treatments-xml/data/03/A7/4C/03A74C6ACA5DFFB4FF118156FEBDFEF2.xml
2024-06-21 12:22:17 +02:00

848 lines
99 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="75030925DE60E9D23BF2B51D3B80F324" ID-DOI="10.5281/zenodo.197522" ID-GBIF-Dataset="57248f86-bb49-45e3-ae37-94bb510769d7" ID-ISSN="1175-5326" ID-Zenodo-Dep="197522" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460272452079" checkinUser="plazi" docAuthor="Ayala-Varela, Fernando P. &amp; Velasco, Julián Andrés" docDate="2010" docId="03A74C6ACA5DFFB4FF118156FEBDFEF2" docLanguage="en" docName="zt02577p056.pdf" docOrigin="Zootaxa 2577" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anolis otongae Ayala-Varela &amp; Velasco, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="55" masterDocId="FF9E3412CA5CFFBDFF86867EFFC8FF9E" masterDocTitle="A new species of dactyloid anole (SQUAMATA: IGUANIDAE) from the western Andes of Ecuador" masterLastPageNumber="56" masterPageNumber="46" pageNumber="47" updateTime="1698247112618" updateUser="plazi">
<mods:mods id="8A07261EC704E54273BA60265B443115" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="31A9126EDECAEFD9F01B6E23EA211846">
<mods:title id="6CE3F607B2716DDFD1D6996D60D14970">A new species of dactyloid anole (SQUAMATA: IGUANIDAE) from the western Andes of Ecuador</mods:title>
</mods:titleInfo>
<mods:name id="9603ECBF1A5F0144BB306F97CAE925B5" type="personal">
<mods:role id="9923360066857D1789F9E2B8E4B10F9E">
<mods:roleTerm id="3BA01C69FD60326587C6BB7B6C3B46B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E40120FA4406266EB8D5F92D496C5A9">Ayala-Varela, Fernando P.</mods:namePart>
</mods:name>
<mods:name id="BEC2BE0E824A7C1BF51D6497D927C95E" type="personal">
<mods:role id="FF15C09DF2D1ADB9C038B37D66636D45">
<mods:roleTerm id="216DC2D667003C8D2A3F9500EC8717DD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19F3FFE8E9E90831CAAADBDC78064848">Velasco, Julián Andrés</mods:namePart>
</mods:name>
<mods:typeOfResource id="4FC9D55F557A98400BCA85A0DAC8B44C">text</mods:typeOfResource>
<mods:relatedItem id="C89310B6E9F715D7E68E9763C8778C2F" type="host">
<mods:titleInfo id="C2D8A071C4DC9CA0A140AFC8076B2837">
<mods:title id="E4D10750A763612D3682C403BEF959D8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7E84707E980644E0B6EEC1C77B3918C2">
<mods:date id="9F468BBB2DB491A60D2C5C65B5824A5C">2010</mods:date>
<mods:detail id="E7FA68DDBE99C24509671F79902CF070" type="volume">
<mods:number id="14AD623B3FAD6E017E5FFDE0FB632203">2577</mods:number>
</mods:detail>
<mods:extent id="3160C5CE5D6BA8D38797F7EBA80CCF98" unit="page">
<mods:start id="4F23551A395F6B86C5541FFA051A7A6E">46</mods:start>
<mods:end id="57D9E43E4376E15D786638F702BDB46F">56</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="503658A9D931CB2DECFA66647E5E74D3">journal article</mods:classification>
<mods:identifier id="F0E20855B03681FC90EDE9E609460BC2" type="DOI">10.5281/zenodo.197522</mods:identifier>
<mods:identifier id="00C9029494D585A16756B686335FC56A" type="GBIF-Dataset">57248f86-bb49-45e3-ae37-94bb510769d7</mods:identifier>
<mods:identifier id="44001BC3B07BEDE64846B61BA2851B52" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="23CFAD705F36872E80F9F9850635A8F5" type="Zenodo-Dep">197522</mods:identifier>
</mods:mods>
<treatment id="03A74C6ACA5DFFB4FF118156FEBDFEF2" ID-DOI="http://doi.org/10.5281/zenodo.5625658" ID-GBIF-Taxon="119395382" ID-Zenodo-Dep="5625658" LSID="urn:lsid:plazi:treatment:03A74C6ACA5DFFB4FF118156FEBDFEF2" httpUri="http://treatment.plazi.org/id/03A74C6ACA5DFFB4FF118156FEBDFEF2" lastPageId="9" lastPageNumber="55" pageId="1" pageNumber="47">
<subSubSection id="C314AEF7CA5DFFBCFF118156FEE1F8FA" pageId="1" pageNumber="47" type="nomenclature">
<paragraph id="8BB1FD7CCA5DFFBCFF118156FE70F8DC" blockId="1.[151,440,1832,1892]" box="[151,440,1832,1858]" pageId="1" pageNumber="47">
<heading id="D0F94A10CA5DFFBCFF118156FE70F8DC" bold="true" box="[151,440,1832,1858]" fontSize="11" level="1" pageId="1" pageNumber="47" reason="1">
<emphasis id="B97A216ECA5DFFBCFF118156FE70F8DC" bold="true" box="[151,440,1832,1858]" pageId="1" pageNumber="47">
<taxonomicName id="4C0E86FFCA5DFFBCFF118156FE84F8DC" box="[151,332,1832,1858]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="47" phylum="Chordata" rank="species" species="otongae" status="sp. nov.">
<emphasis id="B97A216ECA5DFFBCFF118156FE84F8DC" bold="true" box="[151,332,1832,1858]" italics="true" pageId="1" pageNumber="47">Anolis otongae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2499C15CA5DFFBCFEDD8156FE70F8DC" box="[347,440,1832,1858]" pageId="1" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB1FD7CCA5DFFBCFF118134FEE1F8FA" blockId="1.[151,440,1832,1892]" box="[151,297,1866,1892]" pageId="1" pageNumber="47">
<figureCitation id="1335E1F9CA5DFFBCFF118134FECDF8FA" box="[151,261,1866,1892]" captionStart="FIGURE 1" captionStartId="2.[151,255,1880,1904]" captionTargetBox="[159,1436,1202,1860]" captionTargetId="figure@2.[159,1481,1202,1861]" captionTargetPageId="2" captionText="FIGURE 1. Anolis otongae, QCAZ 3872, adult female (SVL 63.3 mm)." httpUri="https://zenodo.org/record/197523/files/figure.png" pageId="1" pageNumber="47">Figures 1</figureCitation>
,
<figureCitation id="1335E1F9CA5DFFBCFE958134FEEDF8FA" box="[275,293,1866,1892]" captionStart="FIGURE 2" captionStartId="3.[151,255,1991,2015]" captionTargetBox="[381,1252,190,1963]" captionTargetId="figure@3.[333,1253,172,1971]" captionTargetPageId="3" captionText="FIGURE 2. Anolis otongae, holotype, QCAZ 2051. Head in dorsal (A), lateral (B), and ventral (C) views. Scale 5 mm." httpUri="https://zenodo.org/record/197524/files/figure.png" pageId="1" pageNumber="47">2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C314AEF7CA5DFFBFFF1181EEFD52FD22" lastPageId="2" lastPageNumber="48" pageId="1" pageNumber="47" type="materials_examined">
<paragraph id="8BB1FD7CCA5DFFBCFF1181EEFB96F86A" blockId="1.[151,1436,1936,2036]" pageId="1" pageNumber="47">
<emphasis id="B97A216ECA5DFFBCFF1181EEFEDFF834" bold="true" box="[151,279,1936,1962]" pageId="1" pageNumber="47">
<typeStatus id="54B543DECA5DFFBCFF1181EEFEDAF834" box="[151,274,1936,1962]" pageId="1" pageNumber="47" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED1F65B9CA5DFFBCFEA681EEFEBBF834" box="[288,371,1936,1962]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="47">QCAZ</collectionCode>
2051, adult male,
<collectingCountry id="F319BDECCA5DFFBCFDD981EEFD01F834" box="[607,713,1936,1962]" name="Ecuador" pageId="1" pageNumber="47">Ecuador</collectingCountry>
, Provincia Cotopaxi, Cantón Sigchos, Reserva de Bosque Integral Otonga, near San Francisco de Las Pampas,
<geoCoordinate id="EE3A9BBBCA5DFFBCFC9181CBFC6FF851" box="[791,935,1973,1999]" direction="south" orientation="latitude" pageId="1" pageNumber="47" precision="1" value="-0.41889998">0º25'8.04&quot;S</geoCoordinate>
,
<geoCoordinate id="EE3A9BBBCA5DFFBCFC3581CBFB93F851" box="[947,1115,1973,1999]" direction="west" orientation="longitude" pageId="1" pageNumber="47" precision="1" value="-79.0039">79º0'14.04&quot;W</geoCoordinate>
,
<quantity id="4CF65099CA5DFFBCFBEE81CBFADBF851" box="[1128,1299,1973,1999]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="1" pageNumber="47" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">20002200 m</quantity>
,
<date id="FFB0DBBCCA5DFFBCFAA681CBFF1CF86A" pageId="1" pageNumber="47" value="1993-08-30">30 August 1993</date>
, collected by Néstor Acosta, Paola Ramón, César Tapia and Luis A. Coloma.
</paragraph>
<paragraph id="8BB1FD7CCA5EFFBFFF4386E9FD52FD22" blockId="2.[151,1437,151,1148]" pageId="2" pageNumber="48">
<emphasis id="B97A216ECA5EFFBFFF4386E9FE84FF2F" bold="true" box="[197,332,151,177]" pageId="2" pageNumber="48">
<typeStatus id="54B543DECA5EFFBFFF4386E9FE80FF2F" box="[197,328,151,177]" pageId="2" pageNumber="48" type="paratype">Paratypes</typeStatus>
.
</emphasis>
(18, all from
<collectingCountry id="F319BDECCA5EFFBFFE7086E6FDA8FF2C" box="[502,608,152,178]" name="Ecuador" pageId="2" pageNumber="48">Ecuador</collectingCountry>
).
<collectionCode id="ED1F65B9CA5EFFBFFDF286E6FD0DFF2C" box="[628,709,152,178]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
1696, Provincia Cotopaxi, Peñas Coloradas,
<date id="FFB0DBBCCA5EFFBFFB6D86E6FA50FF2C" box="[1259,1432,152,178]" pageId="2" pageNumber="48" value="1992-04-20">20 April 1992</date>
, Giovanni Onore;
<collectionCode id="ED1F65B9CA5EFFBFFEE186C3FE7FFF49" box="[359,439,189,215]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
1721, same locality data as
<typeStatus id="54B543DECA5EFFBFFC8F86C3FCBBFF49" box="[777,883,189,215]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
,
<date id="FFB0DBBCCA5EFFBFFC0686C3FBD3FF49" box="[896,1051,189,215]" pageId="2" pageNumber="48" value="1994-04-02">2 April 1994</date>
, Daniela Andrade;
<collectionCode id="ED1F65B9CA5EFFBFFA8786C3FA99FF49" box="[1281,1361,189,215]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
2050, 2052, same collection data as
<typeStatus id="54B543DECA5EFFBFFD85869CFDB8FF62" box="[515,624,226,252]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED1F65B9CA5EFFBFFDFB869CFD06FF62" box="[637,718,226,252]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
2128, near San Francisco de Las Pampas, approx.
<quantity id="4CF65099CA5EFFBFFABF869CFA5FFF62" box="[1337,1431,226,252]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="48" unit="m" value="1800.0">1800 m</quantity>
,
<date id="FFB0DBBCCA5EFFBFFF118776FE9CFEBC" box="[151,340,264,290]" pageId="2" pageNumber="48" value="1989-12">December 1989</date>
, Giovanni Onore;
<collectionCode id="ED1F65B9CA5EFFBFFDA88776FDB6FEBC" box="[558,638,264,290]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
2781, Provincia Pichincha, Los Libres,
<quantity id="4CF65099CA5EFFBFFBD18776FB36FEBC" box="[1111,1278,264,290]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="2" pageNumber="48" unit="m" value="2250.0" valueMax="2500.0" valueMin="2000.0">20002500 m</quantity>
,
<date id="FFB0DBBCCA5EFFBFFA8C8776FF1CFED9" pageId="2" pageNumber="48" value="1994-11-04">4 November 1994</date>
, Dandie Salazar Pico;
<collectionCode id="ED1F65B9CA5EFFBFFE5A8753FDE4FED9" box="[476,556,301,327]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
3129, same locality data as
<typeStatus id="54B543DECA5EFFBFFCF38753FC16FED9" box="[885,990,301,327]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
,
<date id="FFB0DBBCCA5EFFBFFC6F8753FB65FED9" box="[1001,1197,301,327]" pageId="2" pageNumber="48" value="1996-01-28">28 January 1996</date>
, César Tapia, Ernest E. Williams and Luis A. Coloma;
<collectionCode id="ED1F65B9CA5EFFBFFDA4872CFDBAFEF2" box="[546,626,338,364]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
3706, same locality data as
<typeStatus id="54B543DECA5EFFBFFC3F872CFBEAFEF2" box="[953,1058,338,364]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
,
<quantity id="4CF65099CA5EFFBFFBAB872CFB41FEF2" box="[1069,1161,338,364]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="48" unit="m" value="2000.0">2000 m</quantity>
,
<date id="FFB0DBBCCA5EFFBFFB15872CFAECFEF2" box="[1171,1316,338,364]" pageId="2" pageNumber="48" value="1997-03">March 1997</date>
, Giovanni Onore;
<collectionCode id="ED1F65B9CA5EFFBFFF758706FE8CFE0C" box="[243,324,376,402]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
3796, same locality data as
<typeStatus id="54B543DECA5EFFBFFD1A8706FCC0FE0C" box="[668,776,376,402]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
,
<date id="FFB0DBBCCA5EFFBFFC938706FC39FE0C" box="[789,1009,376,402]" pageId="2" pageNumber="48" value="1997-11-02">2 November 1997</date>
, Hugo Mogollón;
<collectionCode id="ED1F65B9CA5EFFBFFB578706FAEAFE0C" box="[1233,1314,376,402]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
387273, same locality data as
<typeStatus id="54B543DECA5EFFBFFE1487E3FE33FE29" box="[402,507,413,439]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
,
<date id="FFB0DBBCCA5EFFBFFD8187E3FCF9FE29" box="[519,817,413,439]" pageId="2" pageNumber="48" value="1998-02-17" valueMax="1998-02-21" valueMin="1998-02-17">17 and 21 February 1998</date>
, respectively, Juan M. Guayasamín, Luis A. Coloma and Alexandra Quiguango;
<collectionCode id="ED1F65B9CA5EFFBFFE5987BCFDE7FE42" box="[479,559,450,476]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
4025, same locality data as
<typeStatus id="54B543DECA5EFFBFFC0687BCFC22FE42" box="[896,1002,450,476]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
,
<quantity id="4CF65099CA5EFFBFFC7087BCFB9BFE42" box="[1014,1107,450,476]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="48" unit="m" value="2000.0">2000 m</quantity>
,
<date id="FFB0DBBCCA5EFFBFFBE687BCFA80FE42" box="[1120,1352,450,476]" pageId="2" pageNumber="48" value="1998-09-19">19 September 1998</date>
, María Gloria Rivas, Martín R. Bustamante and Juan M. Guayasamín;
<collectionCode id="ED1F65B9CA5EFFBFFC0D8796FC13FD9C" box="[907,987,488,514]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
5481, same locality data as
<typeStatus id="54B543DECA5EFFBFFAAB8796FA5FFD9C" box="[1325,1431,488,514]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
,
<date id="FFB0DBBCCA5EFFBFFF118473FE9EFDB9" box="[151,342,525,551]" pageId="2" pageNumber="48" value="2000-02">February 2000</date>
, Martín R. Bustamante;
<collectionCode id="ED1F65B9CA5EFFBFFD128473FD21FDB9" box="[660,745,525,551]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
6219, same data locality as
<typeStatus id="54B543DECA5EFFBFFBDA8473FB18FDB9" box="[1116,1232,525,551]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
,
<geoCoordinate id="EE3A9BBBCA5EFFBFFB5B8473FA5FFDB9" box="[1245,1431,525,551]" direction="south" orientation="latitude" pageId="2" pageNumber="48" precision="1" value="-0.419083">0º25'8.6988&quot;S</geoCoordinate>
,
<geoCoordinate id="EE3A9BBBCA5EFFBFFF11844CFE8EFDD2" box="[151,326,562,588]" direction="west" orientation="longitude" pageId="2" pageNumber="48" precision="1" value="-79.0122">79º0'43.92&quot;W</geoCoordinate>
,
<quantity id="4CF65099CA5EFFBFFEE6844CFE0BFDD2" box="[352,451,562,588]" metricMagnitude="3" metricUnit="m" metricValue="2.214" pageId="2" pageNumber="48" unit="m" value="2214.0">2214 m</quantity>
,
<date id="FFB0DBBCCA5EFFBFFE57844CFD0FFDD2" box="[465,711,562,588]" pageId="2" pageNumber="48" value="2007-11-17">17 November 2007</date>
, David Salazar-V., Elicio Tapia and George Vaca;
<collectionCode id="ED1F65B9CA5EFFBFFACF844CFA55FDD2" box="[1353,1437,562,588]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="48">QCAZ</collectionCode>
639496, Provincia Pichincha, La Victoria,
<geoCoordinate id="EE3A9BBBCA5EFFBFFD208426FCAAFDEC" box="[678,866,600,626]" direction="south" orientation="latitude" pageId="2" pageNumber="48" precision="1" value="-0.47746983">0º28'38.8914&quot;S</geoCoordinate>
,
<geoCoordinate id="EE3A9BBBCA5EFFBFFCE88426FBFAFDEC" box="[878,1074,600,626]" direction="west" orientation="longitude" pageId="2" pageNumber="48" precision="1" value="-79.05336">79º3'12.0954&quot;W</geoCoordinate>
,
<quantity id="4CF65099CA5EFFBFFBB98426FB55FDEC" box="[1087,1181,600,626]" metricMagnitude="3" metricUnit="m" metricValue="2.104" pageId="2" pageNumber="48" unit="m" value="2104.0">2104 m</quantity>
,
<date id="FFB0DBBCCA5EFFBFFB2F8426FA9AFDEC" box="[1193,1362,600,626]" pageId="2" pageNumber="48" value="2003-04-27">27 April 2003</date>
and
<date id="FFB0DBBCCA5EFFBFFA0B8426FEC7FD09" pageId="2" pageNumber="48" value="2003-05-01">1 May 2003</date>
respectively, Italo Tapia, César Tapia and Edwin P. Vargas;
<collectionCode id="ED1F65B9CA5EFFBFFC508403FBF8FD09" box="[982,1072,637,663]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="48">MHNG</collectionCode>
2300.08, same locality data as
<typeStatus id="54B543DECA5EFFBFFF1184DCFEC9FD22" box="[151,257,674,700]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
,
<date id="FFB0DBBCCA5EFFBFFE8A84DCFE01FD22" box="[268,457,674,700]" pageId="2" pageNumber="48" value="1985-12">December 1985</date>
, Giovanni Onore.
</paragraph>
</subSubSection>
<subSubSection id="C314AEF7CA5EFFBFFF4084B9FD04FBE2" pageId="2" pageNumber="48" type="diagnosis">
<paragraph id="8BB1FD7CCA5EFFBFFF4084B9FC79FB92" blockId="2.[151,1437,151,1148]" pageId="2" pageNumber="48">
<emphasis id="B97A216ECA5EFFBFFF4084B9FE8EFD7F" bold="true" box="[198,326,711,737]" pageId="2" pageNumber="48">Diagnosis.</emphasis>
According to
<bibRefCitation id="EF9F808DCA5EFFBFFE7284B6FD74FD7C" author="Etheridge" box="[500,700,712,738]" pageId="2" pageNumber="48" refString="Etheridge, R. (1959) The relationships of the anoles (Reptilia: Sauria: Iguanidae): an interpretation based on skeletal morphology. PhD. Thesis, University of Michigan, Ann Harbor, Michigan, USA." type="book" year="1959">Etheridge (1959)</bibRefCitation>
and
<bibRefCitation id="EF9F808DCA5EFFBFFD7F84B6FC09FD7C" author="Williams" box="[761,961,712,738]" pageId="2" pageNumber="48" refString="Williams, E. E. (1976 a) South American anoles: The species groups. Papeis Avulsos de Zoologia, 29, 259 - 268." type="journal article" year="1976" yearSuffix="a">Williams (1976a</bibRefCitation>
,
<bibRefCitation id="EF9F808DCA5EFFBFFC4084B6FC1EFD7C" author="Williams" box="[966,982,712,738]" pageId="2" pageNumber="48" refString="Williams, E. E. (1976 b) West Indian anoles: a taxonomic and evolutionary summary. I. Introduction and a species list. Breviora, 440, 1 - 21." type="journal article" year="1976" yearSuffix="b">b</bibRefCitation>
) the new species belongs to the alpha section of anoles by lacking transverse processes on most or all of the autotomic caudal vertebrae (distinctive, slender, transverse processes that are usually oriented forwardly lie posterior to the autotomy septum and are present on all or most caudal vertebrae in beta section of anoles), and it belongs to the
<emphasis id="B97A216ECA5EFFBFFB228546FADFFCCF" box="[1188,1303,824,849]" italics="true" pageId="2" pageNumber="48">punctatus</emphasis>
subsection (
<bibRefCitation id="EF9F808DCA5EFFBFFF198523FE95FCE9" author="Williams" box="[159,349,861,887]" pageId="2" pageNumber="48" refString="Williams, E. E. (1976 a) South American anoles: The species groups. Papeis Avulsos de Zoologia, 29, 259 - 268." type="journal article" year="1976" yearSuffix="a">Williams 1976a</bibRefCitation>
) by having an arrow-shaped interclavicle (T-shaped in
<emphasis id="B97A216ECA5EFFBFFC698523FBB6FCE8" box="[1007,1150,861,886]" italics="true" pageId="2" pageNumber="48">carolinensis</emphasis>
-subsection). Within the
<emphasis id="B97A216ECA5EFFBFFF1185FCFEC2FC05" box="[151,266,898,923]" italics="true" pageId="2" pageNumber="48">punctatus</emphasis>
subsection,
<taxonomicName id="4C0E86FFCA5EFFBFFE1885FCFD99FC05" box="[414,593,898,923]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="48" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA5EFFBFFE1885FCFD99FC05" box="[414,593,898,923]" italics="true" pageId="2" pageNumber="48">Anolis otongae</emphasis>
</taxonomicName>
is a member of the
<taxonomicName id="4C0E86FFCA5EFFBFFCC785FCFC6CFC05" box="[833,932,898,923]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="48" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="B97A216ECA5EFFBFFCC785FCFC6CFC05" box="[833,932,898,923]" italics="true" pageId="2" pageNumber="48">latifrons</emphasis>
</taxonomicName>
series (
<bibRefCitation id="EF9F808DCA5EFFBFFC7985FCFB71FC02" author="Etheridge" box="[1023,1209,898,924]" pageId="2" pageNumber="48" refString="Etheridge, R. (1959) The relationships of the anoles (Reptilia: Sauria: Iguanidae): an interpretation based on skeletal morphology. PhD. Thesis, University of Michigan, Ann Harbor, Michigan, USA." type="book" year="1959">Etheridge 1959</bibRefCitation>
) by having at least four parasternal chevrons attached to the dorsal ribs, and the lateral processes of the interclavicle divergent from the proximal parts of the clavicles.
<taxonomicName id="4C0E86FFCA5EFFBFFDF085B3FCE0FC78" box="[630,808,973,998]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="48" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA5EFFBFFDF085B3FCE0FC78" box="[630,808,973,998]" italics="true" pageId="2" pageNumber="48">Anolis otongae</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C0E86FFCA5EFFBFFC5A85B3FBB0FC78" box="[988,1144,973,998]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="48" phylum="Chordata" rank="species" species="aequatorialis">
<emphasis id="B97A216ECA5EFFBFFC5A85B3FBB0FC78" box="[988,1144,973,998]" italics="true" pageId="2" pageNumber="48">aequatorialis</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EF9F808DCA5EFFBFFAB685B3FF28FB92" author="Williams" pageId="2" pageNumber="48" refString="Williams, E. E. (1976 a) South American anoles: The species groups. Papeis Avulsos de Zoologia, 29, 259 - 268." type="journal article" year="1976" yearSuffix="a">Williams 1976a</bibRefCitation>
) by being of moderate to large size with narrow toe lamellae.
</paragraph>
<paragraph id="8BB1FD7CCA5EFFBFFF438266FD04FBE2" blockId="2.[151,1437,151,1148]" pageId="2" pageNumber="48">
According to
<bibRefCitation id="EF9F808DCA5EFFBFFEE38266FD58FBAC" author="Savage" box="[357,656,1048,1074]" pageId="2" pageNumber="48" refString="Savage, J. M. &amp; Guyer, C. (1989) Infrageneric classification and species composition of the anole genera, Anolis, Ctenonotus, Dactyloa, Norops and Semiurus (Sauria: Iguanidae). Amphibia-Reptilia, 10, 105 - 116." type="journal article" year="1989">Savage and Guyer (1989)</bibRefCitation>
<taxonomicName id="4C0E86FFCA5EFFBFFD1E8266FC80FBAF" box="[664,840,1048,1073]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="48" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA5EFFBFFD1E8266FC80FBAF" box="[664,840,1048,1073]" italics="true" pageId="2" pageNumber="48">Anolis otongae</emphasis>
</taxonomicName>
is a member of the
<taxonomicName id="4C0E86FFCA5EFFBFFBA98266FB52FBAF" box="[1071,1178,1048,1073]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="2" pageNumber="48" phylum="Chordata" rank="genus">
<emphasis id="B97A216ECA5EFFBFFBA98266FB52FBAF" box="[1071,1178,1048,1073]" italics="true" pageId="2" pageNumber="48">Dactyloa</emphasis>
</taxonomicName>
clade by the presence of caudal autotomy and ≤ 8 septate caudal vertebrae, and member of the
<taxonomicName id="4C0E86FFCA5EFFBFFBD38243FACBFBC8" box="[1109,1283,1085,1110]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="48" phylum="Chordata" rank="species" species="aequatorialis">
<emphasis id="B97A216ECA5EFFBFFBD38243FACBFBC8" box="[1109,1283,1085,1110]" italics="true" pageId="2" pageNumber="48">aequatorialis</emphasis>
</taxonomicName>
series (i.e.,
<taxonomicName id="4C0E86FFCA5EFFBFFF11821DFEFBFBE2" box="[151,307,1123,1148]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="48" phylum="Chordata" rank="species" species="aequatorialis">
<emphasis id="B97A216ECA5EFFBFFF11821DFEFBFBE2" box="[151,307,1123,1148]" italics="true" pageId="2" pageNumber="48">aequatorialis</emphasis>
</taxonomicName>
species group of
<bibRefCitation id="EF9F808DCA5EFFBFFD87821CFD08FBE2" author="Williams" box="[513,704,1122,1148]" pageId="2" pageNumber="48" refString="Williams, E. E. (1976 a) South American anoles: The species groups. Papeis Avulsos de Zoologia, 29, 259 - 268." type="journal article" year="1976" yearSuffix="a">Williams 1976a</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF71ADF4CA5EFFBFFF118126FC54F8F1" box="[151,924,1880,1904]" httpUri="https://zenodo.org/record/197523/files/figure.png" pageId="2" pageNumber="48" targetBox="[159,1436,1202,1860]" targetPageId="2">
<paragraph id="8BB1FD7CCA5EFFBFFF118126FC54F8F1" blockId="2.[151,924,1880,1904]" box="[151,924,1880,1904]" pageId="2" pageNumber="48">
<emphasis id="B97A216ECA5EFFBFFF118126FED1F8EE" bold="true" box="[151,281,1880,1904]" pageId="2" pageNumber="48">FIGURE 1.</emphasis>
<taxonomicName id="4C0E86FFCA5EFFBFFEA68126FE09F8F1" box="[288,449,1880,1903]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="48" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA5EFFBFFEA68126FE09F8F1" box="[288,449,1880,1903]" italics="true" pageId="2" pageNumber="48">Anolis otongae</emphasis>
</taxonomicName>
, QCAZ 3872, adult female (SVL 63.3 mm).
</paragraph>
</caption>
<caption id="DF71ADF4CA5FFFBEFF1181B9FA5AF841" box="[151,1426,1991,2015]" httpUri="https://zenodo.org/record/197524/files/figure.png" pageId="3" pageNumber="49" targetBox="[381,1252,190,1963]" targetPageId="3">
<paragraph id="8BB1FD7CCA5FFFBEFF1181B9FA5AF841" blockId="3.[151,1426,1991,2015]" box="[151,1426,1991,2015]" pageId="3" pageNumber="49">
<emphasis id="B97A216ECA5FFFBEFF1181B9FED1F841" bold="true" box="[151,281,1991,2015]" pageId="3" pageNumber="49">FIGURE 2.</emphasis>
<taxonomicName id="4C0E86FFCA5FFFBEFEA681B6FE09F841" box="[288,449,1992,2015]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="49" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA5FFFBEFEA681B6FE09F841" box="[288,449,1992,2015]" italics="true" pageId="3" pageNumber="49">Anolis otongae</emphasis>
</taxonomicName>
, holotype, QCAZ 2051. Head in dorsal (A), lateral (B), and ventral (C) views. Scale 5 mm.
</paragraph>
</caption>
<subSubSection id="C314AEF7CA58FFB9FF4386E6FCEFF932" pageId="4" pageNumber="50" type="reference_group">
<paragraph id="8BB1FD7CCA58FFB9FF4386E6FABFFEF2" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="50">
<taxonomicName id="4C0E86FFCA58FFB9FF4386E6FEBEFF2F" box="[197,374,152,177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA58FFB9FF4386E6FEBEFF2F" box="[197,374,152,177]" italics="true" pageId="4" pageNumber="50">Anolis otongae</emphasis>
</taxonomicName>
differs from most species in the
<emphasis id="B97A216ECA58FFB9FD7A86E6FCA6FF2F" box="[764,878,152,177]" italics="true" pageId="4" pageNumber="50">punctatus</emphasis>
and
<emphasis id="B97A216ECA58FFB9FC2F86E6FBCDFF2F" box="[937,1029,152,177]" italics="true" pageId="4" pageNumber="50">tigrinus</emphasis>
species group (Williams 1992) by having relatively small head scales, from the
<taxonomicName id="4C0E86FFCA58FFB9FD3786C3FD3EFF48" box="[689,758,189,214]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B97A216ECA58FFB9FD3786C3FD3EFF48" box="[689,758,189,214]" italics="true" pageId="4" pageNumber="50">laevis</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EF9F808DCA58FFB9FC3686C3FBA6FF49" author="Williams" box="[944,1134,189,215]" pageId="4" pageNumber="50" refString="Williams, E. E. (1976 a) South American anoles: The species groups. Papeis Avulsos de Zoologia, 29, 259 - 268." type="journal article" year="1976" yearSuffix="a">Williams 1976a</bibRefCitation>
) by lacking a soft, multiscaled proboscis (sensu
<bibRefCitation id="EF9F808DCA58FFB9FE4B869CFD45FF62" author="Williams" box="[461,653,226,252]" pageId="4" pageNumber="50" refString="Williams, E. E. (1979) South American anoles: the species groups. 2. The proboscis anoles (Anolis laevis group). Breviora, 449, 1 - 19." type="journal article" year="1979">Williams 1979</bibRefCitation>
), from the
<emphasis id="B97A216ECA58FFB9FC9D869CFCA5FF65" box="[795,877,226,251]" italics="true" pageId="4" pageNumber="50">roquet</emphasis>
species group (
<bibRefCitation id="EF9F808DCA58FFB9FBB1869CFACCFF62" author="Williams" box="[1079,1284,226,252]" pageId="4" pageNumber="50" refString="Williams, E. E. (1976 a) South American anoles: The species groups. Papeis Avulsos de Zoologia, 29, 259 - 268." type="journal article" year="1976" yearSuffix="a">Williams 1976a</bibRefCitation>
) by lacking supraorbital semicircles in contact with each other, interparietal in contact with supraorbital semicircles and posterior border of mental straight (sensu
<bibRefCitation id="EF9F808DCA58FFB9FD0A8753FCC9FED9" author="Poe" box="[652,769,301,327]" pageId="4" pageNumber="50" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89." type="journal article" year="2004">Poe 2004</bibRefCitation>
), and from the
<taxonomicName id="4C0E86FFCA58FFB9FC308753FBD2FED8" box="[950,1050,301,326]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="B97A216ECA58FFB9FC308753FBD2FED8" box="[950,1050,301,326]" italics="true" pageId="4" pageNumber="50">latifrons</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EF9F808DCA58FFB9FB528753FA5BFED9" author="Williams" box="[1236,1427,301,327]" pageId="4" pageNumber="50" refString="Williams, E. E. (1976 a) South American anoles: The species groups. Papeis Avulsos de Zoologia, 29, 259 - 268." type="journal article" year="1976" yearSuffix="a">Williams 1976a</bibRefCitation>
) by having a snout-vent length less than
<quantity id="4CF65099CA58FFB9FDE1872CFD01FEF2" box="[615,713,338,364]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="50" unit="mm" value="100.0">100 mm</quantity>
and lacking a T-shaped interclavicle (sensu
<bibRefCitation id="EF9F808DCA58FFB9FB49872CFAA3FEF2" author="Savage" box="[1231,1387,338,364]" pageId="4" pageNumber="50" refString="Savage, J. M. (2002) The Amphibians and Reptiles of Costa Rica. A Herpetofauna between Two Continents, between two Seas. University of Chicago Press. Chicago, Illinois, USA." type="book" year="2002">Savage 2002</bibRefCitation>
).
</paragraph>
<paragraph id="8BB1FD7CCA58FFB9FF438706FE50FA39" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="50">
We assign
<taxonomicName id="4C0E86FFCA58FFB9FEC58706FE3CFE0F" box="[323,500,376,401]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA58FFB9FEC58706FE3CFE0F" box="[323,500,376,401]" italics="true" pageId="4" pageNumber="50">Anolis otongae</emphasis>
</taxonomicName>
to the
<emphasis id="B97A216ECA58FFB9FDC58706FD73FE0F" box="[579,699,376,401]" italics="true" pageId="4" pageNumber="50">eulaemus-</emphasis>
subgroup (
<bibRefCitation id="EF9F808DCA58FFB9FCB18706FBB7FE0C" author="Williams" box="[823,1151,376,402]" pageId="4" pageNumber="50" refString="Williams, E. E. &amp; Duellman, W. E. (1984) Anolis fitchi, a new species of the Anolis aequatorialis group from Ecuador and Colombia. University of Kansas Publications of the Museum of Natural History, 10, 257 - 266." type="journal article" year="1984">Williams &amp; Duellman 1984</bibRefCitation>
) because it has a typical
<taxonomicName id="4C0E86FFCA58FFB9FF1187E3FF2CFE28" box="[151,228,413,438]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B97A216ECA58FFB9FF1187E3FF2CFE28" box="[151,228,413,438]" italics="true" pageId="4" pageNumber="50">Anolis</emphasis>
</taxonomicName>
digit, in which the distal lamellae of phalanx II distinctly overlap the proximal scales of phalanx I.
<emphasis id="B97A216ECA58FFB9FF1187BCFD8DFE45" box="[151,581,450,475]" italics="true" pageId="4" pageNumber="50">
<taxonomicName id="4C0E86FFCA58FFB9FF1187BCFE5EFE45" box="[151,406,450,475]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="aequatorialis">Anolis aequatorialis</taxonomicName>
,
<taxonomicName id="4C0E86FFCA58FFB9FE2587BCFD8DFE45" authority="Hulebak et al." authorityName="Hulebak et al." box="[419,581,450,475]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="kunayalae">A. kunayalae</taxonomicName>
</emphasis>
Hulebak
<emphasis id="B97A216ECA58FFB9FD3B87BCFD34FE45" box="[701,764,450,475]" italics="true" pageId="4" pageNumber="50">et al.</emphasis>
,
<taxonomicName id="4C0E86FFCA58FFB9FC8B87BCFC3AFE42" authority="Williams" authorityName="Williams" box="[781,1010,450,476]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="mirus">
<emphasis id="B97A216ECA58FFB9FC8B87BCFCBEFE45" box="[781,886,450,475]" italics="true" pageId="4" pageNumber="50">A. mirus</emphasis>
Williams
</taxonomicName>
, and
<taxonomicName id="4C0E86FFCA58FFB9FBB287BCFAD7FE42" authority="Williams" authorityName="Williams" box="[1076,1311,450,476]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="parilis">
<emphasis id="B97A216ECA58FFB9FBB287BCFB60FE45" box="[1076,1192,450,475]" italics="true" pageId="4" pageNumber="50">A. parilis</emphasis>
Williams
</taxonomicName>
belong to
<emphasis id="B97A216ECA58FFB9FF118796FE8EFD9F" box="[151,326,488,513]" italics="true" pageId="4" pageNumber="50">aequatorialis-</emphasis>
subgroup (
<bibRefCitation id="EF9F808DCA58FFB9FE4E8796FCEAFD9C" author="Williams" box="[456,802,488,514]" pageId="4" pageNumber="50" refString="Williams, E. E. &amp; Duellman, W. E. (1984) Anolis fitchi, a new species of the Anolis aequatorialis group from Ecuador and Colombia. University of Kansas Publications of the Museum of Natural History, 10, 257 - 266." type="journal article" year="1984">Williams &amp; Duellman 1984</bibRefCitation>
) by having a
<taxonomicName id="4C0E86FFCA58FFB9FC4B8796FBEEFD9F" box="[973,1062,488,513]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B97A216ECA58FFB9FC4B8796FBEEFD9F" box="[973,1062,488,513]" italics="true" pageId="4" pageNumber="50">Norops</emphasis>
</taxonomicName>
-
<typeStatus id="54B543DECA58FFB9FBA88796FBADFD9C" box="[1070,1125,488,514]" pageId="4" pageNumber="50">type</typeStatus>
digit, in which the distal lamellae of phalanx II not distinct from first phalanx. Among species in the
<taxonomicName id="4C0E86FFCA58FFB9FBA28473FB5CFDB8" box="[1060,1172,525,550]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="eulaemus">
<emphasis id="B97A216ECA58FFB9FBA28473FB5CFDB8" box="[1060,1172,525,550]" italics="true" pageId="4" pageNumber="50">eulaemus</emphasis>
</taxonomicName>
-subgroup,
<taxonomicName id="4C0E86FFCA58FFB9FA9D8473FA54FDB8" box="[1307,1436,525,550]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA58FFB9FA9D8473FA54FDB8" box="[1307,1436,525,550]" italics="true" pageId="4" pageNumber="50">A. otongae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0E86FFCA58FFB9FEAB844CFDF7FDD2" authority="Williams" authorityName="Williams" box="[301,575,562,588]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="antioquiae">
<emphasis id="B97A216ECA58FFB9FEAB844CFE05FDD5" box="[301,461,562,587]" italics="true" pageId="4" pageNumber="50">A. antioquiae</emphasis>
Williams
</taxonomicName>
in lacking a canthal ridge projecting above the loreal region (very sharp canthal ridge projecting above the loreal region in
<taxonomicName id="4C0E86FFCA58FFB9FD6D8426FC41FDEF" box="[747,905,600,625]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="antioquiae">
<emphasis id="B97A216ECA58FFB9FD6D8426FC41FDEF" box="[747,905,600,625]" italics="true" pageId="4" pageNumber="50">A. antioquiae</emphasis>
</taxonomicName>
), and females dewlap absent (female dewlap moderate extending to the level of the arms in
<taxonomicName id="4C0E86FFCA58FFB9FD5E8403FCB5FD08" box="[728,893,637,662]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="antioquiae">
<emphasis id="B97A216ECA58FFB9FD5E8403FCB5FD08" box="[728,893,637,662]" italics="true" pageId="4" pageNumber="50">A. antioquiae</emphasis>
</taxonomicName>
). The new species differs from
<taxonomicName id="4C0E86FFCA58FFB9FA818403FED9FD22" authority="Boulenger" authorityName="Boulenger" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="eulaemus">
<emphasis id="B97A216ECA58FFB9FA818403FA55FD08" box="[1287,1437,637,662]" italics="true" pageId="4" pageNumber="50">A. eulaemus</emphasis>
Boulenger
</taxonomicName>
in having the male dewlap skin white or pale-yellow and greenish-yellow anteriorly (male dewlap skin brown with a pale brown border in
<taxonomicName id="4C0E86FFCA58FFB9FDEC84B6FD31FD7F" box="[618,761,712,737]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="eulaemus">
<emphasis id="B97A216ECA58FFB9FDEC84B6FD31FD7F" box="[618,761,712,737]" italics="true" pageId="4" pageNumber="50">A. eulaemus</emphasis>
</taxonomicName>
), female dewlap absent (female dewlap rudimentary with a dark skin in
<taxonomicName id="4C0E86FFCA58FFB9FEC48493FE1CFC98" box="[322,468,749,774]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="eulaemus">
<emphasis id="B97A216ECA58FFB9FEC48493FE1CFC98" box="[322,468,749,774]" italics="true" pageId="4" pageNumber="50">A. eulaemus</emphasis>
</taxonomicName>
), and adults with snout-vent length &lt;
<quantity id="4CF65099CA58FFB9FC1A8493FC38FC99" box="[924,1008,749,775]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="50" unit="mm" value="70.0">70 mm</quantity>
(adults with snout-vent length&gt;
<quantity id="4CF65099CA58FFB9FAF88493FF0DFCB2" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="50" unit="mm" value="80.0">80 mm</quantity>
in
<taxonomicName id="4C0E86FFCA58FFB9FF6D856CFEB3FCB5" box="[235,379,786,811]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="eulaemus">
<emphasis id="B97A216ECA58FFB9FF6D856CFEB3FCB5" box="[235,379,786,811]" italics="true" pageId="4" pageNumber="50">A. eulaemus</emphasis>
</taxonomicName>
).
<taxonomicName id="4C0E86FFCA58FFB9FE12856CFD8DFCB5" box="[404,581,786,811]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA58FFB9FE12856CFD8DFCB5" box="[404,581,786,811]" italics="true" pageId="4" pageNumber="50">Anolis otongae</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C0E86FFCA58FFB9FC00856CFACCFCB2" authority="Williams and Duellman" authorityName="Williams and Duellman" box="[902,1284,786,812]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="fitchi">
<emphasis id="B97A216ECA58FFB9FC00856CFC2BFCB5" box="[902,995,786,811]" italics="true" pageId="4" pageNumber="50">A. fitchi</emphasis>
Williams and Duellman
</taxonomicName>
by lacking a dewlap in females (females with moderate dewlap size in
<taxonomicName id="4C0E86FFCA58FFB9FCDC8546FC72FCCF" box="[858,954,824,849]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="fitchi">
<emphasis id="B97A216ECA58FFB9FCDC8546FC72FCCF" box="[858,954,824,849]" italics="true" pageId="4" pageNumber="50">A. fitchi</emphasis>
</taxonomicName>
), by having a dewlap with rows of 36 scales separated by naked skin (dewlap with large scales in rows of one scale separated by naked skin in
<taxonomicName id="4C0E86FFCA58FFB9FA058523FF1BFC05" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="fitchi">
<emphasis id="B97A216ECA58FFB9FA058523FF1BFC05" italics="true" pageId="4" pageNumber="50">A. fitchi</emphasis>
</taxonomicName>
).
<taxonomicName id="4C0E86FFCA58FFB9FF6A85FCFE54FC05" box="[236,412,898,923]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA58FFB9FF6A85FCFE54FC05" box="[236,412,898,923]" italics="true" pageId="4" pageNumber="50">Anolis otongae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C0E86FFCA58FFB9FDFC85FCFCDDFC05" box="[634,789,898,923]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="gemmosus">
<emphasis id="B97A216ECA58FFB9FDFC85FCFCDDFC05" box="[634,789,898,923]" italics="true" pageId="4" pageNumber="50">A. gemmosus</emphasis>
</taxonomicName>
but differs from it by the following features (character states of
<taxonomicName id="4C0E86FFCA58FFB9FE8385D6FE6EFC5F" box="[261,422,936,961]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="gemmosus">
<emphasis id="B97A216ECA58FFB9FE8385D6FE6EFC5F" box="[261,422,936,961]" italics="true" pageId="4" pageNumber="50">A. gemmosus</emphasis>
</taxonomicName>
in parentheses): side of the head with a stripe dark coppery-brown (stripe absent), females with dark brown dorsal chevrons extending onto flanks (chevrons absent), male dewlap skin white or pale-yellow and greenish-yellow anteriorly with pale-green gorgetals rows and white sternals rows (male dewlap skin dull yellowish green on the basal area, shading to dull greenish yellow on the outer part; or dewlap skin dull greenish yellow outer part, but the basal part was bluish green with six narrow sharply defined white stripes diverging from a center on the anterior basal portion), dewlap with wide scale rows of three to six scales per row (narrow scale rows of 23 scales per row), if present, interparietal scale surrounded by relatively enlarged flat scales (interparietal surrounded by small swollen scales), and enlarged postanal scales separated by 12 scales (postanal scales separated by 35 scales). Additionally, PCA analyses (
<tableCitation id="C68CC8C7CA58FFB9FADF82ACFF00FA8C" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="5.[151,240,562,586]" captionStartId-1="5.[151,240,887,911]" captionTargetBox-0="[151,1436,680,831]" captionTargetBox-1="[151,1436,1005,1540]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="TABLE 1. Principal component analysis (PCA) of Anolis gemmosus and A. otongae morphology. The rotation sums of squared loadings are given." captionText-1="TABLE 2. Principal component analysis (PCA) of Anolis gemmosus and A. otongae morphology. The loadings for the morphology variables are given." httpUri-0="http://table.plazi.org/id/DF71ADF4CA59FFB8FF11844CFE73FDF7" httpUri-1="http://table.plazi.org/id/DF71ADF4CA59FFB8FF118509FE38FC31" pageId="4" pageNumber="50" tableUuid-0="DF71ADF4CA59FFB8FF11844CFE73FDF7" tableUuid-1="DF71ADF4CA59FFB8FF118509FE38FC31">Table 12</tableCitation>
,
<figureCitation id="1335E1F9CA58FFB9FF508286FED7FA8C" box="[214,287,1272,1298]" captionStart="FIGURE 4" captionStartId="8.[151,255,1302,1326]" captionTargetBox="[159,1435,187,1289]" captionTargetId="figure@8.[151,1436,176,1297]" captionTargetPageId="8" captionText="FIGURE 4. Distribution of Anolis gemmosus and A. otongae along the first and second principal components axes." httpUri="https://zenodo.org/record/197526/files/figure.png" pageId="4" pageNumber="50">Fig. 4</figureCitation>
) with morphological variables showed that PC1 (36.9% of total variation) represented mainly snout-vent length and head length. Specimens of
<taxonomicName id="4C0E86FFCA58FFB9FD668363FCA8FAA8" box="[736,864,1309,1334]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA58FFB9FD668363FCA8FAA8" box="[736,864,1309,1334]" italics="true" pageId="4" pageNumber="50">A. otongae</emphasis>
</taxonomicName>
showed a tendency to have larger bodies (SVL) and longer heads than
<taxonomicName id="4C0E86FFCA58FFB9FE24833DFDF5FAC2" box="[418,573,1347,1372]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="gemmosus">
<emphasis id="B97A216ECA58FFB9FE24833DFDF5FAC2" box="[418,573,1347,1372]" italics="true" pageId="4" pageNumber="50">A. gemmosus</emphasis>
</taxonomicName>
. The PC2 (30.7% of total variation) represented mostly hindlimb length. The Wilks lambda test showed a significant separation in morphological space between both species (Wilks λ = 0.176, P &lt;0.001).
</paragraph>
<paragraph id="8BB1FD7CCA58FFB9FF4383CDFCEFF932" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="50">
<taxonomicName id="4C0E86FFCA58FFB9FF4383CDFE4DFA52" box="[197,389,1459,1484]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA58FFB9FF4383CDFE4DFA52" box="[197,389,1459,1484]" italics="true" pageId="4" pageNumber="50">Anolis otongae</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C0E86FFCA58FFB9FD6483CDFBD8FA52" authority="Williams" authorityName="Williams" box="[738,1040,1458,1484]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="maculigula">
<emphasis id="B97A216ECA58FFB9FD6483CDFC5CFA52" box="[738,916,1459,1484]" italics="true" pageId="4" pageNumber="50">A. maculigula</emphasis>
Williams
</taxonomicName>
and
<taxonomicName id="4C0E86FFCA58FFB9FBC983CDFAF1FA52" box="[1103,1337,1459,1484]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="megalophitecus">
<emphasis id="B97A216ECA58FFB9FBC983CDFAF1FA52" box="[1103,1337,1459,1484]" italics="true" pageId="4" pageNumber="50">A. megalophitecus</emphasis>
</taxonomicName>
Rueda- Almonacid in having the male dewlap skin white or pale-yellow and greenish yellow anteriorly (male dewlap skin in the base with orange stripes on blue gray ground, anterior third pale bluish rose, posterior portion white becoming pale blue toward the belly in
<taxonomicName id="4C0E86FFCA58FFB9FD41805DFCBAF9A2" box="[711,882,1571,1596]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="maculigula">
<emphasis id="B97A216ECA58FFB9FD41805DFCBAF9A2" box="[711,882,1571,1596]" italics="true" pageId="4" pageNumber="50">A. maculigula</emphasis>
</taxonomicName>
; male dewlap skin sepia with red narrow and irregulars stripes to each side of rows in
<taxonomicName id="4C0E86FFCA58FFB9FDFC8036FC9CF9FF" box="[634,852,1608,1633]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="megalophitecus">
<emphasis id="B97A216ECA58FFB9FDFC8036FC9CF9FF" box="[634,852,1608,1633]" italics="true" pageId="4" pageNumber="50">A. megalophitecus</emphasis>
</taxonomicName>
).
<taxonomicName id="4C0E86FFCA58FFB9FCEB8036FBE8F9FF" box="[877,1056,1608,1633]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA58FFB9FCEB8036FBE8F9FF" box="[877,1056,1608,1633]" italics="true" pageId="4" pageNumber="50">Anolis otongae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0E86FFCA58FFB9FB3B8036FED9F919" authority="Boulenger" authorityName="Boulenger" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="ventrimaculatus">
<emphasis id="B97A216ECA58FFB9FB3B8036FA54F9FF" box="[1213,1436,1608,1633]" italics="true" pageId="4" pageNumber="50">A. ventrimaculatus</emphasis>
Boulenger
</taxonomicName>
by having a dewlap with rows of 36 scales separated by naked skin (large scales with rows of 1 or 2 scales separated by naked skin in
<taxonomicName id="4C0E86FFCA58FFB9FDBE80EDFCDDF932" box="[568,789,1683,1708]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="50" phylum="Chordata" rank="species" species="ventrimaculatus">
<emphasis id="B97A216ECA58FFB9FDBE80EDFCDDF932" box="[568,789,1683,1708]" italics="true" pageId="4" pageNumber="50">A. ventrimaculatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C314AEF7CA58FFBAFF4380C6FEC4FEBC" lastPageId="7" lastPageNumber="53" pageId="4" pageNumber="50" type="description">
<paragraph id="8BB1FD7CCA58FFB9FF4380C6FDF2F969" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="50">
Description of
<typeStatus id="54B543DECA58FFB9FEFA80C6FE2EF94C" box="[380,486,1720,1746]" pageId="4" pageNumber="50" type="holotype">holotype</typeStatus>
(scores for
<typeStatus id="54B543DECA58FFB9FDFF80C6FD25F94C" box="[633,749,1720,1746]" pageId="4" pageNumber="50" type="paratype">paratypes</typeStatus>
in parentheses; juvenile specimens [QCAZ 1696, 3706] were excluded from measurements)
</paragraph>
<paragraph id="8BB1FD7CCA58FFB9FF43817CFB78F82C" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="50">
SVL
<quantity id="4CF65099CA58FFB9FE8C817CFEB4F882" box="[266,380,1794,1820]" metricMagnitude="-2" metricUnit="m" metricValue="6.62" pageId="4" pageNumber="50" unit="mm" value="66.2">66.2 mm</quantity>
(
<quantity id="4CF65099CA58FFB9FE08817CFD9CF882" box="[398,596,1794,1820]" metricMagnitude="-2" metricUnit="m" metricValue="6.159999999999999" metricValueMax="6.709999999999999" metricValueMin="5.61" pageId="4" pageNumber="50" unit="mm" value="61.599999999999994" valueMax="67.1" valueMin="56.1">56.167.1 mm</quantity>
); tail length 158.0 mm (118.0170.0 mm); head length
<quantity id="4CF65099CA58FFB9FAAD817CFA55F882" box="[1323,1437,1794,1820]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="4" pageNumber="50" unit="mm" value="18.3">18.3 mm</quantity>
(
<quantity id="4CF65099CA58FFB9FF198156FE9AF8DC" box="[159,338,1832,1858]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" metricValueMax="1.81" metricValueMin="1.51" pageId="4" pageNumber="50" unit="mm" value="16.6" valueMax="18.1" valueMin="15.1">15.118.1 mm</quantity>
); head width
<quantity id="4CF65099CA58FFB9FE778156FD83F8DC" box="[497,587,1832,1858]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="4" pageNumber="50" unit="mm" value="9.6">9.6 mm</quantity>
(
<quantity id="4CF65099CA58FFB9FDDD8156FD39F8DC" box="[603,753,1832,1858]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="9.9" metricValueMin="7.6" pageId="4" pageNumber="50" unit="mm" value="8.75" valueMax="9.9" valueMin="7.6">7.69.9 mm</quantity>
); head height
<quantity id="4CF65099CA58FFB9FC108156FC38F8DC" box="[918,1008,1832,1858]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="4" pageNumber="50" unit="mm" value="8.3">8.3 mm</quantity>
(
<quantity id="4CF65099CA58FFB9FB868156FB5EF8DC" box="[1024,1174,1832,1858]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.5" metricValueMin="6.5" pageId="4" pageNumber="50" unit="mm" value="7.5" valueMax="8.5" valueMin="6.5">6.58.5 mm</quantity>
); forelimb length
<quantity id="4CF65099CA58FFB9FAEF8156FF0DF8F9" metricMagnitude="-2" metricUnit="m" metricValue="3.28" pageId="4" pageNumber="50" unit="mm" value="32.8">32.8 mm</quantity>
(24.931.0 mm); hindlimb length
<quantity id="4CF65099CA58FFB9FDE18133FD1AF8F9" box="[615,722,1869,1895]" metricMagnitude="-2" metricUnit="m" metricValue="5.46" pageId="4" pageNumber="50" unit="mm" value="54.6">54.6 mm</quantity>
(41.555.0 mm); dewlap length
<quantity id="4CF65099CA58FFB9FBD88133FB01F8F9" box="[1118,1225,1869,1895]" metricMagnitude="-2" metricUnit="m" metricValue="3.3299999999999996" pageId="4" pageNumber="50" unit="mm" value="33.3">33.3 mm</quantity>
(
<quantity id="4CF65099CA58FFB9FB5F8133FA47F8F9" box="[1241,1423,1869,1895]" metricMagnitude="-2" metricUnit="m" metricValue="3.075" metricValueMax="3.51" metricValueMin="2.6399999999999997" pageId="4" pageNumber="50" unit="mm" value="30.75" valueMax="35.1" valueMin="26.4">26.435.1 mm</quantity>
); dewlap height
<quantity id="4CF65099CA58FFB9FEC5810CFE64F812" box="[323,428,1906,1932]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="4" pageNumber="50" unit="mm" value="14.3">14.3 mm</quantity>
(
<quantity id="4CF65099CA58FFB9FE3D810CFDA5F812" box="[443,621,1906,1932]" metricMagnitude="-2" metricUnit="m" metricValue="1.505" metricValueMax="1.78" metricValueMin="1.23" pageId="4" pageNumber="50" unit="mm" value="15.05" valueMax="17.8" valueMin="12.3">12.317.8 mm</quantity>
); interparietal length 1.0 (
<quantity id="4CF65099CA58FFB9FC1D810CFBF8F812" box="[923,1072,1906,1932]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="4" pageNumber="50" unit="mm" value="1.05" valueMax="1.3" valueMin="0.8">0.81.3 mm</quantity>
); ear opening maximum length 2.2 (
<quantity id="4CF65099CA58FFB9FF4D81E6FEA9F82C" box="[203,353,1944,1970]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.3" metricValueMin="1.7" pageId="4" pageNumber="50" unit="mm" value="2.0" valueMax="2.3" valueMin="1.7">1.72.3 mm</quantity>
); snout length 8.1 (
<quantity id="4CF65099CA58FFB9FDC681E6FD1CF82C" box="[576,724,1944,1970]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" metricValueMax="8.4" metricValueMin="6.9" pageId="4" pageNumber="50" unit="mm" value="7.65" valueMax="8.4" valueMin="6.9">6.98.4 mm</quantity>
); interorbital distance 2.8 (2.43.0 mm).
</paragraph>
<paragraph id="8BB1FD7CCA59FFB8FF4386E6FBF8FD9C" blockId="5.[151,1437,152,514]" pageId="5" pageNumber="51">Head scales rugose (smooth, unicarinate, multicarinate or striate); 15 (1418) scales between first canthals; 17 (1217) scales between second canthals; seven (58) scales bordering the rostral posteriorly; circumnasal separated from rostral by one scale (circumnasal in contact with rostral, anterior nasal and inferior nasal); supraorbital semicircles separated by four (35) scales; supraocular disk with nine (29) enlarged scales; one elongated supracilliary followed by a series of granules; six (610) loreal rows; 45 loreal scales (4085); interparietal, when present, surrounded by relatively enlarged scales; interparietal smaller than ear opening; two (117) enlarged nape scales posterior to interparietal; 45 (36) scales between interparietal and semicircles; suboculars and supralabials in contact; 6 (69) supralabials counted up to a point below center of eye; 6 (69) infralabials counted up to a point below center of eye; five (47) postmentals; one enlarged sublabial in contact with infralabials (samewise or sublabials absent).</paragraph>
<caption id="DF71ADF4CA59FFB8FF11844CFE73FDF7" ID-Table-UUID="DF71ADF4CA59FFB8FF11844CFE73FDF7" httpUri="http://table.plazi.org/id/DF71ADF4CA59FFB8FF11844CFE73FDF7" pageId="5" pageNumber="51" targetBox="[151,1436,680,831]" targetIsTable="true" targetPageId="5">
<paragraph id="8BB1FD7CCA59FFB8FF11844CFE73FDF7" blockId="5.[151,1436,562,660]" pageId="5" pageNumber="51">
<emphasis id="B97A216ECA59FFB8FF11844CFEC3FDD4" bold="true" box="[151,267,562,586]" pageId="5" pageNumber="51">TABLE 1.</emphasis>
Principal component analysis (PCA) of
<taxonomicName id="4C0E86FFCA59FFB8FD47844CFCB4FDD7" box="[705,892,562,585]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="51" phylum="Chordata" rank="species" species="gemmosus">
<emphasis id="B97A216ECA59FFB8FD47844CFCB4FDD7" box="[705,892,562,585]" italics="true" pageId="5" pageNumber="51">Anolis gemmosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E86FFCA59FFB8FC37844CFBEDFDD7" box="[945,1061,562,585]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="51" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA59FFB8FC37844CFBEDFDD7" box="[945,1061,562,585]" italics="true" pageId="5" pageNumber="51">A. otongae</emphasis>
</taxonomicName>
morphology. The rotation sums of squared loadings are given.
</paragraph>
</caption>
<paragraph id="8BB1FD7CCA59FFB8FC848403FB9AFC47" blockId="5.[151,1436,562,660]" lastBlockId="5.[151,1436,887,985]" pageId="5" pageNumber="51">Raw Rescaled Raw Rescaled</paragraph>
<paragraph id="8BB1FD7CCA59FFB8FF1984ACFB33FCA1" pageId="5" pageNumber="51">
<table id="F90E0FDCCA590042FF1184D6FA54FCA1" box="[151,1436,680,831]" gridcols="5" gridrows="3" pageId="5" pageNumber="51">
<tr id="353EFF3ECA590042FF1184D6FA54FD77" box="[151,1436,680,745]" gridrow="0" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF1184D6FD92FD77" box="[151,602,680,745]" gridcol="0" gridrow="0" pageId="5" pageNumber="51">Eigenvalue</th>
<th id="76EF9642CA590042FC8484D6FC84FD77" box="[770,844,680,745]" gridcol="1" gridrow="0" pageId="5" pageNumber="51">1 16.699</th>
<th id="76EF9642CA590042FC1084D6FC28FD77" box="[918,992,680,745]" gridcol="2" gridrow="0" pageId="5" pageNumber="51">2 22.721</th>
<th id="76EF9642CA590042FBA284D6FBA5FD77" box="[1060,1133,680,745]" gridcol="3" gridrow="0" pageId="5" pageNumber="51">1 4.430</th>
<th id="76EF9642CA590042FB3484D6FA54FD77" box="[1202,1436,680,745]" gridcol="4" gridrow="0" pageId="5" pageNumber="51">2 3.685</th>
</tr>
<tr id="353EFF3ECA590042FF118483FA54FC8A" box="[151,1436,765,788]" gridrow="1" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF118483FD92FC8A" box="[151,602,765,788]" gridcol="0" gridrow="1" pageId="5" pageNumber="51">% of total variance explained</th>
<td id="76EF9642CA590042FC848483FC84FC8A" box="[770,844,765,788]" gridcol="1" gridrow="1" pageId="5" pageNumber="51">37.568</td>
<td id="76EF9642CA590042FC108483FC28FC8A" box="[918,992,765,788]" gridcol="2" gridrow="1" pageId="5" pageNumber="51">51.115</td>
<td id="76EF9642CA590042FBA28483FBA5FC8A" box="[1060,1133,765,788]" gridcol="3" gridrow="1" pageId="5" pageNumber="51">36.920</td>
<td id="76EF9642CA590042FB348483FA54FC8A" box="[1202,1436,765,788]" gridcol="4" gridrow="1" pageId="5" pageNumber="51">30.709</td>
</tr>
<tr id="353EFF3ECA590042FF118556FA54FCA1" box="[151,1436,808,831]" gridrow="2" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF118556FD92FCA1" box="[151,602,808,831]" gridcol="0" gridrow="2" pageId="5" pageNumber="51">Cumulative % of total variance explained</th>
<td id="76EF9642CA590042FC848556FC84FCA1" box="[770,844,808,831]" gridcol="1" gridrow="2" pageId="5" pageNumber="51">37.568</td>
<td id="76EF9642CA590042FC108556FC28FCA1" box="[918,992,808,831]" gridcol="2" gridrow="2" pageId="5" pageNumber="51">88.683</td>
<td id="76EF9642CA590042FBA28556FBA5FCA1" box="[1060,1133,808,831]" gridcol="3" gridrow="2" pageId="5" pageNumber="51">36.920</td>
<td id="76EF9642CA590042FB348556FA54FCA1" box="[1202,1436,808,831]" gridcol="4" gridrow="2" pageId="5" pageNumber="51">67.629</td>
</tr>
</table>
</paragraph>
<caption id="DF71ADF4CA59FFB8FF118509FE38FC31" ID-Table-UUID="DF71ADF4CA59FFB8FF118509FE38FC31" httpUri="http://table.plazi.org/id/DF71ADF4CA59FFB8FF118509FE38FC31" pageId="5" pageNumber="51" targetBox="[151,1436,1005,1540]" targetIsTable="true" targetPageId="5">
<paragraph id="8BB1FD7CCA59FFB8FF118509FE38FC31" blockId="5.[151,1436,887,985]" pageId="5" pageNumber="51">
<emphasis id="B97A216ECA59FFB8FF118509FEC3FC11" bold="true" box="[151,267,887,911]" pageId="5" pageNumber="51">TABLE 2.</emphasis>
Principal component analysis (PCA) of
<taxonomicName id="4C0E86FFCA59FFB8FD458506FCB7FC11" box="[707,895,888,911]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="51" phylum="Chordata" rank="species" species="gemmosus">
<emphasis id="B97A216ECA59FFB8FD458506FCB7FC11" box="[707,895,888,911]" italics="true" pageId="5" pageNumber="51">Anolis gemmosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E86FFCA59FFB8FC328506FBE1FC11" box="[948,1065,888,911]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="51" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA59FFB8FC328506FBE1FC11" box="[948,1065,888,911]" italics="true" pageId="5" pageNumber="51">A. otongae</emphasis>
</taxonomicName>
morphology. The loadings for the morphology variables are given.
</paragraph>
</caption>
<paragraph id="8BB1FD7CCA59FFB8FD368593FB06F99A" pageId="5" pageNumber="51">
<table id="F90E0FDCCA590042FF118593FA54F99A" box="[151,1436,1005,1540]" gridcols="5" gridrows="9" pageId="5" pageNumber="51">
<tr id="353EFF3ECA590042FF118593FA54FB9A" box="[151,1436,1005,1028]" gridrow="0" pageId="5" pageNumber="51" rowspan-0="1">
<th id="76EF9642CA590042FD368593FD24FB9A" box="[688,748,1005,1028]" gridcol="1" gridrow="0" pageId="5" pageNumber="51">1</th>
<th id="76EF9642CA590042FCD58593FC50FB9A" box="[851,920,1005,1028]" gridcol="2" gridrow="0" pageId="5" pageNumber="51">2</th>
<th id="76EF9642CA590042FC748593FBE6FB9A" box="[1010,1070,1005,1028]" gridcol="3" gridrow="0" pageId="5" pageNumber="51">1</th>
<th id="76EF9642CA590042FB148593FA54FB9A" box="[1170,1436,1005,1028]" gridcol="4" gridrow="0" pageId="5" pageNumber="51">2</th>
</tr>
<tr id="353EFF3ECA590042FF118266FA54FBC7" box="[151,1436,1048,1113]" gridrow="1" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF118266FE15FBC7" box="[151,477,1048,1113]" gridcol="0" gridrow="1" pageId="5" pageNumber="51">Head length Head width</th>
<td id="76EF9642CA590042FD368266FD24FBC7" box="[688,748,1048,1113]" gridcol="1" gridrow="1" pageId="5" pageNumber="51">1.159 0.392</td>
<td id="76EF9642CA590042FCD58266FC50FBC7" box="[851,920,1048,1113]" gridcol="2" gridrow="1" pageId="5" pageNumber="51">0.591 0.347</td>
<td id="76EF9642CA590042FC748266FBE6FBC7" box="[1010,1070,1048,1113]" gridcol="3" gridrow="1" pageId="5" pageNumber="51">0.801 0.644</td>
<td id="76EF9642CA590042FB148266FA54FBC7" box="[1170,1436,1048,1113]" gridcol="4" gridrow="1" pageId="5" pageNumber="51">0.409 0.570</td>
</tr>
<tr id="353EFF3ECA590042FF118213FA54FB1A" box="[151,1436,1133,1156]" gridrow="2" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF118213FE15FB1A" box="[151,477,1133,1156]" gridcol="0" gridrow="2" pageId="5" pageNumber="51">Head height</th>
<td id="76EF9642CA590042FD368213FD24FB1A" box="[688,748,1133,1156]" gridcol="1" gridrow="2" pageId="5" pageNumber="51">0.255</td>
<td id="76EF9642CA590042FCD58213FC50FB1A" box="[851,920,1133,1156]" gridcol="2" gridrow="2" pageId="5" pageNumber="51">0.548</td>
<td id="76EF9642CA590042FC748213FBE6FB1A" box="[1010,1070,1133,1156]" gridcol="3" gridrow="2" pageId="5" pageNumber="51">0.345</td>
<td id="76EF9642CA590042FB148213FA54FB1A" box="[1170,1436,1133,1156]" gridcol="4" gridrow="2" pageId="5" pageNumber="51">0.741</td>
</tr>
<tr id="353EFF3ECA590042FF1182E6FA54FB47" box="[151,1436,1176,1241]" gridrow="3" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF1182E6FE15FB47" box="[151,477,1176,1241]" gridcol="0" gridrow="3" pageId="5" pageNumber="51">Forelimb length Hindlimb length</th>
<td id="76EF9642CA590042FD3682E6FD24FB47" box="[688,748,1176,1241]" gridcol="1" gridrow="3" pageId="5" pageNumber="51">0.601 0.181</td>
<td id="76EF9642CA590042FCD582E6FC50FB47" box="[851,920,1176,1241]" gridcol="2" gridrow="3" pageId="5" pageNumber="51">1.762 4.041</td>
<td id="76EF9642CA590042FC7482E6FBE6FB47" box="[1010,1070,1176,1241]" gridcol="3" gridrow="3" pageId="5" pageNumber="51">0.282 0.044</td>
<td id="76EF9642CA590042FB1482E6FA54FB47" box="[1170,1436,1176,1241]" gridcol="4" gridrow="3" pageId="5" pageNumber="51">0.827 0.985</td>
</tr>
<tr id="353EFF3ECA590042FF118293FA54FA9A" box="[151,1436,1261,1284]" gridrow="4" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF118293FE15FA9A" box="[151,477,1261,1284]" gridcol="0" gridrow="4" pageId="5" pageNumber="51">Snout-vent length</th>
<td id="76EF9642CA590042FD368293FD24FA9A" box="[688,748,1261,1284]" gridcol="1" gridrow="4" pageId="5" pageNumber="51">3.286</td>
<td id="76EF9642CA590042FCD58293FC50FA9A" box="[851,920,1261,1284]" gridcol="2" gridrow="4" pageId="5" pageNumber="51">1.465</td>
<td id="76EF9642CA590042FC748293FBE6FA9A" box="[1010,1070,1261,1284]" gridcol="3" gridrow="4" pageId="5" pageNumber="51">0.902</td>
<td id="76EF9642CA590042FB148293FA54FA9A" box="[1170,1436,1261,1284]" gridcol="4" gridrow="4" pageId="5" pageNumber="51">0.402</td>
</tr>
<tr id="353EFF3ECA590042FF118366FA54FAC7" box="[151,1436,1304,1369]" gridrow="5" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF118366FE15FAC7" box="[151,477,1304,1369]" gridcol="0" gridrow="5" pageId="5" pageNumber="51">Jaw lenght Interorbital distance</th>
<td id="76EF9642CA590042FD368366FD24FAC7" box="[688,748,1304,1369]" gridcol="1" gridrow="5" pageId="5" pageNumber="51">0.669 0.171</td>
<td id="76EF9642CA590042FCD58366FC50FAC7" box="[851,920,1304,1369]" gridcol="2" gridrow="5" pageId="5" pageNumber="51">0.239 0.125</td>
<td id="76EF9642CA590042FC748366FBE6FAC7" box="[1010,1070,1304,1369]" gridcol="3" gridrow="5" pageId="5" pageNumber="51">0.663 0.590</td>
<td id="76EF9642CA590042FB148366FA54FAC7" box="[1170,1436,1304,1369]" gridcol="4" gridrow="5" pageId="5" pageNumber="51">0.237 0.432</td>
</tr>
<tr id="353EFF3ECA590042FF118313FA54FA1A" box="[151,1436,1389,1412]" gridrow="6" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF118313FE15FA1A" box="[151,477,1389,1412]" gridcol="0" gridrow="6" pageId="5" pageNumber="51">Ear opening maximum length</th>
<td id="76EF9642CA590042FD368313FD24FA1A" box="[688,748,1389,1412]" gridcol="1" gridrow="6" pageId="5" pageNumber="51">0.007</td>
<td id="76EF9642CA590042FCD58313FC50FA1A" box="[851,920,1389,1412]" gridcol="2" gridrow="6" pageId="5" pageNumber="51">0.185</td>
<td id="76EF9642CA590042FC748313FBE6FA1A" box="[1010,1070,1389,1412]" gridcol="3" gridrow="6" pageId="5" pageNumber="51">0.022</td>
<td id="76EF9642CA590042FB148313FA54FA1A" box="[1170,1436,1389,1412]" gridcol="4" gridrow="6" pageId="5" pageNumber="51">0.605</td>
</tr>
<tr id="353EFF3ECA590042FF1183E6FA54FA47" box="[151,1436,1432,1497]" gridrow="7" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF1183E6FE15FA47" box="[151,477,1432,1497]" gridcol="0" gridrow="7" pageId="5" pageNumber="51">Axilla-groin distance Internasal distance</th>
<td id="76EF9642CA590042FD3683E6FD24FA47" box="[688,748,1432,1497]" gridcol="1" gridrow="7" pageId="5" pageNumber="51">1.801 0.145</td>
<td id="76EF9642CA590042FCD583E6FC50FA47" box="[851,920,1432,1497]" gridcol="2" gridrow="7" pageId="5" pageNumber="51">-0.430 -0.003</td>
<td id="76EF9642CA590042FC7483E6FBE6FA47" box="[1010,1070,1432,1497]" gridcol="3" gridrow="7" pageId="5" pageNumber="51">0.794 0.673</td>
<td id="76EF9642CA590042FB1483E6FA54FA47" box="[1170,1436,1432,1497]" gridcol="4" gridrow="7" pageId="5" pageNumber="51">-0.189 -0.014</td>
</tr>
<tr id="353EFF3ECA590042FF118393FA54F99A" box="[151,1436,1517,1540]" gridrow="8" pageId="5" pageNumber="51">
<th id="76EF9642CA590042FF118393FE15F99A" box="[151,477,1517,1540]" gridcol="0" gridrow="8" pageId="5" pageNumber="51">Snout length</th>
<td id="76EF9642CA590042FD368393FD24F99A" box="[688,748,1517,1540]" gridcol="1" gridrow="8" pageId="5" pageNumber="51">0.455</td>
<td id="76EF9642CA590042FCD58393FC50F99A" box="[851,920,1517,1540]" gridcol="2" gridrow="8" pageId="5" pageNumber="51">0.278</td>
<td id="76EF9642CA590042FC748393FBE6F99A" box="[1010,1070,1517,1540]" gridcol="3" gridrow="8" pageId="5" pageNumber="51">0.698</td>
<td id="76EF9642CA590042FB148393FA54F99A" box="[1170,1436,1517,1540]" gridcol="4" gridrow="8" pageId="5" pageNumber="51">0.427</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB1FD7CCA59FFB8FF438043FCBFF8A9" blockId="5.[151,1437,1597,2034]" pageId="5" pageNumber="51">Dorsal scales swollen and unicarinate, with no enlarged middorsal row, 10 (912) longitudinal rows in 5% of SVL; flank scales separated by granules (juxtaposed); ventral scales imbricate, separated or juxtaposed, smooth, with 10 (811) longitudinal rows in 5% of SVL; ventrals larger than dorsals; toepads overlap the first phalanx in all toes; seventeen (1623) lamellae under second and third phalanges of fourth toe, supradigitals multicarinate; tail weakly compressed; a pair of enlarged postanals in males, separated from cloacal opening by two (12) scales; dewlap large, extending posteriorly to the level of forearms (absent in females), with longitudinal rows of 34 (36) scales separated by naked skin.</paragraph>
<paragraph id="8BB1FD7CCA59FFBBFF43813CFF37FE29" blockId="5.[151,1437,1597,2034]" lastBlockId="6.[151,1437,152,2007]" lastPageId="6" lastPageNumber="52" pageId="5" pageNumber="51">
<emphasis id="B97A216ECA59FFB8FF43813CFE9BF8C2" bold="true" box="[197,339,1858,1884]" pageId="5" pageNumber="51">Osteology.</emphasis>
Skull. Parietal roof flat; parietal crests V-shaped; dorsal surface of skull smooth; no crenulation along lateral edges of parietal; no black pigment on skull; parietal extending over supraoccipital (Etheridge's [1959] &quot;half funnel&quot;); supraoccipital cresting continuous; pineal foramen absent; postfrontal present, reduced; prefrontal in narrow contact with nasal; frontal articulates with nasals anteriorly; anterior edge of nasal forms posterior border of naris; dorsal process of jugal terminates on lateral aspect of postorbital; contact between jugal and squamosal absent; posterior aspect of jugal straight; epipterygoid in contact with parietal; pterygoid and palatine teeth absent; lateral edges of vomer smooth, posterior edges of vomer straight (QCAZ 2052) or pronged (QCAZ 3872); maxilla articulates with ectopterygoid, posterior part of maxilla thick; lateral shelf of quadrate absent; posterolateral process of quadrate present (only in QCAZ 2052), superior fossa of quadrate relatively small, a process of the squamosal fitting into the fossa like a peg in a hole; premaxilla overlaps nasals laterally; basipterygoid with small and well developed spheno-occipital tubercles; large lacrimal in comparison with the lateral aspect of prefrontal; posterior part of postorbital pronged.
</paragraph>
<paragraph id="8BB1FD7CCA5AFFBBFF4387BCFB0DFD09" blockId="6.[151,1437,152,2007]" pageId="6" pageNumber="52">Mandible. No jaw sculpturing in males; mandibular tooth row extends posteriorly to level of anterior inferior alveolar foramen; angular process of articular large; dentary large, extending within mandibular fossa; posterior edge of dentary pronged; splenial large and posterior to posterior edge of dentary (only in QCAZ 2052); anterior mylohyoid foramen absent; ventral aspect of anteromedial process of coronoid projects posteriorly; labial process of coronoid present; surangular foramen completely within surangular; posterolateral aspect of coronoid terminates anterior to surangular foramen; angular absent.</paragraph>
<paragraph id="8BB1FD7CCA5AFFBBFF4384DCFDC9FCB2" blockId="6.[151,1437,152,2007]" pageId="6" pageNumber="52">Postcranial skeleton. Four pairs of postxiphisternal ribs; two pairs of sternal ribs; three pairs of xiphisternal ribs; interclavicles arrow-shaped; clavicle with distal flanges; 2223 presacral vertebrae; three lumbar vertebrae; caudal vertebrae with transverse anterior process lost posteriorly (i.e., alpha condition); caudal autotomy septa present.</paragraph>
<paragraph id="8BB1FD7CCA5AFFBBFF438549FEABFBAC" blockId="6.[151,1437,152,2007]" pageId="6" pageNumber="52">
<emphasis id="B97A216ECA5AFFBBFF438549FDDCFCCF" bold="true" box="[197,532,823,849]" pageId="6" pageNumber="52">Coloration in preservative.</emphasis>
Male pattern I (67% of male specimens examined). Adult male QCAZ 3796: Head, body, limbs and tail grayish purple; four dark brown transverse blotches on flanks; limbs and tail with wide brown blotches; sides of head with a dark brown stripe extending from nares to nape with a white stripe through the ear opening; side of neck with a black blotch containing a small creamy-violet spot in the center; chin and throat mottled with cream and grayish violet; chest and abdomen cream, mottled toward the flanks with light brown; ventral surface of limbs cream, mottled with grayish brown; dewlap cream with grayishviolet scale rows.
</paragraph>
<paragraph id="8BB1FD7CCA5AFFBBFF438243FD5EFA1C" blockId="6.[151,1437,152,2007]" pageId="6" pageNumber="52">
Male pattern II (33% of male specimens examined). Adult male QCAZ 2051 (
<typeStatus id="54B543DECA5AFFBBFB058243FB3DFBC9" box="[1155,1269,1085,1111]" pageId="6" pageNumber="52" type="holotype">holotype</typeStatus>
): Head, body, limbs and tail whitish gray with coppery brown reticulation; limbs with grayish-purple bands and grayishbrown reticulation; tail with coppery brown bands; sides of head with a dark coppery-brown stripe extending from nares to nape and a white stripe through the ear opening; side of neck with a black blotch containing two white spots; flanks with small coppery-brown reticulations; belly cream with grayish-purple blotches; part of ventral surface of limbs yellowish cream with gray reticulations; anterior part of ventral surface of tail pinkish cream with grayish-purple blotches; dewlap cream with pinkish-white scales; ocular scales white. Another adult male (QCAZ 3129) had a pattern similar to the
<typeStatus id="54B543DECA5AFFBBFCC5833CFC7AFAC2" box="[835,946,1346,1372]" pageId="6" pageNumber="52" type="holotype">holotype</typeStatus>
but differed by lacking a black blotch containing white spots on the sides of neck.
</paragraph>
<paragraph id="8BB1FD7CCA5AFFBBFF4383F3FB79F989" blockId="6.[151,1437,152,2007]" pageId="6" pageNumber="52">
Female pattern I (89% of female specimens examined): Adult female QCAZ 6394. Head, body, limbs and tail purplish gray; middorsum with four dark inverted V-shaped blotches, descending to flanks; limbs and tail dark brown banded; side of head with a dark brown stripe, similar to the
<typeStatus id="54B543DECA5AFFBBFC7483A6FB93FA6C" box="[1010,1115,1496,1522]" pageId="6" pageNumber="52" type="holotype">holotype</typeStatus>
, but extending onto flanks; sides of neck cream spotted with purplish brown; belly with purplish-brown reticulations.
</paragraph>
<paragraph id="8BB1FD7CCA5AFFBBFF43805CFBA0F932" blockId="6.[151,1437,152,2007]" pageId="6" pageNumber="52">Female pattern II (11% of female specimens examined). Adult female QCAZ 2781: Head, body, limbs and tail purplish gray; middorsum with a longitudinal cream stripe delineated with brown from nape to tail; limbs with brown bands; side of head with dark brown stripe; sides of neck and flanks spotted dark brown; chin, throat, belly and part ventral of limbs and tail cream with brown reticulations.</paragraph>
<paragraph id="8BB1FD7CCA5AFFBBFF4380C9FE57F812" blockId="6.[151,1437,152,2007]" pageId="6" pageNumber="52">
<emphasis id="B97A216ECA5AFFBBFF4380C9FE6BF94F" bold="true" box="[197,419,1719,1745]" pageId="6" pageNumber="52">Coloration in life.</emphasis>
Adult male QCAZ 4025: Head green with pale brown and gray blotches; white stripe extending from lips to neck; black blotch with whitish cream spots on each side of neck; limbs greenish brown with dark gray bands; flanks greenish brown with pale brown blotches aligned in transverse stripes; throat white with pale green spots; dewlap pale-yellow with pale-green gorgetals rows and white sternals rows; belly whitish with pale gray small spots; tail grayish green with dark gray bands proximally and greenish yellow distally; iris dark blue.
</paragraph>
<paragraph id="8BB1FD7CCA5AFFBBFF4381E6FF13F849" blockId="6.[151,1437,152,2007]" pageId="6" pageNumber="52">Adult male QCAZ 3129: Dewlap white, greenish yellow anteriorly; chin and throat white with pale green spots.</paragraph>
<paragraph id="8BB1FD7CCA5BFFBAFF4386E6FEC4FEBC" blockId="7.[151,1437,152,402]" pageId="7" pageNumber="53">Adult female QCAZ 3872: Each side of the head with a dark brown stripe from nasals to nape; sides of neck and shoulder with dark brown blotches; body, limbs and tail greenish cream; four dorsal, transverse, dark brown blotches descending to flanks at midbody; limbs with dark brown bands; tail incompletely banded; iris dark blue.</paragraph>
</subSubSection>
<subSubSection id="C314AEF7CA5BFFB5FF408753FAC9FA5F" lastPageId="8" lastPageNumber="54" pageId="7" pageNumber="53" type="distribution">
<paragraph id="8BB1FD7CCA5BFFBAFF408753FAC0FE0C" blockId="7.[151,1437,152,402]" pageId="7" pageNumber="53">
<emphasis id="B97A216ECA5BFFBAFF408753FE37FED9" bold="true" box="[198,511,301,327]" pageId="7" pageNumber="53">Distribution and ecology.</emphasis>
<taxonomicName id="4C0E86FFCA5BFFBAFD808753FD7EFED8" box="[518,694,301,326]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="53" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA5BFFBAFD808753FD7EFED8" box="[518,694,301,326]" italics="true" pageId="7" pageNumber="53">Anolis otongae</emphasis>
</taxonomicName>
inhabits the western slopes of the western Andean cordillera in
<collectingCountry id="F319BDECCA5BFFBAFF11872CFF37FEF2" box="[151,255,338,364]" name="Ecuador" pageId="7" pageNumber="53">Ecuador</collectingCountry>
between
<quantity id="4CF65099CA5BFFBAFEFF872CFDE0FEF2" box="[377,552,338,364]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="7" pageNumber="53" unit="m" value="2150.0" valueMax="2500.0" valueMin="1800.0">18002500 m</quantity>
in elevation (
<figureCitation id="1335E1F9CA5BFFBAFD5E872CFCEEFEF2" box="[728,806,338,364]" captionStart="FIGURE 3" captionStartId="7.[151,255,1986,2010]" captionTargetBox="[186,1378,503,1985]" captionTargetId="figure@7.[158,1429,455,2029]" captionTargetPageId="7" captionText="FIGURE 3. Distribution of Anolis otongae (circles) and A. gemmosus (triangles) in Ecuador." httpUri="https://zenodo.org/record/197525/files/figure.png" pageId="7" pageNumber="53">Fig. 3</figureCitation>
). It occurs in Provincias Cotopaxi and Pichincha. According to
<bibRefCitation id="EF9F808DCA5BFFBAFEBE8706FE1EFE0C" author="Sierra" box="[312,470,376,402]" pageId="7" pageNumber="53" refString="Sierra, R. (1999) Propuesta preliminary de un sistema de clasificacion de vegetacion para el Ecuador continental. Proyecto INEFAN / GEF-BIRF y EcoCiencia. Quito, Ecuador." type="book" year="1999">Sierra (1999)</bibRefCitation>
, the
<typeStatus id="54B543DECA5BFFBAFD8B8706FD88FE0C" box="[525,576,376,402]" pageId="7" pageNumber="53">type</typeStatus>
locality of
<taxonomicName id="4C0E86FFCA5BFFBAFD408706FC8CFE0F" box="[710,836,376,401]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="53" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA5BFFBAFD408706FC8CFE0F" box="[710,836,376,401]" italics="true" pageId="7" pageNumber="53">A. otongae</emphasis>
</taxonomicName>
corresponds to mountain cloud forest.
</paragraph>
<caption id="DF71ADF4CA5BFFBAFF1181BCFBB1F844" box="[151,1145,1986,2010]" httpUri="https://zenodo.org/record/197525/files/figure.png" pageId="7" pageNumber="53" targetBox="[186,1378,503,1985]" targetPageId="7">
<paragraph id="8BB1FD7CCA5BFFBAFF1181BCFBB1F844" blockId="7.[151,1145,1986,2010]" box="[151,1145,1986,2010]" pageId="7" pageNumber="53">
<emphasis id="B97A216ECA5BFFBAFF1181BCFED1F844" bold="true" box="[151,281,1986,2010]" pageId="7" pageNumber="53">FIGURE 3.</emphasis>
Distribution of
<taxonomicName id="4C0E86FFCA5BFFBAFE4281BDFDADF844" box="[452,613,1987,2010]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="53" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA5BFFBAFE4281BDFDADF844" box="[452,613,1987,2010]" italics="true" pageId="7" pageNumber="53">Anolis otongae</emphasis>
</taxonomicName>
(circles) and
<taxonomicName id="4C0E86FFCA5BFFBAFD7081BDFC4BF844" box="[758,899,1987,2010]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="53" phylum="Chordata" rank="species" species="gemmosus">
<emphasis id="B97A216ECA5BFFBAFD7081BDFC4BF844" box="[758,899,1987,2010]" italics="true" pageId="7" pageNumber="53">A. gemmosus</emphasis>
</taxonomicName>
(triangles) in Ecuador.
</paragraph>
</caption>
<caption id="DF71ADF4CA54FFB5FF118368FAA0FAB0" box="[151,1384,1302,1326]" httpUri="https://zenodo.org/record/197526/files/figure.png" pageId="8" pageNumber="54" targetBox="[159,1435,187,1289]" targetPageId="8">
<paragraph id="8BB1FD7CCA54FFB5FF118368FAA0FAB0" blockId="8.[151,1384,1302,1326]" box="[151,1384,1302,1326]" pageId="8" pageNumber="54">
<emphasis id="B97A216ECA54FFB5FF118368FED1FAB0" bold="true" box="[151,281,1302,1326]" pageId="8" pageNumber="54">FIGURE 4.</emphasis>
Distribution of
<taxonomicName id="4C0E86FFCA54FFB5FE428369FDB6FAB0" box="[452,638,1303,1326]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="gemmosus">
<emphasis id="B97A216ECA54FFB5FE428369FDB6FAB0" box="[452,638,1303,1326]" italics="true" pageId="8" pageNumber="54">Anolis gemmosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E86FFCA54FFB5FD348369FCEDFAB0" box="[690,805,1303,1326]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA54FFB5FD348369FCEDFAB0" box="[690,805,1303,1326]" italics="true" pageId="8" pageNumber="54">A. otongae</emphasis>
</taxonomicName>
along the first and second principal components axes.
</paragraph>
</caption>
<paragraph id="8BB1FD7CCA54FFB5FF438323FAC9FA5F" blockId="8.[151,1438,1372,2028]" pageId="8" pageNumber="54">
<taxonomicName id="4C0E86FFCA54FFB5FF438323FEB2FAE8" box="[197,378,1373,1398]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA54FFB5FF438323FEB2FAE8" box="[197,378,1373,1398]" italics="true" pageId="8" pageNumber="54">Anolis otongae</emphasis>
</taxonomicName>
was found in primary and secondary forest, grasslands, open forest, and riparian forest. Many specimens were collected at night on low vegetation, sleeping on leaves (
<taxonomicName id="4C0E86FFCA54FFB5FBC983FCFB19FA02" box="[1103,1233,1410,1436]" class="Magnoliopsida" family="Piperaceae" kingdom="Plantae" order="Piperales" pageId="8" pageNumber="54" phylum="Tracheophyta" rank="family">Piperaceae</taxonomicName>
and
<taxonomicName id="4C0E86FFCA54FFB5FA8B83FCFA5CFA02" box="[1293,1428,1410,1436]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="8" pageNumber="54" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
) and ferns. One specimen was collected on a cloudy day on
<taxonomicName id="4C0E86FFCA54FFB5FCCA83D9FB35FA5F" box="[844,1277,1447,1473]" class="Polypodiopsida" family="Thelypteridaceae" genus="Macrothelypteris" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="54" phylum="Tracheophyta" rank="genus">
<emphasis id="B97A216ECA54FFB5FCCA83D9FBDDFA5E" box="[844,1045,1447,1472]" italics="true" pageId="8" pageNumber="54">Macrothelypteris</emphasis>
(Thelypteridaceae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C314AEF7CA54FFB5FF4383B3FE46F872" pageId="8" pageNumber="54" type="reference_group">
<paragraph id="8BB1FD7CCA54FFB5FF4383B3FA82F8A8" blockId="8.[151,1438,1372,2028]" pageId="8" pageNumber="54">
<taxonomicName id="4C0E86FFCA54FFB5FF4383B3FEB6FA78" box="[197,382,1485,1510]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA54FFB5FF4383B3FEB6FA78" box="[197,382,1485,1510]" italics="true" pageId="8" pageNumber="54">Anolis otongae</emphasis>
</taxonomicName>
occurs in sympatry with
<taxonomicName id="4C0E86FFCA54FFB5FD3183B3FC90FA78" box="[695,856,1485,1510]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="gemmosus">
<emphasis id="B97A216ECA54FFB5FD3183B3FC90FA78" box="[695,856,1485,1510]" italics="true" pageId="8" pageNumber="54">A. gemmosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E86FFCA54FFB5FC1383B3FB93FA78" box="[917,1115,1485,1510]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="aequatorialis">
<emphasis id="B97A216ECA54FFB5FC1383B3FB93FA78" box="[917,1115,1485,1510]" italics="true" pageId="8" pageNumber="54">A. aequatorialis</emphasis>
</taxonomicName>
in the Reserva de Bosque Integral Otonga, Provincia Cotopaxi,
<collectingCountry id="F319BDECCA54FFB5FDD1838CFD74F992" box="[599,700,1522,1548]" name="Ecuador" pageId="8" pageNumber="54">Ecuador</collectingCountry>
.
<taxonomicName id="4C0E86FFCA54FFB5FD4E838CFC5EF995" box="[712,918,1522,1547]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="gemmosus">
<emphasis id="B97A216ECA54FFB5FD4E838CFC5EF995" box="[712,918,1522,1547]" italics="true" pageId="8" pageNumber="54">Anolis gemmosus</emphasis>
</taxonomicName>
is distributed from southwestern
<collectingCountry id="F319BDECCA54FFB5FAA1838CFA54F992" box="[1319,1436,1522,1548]" name="Colombia" pageId="8" pageNumber="54">Colombia</collectingCountry>
(Departmento Nariño, Reserva Natural La Planada) to the north part of the western Andes of
<collectingCountry id="F319BDECCA54FFB5FAB38069FA55F9AF" box="[1333,1437,1559,1585]" name="Ecuador" pageId="8" pageNumber="54">Ecuador</collectingCountry>
(Provincias Carchi, Imbabura, Pichincha, and Cotopaxi), between
<quantity id="4CF65099CA54FFB5FB878042FB7EF9C8" box="[1025,1206,1596,1622]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.3" metricValueMin="1.3" pageId="8" pageNumber="54" unit="m" value="1800.0" valueMax="2300.0" valueMin="1300.0">13002300 m</quantity>
elevation.
<taxonomicName id="4C0E86FFCA54FFB5FACC8043FEF0F9E5" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="aequatorialis">
<emphasis id="B97A216ECA54FFB5FACC8043FEF0F9E5" italics="true" pageId="8" pageNumber="54">Anolis aequatorialis</emphasis>
</taxonomicName>
was described by Werner in 1894 with no more specific locality than “
<collectingCountry id="F319BDECCA54FFB5FB1C801CFAC0F9E2" box="[1178,1288,1634,1660]" name="Ecuador" pageId="8" pageNumber="54">Ecuador</collectingCountry>
”.
<bibRefCitation id="EF9F808DCA54FFB5FA98801CFE60F93F" author="Peters" pageId="8" pageNumber="54" refString="Peters, J. A. &amp; Donoso-Barros, R. (1970) Catalogue of the Neotropical Squamata: Part II. Lizards and Amphisbaenians. Bulletin of United States National Museum, 297, 293 pp." type="book" year="1970">Peters and Donoso-Barros (1970)</bibRefCitation>
described its geographic range as “middle altitudes of western slopes in
<collectingCountry id="F319BDECCA54FFB5FAA480F9FA45F93F" box="[1314,1421,1671,1697]" name="Ecuador" pageId="8" pageNumber="54">Ecuador</collectingCountry>
”, whereas
<bibRefCitation id="EF9F808DCA54FFB5FF7880D2FD98F958" author="Uetz" box="[254,592,1708,1734]" pageId="8" pageNumber="54" refString="Uetz, P. &amp; Hallermann, J. (2008) The Reptile Database. Available from http: // www. reptile-database. org (accessed 21 April 2009)." type="journal article" year="2008">Uetz and Hallermann (2008)</bibRefCitation>
extended the range of
<taxonomicName id="4C0E86FFCA54FFB5FCDB80D3FBDAF958" box="[861,1042,1709,1734]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="aequatoralis">
<emphasis id="B97A216ECA54FFB5FCDB80D3FBDAF958" box="[861,1042,1709,1734]" italics="true" pageId="8" pageNumber="54">A. aequatoralis</emphasis>
</taxonomicName>
into “
<collectingCountry id="F319BDECCA54FFB5FBDF80D2FB18F958" box="[1113,1232,1708,1734]" name="Colombia" pageId="8" pageNumber="54">Colombia</collectingCountry>
[Castro, F. (pers. comm.)]”. One of us (J.A.V.), examined material of
<taxonomicName id="4C0E86FFCA54FFB5FC9A80ACFCA2F975" box="[796,874,1746,1771]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="genus">
<emphasis id="B97A216ECA54FFB5FC9A80ACFCA2F975" box="[796,874,1746,1771]" italics="true" pageId="8" pageNumber="54">Anolis</emphasis>
</taxonomicName>
deposited in the collection of Universidad de Nariño,
<collectingCountry id="F319BDECCA54FFB5FF718089FEA6F88F" box="[247,366,1783,1809]" name="Colombia" pageId="8" pageNumber="54">Colombia</collectingCountry>
, and found several specimens of
<taxonomicName id="4C0E86FFCA54FFB5FD748089FC29F88E" box="[754,993,1783,1808]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="aequatorialis">
<emphasis id="B97A216ECA54FFB5FD748089FC29F88E" box="[754,993,1783,1808]" italics="true" pageId="8" pageNumber="54">Anolis aequatorialis</emphasis>
</taxonomicName>
from Reserva Ñambi and Reserva La Planada, confirming the occurrence of
<taxonomicName id="4C0E86FFCA54FFB5FDD88163FCD3F8A8" box="[606,795,1821,1846]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="54" phylum="Chordata" rank="species" species="aequatorialis">
<emphasis id="B97A216ECA54FFB5FDD88163FCD3F8A8" box="[606,795,1821,1846]" italics="true" pageId="8" pageNumber="54">A. aequatorialis</emphasis>
</taxonomicName>
in
<collectingCountry id="F319BDECCA54FFB5FCC78162FC7DF8A8" box="[833,949,1820,1846]" name="Colombia" pageId="8" pageNumber="54">Colombia</collectingCountry>
(See appendix 1 for locality data).
</paragraph>
<paragraph id="8BB1FD7CCA54FFB5FF40813FFE46F872" blockId="8.[151,1438,1372,2028]" pageId="8" pageNumber="54">
<emphasis id="B97A216ECA54FFB5FF40813FFE45F8C5" bold="true" box="[198,397,1857,1883]" pageId="8" pageNumber="54">Natural history.</emphasis>
An adult female (QCAZ 6396) collected on
<date id="FFB0DBBCCA54FFB5FC18813CFB88F8C2" box="[926,1088,1858,1884]" pageId="8" pageNumber="54" value="2003-05-01">May 1st 2003</date>
had two oviductal eggs (16.9 x
<quantity id="4CF65099CA54FFB5FF298119FEC6F81F" box="[175,270,1895,1921]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="54" unit="mm" value="7.5">7.5 mm</quantity>
and 8.3 x
<quantity id="4CF65099CA54FFB5FE108119FE32F81F" box="[406,506,1895,1921]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="54" unit="mm" value="6.2">6.2 mm</quantity>
; 496.7 and
<quantity id="4CF65099CA54FFB5FD0A8119FCC3F81F" box="[652,779,1895,1921]" metricMagnitude="-1" metricUnit="m" metricValue="1.6669999999999998" pageId="8" pageNumber="54" unit="mm" value="166.7">166.7 mm</quantity>
3, respectively). Another adult female (QCAZ 6219) collected on
<date id="FFB0DBBCCA54FFB5FEAB81F4FDE0F83A" box="[301,552,1930,1956]" pageId="8" pageNumber="54" value="2007-11-17">November 17th 2007</date>
also had two oviductal eggs (18.6 x
<quantity id="4CF65099CA54FFB5FC5F81F4FBFBF83A" box="[985,1075,1930,1956]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="8" pageNumber="54" unit="mm" value="8.3">8.3 mm</quantity>
and 18.2 x
<quantity id="4CF65099CA54FFB5FB3881F4FAD4F83A" box="[1214,1308,1930,1956]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="8" pageNumber="54" unit="mm" value="7.1">7.1 mm</quantity>
; 669.5 and
<quantity id="4CF65099CA54FFB5FF1181CEFEC7F854" box="[151,271,1968,1994]" metricMagnitude="-1" metricUnit="m" metricValue="4.794" pageId="8" pageNumber="54" unit="mm" value="479.4">479.4 mm</quantity>
3, respectively). The smallest juvenile (QCAZ 3706) was collected in
<date id="FFB0DBBCCA54FFB5FBCC81CEFB11F854" box="[1098,1241,1968,1994]" pageId="8" pageNumber="54" value="1997-03">March 1997</date>
(
<quantity id="4CF65099CA54FFB5FB6C81CEFA9DF854" box="[1258,1365,1968,1994]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="8" pageNumber="54" unit="mm" value="28.3">28.3 mm</quantity>
SVL,
<quantity id="4CF65099CA54FFB5FF1181ACFF37F872" box="[151,255,2002,2028]" metricMagnitude="-2" metricUnit="m" metricValue="6.44" pageId="8" pageNumber="54" unit="mm" value="64.4">64.4 mm</quantity>
tail length).
</paragraph>
</subSubSection>
<subSubSection id="C314AEF7CA55FFB4FF4086E9FEBDFEF2" pageId="9" pageNumber="55" type="etymology">
<paragraph id="8BB1FD7CCA55FFB4FF4086E9FEBDFEF2" blockId="9.[151,1437,151,364]" pageId="9" pageNumber="55">
<emphasis id="B97A216ECA55FFB4FF4086E9FE9BFF2F" bold="true" box="[198,339,151,177]" pageId="9" pageNumber="55">Etymology.</emphasis>
The epithet
<taxonomicName id="4C0E86FFCA55FFB4FE7E86E6FD9EFF2F" box="[504,598,152,177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="55" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA55FFB4FE7E86E6FD9EFF2F" box="[504,598,152,177]" italics="true" pageId="9" pageNumber="55">otongae</emphasis>
</taxonomicName>
refers to Reserva de Bosque Integral Otonga, located in Provincia Cotopaxi,
<collectingCountry id="F319BDECCA55FFB4FE9186C3FEB7FF49" box="[279,383,189,215]" name="Ecuador" pageId="9" pageNumber="55">Ecuador</collectingCountry>
. The majority of specimens of
<taxonomicName id="4C0E86FFCA55FFB4FD7886C3FC4BFF48" box="[766,899,189,214]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="55" phylum="Chordata" rank="species" species="otongae">
<emphasis id="B97A216ECA55FFB4FD7886C3FC4BFF48" box="[766,899,189,214]" italics="true" pageId="9" pageNumber="55">A. otongae</emphasis>
</taxonomicName>
were collected there. The indigenous name “Otonga” was inspired by either the giant earthworm
<emphasis id="B97A216ECA55FFB4FC95869CFC70FF65" box="[787,952,226,251]" italics="true" pageId="9" pageNumber="55">Onoreodrillus</emphasis>
spp. or by limbless amphibians
<taxonomicName id="4C0E86FFCA55FFB4FABE869CFE44FEBC" authority="Gunther" authorityName="Gunther" class="Amphibia" family="Caeciliidae" genus="Caecilia" kingdom="Animalia" order="Gymnophiona" pageId="9" pageNumber="55" phylum="Chordata" rank="species" species="pachynema">
<emphasis id="B97A216ECA55FFB4FABE869CFEEAFEBF" italics="true" pageId="9" pageNumber="55">Caecilia pachynema</emphasis>
Günther
</taxonomicName>
,
<taxonomicName id="4C0E86FFCA55FFB4FE1A8776FDB1FEBC" authority="Dunn" authorityName="Dunn" box="[412,633,264,290]" class="Amphibia" family="Caeciliidae" genus="Caecilia" kingdom="Animalia" order="Gymnophiona" pageId="9" pageNumber="55" phylum="Chordata" rank="species" species="guntheri">
<emphasis id="B97A216ECA55FFB4FE1A8776FDE1FEBF" box="[412,553,264,289]" italics="true" pageId="9" pageNumber="55">C. guntheri</emphasis>
Dunn
</taxonomicName>
, and
<taxonomicName id="4C0E86FFCA55FFB4FD3D8776FB5BFEBC" authority="Boulenger. These" authorityName="Boulenger. These" box="[699,1171,264,290]" class="Amphibia" family="Rhinatrematidae" genus="Epicrionops" kingdom="Animalia" order="Gymnophiona" pageId="9" pageNumber="55" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B97A216ECA55FFB4FD3D8776FC7BFEBF" box="[699,947,264,289]" italics="true" pageId="9" pageNumber="55">Epicrionops bicolor</emphasis>
Boulenger. These
</taxonomicName>
animals are the most conspicuous and striking of this wildlife reserve, sharing burrowing habits and considered uneatable food by indigenous people.
</paragraph>
</subSubSection>
</treatment>
</document>