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<document id="0D6392603A1891C0E84B265A332E3E63" ID-CLB-Dataset="7383" ID-DOI="10.11646/zootaxa.4951.3.7" ID-GBIF-Dataset="a6ee038f-4cda-4f38-8fde-8ad0efec8f0f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4668453" ID-ZooBank="EB876346-4282-4B4E-8585-ED50F1A616E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617782094822" checkinUser="plazi" docAuthor="Martens, Jochen, Maghradze, Eter &amp; Barjadze, Shalva" docDate="2021" docId="03A787A4FFF8FFDE3584F6521F1DF8E9" docLanguage="en" docName="zootaxa.4951.3.7.pdf" docOrigin="Zootaxa 4951 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nemaspela prometheus Martens &amp; Maghradze &amp; Barjadze 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="549" masterDocId="FF9EFFDCFFFFFFD63513F30F1D1AFFA4" masterDocTitle="Two new species of the genus Nemaspela Šilhavý from caves in Georgia (Opiliones Nemastomatidae)" masterLastPageNumber="558" masterPageNumber="541" pageNumber="548" updateTime="1698936134096" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="DD4F4BEF8014BF90AEDF3F720067ACDD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DE5A981CE2D52B1301C3534034191233">
<mods:title id="6EBC0F5D6B73EAE309030CDB7B1A69C6">Two new species of the genus Nemaspela Šilhavý from caves in Georgia (Opiliones Nemastomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="D2B46E6FD3B9C7EF3FF87C741AFDF435" type="personal">
<mods:role id="C59C0B56BFCA11A6D588E7809FDE482E">
<mods:roleTerm id="A4888792343055C9058499F9201414CF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2E690BEC0C69D6CADDCD472AC15CCFD0">Martens, Jochen</mods:namePart>
<mods:affiliation id="8D1EE043DF5B2C31CD4132982893DDD7">Johannes Gutenberg-Universität, Institut für Organismische und Molekulare Evolutionsbiologie (iomE), D- 55099 Mainz, Germany; Senckenberg Research Institute, Arachnology, D- 60325 Frankfurt am Main, Germany</mods:affiliation>
</mods:name>
<mods:name id="B9D5402F503FD07F6D7B0BF4E22D5FC3" type="personal">
<mods:role id="56AEE36543DC806E33E423319EDDB3C6">
<mods:roleTerm id="C688CF3DD337379C19BD5C295538E593">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A358ACBB1E0146EDD6021368F642BD01">Maghradze, Eter</mods:namePart>
<mods:affiliation id="B28E0CF6506C7947F75FF80A681CD6E8">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia</mods:affiliation>
</mods:name>
<mods:name id="1E4CBBD1493C7B640792B8C0E75B3817" type="personal">
<mods:role id="09817F4F4E2BC3DFE4ED4473C4B216A6">
<mods:roleTerm id="8C2C680125F808C3A9E1490DB1F42A08">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C459254DC0CBE830CDBC8D6576A4AE99">Barjadze, Shalva</mods:namePart>
<mods:affiliation id="EF99A69D50B1C9E9A4FBBD56F18D6D95">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="7001B0A4555D27E40AED4178C21986A6">text</mods:typeOfResource>
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<mods:titleInfo id="4F90A2F7012ACA3F4765272D386C8274">
<mods:title id="288D725DDAAFBC311FFC91D7AD450C18">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CF417FC1B30990198BAFCD22B807AEAB">
<mods:date id="816189920638E7AFEAA1C7516B799527">2021</mods:date>
<mods:detail id="DD1C83DD4388602056E2B633444066A7" type="pubDate">
<mods:number id="7CEDA2B10FD9FA6FDE8853C21DE1A1B3">2021-04-07</mods:number>
</mods:detail>
<mods:detail id="FC4F93B600E9C45EE96B0273E2589BBA" type="volume">
<mods:number id="398EC674C80E82345954EEC7B636582D">4951</mods:number>
</mods:detail>
<mods:detail id="2843511473F011FB26D99613E889A6E9" type="issue">
<mods:number id="AF53358F0A1D00F098557491DB067DDE">3</mods:number>
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<mods:extent id="DC5C899A3A8E5453B390FF286C264E35" unit="page">
<mods:start id="2DFF49D25886D6D4361C77802A40294F">541</mods:start>
<mods:end id="D5736ED91551543F337B9B55BBCFE665">558</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="973A317BB2C90D257382AA1B76581DB1">journal article</mods:classification>
<mods:identifier id="6202491184B954AECEA9BE5D12BFBA50" type="CLB-Dataset">7383</mods:identifier>
<mods:identifier id="C9F455A3A179BACD11200277CAF21853" type="DOI">10.11646/zootaxa.4951.3.7</mods:identifier>
<mods:identifier id="5605959493565D7A60E8DA91F22920B2" type="GBIF-Dataset">a6ee038f-4cda-4f38-8fde-8ad0efec8f0f</mods:identifier>
<mods:identifier id="F4067CD2B5AAFBBABE44B7F646359910" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F78EC1CBA9BA2D89F948EC2DCDA5E980" type="Zenodo-Dep">4668453</mods:identifier>
<mods:identifier id="BEFAFAB6F8AEB3FC62507C76139EEEC8" type="ZooBank">EB876346-4282-4B4E-8585-ED50F1A616E4</mods:identifier>
</mods:mods>
<treatment id="03A787A4FFF8FFDE3584F6521F1DF8E9" ID-DOI="http://doi.org/10.5281/zenodo.4668477" ID-GBIF-Taxon="181003515" ID-Zenodo-Dep="4668477" LSID="urn:lsid:plazi:treatment:03A787A4FFF8FFDE3584F6521F1DF8E9" httpUri="http://treatment.plazi.org/id/03A787A4FFF8FFDE3584F6521F1DF8E9" lastPageId="8" lastPageNumber="549" pageId="7" pageNumber="548">
<subSubSection id="C3146539FFF8FFD13584F6521F0DFAD3" box="[151,535,1373,1400]" pageId="7" pageNumber="548" type="nomenclature">
<paragraph id="8BB136B2FFF8FFD13584F6521F0DFAD3" blockId="7.[151,535,1373,1435]" box="[151,535,1373,1400]" pageId="7" pageNumber="548">
<heading id="D0F981DEFFF8FFD13584F6521F0DFAD3" bold="true" box="[151,535,1373,1400]" fontSize="11" level="1" pageId="7" pageNumber="548" reason="1">
<emphasis id="B97AEAA0FFF8FFD13584F6521F0DFAD3" bold="true" box="[151,535,1373,1400]" pageId="7" pageNumber="548">
<taxonomicName id="4C0E4D31FFF8FFD13584F6521CA8FADC" ID-CoL="9X5JB" authority="Martens &amp; Maghradze &amp; Barjadze, 2021" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2021" box="[151,434,1373,1400]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="548" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B97AEAA0FFF8FFD13584F6521CA8FADC" bold="true" box="[151,434,1373,1400]" italics="true" pageId="7" pageNumber="548">Nemaspela prometheus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957DBFFF8FFD134AAF6521F0DFAD3" box="[441,535,1373,1399]" pageId="7" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFF8FFD13584F68E1C92FA3F" box="[151,392,1409,1435]" pageId="7" pageNumber="548" type="description">
<paragraph id="8BB136B2FFF8FFD13584F68E1C92FA3F" blockId="7.[151,535,1373,1435]" box="[151,392,1409,1435]" pageId="7" pageNumber="548">
(
<figureCitation id="13352A37FFF8FFD1358CF68E1C3BFA3F" box="[159,289,1409,1435]" captionStart-0="FIGURES 19–21" captionStart-1="FIGURES 22–29" captionStart-2="FIGURES 30–32" captionStartId-0="9.[151,264,1943,1967]" captionStartId-1="10.[151,264,1943,1967]" captionStartId-2="12.[151,264,1306,1330]" captionTargetBox-0="[263,1326,181,1919]" captionTargetBox-1="[193,1385,211,1919]" captionTargetBox-2="[165,1415,181,1275]" captionTargetId-0="figure-17@9.[259,1327,181,1919]" captionTargetId-1="figure-23@10.[176,1412,181,1919]" captionTargetId-2="figure-23@12.[151,1436,181,1282]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="FIGURES 19–21. Nemaspela prometheus sp. nov. Habitus in lateral view. 19. Female (bar: 1 mm); 20. Female (bar: 2 mm); 21. Male (bar: 2 mm). Photographs by J. Severin." captionText-1="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." captionText-2="FIGURES 30–32. Nemaspela prometheus sp. nov. Chelicerae in retro-lateral view. 30. Male (bar: 0.8 mm); 31. Female (bar: 0.8 mm). Chelicera in pro-lateral view. 32. Male (bar: 0.8 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.4668467" figureDoi-1="http://doi.org/10.5281/zenodo.4668469" figureDoi-2="http://doi.org/10.5281/zenodo.4668471" httpUri-0="https://zenodo.org/record/4668467/files/figure.png" httpUri-1="https://zenodo.org/record/4668469/files/figure.png" httpUri-2="https://zenodo.org/record/4668471/files/figure.png" pageId="7" pageNumber="548">Figs 19–32</figureCitation>
,
<tableCitation id="C68C0309FFF8FFD13438F68E1C65FA3F" box="[299,383,1409,1435]" captionStart="TABLE 2" captionStartId="11.[151,239,151,175]" captionTargetPageId="11" captionText="TABLE 2. Measurements of Nemaspela prometheus sp. nov. Measurements are given in millimeters. Note: body length is measured in all available specimens, while length of appendages was measured only in holotype male and single female." httpUri="http://table.plazi.org/id/DF71663AFFF4FFDD3584F3981DC8FF53" pageId="7" pageNumber="548" tableUuid="DF71663AFFF4FFDD3584F3981DC8FF53">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFF8FFD13584F6C61941F9F1" pageId="7" pageNumber="548" type="materials_examined">
<paragraph id="8BB136B2FFF8FFD13584F6C619C6F9A3" blockId="7.[151,1437,1481,2017]" pageId="7" pageNumber="548">
<emphasis id="B97AEAA0FFF8FFD13584F6C61C5DFA47" bold="true" box="[151,327,1481,1507]" pageId="7" pageNumber="548">Type material:</emphasis>
<materialsCitation id="3B663CEFFFF8FFD1345EF6C619C6F9A3" ID-GBIF-Occurrence="3071286302" collectingDate="2019-12-27" collectionCode="IZISU" country="Georgia" county="Western" elevation="174" latitude="42.377167" location="Prometheus Cave" longLatPrecision="1" longitude="42.60086" municipality="Sataplia-Tskaltubo" pageId="7" pageNumber="548" specimenCount="1" specimenCount-male="1" stateProvince="Imereti" typeStatus="holotype">
<emphasis id="B97AEAA0FFF8FFD1345EF6C61CD9FA47" bold="true" box="[333,451,1481,1507]" pageId="7" pageNumber="548">
<typeStatus id="54B58810FFF8FFD1345EF6C61CA4FA47" box="[333,446,1481,1507]" pageId="7" pageNumber="548" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D08FD3BFFF8FFD134D9F6C51CC5FA40" box="[458,479,1482,1508]" pageId="7" pageNumber="548" type="male">♂</specimenCount>
,
<date id="FFB01072FFF8FFD134FFF6C61F75FA47" box="[492,623,1481,1507]" pageId="7" pageNumber="548" value="2019-12-27">
<collectingDate id="EFF4E99AFFF8FFD134FFF6C61F75FA47" box="[492,623,1481,1507]" pageId="7" pageNumber="548" value="2019-12-27">27.xii.2019</collectingDate>
</date>
, dark zone,
<location id="8ED16069FFF8FFD137E7F6C61EA2FA47" LSID="urn:lsid:plazi:treatment:03A787A4FFF8FFDE3584F6521F1DF8E9:8ED16069FFF8FFD137E7F6C61EA2FA47" box="[756,952,1481,1507]" country="Georgia" county="Western" latitude="42.377167" longLatPrecision="1" longitude="42.60086" municipality="Sataplia-Tskaltubo" name="Prometheus Cave" pageId="7" pageNumber="548" stateProvince="Imereti">Prometheus Cave</location>
(
<geoCoordinate id="EE3A5075FFF8FFD136D5F6C61972FA47" box="[966,1128,1481,1507]" degrees="42" direction="north" minutes="22" orientation="latitude" pageId="7" pageNumber="548" precision="1" seconds="37.8" value="42.377167">42°22’37.8”N</geoCoordinate>
,
<geoCoordinate id="EE3A5075FFF8FFD13160F6C6180BFA47" box="[1139,1297,1481,1507]" degrees="42" direction="east" minutes="36" orientation="longitude" pageId="7" pageNumber="548" precision="1" seconds="03.1" value="42.60086">42°36’03.1”E</geoCoordinate>
;
<quantity id="4CF69B57FFF8FFD1300FF6C61879FA40" box="[1308,1379,1481,1508]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="7" pageNumber="548" unit="m" value="174.0">
<elevation id="0023D181FFF8FFD1300FF6C61879FA40" box="[1308,1379,1481,1508]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="7" pageNumber="548" unit="m" value="174.0">174 m</elevation>
</quantity>
alt.),
<location id="8ED16069FFF8FFD13584F6E21C84F9A3" LSID="urn:lsid:plazi:treatment:03A787A4FFF8FFDE3584F6521F1DF8E9:8ED16069FFF8FFD13584F6E21C84F9A3" box="[151,414,1517,1543]" country="Georgia" county="Western" latitude="42.377167" longLatPrecision="1" longitude="42.60086" municipality="Sataplia-Tskaltubo" name="Tskaltubo Municipality" pageId="7" pageNumber="548" stateProvince="Imereti">Tskaltubo Municipality</location>
,
<collectingMunicipality id="6BD5ACC8FFF8FFD134BAF6E21F61F9A3" box="[425,635,1517,1543]" pageId="7" pageNumber="548">Sataplia-Tskaltubo</collectingMunicipality>
karst massif,
<collectingRegion id="49CAF850FFF8FFD13605F6E21EA2F9A3" box="[790,952,1517,1543]" country="Georgia" name="Imereti" pageId="7" pageNumber="548">Imereti region</collectingRegion>
,
<collectingCounty id="62D04E3EFFF8FFD136D0F6E11904F9AC" box="[963,1054,1518,1544]" pageId="7" pageNumber="548">Western</collectingCounty>
Georgia,
<collectionCode id="ED1FAE77FFF8FFD1319EF6E219C2F9A3" box="[1165,1240,1517,1543]" pageId="7" pageNumber="548">IZISU</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB136B2FFF8FFD135D4F5181941F9F1" blockId="7.[151,1437,1481,2017]" pageId="7" pageNumber="548">
<materialsCitation id="3B663CEFFFF8FFD135D4F5181F8FF995" ID-GBIF-Occurrence="3071286304" box="[199,661,1559,1585]" collectingDate="2019-12-27" collectionCode="IZISU" country="Georgia" county="Western" elevation="174" latitude="42.377167" location="The" longLatPrecision="1" longitude="42.60086" municipality="Sataplia-Tskaltubo" pageId="7" pageNumber="548" specimenCount="1" stateProvince="Imereti" typeStatus="paratype">
<emphasis id="B97AEAA0FFF8FFD135D4F5181C25F995" bold="true" box="[199,319,1559,1585]" pageId="7" pageNumber="548">
<typeStatus id="54B58810FFF8FFD135D4F5181C25F995" box="[199,319,1559,1585]" pageId="7" pageNumber="548" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<location id="8ED16069FFF8FFD13458F5181C6DF995" LSID="urn:lsid:plazi:treatment:03A787A4FFF8FFDE3584F6521F1DF8E9:8ED16069FFF8FFD13458F5181C6DF995" box="[331,375,1559,1585]" name="The" pageId="7" pageNumber="548">The</location>
same locality as holotype
</materialsCitation>
.
<specimenCount id="9D08FD3BFFF8FFD1378EF5181FDFF995" box="[669,709,1559,1585]" pageId="7" pageNumber="548" type="male">1 ♂</specimenCount>
,
<date id="FFB01072FFF8FFD137C2F5181E4EF995" box="[721,852,1559,1585]" pageId="7" pageNumber="548" value="2019-12-27">27.xii.2019</date>
, CJM 8707;
<materialsCitation id="3B663CEFFFF8FFD136CEF51819C0F995" ID-GBIF-Occurrence="3071286306" box="[989,1242,1559,1585]" collectingDate="2020-02-02" collectionCode="ISIZU" country="Georgia" pageId="7" pageNumber="548" specimenCount="1" specimenCount-male="1" stateProvince="Imereti" typeStatus="paratype">
<specimenCount id="9D08FD3BFFF8FFD136CEF518191FF995" box="[989,1029,1559,1585]" pageId="7" pageNumber="548" type="male">1 ♂</specimenCount>
,
<date id="FFB01072FFF8FFD13102F518199FF995" box="[1041,1157,1559,1585]" pageId="7" pageNumber="548" value="2020-02-02">
<collectingDate id="EFF4E99AFFF8FFD13102F518199FF995" box="[1041,1157,1559,1585]" pageId="7" pageNumber="548" value="2020-02-02">02.ii.2020</collectingDate>
</date>
,
<collectionCode id="ED1FAE77FFF8FFD1319CF51819C0F995" box="[1167,1242,1559,1585]" pageId="7" pageNumber="548">ISIZU</collectionCode>
</materialsCitation>
;
<specimenCount id="9D08FD3BFFF8FFD131F0F518180AF997" box="[1251,1296,1556,1587]" pageId="7" pageNumber="548" type="female">1 ♀</specimenCount>
,
<date id="FFB01072FFF8FFD1300FF5181882F995" box="[1308,1432,1559,1585]" pageId="7" pageNumber="548" value="2020-03-01">01.iii.2020</date>
, CJM 8708;
<materialsCitation id="3B663CEFFFF8FFD13408F5341941F9F1" ID-GBIF-Occurrence="3071286305" box="[283,1115,1595,1621]" collectingDate="2020-06-29" collectionCode="ISIZU" collectorName="All &amp; Eter Maghradze." country="Georgia" pageId="7" pageNumber="548" specimenCount="1" specimenCount-male="1" stateProvince="Imereti" typeStatus="paratype">
<specimenCount id="9D08FD3BFFF8FFD13408F5341C5FF9F1" box="[283,325,1595,1621]" pageId="7" pageNumber="548" type="male">1 ♂</specimenCount>
,
<date id="FFB01072FFF8FFD13440F5341CD7F9F1" box="[339,461,1595,1621]" pageId="7" pageNumber="548" value="2020-06-29">
<collectingDate id="EFF4E99AFFF8FFD13440F5341CD7F9F1" box="[339,461,1595,1621]" pageId="7" pageNumber="548" value="2020-06-29">29.vi.2020</collectingDate>
</date>
,
<collectionCode id="ED1FAE77FFF8FFD134CAF5341F3EF9F1" box="[473,548,1595,1621]" pageId="7" pageNumber="548">ISIZU</collectionCode>
.
<collectorName id="26FB5364FFF8FFD1373EF5341F4BF9F1" box="[557,593,1595,1621]" pageId="7" pageNumber="548">All</collectorName>
specimens were collected by
<collectorName id="26FB5364FFF8FFD136B3F5341941F9F1" box="[928,1115,1595,1621]" pageId="7" pageNumber="548">Eter Maghradze.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFF8FFD135D4F5501F63F88A" pageId="7" pageNumber="548" type="diagnosis">
<paragraph id="8BB136B2FFF8FFD135D4F5501F63F88A" blockId="7.[151,1437,1481,2017]" pageId="7" pageNumber="548">
<emphasis id="B97AEAA0FFF8FFD135D4F5501C58F9DD" bold="true" box="[199,322,1631,1657]" pageId="7" pageNumber="548">Diagnosis.</emphasis>
Generally similar in size and habitus to the other
<taxonomicName id="4C0E4D31FFF8FFD1366EF5501EE3F9DD" authorityName="Silhavy" authorityYear="1966" box="[893,1017,1631,1657]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEAA0FFF8FFD1366EF5501EE3F9DD" box="[893,1017,1631,1657]" italics="true" pageId="7" pageNumber="548">Nemaspela</emphasis>
</taxonomicName>
species from Georgian caves, i.e.
<taxonomicName id="4C0E4D31FFF8FFD13090F56F1C23F939" authorityName="Martens" authorityYear="2006" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="548" phylum="Arthropoda" rank="species" species="femorecurvata">
<emphasis id="B97AEAA0FFF8FFD13090F56F1C23F939" italics="true" pageId="7" pageNumber="548">N. femorecurvata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E4D31FFF8FFD1347DF58B1CFFF939" authority="Martens &amp; Maghradze &amp; Barjadze, 2021" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2021" box="[366,485,1667,1693]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="548" phylum="Arthropoda" rank="species" species="melouri" status="sp. nov.">
<emphasis id="B97AEAA0FFF8FFD1347DF58B1CFFF939" box="[366,485,1667,1693]" italics="true" pageId="7" pageNumber="548">N. melouri</emphasis>
</taxonomicName>
<emphasis id="B97AEAA0FFF8FFD134F8F58C1F59F939" bold="true" box="[491,579,1667,1693]" pageId="7" pageNumber="548">
<taxonomicNameLabel id="A24957DBFFF8FFD134F8F58C1F59F939" box="[491,579,1667,1693]" pageId="7" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, characterized by specific genital morphology (slenderer form of truncus and smaller size of basal inflated part of truncus, glans penis in lateral view less slender than in
<taxonomicName id="4C0E4D31FFF8FFD131BCF5A71833F965" authority="Martens &amp; Maghradze &amp; Barjadze, 2021" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2021" box="[1199,1321,1703,1729]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="548" phylum="Arthropoda" rank="species" species="melouri" status="sp. nov.">
<emphasis id="B97AEAA0FFF8FFD131BCF5A71833F965" box="[1199,1321,1703,1729]" italics="true" pageId="7" pageNumber="548">N. melouri</emphasis>
</taxonomicName>
<emphasis id="B97AEAA0FFF8FFD13022F5A81896F965" bold="true" box="[1329,1420,1703,1729]" pageId="7" pageNumber="548">
<taxonomicNameLabel id="A24957DBFFF8FFD13022F5A81896F965" box="[1329,1420,1703,1729]" pageId="7" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), small body size below
<quantity id="4CF69B57FFF8FFD1348CF5C41CFBF941" box="[415,481,1739,1765]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="548" unit="mm" value="2.0">2 mm</quantity>
(species slightly larger than
<taxonomicName id="4C0E4D31FFF8FFD1363FF5C31E87F941" box="[812,925,1739,1765]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="548" phylum="Arthropoda" rank="species" species="melour">
<emphasis id="B97AEAA0FFF8FFD1363FF5C31E87F941" box="[812,925,1739,1765]" italics="true" pageId="7" pageNumber="548">N. melour</emphasis>
</taxonomicName>
i
<emphasis id="B97AEAA0FFF8FFD136BDF5C41913F941" bold="true" box="[942,1033,1739,1765]" pageId="7" pageNumber="548">sp. nov.</emphasis>
) and appendages (longer pedipalps and legs) and armament of male chelicera (without apophysis of basal article), distinct sexual dimorphism (slightly longer pedipalps in females than in males).
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFF8FFD135D4F4381C6CF8D1" pageId="7" pageNumber="548" type="etymology">
<paragraph id="8BB136B2FFF8FFD135D4F4381C6CF8D1" blockId="7.[151,1437,1481,2017]" pageId="7" pageNumber="548">
<emphasis id="B97AEAA0FFF8FFD135D4F4381C54F8F5" bold="true" box="[199,334,1847,1873]" pageId="7" pageNumber="548">Etymology.</emphasis>
The name is given in accordance with the
<typeStatus id="54B58810FFF8FFD13625F4381E7CF8F5" box="[822,870,1847,1873]" pageId="7" pageNumber="548">type</typeStatus>
locality, the single known locality of this species. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFF8FFDE35D4F4701F33F8A1" lastPageId="8" lastPageNumber="549" pageId="7" pageNumber="548" type="description">
<paragraph id="8BB136B2FFF8FFDE35D4F4701DFCFF5C" blockId="7.[151,1437,1481,2017]" lastBlockId="8.[151,1437,151,1869]" lastPageId="8" lastPageNumber="549" pageId="7" pageNumber="548">
<emphasis id="B97AEAA0FFF8FFD135D4F4701CF9F83D" bold="true" box="[199,483,1919,1945]" pageId="7" pageNumber="548">Description. Male (n=4)</emphasis>
.
<emphasis id="B97AEAA0FFF8FFD134E3F4701FABF83D" box="[496,689,1919,1945]" italics="true" pageId="7" pageNumber="548">Body, dorsal side</emphasis>
(
<figureCitation id="13352A37FFF8FFD137D3F4701E0CF83D" box="[704,790,1919,1945]" captionStart="FIGURES 19–21" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[263,1326,181,1919]" captionTargetId="figure-17@9.[259,1327,181,1919]" captionTargetPageId="9" captionText="FIGURES 19–21. Nemaspela prometheus sp. nov. Habitus in lateral view. 19. Female (bar: 1 mm); 20. Female (bar: 2 mm); 21. Male (bar: 2 mm). Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668467" httpUri="https://zenodo.org/record/4668467/files/figure.png" pageId="7" pageNumber="548">Figs 19</figureCitation>
,
<figureCitation id="13352A37FFF8FFD13631F4701E58F83D" box="[802,834,1919,1945]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="7" pageNumber="548">28</figureCitation>
): Prosoma and opisthosoma forming two independent strongly globular body parts with deep depression between them; prosoma with a small additional elevation similar to a Tu oc extending from the frontal part of opisthosoma, dorsal scutal parts of prosoma and opisthosoma uniformly light yellowish, smooth, difficult to distinguish from neighbouring exoskeletal parts, without any darker spots or markings, no setae present on dorsal side except for few minute ones on rear end of opisthosoma, Tu oc and eyes absent.
</paragraph>
<paragraph id="8BB136B2FFF7FFDE35D4F20C189DFEC1" blockId="8.[151,1437,151,1869]" pageId="8" pageNumber="549">
<emphasis id="B97AEAA0FFF7FFDE35D4F20C1C88FEB9" box="[199,402,259,285]" italics="true" pageId="8" pageNumber="549">Body, ventral side</emphasis>
(
<figureCitation id="13352A37FFF7FFDE34B2F20C1CE3FEB9" box="[417,505,259,285]" captionStart="FIGURES 19–21" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[263,1326,181,1919]" captionTargetId="figure-17@9.[259,1327,181,1919]" captionTargetPageId="9" captionText="FIGURES 19–21. Nemaspela prometheus sp. nov. Habitus in lateral view. 19. Female (bar: 1 mm); 20. Female (bar: 2 mm); 21. Male (bar: 2 mm). Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668467" httpUri="https://zenodo.org/record/4668467/files/figure.png" pageId="8" pageNumber="549">Figs 19</figureCitation>
,
<figureCitation id="13352A37FFF7FFDE3716F20C1F3FFEB9" box="[517,549,259,285]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">28</figureCitation>
): Light yellowish to light brown. Irregular dark brown to blackish spots on the Op gen and the Cx I–IV, the upper end of Cx with a small brown margin contrasting with the whitish joint membrane and all sternites with a single row of black setae contrasting with light cuticle, Op gen with a few scattered setae.
</paragraph>
<paragraph id="8BB136B2FFF7FFDE35D4F260195FFDBD" blockId="8.[151,1437,151,1869]" pageId="8" pageNumber="549">
<emphasis id="B97AEAA0FFF7FFDE35D4F2601C2FFE2D" box="[199,309,367,393]" italics="true" pageId="8" pageNumber="549">Chelicera</emphasis>
(
<figureCitation id="13352A37FFF7FFDE3457F2601CDDFE2D" box="[324,455,367,393]" captionStart="FIGURES 30–32" captionStartId="12.[151,264,1306,1330]" captionTargetBox="[165,1415,181,1275]" captionTargetId="figure-23@12.[151,1436,181,1282]" captionTargetPageId="12" captionText="FIGURES 30–32. Nemaspela prometheus sp. nov. Chelicerae in retro-lateral view. 30. Male (bar: 0.8 mm); 31. Female (bar: 0.8 mm). Chelicera in pro-lateral view. 32. Male (bar: 0.8 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668471" httpUri="https://zenodo.org/record/4668471/files/figure.png" pageId="8" pageNumber="549">Figs 30, 32</figureCitation>
): Very slender and elongated; basal article slightly constricted close to its proximal end, from there slightly and continuously widened distally, distal third slightly bent downward, one bristle on dorsal side at mid-part, two long bristles dorsal sub-distally, no do Apo on basal article present; 2
<superScript id="7C7B9BFAFFF7FFDE315BF2B91942FE60" attach="left" box="[1096,1112,438,452]" fontSize="6" pageId="8" pageNumber="549">nd</superScript>
cheliceral article remarkably slender and longish, very slightly inflated, nearly parallel-sided (in lateral view,
<figureCitation id="13352A37FFF7FFDE310DF2D41983FE51" box="[1054,1177,475,501]" captionStart="FIGURES 30–32" captionStartId="12.[151,264,1306,1330]" captionTargetBox="[165,1415,181,1275]" captionTargetId="figure-23@12.[151,1436,181,1282]" captionTargetPageId="12" captionText="FIGURES 30–32. Nemaspela prometheus sp. nov. Chelicerae in retro-lateral view. 30. Male (bar: 0.8 mm); 31. Female (bar: 0.8 mm). Chelicera in pro-lateral view. 32. Male (bar: 0.8 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668471" httpUri="https://zenodo.org/record/4668471/files/figure.png" pageId="8" pageNumber="549">Fig. 30, 32</figureCitation>
) with several scattered long and strong bristles mostly on frontal surface, some shorter ones ventro-distally.
</paragraph>
<paragraph id="8BB136B2FFF7FFDE35D4F12C1C3FFD55" blockId="8.[151,1437,151,1869]" pageId="8" pageNumber="549">
<emphasis id="B97AEAA0FFF7FFDE35D4F12C1C36FD99" box="[199,300,547,573]" italics="true" pageId="8" pageNumber="549">Pedipalp</emphasis>
(
<figureCitation id="13352A37FFF7FFDE342FF12C1CA4FD99" box="[316,446,547,573]" captionStart="FIGURES 19–21" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[263,1326,181,1919]" captionTargetId="figure-17@9.[259,1327,181,1919]" captionTargetPageId="9" captionText="FIGURES 19–21. Nemaspela prometheus sp. nov. Habitus in lateral view. 19. Female (bar: 1 mm); 20. Female (bar: 2 mm); 21. Male (bar: 2 mm). Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668467" httpUri="https://zenodo.org/record/4668467/files/figure.png" pageId="8" pageNumber="549">Figs 19–20</figureCitation>
,
<figureCitation id="13352A37FFF7FFDE34D9F12C1CF0FD99" box="[458,490,547,573]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">22</figureCitation>
): Very slender and extremely long (in terms of
<taxonomicName id="4C0E4D31FFF7FFDE3111F12C1964FD99" authorityName="Silhavy" authorityYear="1966" box="[1026,1150,547,573]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="549" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEAA0FFF7FFDE3111F12C1964FD99" box="[1026,1150,547,573]" italics="true" pageId="8" pageNumber="549">Nemaspela</emphasis>
</taxonomicName>
morphology), all articles parallel-sided, distal end of Fe slightly swollen, Ta slightly curved downward, no article except Cx noticeably inflated; all articles except Tr bearing scattered clavate setae, these most conspicuous on Fe ventrally, on Pt mostly ventrally on distal half, on Ti a few at basal end ventrally and on Ta on all sides but scattered as well. Setae on Fe and on Pt on ventral side longer than breadth of article at the position. No article with special armament in males and females.
</paragraph>
<paragraph id="8BB136B2FFF7FFDE35D4F1F41926FC01" blockId="8.[151,1437,151,1869]" pageId="8" pageNumber="549">
<emphasis id="B97AEAA0FFF7FFDE35D4F1F41DE6FCB0" box="[199,252,763,788]" italics="true" pageId="8" pageNumber="549">Legs</emphasis>
(
<figureCitation id="13352A37FFF7FFDE3418F1F41C95FCB1" box="[267,399,763,789]" captionStart="FIGURES 19–21" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[263,1326,181,1919]" captionTargetId="figure-17@9.[259,1327,181,1919]" captionTargetPageId="9" captionText="FIGURES 19–21. Nemaspela prometheus sp. nov. Habitus in lateral view. 19. Female (bar: 1 mm); 20. Female (bar: 2 mm); 21. Male (bar: 2 mm). Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668467" httpUri="https://zenodo.org/record/4668467/files/figure.png" pageId="8" pageNumber="549">Figs 19–20</figureCitation>
): Light brownish tending to yellowish brownish and yellowish toward distal end, Tr mottled dark brownish, articles extremely long and slender, thread-like, round in cross-section, armed with some scattered and short black setae. Many pseudo-articulations on Fe I-IV (
<figureCitation id="13352A37FFF7FFDE362DF04C1E95FCF9" box="[830,911,835,861]" captionStart="FIGURES 19–21" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[263,1326,181,1919]" captionTargetId="figure-17@9.[259,1327,181,1919]" captionTargetPageId="9" captionText="FIGURES 19–21. Nemaspela prometheus sp. nov. Habitus in lateral view. 19. Female (bar: 1 mm); 20. Female (bar: 2 mm); 21. Male (bar: 2 mm). Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668467" httpUri="https://zenodo.org/record/4668467/files/figure.png" pageId="8" pageNumber="549">Fig. 19</figureCitation>
), densely packed near basis of articles, towards distal end of article distances between them consecutively more extended; no pseudo-articulations near distal end. Number of pseudo-articulations of Fe of legs: Fe I 15, Fe II 19, Fe III 13, Fe IV 18.
</paragraph>
<paragraph id="8BB136B2FFF7FFDE35D4F0A01CDFFA65" blockId="8.[151,1437,151,1869]" pageId="8" pageNumber="549">
<emphasis id="B97AEAA0FFF7FFDE35D4F0A01CFCFC6D" box="[199,486,943,969]" italics="true" pageId="8" pageNumber="549">Male genital morphology</emphasis>
(
<figureCitation id="13352A37FFF7FFDE34E5F0A01F61FC6D" box="[502,635,943,969]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">Figs 24–27</figureCitation>
): Truncus penis (
<figureCitation id="13352A37FFF7FFDE362DF0A01EA5FC6D" box="[830,959,942,969]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">Figs 24–25</figureCitation>
) extremely slender; basis forming a small inflated part (occupying about one sixth of whole penis length) and only slightly separated from the rest of the truncus; inflated part slender, broadest at proximal end and deeply split medially, completely filled by two penial muscles, their tendons spanning the hole truncus length up to the basis of the glans. Truncus nearly parallel-sided in dorsal/ventral view all over its length (dorsal/ventral view,
<figureCitation id="13352A37FFF7FFDE3656F7301E8DFBFD" box="[837,919,1087,1113]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">Fig. 24</figureCitation>
), but slightly and continuously tapering to just below transition to the glans; in lateral view (
<figureCitation id="13352A37FFF7FFDE378DF76C1FEBFBD9" box="[670,753,1122,1149]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">Fig. 25</figureCitation>
) truncus slightly narrower, strictly parallel-sided and hardly tapering toward glans. Glans (
<figureCitation id="13352A37FFF7FFDE34FEF7881F74FB06" box="[493,622,1159,1186]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">Figs 26–27</figureCitation>
) only inconspicuously separated from distal part of truncus as outlined, though distinctly widened at basis, but only very slightly broader than truncus (
<figureCitation id="13352A37FFF7FFDE3109F7A41974FB62" box="[1050,1134,1195,1222]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">Fig. 26</figureCitation>
), short, starting where two tendons are attached to inner truncus wall, lateral outline only slightly rounded, then continuously tapering toward stylus, stylus short, a continuation of the glans, tapering to a distinctly pointed distal tip. Glans in lateral view (
<figureCitation id="13352A37FFF7FFDE307CF7FC1DAEFA95" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">Fig. 27</figureCitation>
) a continuation of the distal part of truncus, but slightly enlarged dorsally for most of dorsal length, stylus distinctly bent to dorsal side. A few rather long, stiff and unspecialized spicules forming the armament of glans, their arrangement symmetrical in ventral, dorsal and lateral view; one double row each on lateral, ventral and lateral sides, spicules of the same size at the basis of stylus, slightly more densely packed than on glans proper. Measurements are given in
<tableCitation id="C68C0309FFF7FFDE347FF6A81CD8FA65" box="[364,450,1447,1473]" captionStart="TABLE 2" captionStartId="11.[151,239,151,175]" captionTargetPageId="11" captionText="TABLE 2. Measurements of Nemaspela prometheus sp. nov. Measurements are given in millimeters. Note: body length is measured in all available specimens, while length of appendages was measured only in holotype male and single female." httpUri="http://table.plazi.org/id/DF71663AFFF4FFDD3584F3981DC8FF53" pageId="8" pageNumber="549" tableUuid="DF71663AFFF4FFDD3584F3981DC8FF53">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BB136B2FFF7FFDE35D4F6C51C61F93D" blockId="8.[151,1437,151,1869]" pageId="8" pageNumber="549">
<emphasis id="B97AEAA0FFF7FFDE35D4F6C51F12FA40" bold="true" box="[199,520,1482,1509]" pageId="8" pageNumber="549">Description. Female (n=1).</emphasis>
External morphology of body generally similar to male (
<figureCitation id="13352A37FFF7FFDE3187F6C419F6FA41" box="[1172,1260,1483,1509]" captionStart="FIGURES 19–21" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[263,1326,181,1919]" captionTargetId="figure-17@9.[259,1327,181,1919]" captionTargetPageId="9" captionText="FIGURES 19–21. Nemaspela prometheus sp. nov. Habitus in lateral view. 19. Female (bar: 1 mm); 20. Female (bar: 2 mm); 21. Male (bar: 2 mm). Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668467" httpUri="https://zenodo.org/record/4668467/files/figure.png" pageId="8" pageNumber="549">Figs 21</figureCitation>
,
<figureCitation id="13352A37FFF7FFDE31EAF6C41803FA41" box="[1273,1305,1483,1509]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">23</figureCitation>
).
<emphasis id="B97AEAA0FFF7FFDE303DF6C41886FA41" box="[1326,1436,1483,1509]" italics="true" pageId="8" pageNumber="549">Chelicera</emphasis>
(
<figureCitation id="13352A37FFF7FFDE35B3F6E01DEBF9AD" box="[160,241,1519,1545]" captionStart="FIGURES 30–32" captionStartId="12.[151,264,1306,1330]" captionTargetBox="[165,1415,181,1275]" captionTargetId="figure-23@12.[151,1436,181,1282]" captionTargetPageId="12" captionText="FIGURES 30–32. Nemaspela prometheus sp. nov. Chelicerae in retro-lateral view. 30. Male (bar: 0.8 mm); 31. Female (bar: 0.8 mm). Chelicera in pro-lateral view. 32. Male (bar: 0.8 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668471" httpUri="https://zenodo.org/record/4668471/files/figure.png" pageId="8" pageNumber="549">Fig. 31</figureCitation>
) similar to male, but 1
<superScript id="7C7B9BFAFFF7FFDE34E2F6E01CE6FA59" attach="left" box="[497,508,1519,1533]" fontSize="6" pageId="8" pageNumber="549">st</superScript>
and 2
<superScript id="7C7B9BFAFFF7FFDE3752F6E11F4BFA58" attach="left" box="[577,593,1518,1532]" fontSize="6" pageId="8" pageNumber="549">nd</superScript>
articles even longer and thus more slender, setation of both articles similar to male, but front setae of 2
<superScript id="7C7B9BFAFFF7FFDE34C2F51D1CFBF984" attach="left" box="[465,481,1554,1568]" fontSize="6" pageId="8" pageNumber="549">nd</superScript>
article much longer than in male, the length about the diameter (in lateral view,
<figureCitation id="13352A37FFF7FFDE3063F51C1DAEF9F5" captionStart="FIGURES 30–32" captionStartId="12.[151,264,1306,1330]" captionTargetBox="[165,1415,181,1275]" captionTargetId="figure-23@12.[151,1436,181,1282]" captionTargetPageId="12" captionText="FIGURES 30–32. Nemaspela prometheus sp. nov. Chelicerae in retro-lateral view. 30. Male (bar: 0.8 mm); 31. Female (bar: 0.8 mm). Chelicera in pro-lateral view. 32. Male (bar: 0.8 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668471" httpUri="https://zenodo.org/record/4668471/files/figure.png" pageId="8" pageNumber="549">Fig. 31</figureCitation>
) of 2
<superScript id="7C7B9BFAFFF7FFDE35E0F5391C19F9E0" attach="left" box="[243,259,1590,1604]" fontSize="6" pageId="8" pageNumber="549">nd</superScript>
article. Pedipalp (
<figureCitation id="13352A37FFF7FFDE34C6F5381F32F9F5" box="[469,552,1591,1617]" captionStart="FIGURES 22–29" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[193,1385,211,1919]" captionTargetId="figure-23@10.[176,1412,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–29. Nemaspela prometheus sp. nov. Pedipalps in retro-lateral view. 22. Male (bar: 0.8 mm); 23. Female (bar: 0.8 mm). Penis. 24. Ventral view (bar: 0.2 mm); 25. Lateral view (bar: 0.2 mm). Glans penis. 26. Ventral view (bar: 0.05 mm); 27. Lateral view (bar: 0.05 mm). Habitus in lateral view. 28. Female (bar: 0.5 mm); 29. Male (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668469" httpUri="https://zenodo.org/record/4668469/files/figure.png" pageId="8" pageNumber="549">Fig. 23</figureCitation>
) very similar to male including setation, but all articles markedly longer (see Measurements). Number of pseudo-articulations of Fe of legs: Fe I 17, Fe II 21, Fe III 18, Fe IV 21. Measurements are given in
<tableCitation id="C68C0309FFF7FFDE3431F5701C62F93D" box="[290,376,1663,1689]" captionStart="TABLE 2" captionStartId="11.[151,239,151,175]" captionTargetPageId="11" captionText="TABLE 2. Measurements of Nemaspela prometheus sp. nov. Measurements are given in millimeters. Note: body length is measured in all available specimens, while length of appendages was measured only in holotype male and single female." httpUri="http://table.plazi.org/id/DF71663AFFF4FFDD3584F3981DC8FF53" pageId="8" pageNumber="549" tableUuid="DF71663AFFF4FFDD3584F3981DC8FF53">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BB136B2FFF7FFDE35D4F5AD1CAEF945" blockId="8.[151,1437,151,1869]" pageId="8" pageNumber="549">
<emphasis id="B97AEAA0FFF7FFDE35D4F5AD1C25F918" bold="true" box="[199,319,1698,1724]" pageId="8" pageNumber="549">Variation.</emphasis>
The available specimens are rather homogeneous in size, coloration and appendages. For differences between sexes see above.
</paragraph>
<paragraph id="8BB136B2FFF7FFDE35D4F5E51F33F8A1" blockId="8.[151,1437,151,1869]" box="[199,553,1770,1797]" pageId="8" pageNumber="549">
<emphasis id="B97AEAA0FFF7FFDE35D4F5E51C6AF8A0" bold="true" box="[199,368,1770,1796]" pageId="8" pageNumber="549">Relationships.</emphasis>
See Discussion.
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFF7FFDE35D4F4011F1DF8E9" pageId="8" pageNumber="549" type="distribution">
<paragraph id="8BB136B2FFF7FFDE35D4F4011F1DF8E9" blockId="8.[151,1437,151,1869]" pageId="8" pageNumber="549">
<emphasis id="B97AEAA0FFF7FFDE35D4F4011C7BF88C" bold="true" box="[199,353,1806,1832]" pageId="8" pageNumber="549">Distribution.</emphasis>
Known only from Prometheus Cave in Tskaltubo Municipality, Sataplia-Tskaltubo karst massif,
<collectingRegion id="49CAF850FFF7FFDE3584F43C1C23F8E9" box="[151,313,1843,1869]" country="Georgia" name="Imereti" pageId="8" pageNumber="549">Imereti region</collectingRegion>
, Western
<collectingCountry id="F3197622FFF7FFDE34B5F43C1F19F8E9" box="[422,515,1843,1869]" name="Georgia" pageId="8" pageNumber="549">Georgia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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