240 lines
39 KiB
XML
240 lines
39 KiB
XML
<document id="EC24FAFB4007A19C57AB4BB82D57905E" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFE5E51650C685BB60EAF835" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) parvicollis Assing 2017, spec. nov." docType="treatment" docUuid="317676A6-1E6A-495F-9864-B7DFA85717F4" docUuidSource="ZooBank" docVersion="11" lastPageNumber="153" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="153" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="FE651EE4DF9C10FCCB2BFDD4B6D03BD6">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="3B5F508847A1D67CE2E80645DB538D35">Assing, Volker</mods:namePart>
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<mods:title id="1443DC90A4CBE6FC391D83DC90E44C6B">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="E99878273167C6428D1D5615DF31230D">
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<mods:date id="856477EFCC8ABEDFCA6684862C812749">2017</mods:date>
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<mods:number id="EE397E55B0FF2A7F2D7DADE5CF4B1F44">2017-06-30</mods:number>
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<mods:number id="B100C014B73D423CD03786990E6E3318">67</mods:number>
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<treatment id="03A787BAFFE5E51650C685BB60EAF835" ID-DOI="http://doi.org/10.5281/zenodo.5888483" ID-GBIF-Taxon="192498197" ID-Zenodo-Dep="5888483" LSID="urn:lsid:zoobank.org:act:317676A6-1E6A-495F-9864-B7DFA85717F4" httpUri="http://treatment.plazi.org/id/03A787BAFFE5E51650C685BB60EAF835" lastPageNumber="153" pageId="36" pageNumber="153">
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<subSubSection id="C3146527FFE5E51650C685BB61EFFD5B" pageId="36" pageNumber="153" type="nomenclature">
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<paragraph id="8BB136ACFFE5E51650C685BB614BFD7A" blockId="36.[227,740,691,783]" box="[227,576,691,718]" pageId="36" pageNumber="153">
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<heading id="D0F981C0FFE5E51650C685BB614BFD7A" box="[227,576,691,718]" fontSize="12" level="2" pageId="36" pageNumber="153" reason="2">
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<taxonomicName id="4C0E4D2FFFE5E51650C685BB62DBFD7A" ID-CoL="7GHXB" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[227,464,691,718]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="parvicollis" status="sp. nov." subGenus="Zyras">
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<emphasis id="B97AEABEFFE5E51650C685BB6212FD7A" box="[227,281,691,718]" italics="true" pageId="36" pageNumber="153">Zyras</emphasis>
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(
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<emphasis id="B97AEABEFFE5E516510285BB6256FD7A" box="[295,349,691,718]" italics="true" pageId="36" pageNumber="153">Zyras</emphasis>
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)
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<emphasis id="B97AEABEFFE5E516514F85BB62DBFD7A" box="[362,464,691,718]" italics="true" pageId="36" pageNumber="153">parvicollis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24957C5FFE5E51651F285BB614BFD7A" box="[471,576,691,718]" pageId="36" pageNumber="153" rank="species">spec. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BB136ACFFE5E51650C685D361EFFD5B" blockId="36.[227,740,691,783]" box="[227,740,731,751]" pageId="36" pageNumber="153">
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<uri id="FF9F3AAEFFE5E51650C685D361EFFD5B" box="[227,740,731,751]" pageId="36" pageNumber="153">
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urn:lsid:zoobank.org:act:
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146527FFE5E51650C685F362F8FCBB" box="[227,499,763,783]" pageId="36" pageNumber="153" type="description">
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<paragraph id="8BB136ACFFE5E51650C685F362F8FCBB" blockId="36.[227,740,691,783]" box="[227,499,763,783]" pageId="36" pageNumber="153">
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(
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<figureCitation id="13352A29FFE5E51650C285F3622BFCBB" box="[231,288,763,783]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="36" pageNumber="153">Figs 33</figureCitation>
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,
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<figureCitation id="13352A29FFE5E516510285F36237FCBB" box="[295,316,763,783]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="36" pageNumber="153">69</figureCitation>
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,
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<figureCitation id="13352A29FFE5E516516685F3626AFCBB" box="[323,353,763,783]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="36" pageNumber="153">104</figureCitation>
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,
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<figureCitation id="13352A29FFE5E516514D85F362A6FCBB" box="[360,429,763,783]" captionStart-0="Figs 191–207" captionStart-1="Figs 208–226" captionStartId-0="59.[151,191,2011,2033]" captionStartId-1="60.[189,229,1991,2013]" captionTargetBox-0="[153,1385,236,1953]" captionTargetBox-1="[196,1429,238,1961]" captionTargetId-0="figure-115@59.[151,1398,231,1970]" captionTargetId-1="figure-130@60.[189,1436,232,1971]" captionTargetPageId-0="59" captionText-0="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." captionText-1="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5888374" figureDoi-1="http://doi.org/10.5281/zenodo.5888376" httpUri-0="https://zenodo.org/record/5888374/files/figure.png" httpUri-1="https://zenodo.org/record/5888376/files/figure.png" pageId="36" pageNumber="153">206–211</figureCitation>
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,
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<figureCitation id="13352A29FFE5E516519185F362E4FCBB" box="[436,495,763,783]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="36" pageNumber="153">Map 10</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3146527FFE5E516509884216268FBB5" pageId="36" pageNumber="153" type="materials_examined">
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<paragraph id="8BB136ACFFE5E516509884216268FBB5" blockId="36.[189,794,809,1025]" pageId="36" pageNumber="153">
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<emphasis id="B97AEABEFFE5E516509884216250FCF5" bold="true" box="[189,347,809,833]" pageId="36" pageNumber="153">Type material</emphasis>
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:
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<materialsCitation id="3B663CF1FFE5E516514A84216263FC55" ID-GBIF-Occurrence="3450162359" collectingDate="1988-04-18" collectingDateMax="1988-04-27" collectingDateMin="1988-04-18" collectionCode="BMNH" collectorName="M. J. D. Brendell." country="Thailand" determinerName="V. Assing" elevation="300" latitude="15.466666" location="Thung Yai Wildlife Sanctuary" longLatPrecision="1285" longitude="98.8" pageId="36" pageNumber="153" specimenCount="1" stateProvince="Tak Province" typeStatus="holotype">
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<typeStatus id="54B5880EFFE5E516514A842162D9FCF5" box="[367,466,809,833]" pageId="36" pageNumber="153" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D08FD25FFE5E51651C5842262FEFCF5" box="[480,501,810,833]" pageId="36" pageNumber="153">♂</specimenCount>
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: “W.
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<collectingCountry id="F319763CFFE5E516520A842161B0FCF5" box="[559,699,809,833]" name="Thailand" pageId="36" pageNumber="153">THAILAND</collectingCountry>
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:
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</quantity>
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.,
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<location id="8ED16077FFE5E516509884416109FCD5" LSID="urn:lsid:plazi:treatment:03A787BAFFE5E51650C685BB60EAF835:8ED16077FFE5E516509884416109FCD5" box="[189,514,841,865]" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Thung Yai Wildlife Sanctuary" pageId="36" pageNumber="153" stateProvince="Tak Province">Thung Yai Wildlife Sanctuary</location>
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.
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<geoCoordinate id="EE3A506BFFE5E516522884416169FCD5" box="[525,610,841,865]" degrees="15" direction="north" minutes="28" orientation="latitude" pageId="36" pageNumber="153" precision="925" value="15.466666">15°28'N</geoCoordinate>
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–
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<geoCoordinate id="EE3A506BFFE5E51652A7844161DFFCD5" box="[642,724,841,865]" degrees="98" direction="east" minutes="48" orientation="longitude" pageId="36" pageNumber="153" precision="925" value="98.8">98°48'E</geoCoordinate>
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. /
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<collectingRegion id="49CAF84EFFE5E51652D684416216FC35" country="Thailand" name="Tak" pageId="36" pageNumber="153">Tak Province</collectingRegion>
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,
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<location id="8ED16077FFE5E516510D846162E5FC35" LSID="urn:lsid:plazi:treatment:03A787BAFFE5E51650C685BB60EAF835:8ED16077FFE5E516510D846162E5FC35" box="[296,494,873,897]" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Umphang District" pageId="36" pageNumber="153" stateProvince="Tak Province">Umphang District</location>
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,
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<taxonomicName id="4C0E4D2FFFE5E51651DD846161BEFC35" authority="Bae Stream." authorityName="Bae Stream." box="[504,693,873,897]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="song">
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<location id="8ED16077FFE5E51651DD846161BAFC35" LSID="urn:lsid:plazi:treatment:03A787BAFFE5E51650C685BB60EAF835:8ED16077FFE5E51651DD846161BAFC35" box="[504,689,873,897]" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Song Bae Stream" pageId="36" pageNumber="153" stateProvince="Tak Province">Song Bae Stream</location>
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||
.
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</taxonomicName>
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||
<date id="FFB0106CFFE5E516529E846163F8FC15" pageId="36" pageNumber="153" value="1988-04-18" valueMax="1988-04-27" valueMin="1988-04-18">
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<collectingDate id="EFF4E984FFE5E516529E846163F8FC15" pageId="36" pageNumber="153" value="1988-04-18" valueMax="1988-04-27" valueMin="1988-04-18">18–27.iv. 1988</collectingDate>
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</date>
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. / Evergreen rain forest.
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<collectorName id="26FB537AFFE5E51651D184826194FC15" box="[500,671,905,929]" pageId="36" pageNumber="153">M.J.D. Brendell.</collectorName>
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B.M. 1988- 183 / Holotypus ♂
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<taxonomicName id="4C0E4D2FFFE5E51651B784A26134FC75" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[402,575,937,961]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="parvicollis" status="sp. nov.">
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||
<emphasis id="B97AEABEFFE5E51651B784A26134FC75" box="[402,575,937,961]" italics="true" pageId="36" pageNumber="153">Zyras parvicollis</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A24957C5FFE5E516526284A26174FC75" box="[583,639,938,961]" pageId="36" pageNumber="153" rank="species">sp. n.</taxonomicNameLabel>
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||
det.
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||
<determinerName id="7E6DDA68FFE5E516529284A26012FC75" box="[695,793,937,961]" pageId="36" pageNumber="153">V. Assing</determinerName>
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||
2016” (
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||
<collectionCode id="ED1FAE69FFE5E516512F84C1626BFC55" box="[266,352,969,993]" pageId="36" pageNumber="153">BMNH</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B663CF1FFE5E516515784C26255FBB5" ID-GBIF-Occurrence="3450162364" collectingDate="1988-04-18" collectingDateMax="1988-04-27" collectingDateMin="1988-04-18" collectionCode="BMNH" collectorName="M. J. D. Brendell." country="Thailand" determinerName="V. Assing" elevation="300" latitude="15.466666" location="Umphang District" longLatPrecision="1285" longitude="98.8" pageId="36" pageNumber="153" specimenCount="1" stateProvince="Tak" typeStatus="paratype">
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||
<typeStatus id="54B5880EFFE5E516515784C262D7FC56" box="[370,476,970,994]" pageId="36" pageNumber="153" type="paratype">Paratypes</typeStatus>
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: 3 ♂♂: same data as holotype (
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<collectionCode id="ED1FAE69FFE5E51650E184E16213FBB5" box="[196,280,1001,1025]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="36" pageNumber="153">BMNH</collectionCode>
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||
, cAss)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3146527FFE5E5165098832161CEFBD5" pageId="36" pageNumber="153" type="etymology">
|
||
<paragraph id="8BB136ACFFE5E5165098832161CEFBD5" blockId="36.[189,794,1065,1121]" pageId="36" pageNumber="153">
|
||
<emphasis id="B97AEABEFFE5E51650988321623FFBF5" bold="true" box="[189,308,1065,1089]" pageId="36" pageNumber="153">Etymology</emphasis>
|
||
: The specific epithet (adjective) alludes to the relatively small and weakly transverse pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFE5E51650988381608BF915" pageId="36" pageNumber="153" type="description">
|
||
<paragraph id="8BB136ACFFE5E516509883816179F9D5" blockId="36.[189,794,1161,2081]" pageId="36" pageNumber="153">
|
||
<emphasis id="B97AEABEFFE5E516509883816234FB15" bold="true" box="[189,319,1161,1185]" pageId="36" pageNumber="153">Description</emphasis>
|
||
: Size highly variable: body length
|
||
<quantity id="4CF69B49FFE5E516509883A16235FB75" box="[189,318,1193,1217]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" metricValueMax="6.8" metricValueMin="5.8" pageId="36" pageNumber="153" unit="mm" value="6.3" valueMax="6.8" valueMin="5.8">5.8–6.8 mm</quantity>
|
||
; length of forebody 2.5–3.0 mm. Coloration (
|
||
<figureCitation id="13352A29FFE5E51650D283C1624DFB55" box="[247,326,1225,1249]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="36" pageNumber="153">Figs 33</figureCitation>
|
||
,
|
||
<figureCitation id="13352A29FFE5E516517683C16264FB55" box="[339,367,1225,1249]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="36" pageNumber="153">69</figureCitation>
|
||
,
|
||
<figureCitation id="13352A29FFE5E516515F83C162AFFB55" box="[378,420,1225,1249]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="36" pageNumber="153">104</figureCitation>
|
||
): head and pronotum black; elytra dark-yellowish with the postero-lateral portion more or less extensively and rather sharply blackish; abdomen with tergites III–V dark-brown to blackish-brown with the posterior margins and the paratergites more or less extensively dark-reddish, tergite VI blackish-brown to black with the posterior margin and the paratergites dark-reddish, tergites VII–VIII and segments IX–X blackish-brown to black; legs pale-yellowish; antennae blackish with antennomeres I–II yellowish-brown to brown and the apex of antennomere XI dark-reddish; maxillary palpi reddish-brown to dark-brown, with the terminal palpomere yellowish to reddish.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE5E516509881616171F815" blockId="36.[189,794,1161,2081]" pageId="36" pageNumber="153">
|
||
Head (
|
||
<figureCitation id="13352A29FFE5E51651238161625AF935" box="[262,337,1641,1665]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="36" pageNumber="153">Fig. 69</figureCitation>
|
||
) distinctly transverse, broadly impunctate along middle, in lateral portions with sparse and moderately coarse punctation. Eyes large and bulging, much longer than postocular region in dorsal view. Antenna (
|
||
<figureCitation id="13352A29FFE5E51650E081E16204F8B5" box="[197,271,1769,1793]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="36" pageNumber="153">Fig. 33</figureCitation>
|
||
) slender,
|
||
<quantity id="4CF69B49FFE5E516515E81E162F0F8B5" box="[379,507,1769,1793]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="36" pageNumber="153" unit="mm" value="2.2" valueMax="2.3" valueMin="2.1">2.1–2.3 mm</quantity>
|
||
long; antennomeres IV–V weakly oblong, VI approximately as long as broad; VII approximately as long as broad or weakly transverse; VIII–X weakly transverse, X approximately 1.5 times as broad as long, and XI distinctly elongate, approximately as long as the combined length of VIII–X.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE5E516509880A2671DFE35" blockId="36.[189,794,1161,2081]" lastBlockId="36.[831,1437,233,1313]" pageId="36" pageNumber="153">
|
||
Pronotum (
|
||
<figureCitation id="13352A29FFE5E516511C80A1628FF875" box="[313,388,1961,1985]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="36" pageNumber="153">Fig. 69</figureCitation>
|
||
) relatively small (in relation to head), approximately 1.2 times as broad as long and 1.10–1.18 times as broad as head, broadest near anterior angles; lateral margins straight or weakly convex in posterior two-thirds (dorsal view), strongly converging posteriad; surface smooth, without impressions; punctation coarse, moderately sparse, and moderately irregularly distributed; median line narrowly impunctate or anteriorly with scattered punctures.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE5E51653658681608BFD95" blockId="36.[831,1437,233,1313]" pageId="36" pageNumber="153">
|
||
Elytra (
|
||
<figureCitation id="13352A29FFE5E51653B1868160EAFE15" box="[916,993,393,417]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="36" pageNumber="153">Fig. 69</figureCitation>
|
||
) 0.88–0.95 times as long as pronotum; punctation coarse and dense, somewhat sparser in posterior fourth. Hind wings fully developed. Metatarsomere I slightly shorter than the combined length of II–IV.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE5E516531A85216708FBD5" blockId="36.[831,1437,233,1313]" pageId="36" pageNumber="153">
|
||
Abdomen (
|
||
<figureCitation id="13352A29FFE5E516539E8521671EFDF5" box="[955,1045,553,577]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="36" pageNumber="153">Fig. 104</figureCitation>
|
||
) slightly narrower than elytra, with moderately deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a row or with a transverse band of coarse non-setiferous punctures; tergite III with a lateral setiferous puncture on either side and with four setiferous punctures at posterior margin (median pair somewhat separated from margin); tergites IV and V with a lateral setiferous puncture on either side and with six setiferous punctures at posterior margin; tergite VI with a narrow transverse row of non-setiferous punctures at anterior margin, with a lateral setiferous puncture on either side, and with six setiferous punctures at or near posterior margin; tergite VII with a transverse band of non-setiferous punctures anteriorly, and with two transverse rows of setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII with sparse long and black setae in posterior portion.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE5E516531A836267ABFA95" blockId="36.[831,1437,233,1313]" pageId="36" pageNumber="153">
|
||
♂: posterior margin of tergite VIII (
|
||
<figureCitation id="13352A29FFE5E51654C283616634FB35" box="[1255,1343,1129,1153]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="36" pageNumber="153">Fig. 206</figureCitation>
|
||
) with a median pair of blunt teeth; sternite VIII (
|
||
<figureCitation id="13352A29FFE5E51655188381669EFB15" box="[1341,1429,1161,1185]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="36" pageNumber="153">Fig. 207</figureCitation>
|
||
) weakly transverse, posterior margin convex; median lobe of aedeagus
|
||
<quantity id="4CF69B49FFE5E516542783C16790FB55" box="[1026,1179,1225,1249]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="36" pageNumber="153" unit="mm" value="0.825" valueMax="0.85" valueMin="0.8">0.80–0.85 mm</quantity>
|
||
long and shaped as in
|
||
<figureCitation id="13352A29FFE5E516531A83E160C5FAB5" box="[831,974,1257,1281]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="36" pageNumber="153">Figs 208–211</figureCitation>
|
||
; paramere approximately
|
||
<quantity id="4CF69B49FFE5E51654C083E1663CFAB5" box="[1253,1335,1257,1281]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="36" pageNumber="153" unit="mm" value="0.9">0.9 mm</quantity>
|
||
long and with moderately long apical lobe.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE5E516531A8241608BF915" blockId="36.[831,1437,1353,1697]" pageId="36" pageNumber="153">
|
||
<emphasis id="B97AEABEFFE5E516531A8241671CFAD5" bold="true" box="[831,1047,1353,1377]" pageId="36" pageNumber="153">Comparative notes</emphasis>
|
||
:
|
||
<taxonomicName id="4C0E4D2FFFE5E516540E824267D4FAD5" authorityName="Assing" authorityYear="2017" box="[1067,1247,1353,1377]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="parvicollis">
|
||
<emphasis id="B97AEABEFFE5E516540E824267D4FAD5" box="[1067,1247,1353,1377]" italics="true" pageId="36" pageNumber="153">Zyras parvicollis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from the similarly coloured
|
||
<taxonomicName id="4C0E4D2FFFE5E5165451826267FAFA35" authorityName="CAMERON" authorityYear="1939" box="[1140,1265,1386,1409]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="proximus">
|
||
<emphasis id="B97AEABEFFE5E5165451826267FAFA35" box="[1140,1265,1386,1409]" italics="true" pageId="36" pageNumber="153">Z. proximus</emphasis>
|
||
</taxonomicName>
|
||
by a pronotum without impressions and with less irregular punctation, by the presence of only two blunt teeth at the posterior margin of the male tergite VIII, by a transverse male sternite VIII, by the shape of the ventral process of the aedeagus, and by a shorter apical lobe of the paramere. From other sympatric species of similar habitus and punctation (
|
||
<taxonomicName id="4C0E4D2FFFE5E51653E381426755F9D5" authorityName="Assing" authorityYear="2017" box="[966,1118,1610,1633]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="novinversus">
|
||
<emphasis id="B97AEABEFFE5E51653E381426755F9D5" box="[966,1118,1610,1633]" italics="true" pageId="36" pageNumber="153">Z. novinversus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0E4D2FFFE5E516544881426603F9D5" authorityName="Assing" authorityYear="2017" box="[1133,1288,1609,1633]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="brevilobatus">
|
||
<emphasis id="B97AEABEFFE5E516544881426603F9D5" box="[1133,1288,1609,1633]" italics="true" pageId="36" pageNumber="153">Z. brevilobatus</emphasis>
|
||
</taxonomicName>
|
||
), it is readily separated by the different coloration of the abdomen alone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFE5E516531A81C160EAF835" pageId="36" pageNumber="153" type="distribution">
|
||
<paragraph id="8BB136ACFFE5E516531A81C160EAF835" blockId="36.[831,1436,1737,1921]" pageId="36" pageNumber="153">
|
||
<emphasis id="B97AEABEFFE5E516531A81C167B9F955" bold="true" box="[831,1202,1737,1761]" pageId="36" pageNumber="153">Distribution and natural history</emphasis>
|
||
: The
|
||
<typeStatus id="54B5880EFFE5E51654DD81C2662DF956" box="[1272,1318,1738,1762]" pageId="36" pageNumber="153">type</typeStatus>
|
||
locality is situated in West
|
||
<collectingCountry id="F319763CFFE5E51653D581E1675BF8B5" box="[1008,1104,1769,1793]" name="Thailand" pageId="36" pageNumber="153">Thailand</collectingCountry>
|
||
(
|
||
<figureCitation id="13352A29FFE5E516547881E167B8F8B5" box="[1117,1203,1769,1793]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="36" pageNumber="153">Map 10</figureCitation>
|
||
). The specimens were collected in an evergreen rain forest at an altitude of
|
||
<quantity id="4CF69B49FFE5E516531A8021608FF8F5" box="[831,900,1833,1857]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="36" pageNumber="153" unit="m" value="300.0">300 m</quantity>
|
||
, most likely with flight (Malaise?) traps, together with
|
||
<taxonomicName id="4C0E4D2FFFE5E51653A18042670AF8D5" authorityName="CAMERON" authorityYear="1939" box="[900,1025,1866,1889]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="proximus">
|
||
<emphasis id="B97AEABEFFE5E51653A18042670AF8D5" box="[900,1025,1866,1889]" italics="true" pageId="36" pageNumber="153">Z. proximus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0E4D2FFFE5E5165438804267BFF8D5" authorityName="Assing" authorityYear="2017" box="[1053,1204,1866,1889]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="novinversus">
|
||
<emphasis id="B97AEABEFFE5E5165438804267BFF8D5" box="[1053,1204,1866,1889]" italics="true" pageId="36" pageNumber="153">Z. novinversus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0E4D2FFFE5E51654F580426652F8D5" authorityName="Cameron" authorityYear="1930" box="[1232,1369,1865,1889]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="bettotanus">
|
||
<emphasis id="B97AEABEFFE5E51654F580426652F8D5" box="[1232,1369,1865,1889]" italics="true" pageId="36" pageNumber="153">Z. bettotanus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0E4D2FFFE5E516531A806260D0F835" authorityName="Assing" authorityYear="2017" box="[831,987,1897,1921]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="brevilobatus">
|
||
<emphasis id="B97AEABEFFE5E516531A806260D0F835" box="[831,987,1897,1921]" italics="true" pageId="36" pageNumber="153">Z. brevilobatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |