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<document id="BE9B9518B2AD72ACC42E216D73889F08" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFD3E52050C686FB670BF815" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) morvani Pace 1986" docType="treatment" docVersion="11" lastPageNumber="135" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="135" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="DC2A6CDE3FD3B8946A551FA50150097D">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="289572198893F17D77F060E04DE07E9B">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="1061911A107C4BAE6DB9C965036A23C2">text</mods:typeOfResource>
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<mods:title id="53FC9749DE4A63E7C56DE6947FE98934">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="34F074F43422357CBDEA39CAB89C6EFC">
<mods:date id="946E63BCBEF9D24E2A73EE4980FB1F38">2017</mods:date>
<mods:detail id="A35276C93FA8E3907C49218C09C85FEC" type="pubDate">
<mods:number id="B9DF4DF4D1A701174401CB36D54316AB">2017-06-30</mods:number>
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<mods:number id="A06B27AD6161F80B6E98E79708B5F225">67</mods:number>
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<mods:identifier id="994ECC7581AE17C9D9A1CC193CDAF605" type="CLB-Dataset">2472</mods:identifier>
<mods:identifier id="1BB827859C6701D0DF6CD1AEF3F010AE" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
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</mods:mods>
<treatment id="03A787BAFFD3E52050C686FB670BF815" ID-DOI="http://doi.org/10.5281/zenodo.5888465" ID-GBIF-Taxon="192498167" ID-Zenodo-Dep="5888465" LSID="urn:lsid:plazi:treatment:03A787BAFFD3E52050C686FB670BF815" httpUri="http://treatment.plazi.org/id/03A787BAFFD3E52050C686FB670BF815" lastPageNumber="135" pageId="18" pageNumber="135">
<subSubSection id="C3146527FFD3E52050C686FB613EFDBA" box="[227,565,499,526]" pageId="18" pageNumber="135" type="nomenclature">
<paragraph id="8BB136ACFFD3E52050C686FB613EFDBA" blockId="18.[227,565,499,559]" box="[227,565,499,526]" pageId="18" pageNumber="135">
<heading id="D0F981C0FFD3E52050C686FB613EFDBA" box="[227,565,499,526]" centered="true" fontSize="12" level="2" pageId="18" pageNumber="135" reason="2">
<taxonomicName id="4C0E4D2FFFD3E52050C686FB613EFDBA" ID-CoL="5DDDV" authority="PACE, 1986" authorityName="Pace" authorityYear="1986" box="[227,565,499,526]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="morvani" subGenus="Zyras">
<emphasis id="B97AEABEFFD3E52050C686FB6212FDBA" box="[227,281,499,526]" italics="true" pageId="18" pageNumber="135">Zyras</emphasis>
(
<emphasis id="B97AEABEFFD3E520510286FB6256FDBA" box="[295,349,499,526]" italics="true" pageId="18" pageNumber="135">Zyras</emphasis>
)
<emphasis id="B97AEABEFFD3E520514F86FB62B7FDBA" box="[362,444,499,526]" italics="true" pageId="18" pageNumber="135">morvani</emphasis>
PACE, 1986
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD3E52050C6851362EAFD9B" box="[227,481,539,559]" pageId="18" pageNumber="135" type="description">
<paragraph id="8BB136ACFFD3E52050C6851362EAFD9B" blockId="18.[227,565,499,559]" box="[227,481,539,559]" pageId="18" pageNumber="135">
(
<figureCitation id="13352A29FFD3E52050C28513622BFD9B" box="[231,288,539,559]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="18" pageNumber="135">Figs 25</figureCitation>
,
<figureCitation id="13352A29FFD3E520510285136237FD9B" box="[295,316,539,559]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="18" pageNumber="135">51</figureCitation>
,
<figureCitation id="13352A29FFD3E520516685136253FD9B" box="[323,344,539,559]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="18" pageNumber="135">93</figureCitation>
,
<figureCitation id="13352A29FFD3E520517A851362A8FD9B" box="[351,419,539,559]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="18" pageNumber="135">148–149</figureCitation>
,
<figureCitation id="13352A29FFD3E520518F851362D7FD9B" box="[426,476,539,559]" captionStart="Map 3" captionStartId="64.[189,233,2058,2079]" captionTargetBox="[189,1436,1492,2047]" captionTargetId="figure-120@64.[189,1436,1492,2047]" captionTargetPageId="64" captionText="Map 3: Revised distributionsof Zyras perforatus (blackcircles),Z. morvani(white circles),andZ. truncatus(white star)in theHimalaya." figureDoi="http://doi.org/10.5281/zenodo.5742399" httpUri="https://zenodo.org/record/5742399/files/figure.png" pageId="18" pageNumber="135">Map 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD3E520509885446135FDD4" box="[189,574,587,609]" pageId="18" pageNumber="135" type="reference_group">
<paragraph id="8BB136ACFFD3E520509885446135FDD4" blockId="18.[189,574,587,609]" box="[189,574,587,609]" pageId="18" pageNumber="135">
<treatmentCitationGroup id="AB1E1182FFD3E520509885446135FDD4" box="[189,574,587,609]" pageId="18" pageNumber="135">
<treatmentCitation id="0AAF10BDFFD3E520509885446131FDD4" author="PACE, R." box="[189,570,587,609]" page="182" pageId="18" pageNumber="135" year="1986">
<taxonomicName id="4C0E4D2FFFD3E520509885446131FDD4" ID-CoL="5DDDV" authority="PACE, 1986 c: 182" authorityName="Pace" authorityPageNumber="182" authorityYear="1986" box="[189,570,587,609]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="morvani" subGenus="Zyras">
<emphasis id="B97AEABEFFD3E5205098854463FAFDD5" box="[189,241,588,609]" italics="true" pageId="18" pageNumber="135">Zyras</emphasis>
(
<emphasis id="B97AEABEFFD3E52050DA85446238FDD5" box="[255,307,588,609]" italics="true" pageId="18" pageNumber="135">Zyras</emphasis>
)
<emphasis id="B97AEABEFFD3E52051648544629AFDD5" box="[321,401,588,609]" italics="true" pageId="18" pageNumber="135">morvani</emphasis>
<bibRefCitation id="EF9F4B5DFFD3E52051B385436131FDD4" author="PACE, R." box="[406,570,587,609]" pageId="18" pageNumber="135" pagination="165 - 191" refId="ref58188" refString="PACE, R. 1986 c: Aleocharinae dell'Himalaya raccolte da Guillaume de Rougemont (Coleoptera Staphylinidae). - Bollettino del Museo Civico di Storia Naturale di Verona 12: 165 - 191." type="journal article" year="1986">PACE, 1986c: 182</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD3E52050988581627CFC95" pageId="18" pageNumber="135" type="materials_examined">
<paragraph id="8BB136ACFFD3E52050988581627CFC95" blockId="18.[189,794,649,801]" pageId="18" pageNumber="135">
<emphasis id="B97AEABEFFD3E5205098858162DAFD15" bold="true" box="[189,465,649,673]" pageId="18" pageNumber="135">Type material examined</emphasis>
:
<materialsCitation id="3B663CF1FFD3E52051C68581627CFC95" ID-GBIF-Occurrence="3450162474" collectingDate="1973-04" collectionCode="MCSNV" collectorName="P. Morvan" determinerName="V. Assing" elevation="1750" location="Takumsibang" municipality="Takumsibang" pageId="18" pageNumber="135" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B5880EFFD3E52051C68581614DFD15" box="[483,582,649,673]" pageId="18" pageNumber="135" type="holotype">Holotype</typeStatus>
<specimenCount id="9D08FD25FFD3E52052778582616CFD15" box="[594,615,650,673]" pageId="18" pageNumber="135" type="generic">♂</specimenCount>
: “
<collectingMunicipality id="6BD5ACD6FFD3E52052A78581601DFD15" box="[642,790,649,673]" pageId="18" pageNumber="135">Takumsibang</collectingMunicipality>
,
<quantity id="4CF69B49FFD3E520509885A16205FD75" box="[189,270,681,705]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="18" pageNumber="135" unit="m" value="1750.0">
<elevation id="0023D19FFFD3E520509885A16205FD75" box="[189,270,681,705]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="18" pageNumber="135" unit="m" value="1750.0">1750 m</elevation>
</quantity>
.
<date id="FFB0106CFFD3E520513C85A16259FD75" box="[281,338,681,705]" pageId="18" pageNumber="135" value="1973-04">
<collectingDate id="EFF4E984FFD3E520513C85A16259FD75" box="[281,338,681,705]" pageId="18" pageNumber="135" value="1973-04">IV.73</collectingDate>
</date>
,
<collectorName id="26FB537AFFD3E520517885A262C2FD75" box="[349,457,682,705]" pageId="18" pageNumber="135">P. Morvan</collectorName>
/ Coll. Rougemont / Holotypus
<taxonomicName id="4C0E4D2FFFD3E520509885C2625FFD55" authorityName="Pace" authorityYear="1986" box="[189,340,714,737]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="morvani">
<emphasis id="B97AEABEFFD3E520509885C2625FFD55" box="[189,340,714,737]" italics="true" pageId="18" pageNumber="135">Zyras morvani</emphasis>
</taxonomicName>
m., det.
<originalDetermination id="A8108E50FFD3E520518E85C262FCFD55" box="[427,503,714,737]" pageId="18" pageNumber="135">R. Pace</originalDetermination>
1983 /
<taxonomicName id="4C0E4D2FFFD3E520526485C261D3FD55" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[577,728,714,737]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="morvani" status="sp. nov.">
<emphasis id="B97AEABEFFD3E520526485C261D3FD55" box="[577,728,714,737]" italics="true" pageId="18" pageNumber="135">Zyras morvani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFD3E52052F985C2601EFD55" box="[732,789,714,737]" pageId="18" pageNumber="135" rank="species">n. sp.</taxonomicNameLabel>
, det. R. Pace, 1983 /
<taxonomicName id="4C0E4D2FFFD3E52051B885E2617CFCB5" authority="Pace" authorityName="Pace" authorityYear="1986" box="[413,631,746,769]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="morvani">
<emphasis id="B97AEABEFFD3E52051B885E26131FCB5" box="[413,570,746,769]" italics="true" pageId="18" pageNumber="135">Zyras morvani</emphasis>
Pace
</taxonomicName>
, det.
<determinerName id="7E6DDA68FFD3E520529285E26012FCB5" box="[695,793,745,769]" pageId="18" pageNumber="135">V. Assing</determinerName>
2016” (
<collectionCode id="ED1FAE69FFD3E520512D84016260FC95" box="[264,363,777,801]" collectionName="MCSNV" pageId="18" pageNumber="135">MCSNV</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD3E52050988441626DFBB5" pageId="18" pageNumber="135" type="discussion">
<paragraph id="8BB136ACFFD3E5205098844161C8FC35" blockId="18.[189,794,841,897]" pageId="18" pageNumber="135">
<emphasis id="B97AEABEFFD3E520509884416227FCD5" bold="true" box="[189,300,841,865]" pageId="18" pageNumber="135">Comment</emphasis>
: The original description is based on a unique male from “
<collectingCountry id="F319763CFFD3E520511E84616276FC35" box="[315,381,873,897]" name="Nepal" pageId="18" pageNumber="135">Nepal</collectingCountry>
, Takum Sibang” (
<bibRefCitation id="EF9F4B5DFFD3E520521C846161B3FC35" author="PACE, R." box="[569,696,873,897]" pageId="18" pageNumber="135" pagination="165 - 191" refId="ref58188" refString="PACE, R. 1986 c: Aleocharinae dell'Himalaya raccolte da Guillaume de Rougemont (Coleoptera Staphylinidae). - Bollettino del Museo Civico di Storia Naturale di Verona 12: 165 - 191." type="journal article" year="1986">PACE 1986c</bibRefCitation>
).
</paragraph>
<paragraph id="8BB136ACFFD3E520509884A1626DFBB5" blockId="18.[189,794,937,1025]" pageId="18" pageNumber="135">
<emphasis id="B97AEABEFFD3E520509884A16102FC75" bold="true" box="[189,521,937,961]" pageId="18" pageNumber="135">Additional material examined</emphasis>
:
<materialsCitation id="3B663CF1FFD3E520523284A16268FBB5" ID-GBIF-Occurrence="3450162427" collectingDate="1993-05-30" collectorName="Schmidt" country="Nepal" county="Syangde" elevation="1100" location="Annapurna" municipality="Marsyangdi valley" pageId="18" pageNumber="135" specimenCount="1">
<emphasis id="B97AEABEFFD3E520523284A16153FC75" bold="true" box="[535,600,937,961]" pageId="18" pageNumber="135">
<collectingCountry id="F319763CFFD3E520523284A16153FC75" box="[535,600,937,961]" name="Nepal" pageId="18" pageNumber="135">Nepal</collectingCountry>
</emphasis>
: 1 ♀,
<location id="8ED16077FFD3E52052BC84A1601DFC75" LSID="urn:lsid:plazi:treatment:03A787BAFFD3E52050C686FB670BF815:8ED16077FFD3E52052BC84A1601DFC75" box="[665,790,937,961]" country="Nepal" county="Syangde" municipality="Marsyangdi valley" name="Annapurna" pageId="18" pageNumber="135">Annapurna</location>
,
<collectingMunicipality id="6BD5ACD6FFD3E520509884C1628BFC55" box="[189,384,969,993]" pageId="18" pageNumber="135">Marsyangdi valley</collectingMunicipality>
,
<collectingCounty id="62D04E20FFD3E52051A984C162EFFC55" box="[396,484,969,993]" pageId="18" pageNumber="135">Syangde</collectingCounty>
env.,
<quantity id="4CF69B49FFD3E520520684C1617FFC55" box="[547,628,969,993]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="135" unit="m" value="1100.0">
<elevation id="0023D19FFFD3E520520684C1617FFC55" box="[547,628,969,993]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="135" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date id="FFB0106CFFD3E520525A84C161E1FC55" box="[639,746,969,993]" pageId="18" pageNumber="135" value="1993-05-30">
<collectingDate id="EFF4E984FFD3E520525A84C161E1FC55" box="[639,746,969,993]" pageId="18" pageNumber="135" value="1993-05-30">30.V.1993</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFD3E520509884E1621DFBB5" box="[189,278,1001,1025]" pageId="18" pageNumber="135">Schmidt</collectorName>
(cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD3E5205098832160A1F955" pageId="18" pageNumber="135" type="description">
<paragraph id="8BB136ACFFD3E520509883216221FA35" blockId="18.[189,794,1065,2049]" pageId="18" pageNumber="135">
<emphasis id="B97AEABEFFD3E520509883216251FBF5" bold="true" box="[189,346,1065,1089]" pageId="18" pageNumber="135">Redescription</emphasis>
: Body length 5.6–6.0 mm; length of forebody
<quantity id="4CF69B49FFD3E52051258341628EFBD5" box="[256,389,1097,1121]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="18" pageNumber="135" unit="mm" value="2.65" valueMax="2.7" valueMin="2.6">2.6–2.7 mm</quantity>
. Coloration (
<figureCitation id="13352A29FFD3E52052058341617AFBD5" box="[544,625,1097,1121]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="18" pageNumber="135">Figs 25</figureCitation>
,
<figureCitation id="13352A29FFD3E52052A783416195FBD5" box="[642,670,1097,1121]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="18" pageNumber="135">51</figureCitation>
,
<figureCitation id="13352A29FFD3E520528A834161C6FBD5" box="[687,717,1097,1121]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="18" pageNumber="135">93</figureCitation>
): head black; pronotum and elytra dark-brown to blackishbrown; abdomen dark-brown to blackish-brown with the anterior portions of the paratergites paler; legs yellow with the apices of all femora blackish; antennae with the basal 3–4 antennomeres dark-brown to blackish-brown and the remainder gradually becoming paler towards apex, apical 3–5 antennomers dark-yellowish; maxillary palpi brown to dark-brown with palpomere IV yellowish.
</paragraph>
<paragraph id="8BB136ACFFD3E5205098828162F2F955" blockId="18.[189,794,1065,2049]" pageId="18" pageNumber="135">
Head (
<figureCitation id="13352A29FFD3E520512F8281625EFA15" box="[266,341,1417,1441]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="18" pageNumber="135">Fig. 51</figureCitation>
) moderately transverse, median portion extensively impunctate; punctures in lateral portions sparse and fine. Eyes large and bulging, more than twice as long as distance from posterior margin of eye to posterior constriction of head in dorsal view. Antenna (
<figureCitation id="13352A29FFD3E52052E281016019F995" box="[711,786,1545,1569]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="18" pageNumber="135">Fig. 25</figureCitation>
) approximately 2.0 mm long and moderately slender; antennomeres IV weakly oblong, V approximately as broad as long, VI–X of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long or nearly so, and XI barely as long as the combined length of IX and X.
</paragraph>
<paragraph id="8BB136ACFFD3E520509881E262C3F7B5" blockId="18.[189,794,1065,2049]" pageId="18" pageNumber="135">
Pronotum (
<figureCitation id="13352A29FFD3E520511B81E16282F8B5" box="[318,393,1769,1793]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="18" pageNumber="135">Fig. 51</figureCitation>
) weakly transverse, 1.03–1.07 times as broad as long and 1.13–1.15 times as broad as head, broadest in anterior half, distinctly tapering posteriad; lateral margins distinctly sinuate in posterior half in dorsal view; punctation fine and slightly irregularly distributed; lateral margins each with five long and erect black setae, anterior margins with an additional long and erect black seta on either side; pubescence of disc rather long, sub-erect, and pale.
</paragraph>
<paragraph id="8BB136ACFFD3E520531A87E1663AFE15" blockId="18.[831,1437,233,1281]" pageId="18" pageNumber="135">
Elytra (
<figureCitation id="13352A29FFD3E52053BE87E160ECFEB5" box="[923,999,233,257]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="18" pageNumber="135">Fig. 51</figureCitation>
) 0.78–0.82 times as long as pronotum; punctation fine, moderately dense, and regularly distributed; whole disc with dense, long, and sub-erect pale pubescence, laterally with long, stout, and erect setae. Hind wings fully developed. Metatarsomere I longer than the combined length of II and III.
</paragraph>
<paragraph id="8BB136ACFFD3E520531A86A160A4FBB5" blockId="18.[831,1437,233,1281]" pageId="18" pageNumber="135">
Abdomen (
<figureCitation id="13352A29FFD3E520539886A16701FE75" box="[957,1034,425,449]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="18" pageNumber="135">Fig. 93</figureCitation>
) slightly narrower than elytra, with rather deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of numerous rather weakly defined grooves; tergite III with a lateral setiferous puncture on either side and with numerous fine setiferous punctures at posterior margin; tergites IV and V with a median pair of setiferous punctures, with a lateral setiferous puncture on either side, and with numerous fine setiferous punctures at posterior margin; tergite VI with a transverse row of non-setiferous punctures anteriorly, with a transverse row of rather numerous setiferous punctures at posterior third, and with numerous setiferous punctures at posterior margin; tergite VII with a transverse band of non-setiferous punctures anteriorly and with two transverse rows of setiferous punctures posteriorly, posterior margin with palisade fringe; posterior margin of tergite VIII with a shallow median concavity (
<figureCitation id="13352A29FFD3E520536284E160A8FBB5" box="[839,931,1001,1025]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="18" pageNumber="135">Fig. 148</figureCitation>
).
</paragraph>
<paragraph id="8BB136ACFFD3E520531A8302608BFB75" blockId="18.[831,1437,233,1281]" pageId="18" pageNumber="135">
♂: sternite VIII (
<figureCitation id="13352A29FFD3E520543183016765FB95" box="[1044,1134,1033,1057]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="18" pageNumber="135">Fig. 149</figureCitation>
) oblong, strongly convex posteriorly; median lobe of aedeagus approximately
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long and shaped as figured by
<bibRefCitation id="EF9F4B5DFFD3E520552483416693FBD5" author="PACE, R." box="[1281,1432,1097,1121]" pageId="18" pageNumber="135" pagination="165 - 191" refId="ref58188" refString="PACE, R. 1986 c: Aleocharinae dell'Himalaya raccolte da Guillaume de Rougemont (Coleoptera Staphylinidae). - Bollettino del Museo Civico di Storia Naturale di Verona 12: 165 - 191." type="journal article" year="1986">PACE (1986c)</bibRefCitation>
; paramere slightly longer than median lobe, apical lobe not distinctly modified, moderately long and apically acute.
</paragraph>
<paragraph id="8BB136ACFFD3E520531A83C26711FAB5" blockId="18.[831,1437,233,1281]" pageId="18" pageNumber="135">♀: posterior margin of sternite VIII convex, in the middle indistinctly concave.</paragraph>
<paragraph id="8BB136ACFFD3E520531A822160A1F955" blockId="18.[831,1436,1321,1761]" pageId="18" pageNumber="135">
<emphasis id="B97AEABEFFD3E520531A8221671FFAF5" bold="true" box="[831,1044,1321,1345]" pageId="18" pageNumber="135">Comparative notes</emphasis>
: Though similar in size and habitus to
<taxonomicName id="4C0E4D2FFFD3E5205347824260E0FAD5" authorityName="Last" authorityYear="1969" box="[866,1003,1353,1377]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="condignus">
<emphasis id="B97AEABEFFD3E5205347824260E0FAD5" box="[866,1003,1353,1377]" italics="true" pageId="18" pageNumber="135">Z. condignus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0E4D2FFFD3E52053DA8242677CFAD5" authorityName="Pace" authorityYear="1986" box="[1023,1143,1354,1377]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="morvani">
<emphasis id="B97AEABEFFD3E52053DA8242677CFAD5" box="[1023,1143,1354,1377]" italics="true" pageId="18" pageNumber="135">Z. morvani</emphasis>
</taxonomicName>
differs from this species in numerous characters, such as much shorter and less slender antennae with more transverse antennomeres VI–X, the coloration of the antennae and the legs, much larger eyes, a more slender pronotum with sinuate lateral margins and a smooth surface, finer and practically regular punctation of the pronotum and the elytra, a different punctation pattern of the abdomen (particularly the much more numerous setiferous punctures at the posterior margins of the tergites), a completely different shape of the male sternite VIII, and the shapes of the median lobe and the parameres of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD3E520531A8001670BF815" pageId="18" pageNumber="135" type="distribution">
<paragraph id="8BB136ACFFD3E520531A8001670BF815" blockId="18.[831,1436,1801,1953]" pageId="18" pageNumber="135">
<emphasis id="B97AEABEFFD3E520531A800167A2F895" bold="true" box="[831,1193,1801,1825]" pageId="18" pageNumber="135">Distribution and natural history</emphasis>
: The currently known distribution is confined to two localities in the
<collectingRegion id="49CAF84EFFD3E52055668021606EF8D5" country="Nepal" name="Dhawalagiri" pageId="18" pageNumber="135">Dhaulagiri</collectingRegion>
and Annapurna ranges,
<collectingCountry id="F319763CFFD3E5205458804167B6F8D5" box="[1149,1213,1865,1889]" name="Nepal" pageId="18" pageNumber="135">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFD3E52054EA8041661DF8D5" box="[1231,1302,1865,1889]" captionStart="Map 3" captionStartId="64.[189,233,2058,2079]" captionTargetBox="[189,1436,1492,2047]" captionTargetId="figure-120@64.[189,1436,1492,2047]" captionTargetPageId="64" captionText="Map 3: Revised distributionsof Zyras perforatus (blackcircles),Z. morvani(white circles),andZ. truncatus(white star)in theHimalaya." figureDoi="http://doi.org/10.5281/zenodo.5742399" httpUri="https://zenodo.org/record/5742399/files/figure.png" pageId="18" pageNumber="135">Map 3</figureCitation>
), where the two known specimens were collected at altitudes of 1100 and
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.
</paragraph>
</subSubSection>
</treatment>
</document>
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