueMin="1988-04-27">
<collectingDate id="EFF4E984FFE9E51A5105860162CEFE95" box="[288,453,265,289]" pageId="40" pageNumber="157" value="1988-04-27" valueMax="1988-05-06" valueMin="1988-04-27">27.IV.6.V.1988</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE9E51A522486016152FE95" box="[513,601,265,289]" pageId="40" pageNumber="157">Brendell</collectorName>
(cAss)
</materialsCitation>
.
<materialsCitation id="3B663CF1FFE9E51A52908601620CFED5" ID-GBIF-Occurrence="3450162380" collectingDate="1984-03-22" collectingDateMax="1984-03-30" collectingDateMin="1984-03-22" collectorName="Rougemont" country="Malaysia" location="Cameron Highlands" pageId="40" pageNumber="157" specimenCount="1">
<emphasis id="B97AEABEFFE9E51A52908601601FFE95" bold="true" box="[693,788,265,289]" pageId="40" pageNumber="157">
<collectingCountry id="F319763CFFE9E51A52908601601FFE95" box="[693,788,265,289]" name="Malaysia" pageId="40" pageNumber="157">Malaysia</collectingCountry>
</emphasis>
: 1 ♀,
<location id="8ED16077FFE9E51A50CB862162CDFEF5" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE9E51A50CB862162CDFEF5" box="[238,454,297,321]" country="Malaysia" name="Cameron Highlands" pageId="40" pageNumber="157">Cameron Highlands</location>
,
<date id="FFB0106CFFE9E51A51F586216161FEF5" box="[464,618,297,321]" pageId="40" pageNumber="157" value="1984-03-22" valueMax="1984-03-30" valueMin="1984-03-22">
<collectingDate id="EFF4E984FFE9E51A51F586216161FEF5" box="[464,618,297,321]" pageId="40" pageNumber="157" value="1984-03-22" valueMax="1984-03-30" valueMin="1984-03-22">2230.III.1984</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE9E51A52BA86226011FEF6" box="[671,794,298,322]" pageId="40" pageNumber="157">Rougemont</collectorName>
(cRou)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFE9E51A5135864262CCFE15" ID-GBIF-Occurrence="3455147310" collectingDate="2004-03" collectingMethod="flight interception trap" collectorName="Maruyama" country="Malaysia" elevation="250" location="Ulu Gombak Field Studies Centre" pageId="40" pageNumber="157" specimenCount="7" stateProvince="Selangor">
<specimenCount id="9D08FD25FFE9E51A513586426247FED5" box="[272,332,330,353]" pageId="40" pageNumber="157" type="generic">7 exs.</specimenCount>
,
<collectingRegion id="49CAF84EFFE9E51A5171864162A4FED5" box="[340,431,329,353]" country="Malaysia" name="Selangor" pageId="40" pageNumber="157">Selangor</collectingRegion>
,
<location id="8ED16077FFE9E51A51928641601DFED5" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE9E51A51928641601DFED5" box="[439,790,329,353]" country="Malaysia" name="Ulu Gombak Field Studies Centre" pageId="40" pageNumber="157" stateProvince="Selangor">Ulu Gombak Field Studies Centre</location>
,
<quantity id="4CF69B49FFE9E51A5098866163F4FE35" box="[189,255,361,385]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="40" pageNumber="157" unit="m" value="250.0">
<elevation id="0023D19FFFE9E51A5098866163F4FE35" box="[189,255,361,385]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="40" pageNumber="157" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<collectingMethod id="524F4EBBFFE9E51A512A8661610CFE35" box="[271,519,361,385]" pageId="40" pageNumber="157">flight interception trap</collectingMethod>
,
<date id="FFB0106CFFE9E51A523086616161FE35" box="[533,618,361,385]" pageId="40" pageNumber="157" value="2004-03">
<collectingDate id="EFF4E984FFE9E51A523086616161FE35" box="[533,618,361,385]" pageId="40" pageNumber="157" value="2004-03">III.2004</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE9E51A528286626012FE36" box="[679,793,362,386]" pageId="40" pageNumber="157">Maruyama</collectorName>
et al. (cHla, cMar, cAss)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFE9E51A51F286816251FE55" ID-GBIF-Occurrence="3450162331" collectingDate="1997-03" collectingMethod="flight interception trap" collectionCode="BMNH" collectorName="S Lokan &amp; Chung" country="Malaysia" location="Sandakan" municipality="Borneo" pageId="40" pageNumber="157" specimenCount="8" stateProvince="Sabah">
<specimenCount id="9D08FD25FFE9E51A51F286816118FE15" box="[471,531,393,417]" pageId="40" pageNumber="157" type="generic">8 exs.</specimenCount>
,
<collectingMunicipality id="6BD5ACD6FFE9E51A52078682617AFE15" box="[546,625,394,417]" pageId="40" pageNumber="157">Borneo</collectingMunicipality>
,
<collectingRegion id="49CAF84EFFE9E51A52A4868161C9FE15" box="[641,706,393,417]" country="Malaysia" name="Sabah" pageId="40" pageNumber="157">Sabah</collectingRegion>
,
<location id="8ED16077FFE9E51A52F4868163EDFE75" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE9E51A52F4868163EDFE75" country="Malaysia" municipality="Borneo" name="Sandakan" pageId="40" pageNumber="157" stateProvince="Sabah">Sandakan</location>
,
<collectorName id="26FB537AFFE9E51A50D586A1624CFE75" box="[240,327,425,449]" pageId="40" pageNumber="157">S Lokan</collectorName>
,
<collectingMethod id="524F4EBBFFE9E51A517486A1614BFE75" box="[337,576,425,449]" pageId="40" pageNumber="157">flight interception trap</collectingMethod>
,
<date id="FFB0106CFFE9E51A526C86A16195FE75" box="[585,670,425,449]" pageId="40" pageNumber="157" value="1997-03">
<collectingDate id="EFF4E984FFE9E51A526C86A16195FE75" box="[585,670,425,449]" pageId="40" pageNumber="157" value="1997-03">III.1997</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE9E51A52F786A16011FE75" box="[722,794,425,449]" pageId="40" pageNumber="157">Chung</collectorName>
(
<collectionCode id="ED1FAE69FFE9E51A50E186C1621DFE55" box="[196,278,457,481]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="40" pageNumber="157">BMNH</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFE9E51A514686C161DAFE55" ID-GBIF-Occurrence="3450162309" box="[355,721,457,481]" collectingDate="1996-09" collectionCode="BMNH" collectorName="S Lokan &amp; Chung" country="Malaysia" location="Sandakan" municipality="Borneo" pageId="40" pageNumber="157" specimenCount="1" stateProvince="Sabah">
<specimenCount id="9D08FD25FFE9E51A514686C1629EFE55" box="[355,405,457,481]" pageId="40" pageNumber="157" type="generic">1 ex.</specimenCount>
, same data, but
<date id="FFB0106CFFE9E51A521286C16182FE55" box="[567,649,457,481]" pageId="40" pageNumber="157" value="1996-09">
<collectingDate id="EFF4E984FFE9E51A521286C16182FE55" box="[567,649,457,481]" pageId="40" pageNumber="157" value="1996-09">IX.1996</collectingDate>
</date>
(cAss)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFE9E51A52FE86C162D7FD95" ID-GBIF-Occurrence="3450162407" collectingDate="1996-10" collectingMethod="flight interception trap" collectionCode="BMNH" collectorName="Chung" country="Malaysia" location="Sepilok" municipality="Sandakan" pageId="40" pageNumber="157" specimenCount="4" stateProvince="Sabah">
<specimenCount id="9D08FD25FFE9E51A52FE86C1601DFE55" box="[731,790,457,481]" pageId="40" pageNumber="157" type="generic">4 exs.</specimenCount>
,
<collectingRegion id="49CAF84EFFE9E51A509886E163F5FDB5" box="[189,254,489,513]" country="Malaysia" name="Sabah" pageId="40" pageNumber="157">Sabah</collectingRegion>
,
<collectingMunicipality id="6BD5ACD6FFE9E51A512D86E1627BFDB5" box="[264,368,489,513]" pageId="40" pageNumber="157">Sandakan</collectingMunicipality>
,
<location id="8ED16077FFE9E51A515F86E162C2FDB5" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE9E51A515F86E162C2FDB5" box="[378,457,489,513]" country="Malaysia" municipality="Sandakan" name="Sepilok" pageId="40" pageNumber="157" stateProvince="Sabah">Sepilok</location>
,
<collectingMethod id="524F4EBBFFE9E51A51F686E161CAFDB5" box="[467,705,489,513]" pageId="40" pageNumber="157">flight interception trap</collectingMethod>
,
<date id="FFB0106CFFE9E51A52EF86E1601DFDB5" box="[714,790,489,513]" pageId="40" pageNumber="157" value="1996-10">
<collectingDate id="EFF4E984FFE9E51A52EF86E1601DFDB5" box="[714,790,489,513]" pageId="40" pageNumber="157" value="1996-10">X.1996</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE9E51A50CF85016239FD95" box="[234,306,521,545]" pageId="40" pageNumber="157">Chung</collectorName>
(
<collectionCode id="ED1FAE69FFE9E51A51678501629FFD95" box="[322,404,521,545]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="40" pageNumber="157">BMNH</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFE9E51A51CF85016243FDD5" ID-GBIF-Occurrence="3450162324" collectingDate="2007-02-14" collectingDateMax="2007-02-16" collectingDateMin="2007-02-14" collectingMethod="flight interception trap" collectorName="B. R. &amp; Rougemont" country="Malaysia" location="Danum valley" pageId="40" pageNumber="157" specimenCount="2" stateProvince="Sabah">
<specimenCount id="9D08FD25FFE9E51A51CF8501612EFD95" box="[490,549,521,545]" pageId="40" pageNumber="157" type="generic">2 exs.</specimenCount>
,
<collectingRegion id="49CAF84EFFE9E51A521785016178FD95" box="[562,627,521,545]" country="Malaysia" name="Sabah" pageId="40" pageNumber="157">Sabah</collectingRegion>
,
<location id="8ED16077FFE9E51A52A58502601CFD95" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE9E51A52A58502601CFD95" box="[640,791,521,545]" country="Malaysia" name="Danum valley" pageId="40" pageNumber="157" stateProvince="Sabah">Danum valley</location>
,
<collectorName id="26FB537AFFE9E51A5098852263E2FDF5" box="[189,233,554,577]" name="M. BERNHAUER" pageId="40" pageNumber="157">B.R.</collectorName>
L.,
<collectingMethod id="524F4EBBFFE9E51A512F852162F0FDF5" box="[266,507,553,577]" pageId="40" pageNumber="157">flight interception trap</collectingMethod>
,
<date id="FFB0106CFFE9E51A52238521619CFDF5" box="[518,663,553,577]" pageId="40" pageNumber="157" value="2007-02-14" valueMax="2007-02-16" valueMin="2007-02-14">
<collectingDate id="EFF4E984FFE9E51A52238521619CFDF5" box="[518,663,553,577]" pageId="40" pageNumber="157" value="2007-02-14" valueMax="2007-02-16" valueMin="2007-02-14">1416.II.2007</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE9E51A52EB852263FEFDD5" pageId="40" pageNumber="157">Rougemont</collectorName>
(cRou)
</materialsCitation>
.
<materialsCitation id="3B663CF1FFE9E51A517685416130FD15" ID-GBIF-Occurrence="3450162405" collectingDate="2004-01-20" collectingDateMax="2004-02-07" collectingDateMin="2004-01-20" collectingMethod="flight interception trap" collectorName="Maruyama" country="Indonesia" elevation="250" location="Kebun Raya Bogor" municipality="Bogor" pageId="40" pageNumber="157" specimenCount="1" stateProvince="Jawa Barat">
<emphasis id="B97AEABEFFE9E51A5176854162B6FDD5" bold="true" box="[339,445,585,609]" pageId="40" pageNumber="157">
<collectingCountry id="F319763CFFE9E51A5176854162B6FDD5" box="[339,445,585,609]" name="Indonesia" pageId="40" pageNumber="157">Indonesia</collectingCountry>
</emphasis>
: 2 ♂♂, 3♀♀,
<collectingRegion id="49CAF84EFFE9E51A527D854261C1FDD5" box="[600,714,586,609]" country="Indonesia" name="Jawa Barat" pageId="40" pageNumber="157">Jawa Barat</collectingRegion>
,
<collectingMunicipality id="6BD5ACD6FFE9E51A52F08542601DFDD6" box="[725,790,586,610]" pageId="40" pageNumber="157">Bogor</collectingMunicipality>
,
<location id="8ED16077FFE9E51A50988561628AFD36" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE9E51A50988561628AFD36" box="[189,385,617,642]" country="Indonesia" municipality="Bogor" name="Kebun Raya Bogor" pageId="40" pageNumber="157" stateProvince="Jawa Barat">Kebun Raya Bogor</location>
,
<quantity id="4CF69B49FFE9E51A51AF856162C7FD35" box="[394,460,617,641]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="40" pageNumber="157" unit="m" value="250.0">
<elevation id="0023D19FFFE9E51A51AF856162C7FD35" box="[394,460,617,641]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="40" pageNumber="157" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<collectingMethod id="524F4EBBFFE9E51A51F3856161C8FD35" box="[470,707,617,641]" pageId="40" pageNumber="157">flight interception trap</collectingMethod>
,
<date id="FFB0106CFFE9E51A52E985616202FD15" pageId="40" pageNumber="157" value="2004-01-20" valueMax="2004-02-07" valueMin="2004-01-20">
<collectingDate id="EFF4E984FFE9E51A52E985616202FD15" pageId="40" pageNumber="157" value="2004-01-20" valueMax="2004-02-07" valueMin="2004-01-20">20.I.7. II.2004</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE9E51A5118858262A4FD16" box="[317,431,650,674]" pageId="40" pageNumber="157">Maruyama</collectorName>
(cMar, cAss)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFE9E51A5260858163F3FCB5" ID-GBIF-Occurrence="3455147304" collectingDate="2004-01-29" collectingDateMax="2004-02-04" collectingDateMin="2004-01-29" collectingMethod="flight interception trap" collectorName="Maruyama" country="Indonesia" county="Cianjur" elevation="1400" location="Cibodas" municipality="Kebun Raya" pageId="40" pageNumber="157" specimenCount="1" stateProvince="Jawa Barat">
1 ♂, 1 ♀,
<collectingRegion id="49CAF84EFFE9E51A52838582601DFD15" box="[678,790,650,673]" country="Indonesia" name="Jawa Barat" pageId="40" pageNumber="157">Jawa Barat</collectingRegion>
,
<collectingCounty id="62D04E20FFE9E51A509885A16206FD75" box="[189,269,681,705]" pageId="40" pageNumber="157">Cianjur</collectingCounty>
,
<collectingMunicipality id="6BD5ACD6FFE9E51A513985A162A9FD76" box="[284,418,681,706]" pageId="40" pageNumber="157">Kebun Raya</collectingMunicipality>
,
<location id="8ED16077FFE9E51A519785A16101FD75" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE9E51A519785A16101FD75" box="[434,522,681,705]" country="Indonesia" county="Cianjur" municipality="Kebun Raya" name="Cibodas" pageId="40" pageNumber="157" stateProvince="Jawa Barat">Cibodas</location>
,
<quantity id="4CF69B49FFE9E51A523F85A16162FD75" box="[538,617,681,705]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="40" pageNumber="157" unit="m" value="1400.0">
<elevation id="0023D19FFFE9E51A523F85A16162FD75" box="[538,617,681,705]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="40" pageNumber="157" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<collectingMethod id="524F4EBBFFE9E51A525C85A16228FD55" pageId="40" pageNumber="157">flight interception trap</collectingMethod>
,
<date id="FFB0106CFFE9E51A511285C162C0FD55" box="[311,459,713,737]" pageId="40" pageNumber="157" value="2004-01-29" valueMax="2004-02-04" valueMin="2004-01-29">
<collectingDate id="EFF4E984FFE9E51A511285C162C0FD55" box="[311,459,713,737]" pageId="40" pageNumber="157" value="2004-01-29" valueMax="2004-02-04" valueMin="2004-01-29">29.I.-4.II.2004</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE9E51A523685C2618EFD56" box="[531,645,714,738]" pageId="40" pageNumber="157">Maruyama</collectorName>
et al. (cMar, cAss)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFE9E51A512185E161CAFCF5" ID-GBIF-Occurrence="3450162329" collectingDate="2001-08" collectingMethod="flight interception trap" collectionCode="BMNH" collectorName="Brendell &amp; Mendel" country="Indonesia" latitude="-0.05" location="Busang - Rekut" longLatPrecision="1308" longitude="113.98333" municipality="Borneo" pageId="40" pageNumber="157" specimenCount="9" stateProvince="Kalimantan Tengah">
<specimenCount id="9D08FD25FFE9E51A512185E16234FCB5" box="[260,319,745,769]" pageId="40" pageNumber="157" type="generic">9 exs.</specimenCount>
,
<collectingMunicipality id="6BD5ACD6FFE9E51A516E85E26291FCB5" box="[331,410,746,769]" pageId="40" pageNumber="157">Borneo</collectingMunicipality>
,
<collectingRegion id="49CAF84EFFE9E51A518085E1617CFCB5" box="[421,631,745,769]" country="Indonesia" name="Kalimantan Tengah" pageId="40" pageNumber="157">Kalimantan Tengah</collectingRegion>
,
<location id="8ED16077FFE9E51A52A785E16011FCB5" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE9E51A52A785E16011FCB5" box="[642,794,745,769]" country="Indonesia" latitude="-0.05" longLatPrecision="1308" longitude="113.98333" municipality="Borneo" name="Busang - Rekut" pageId="40" pageNumber="157" stateProvince="Kalimantan Tengah">BusangRekut</location>
confluence,
<geoCoordinate id="EE3A506BFFE9E51A51628401628CFC95" box="[327,391,777,801]" degrees="0" direction="south" minutes="03" orientation="latitude" pageId="40" pageNumber="157" precision="925" value="-0.05">0°03'S</geoCoordinate>
,
<geoCoordinate id="EE3A506BFFE9E51A51BB840162F1FC95" box="[414,506,777,801]" degrees="113" direction="east" minutes="59" orientation="longitude" pageId="40" pageNumber="157" precision="925" value="113.98333">113°59'E</geoCoordinate>
,
<collectingMethod id="524F4EBBFFE9E51A52358401601DFC95" box="[528,790,777,801]" pageId="40" pageNumber="157">flight interception trap</collectingMethod>
,
<date id="FFB0106CFFE9E51A50988421622FFCF5" box="[189,292,809,833]" pageId="40" pageNumber="157" value="2001-08">
<collectingDate id="EFF4E984FFE9E51A50988421622FFCF5" box="[189,292,809,833]" pageId="40" pageNumber="157" value="2001-08">VIII.2001</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE9E51A5172842162A4FCF5" box="[343,431,809,833]" pageId="40" pageNumber="157">Brendell</collectorName>
&amp;
<collectorName id="26FB537AFFE9E51A51E884216116FCF5" box="[461,541,809,833]" pageId="40" pageNumber="157">Mendel</collectorName>
(
<collectionCode id="ED1FAE69FFE9E51A520F84216177FCF5" box="[554,636,809,833]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="40" pageNumber="157">BMNH</collectionCode>
, cAss)
</materialsCitation>
.
<materialsCitation id="3B663CF1FFE9E51A52EF84216264FC15" ID-GBIF-Occurrence="3450162478" collectingDate="1992-10" collectingMethod="Malaise trap" collectionCode="BMNH" collectorName="Martin" country="Brunei" elevation="150" latitude="4.5666666" location="Kuala Belalong" longLatPrecision="1306" longitude="115.11667" pageId="40" pageNumber="157" specimenCount="1" stateProvince="Temburong District">
<emphasis id="B97AEABEFFE9E51A52EF8421601FFCF5" bold="true" box="[714,788,809,833]" pageId="40" pageNumber="157">
<collectingCountry id="F319763CFFE9E51A52EF8421601FFCF5" box="[714,788,809,833]" name="Brunei" pageId="40" pageNumber="157">Brunei</collectingCountry>
</emphasis>
:
<specimenCount id="9D08FD25FFE9E51A5098844163E4FCD5" box="[189,239,841,865]" pageId="40" pageNumber="157" type="generic">1 ex.</specimenCount>
,
<collectingRegion id="49CAF84EFFE9E51A50DB844162DFFCD5" box="[254,468,841,865]" country="Brunei" name="Temburong" pageId="40" pageNumber="157">Temburong District</collectingRegion>
,
<location id="8ED16077FFE9E51A51C68441618EFCD5" LSID="urn:lsid:plazi:treatment:03A787BAFFE6E51A531A86DB618DFB15:8ED16077FFE9E51A51C68441618EFCD5" box="[483,645,841,865]" country="Brunei" latitude="4.5666666" longLatPrecision="1306" longitude="115.11667" name="Kuala Belalong" pageId="40" pageNumber="157" stateProvince="Temburong District">Kuala Belalong</location>
FSC,
<geoCoordinate id="EE3A506BFFE9E51A52E88441601EFCD5" box="[717,789,841,865]" degrees="4" direction="north" minutes="34" orientation="latitude" pageId="40" pageNumber="157" precision="925" value="4.5666666">4°34'N</geoCoordinate>
,
<geoCoordinate id="EE3A506BFFE9E51A509884616211FC35" box="[189,282,873,897]" degrees="115" direction="east" minutes="07" orientation="longitude" pageId="40" pageNumber="157" precision="925" value="115.11667">115°07'E</geoCoordinate>
,
<quantity id="4CF69B49FFE9E51A510C84616260FC35" box="[297,363,873,897]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="40" pageNumber="157" unit="m" value="150.0">
<elevation id="0023D19FFFE9E51A510C84616260FC35" box="[297,363,873,897]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="40" pageNumber="157" unit="m" value="150.0">150 m</elevation>
</quantity>
, steep forest,
<collectingMethod id="524F4EBBFFE9E51A522384616186FC35" box="[518,653,873,897]" pageId="40" pageNumber="157">Malaise trap</collectingMethod>
,
<date id="FFB0106CFFE9E51A52B9846161ECFC35" box="[668,743,873,897]" pageId="40" pageNumber="157" value="1992-10">
<collectingDate id="EFF4E984FFE9E51A52B9846161ECFC35" box="[668,743,873,897]" pageId="40" pageNumber="157" value="1992-10">X.1992</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE9E51A50988481620CFC15" box="[189,263,905,929]" pageId="40" pageNumber="157">Martin</collectorName>
(
<collectionCode id="ED1FAE69FFE9E51A513184816262FC15" box="[276,361,905,929]" pageId="40" pageNumber="157">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE9E51A509884C1618DFB15" pageId="40" pageNumber="157" type="distribution">
<paragraph id="8BB136ACFFE9E51A509884C1618DFB15" blockId="40.[189,794,969,1185]" pageId="40" pageNumber="157">
<emphasis id="B97AEABEFFE9E51A509884C1613FFC55" bold="true" box="[189,564,969,993]" pageId="40" pageNumber="157">Distribution and natural history</emphasis>
:
<taxonomicName id="4C0E4D2FFFE9E51A526284C261F0FC55" authorityName="Cameron" authorityYear="1930" box="[583,763,969,993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="157" phylum="Arthropoda" rank="species" species="bettotanus">
<emphasis id="B97AEABEFFE9E51A526284C261F0FC55" box="[583,763,969,993]" italics="true" pageId="40" pageNumber="157">Zyras bettotanus</emphasis>
</taxonomicName>
is evidently widespread in the Oriental region (
<figureCitation id="13352A29FFE9E51A528984E161F8FBB5" box="[684,755,1001,1025]" captionStart="Map 9" captionStartId="69.[151,196,1235,1256]" captionTargetBox="[151,1398,232,1226]" captionTargetId="figure-70@69.[151,1398,232,1226]" captionTargetPageId="69" captionText="Map 9: Revised distributions of Zyras bettotanus (black circles), Z. nilgiriensis (star), and Z. longilobatus (white circles) in the" figureDoi="http://doi.org/10.5281/zenodo.5742419" httpUri="https://zenodo.org/record/5742419/files/figure.png" pageId="40" pageNumber="157">Map 9</figureCitation>
). It is now known from South
<collectingCountry id="F319763CFFE9E51A51CD83016127FB95" box="[488,556,1033,1057]" name="China" pageId="40" pageNumber="157">China</collectingCountry>
,
<collectingCountry id="F319763CFFE9E51A521C83016190FB95" box="[569,667,1033,1057]" name="Thailand" pageId="40" pageNumber="157">Thailand</collectingCountry>
, Peninsular
<collectingCountry id="F319763CFFE9E51A509883216216FBF5" box="[189,285,1065,1089]" name="Malaysia" pageId="40" pageNumber="157">Malaysia</collectingCountry>
, Java, and Borneo (
<collectingCountry id="F319763CFFE9E51A522583216168FBF5" box="[512,611,1065,1089]" name="Malaysia" pageId="40" pageNumber="157">Malaysia</collectingCountry>
:
<collectingRegion id="49CAF84EFFE9E51A5257832161BFFBF5" box="[626,692,1065,1089]" country="Malaysia" name="Sabah" pageId="40" pageNumber="157">Sabah</collectingRegion>
;
<collectingCountry id="F319763CFFE9E51A52E1832163D7FBD5" name="Indonesia" pageId="40" pageNumber="157">Indonesia</collectingCountry>
:
<collectingRegion id="49CAF84EFFE9E51A50C2834162B1FBD5" box="[231,442,1097,1121]" country="Indonesia" name="Kalimantan Tengah" pageId="40" pageNumber="157">Kalimantan Tengah</collectingRegion>
;
<collectingCountry id="F319763CFFE9E51A51E083416119FBD5" box="[453,530,1097,1121]" name="Brunei" pageId="40" pageNumber="157">Brunei</collectingCountry>
). At least the majority of the specimens was collected with flight interception and Malaise traps at low altitudes (
<quantity id="4CF69B49FFE9E51A51DB83816172FB15" box="[510,633,1161,1185]" metricMagnitude="2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="40" pageNumber="157" unit="m" value="225.0" valueMax="300.0" valueMin="150.0">150300 m</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>