366 lines
44 KiB
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366 lines
44 KiB
XML
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<mods:title id="2121B89256C18264474927C05E5F5207">The taxonomy of Heteropanope indica De Man, 1887 from the Indian Ocean, with a note on Pilumnopeus pearsei (Rathbun, 1932) (Crustacea: Decapoda: Brachyura: Pilumnidae)</mods:title>
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<mods:namePart id="30C08D9B276DA138E6B64B338EE3D950">Patel, Krupal</mods:namePart>
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<mods:title id="5FC3AF20B4B706AED56195AAEA932760">Zootaxa</mods:title>
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<heading id="D0F8BC48FF81C20AFF6CFCA692F5FCC3" bold="true" box="[151,709,799,826]" fontSize="11" level="1" pageId="1" pageNumber="385" reason="1">
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<emphasis id="B97BD736FF81C20AFF6CFCA692F5FCC3" bold="true" box="[151,709,799,826]" pageId="1" pageNumber="385">
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Genus
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<taxonomicName id="4C0F70A7FF81C20AFF15FCA692F5FCC3" ID-CoL="7PTLN" authority="A. Milne-Edwards, 1867" authorityName="A. Milne-Edwards" authorityYear="1867" box="[238,709,799,826]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
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<emphasis id="B97BD736FF81C20AFF15FCA691BEFCC0" bold="true" box="[238,398,799,826]" italics="true" pageId="1" pageNumber="385">Pilumnopeus</emphasis>
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A.
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<bibRefCitation id="EF9E76D5FF81C20AFE43FCA692F5FCC3" author="Milne-Edwards, A." box="[440,709,799,825]" pageId="1" pageNumber="385" pagination="263 - 288" refId="ref7922" refString="Milne-Edwards, A. (1867) Descriptions de quelques especes nouvelles de Crustaces Brachyoures, Tribu des Oxyrhinques. Familie des Maiens. Annales de la Societe Entomologique de France, Series 4, 7, 263 - 288." type="journal article" year="1867">Milne-Edwards, 1867</bibRefCitation>
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<taxonomicName id="4C0F70A7FF81C20AFF6CFCDD9267FC86" ID-CoL="7PTLN" authority="A. Milne Edwards, 1867: 277" authorityName="A. Milne Edwards" authorityPageNumber="277" authorityYear="1867" box="[151,599,867,892]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
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<emphasis id="B97BD736FF81C20AFF6CFCDD912DFC86" box="[151,285,868,892]" italics="true" pageId="1" pageNumber="385">Pilumnopeus</emphasis>
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A. Milne Edwards, 1867: 277
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(
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<paragraph id="8BB00B24FF81C20AFD9FFCDD91D7FC22" blockId="1.[151,1437,867,1045]" pageId="1" pageNumber="385">
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<typeStatus id="54B4B586FF81C20AFD9FFCDD92A2FC86" box="[612,658,868,892]" pageId="1" pageNumber="385">type</typeStatus>
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species:
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<taxonomicName id="4C0F70A7FF81C20AFD09FCDD94CDFC81" ID-CoL="4HVQB" authority="A. Milne-Edwards, 1867" authorityName="A. Milne-Edwards" authorityYear="1867" box="[754,1277,867,892]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="crassimanus">
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<emphasis id="B97BD736FF81C20AFD09FCDD93CBFC81" box="[754,1019,868,892]" italics="true" pageId="1" pageNumber="385">Pilumnopeus crassimanus</emphasis>
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A.
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<bibRefCitation id="EF9E76D5FF81C20AFBE4FCDD94CDFC81" author="Milne-Edwards, A." box="[1055,1277,867,892]" pageId="1" pageNumber="385" pagination="263 - 288" refId="ref7922" refString="Milne-Edwards, A. (1867) Descriptions de quelques especes nouvelles de Crustaces Brachyoures, Tribu des Oxyrhinques. Familie des Maiens. Annales de la Societe Entomologique de France, Series 4, 7, 263 - 288." type="journal article" year="1867">Milne-Edwards, 1867</bibRefCitation>
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</taxonomicName>
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=
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<taxonomicName id="4C0F70A7FF81C20AFAE3FCDD91DBFC60" ID-CoL="4HVQN" authority="(Kinahan, 1856)" baseAuthorityName="Kinahan" baseAuthorityYear="1856" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="serratifrons">
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<emphasis id="B97BD736FF81C20AFAE3FCDD910EFC60" italics="true" pageId="1" pageNumber="385">Pilumnopeus serratifrons</emphasis>
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(
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<bibRefCitation id="EF9E76D5FF81C20AFEB0FC3B91D4FC60" author="Kinahan, J. R." box="[331,484,898,922]" pageId="1" pageNumber="385" pagination="111 - 134" refId="ref7752" refString="Kinahan, J. R. (1856) Remarks on the Habits and Distribution of Marine Crustacea on the eastern shores of Port Phillip, Victoria, Australia; with Descriptions of undescribed Species and Genera. Journal of the Royal Dublin Society, 1 (3), 111 - 134, pls. 3 + 4." type="journal article" year="1856">Kinahan, 1856</bibRefCitation>
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)
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, subsequent designation by
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<bibRefCitation id="EF9E76D5FF81C20AFCFCFC3B93BFFC60" author="Balss, H." box="[775,911,898,922]" pageId="1" pageNumber="385" pagination="1 - 47" refId="ref6577" refString="Balss, H. (1933) Beitrage zur Kenntnis der Gattung Pilumnus (Crustacea, Decapoda) und verwandter Gattungen. Capita Zoologica, 4, 1 - 47." type="journal article" year="1933">Balss (1933))</bibRefCitation>
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.—
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<treatmentCitation id="0AAE2D35FF81C20AFC55FC3B9477FC60" author="Balss, H." box="[942,1095,898,922]" page="33" pageId="1" pageNumber="385" year="1933">
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<bibRefCitation id="EF9E76D5FF81C20AFC55FC3B9477FC60" author="Balss, H." box="[942,1095,898,922]" pageId="1" pageNumber="385" pagination="1 - 47" refId="ref6577" refString="Balss, H. (1933) Beitrage zur Kenntnis der Gattung Pilumnus (Crustacea, Decapoda) und verwandter Gattungen. Capita Zoologica, 4, 1 - 47." type="journal article" year="1933">Balss 1933: 33</bibRefCitation>
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</treatmentCitation>
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.—
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<treatmentCitation id="0AAE2D35FF81C20AFB9DFC3B953FFC60" author="Sakai, T." box="[1126,1295,898,922]" page="542" pageId="1" pageNumber="385" year="1939">
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<bibRefCitation id="EF9E76D5FF81C20AFB9DFC3B953FFC60" author="Sakai, T." box="[1126,1295,898,922]" pageId="1" pageNumber="385" pagination="42 - 111" refId="ref8476" refString="Sakai, T. (1939) n. k. In: Studies on the crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo Co., Ltd., Tokyo, pp. 365 - 741 + 1 - 23 (index), pls. 42 - 111." type="book chapter" year="1939">Sakai 1939: 542</bibRefCitation>
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</treatmentCitation>
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.—
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<treatmentCitation id="0AAE2D35FF81C20AFAD5FC3B90EEFC43" author="Dell, R. K." page="18" pageId="1" pageNumber="385" year="1968">
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<bibRefCitation id="EF9E76D5FF81C20AFAD5FC3B90EEFC43" author="Dell, R. K." pageId="1" pageNumber="385" pagination="13 - 28" refId="ref7096" refString="Dell, R. K. (1968) Notes on New Zealand Crabs. Records of the Dominion Museum, 6 (3), 13 - 28." type="journal article" year="1968">Dell 1968: 18</bibRefCitation>
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</treatmentCitation>
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.—
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<treatmentCitation id="0AAE2D35FF81C20AFF07FC189201FC43" author="Takeda, M. & Miyake, S." box="[252,561,929,953]" page="120" pageId="1" pageNumber="385" year="1969">
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<bibRefCitation id="EF9E76D5FF81C20AFF07FC189201FC43" author="Takeda, M. & Miyake, S." box="[252,561,929,953]" pageId="1" pageNumber="385" pagination="93 - 156" refId="ref8735" refString="Takeda, M. & Miyake, S. (1969) Pilumnid crabs of the family Xanthidae from the West Pacific. II. Twenty-one species of four genera, with descriptions of four new species. OHMU, Occasional Papers of the Zoological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka, Japan, 2, 93 - 156." type="journal article" year="1969">Takeda & Miyake 1969: 120</bibRefCitation>
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</treatmentCitation>
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(part).—
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<treatmentCitation id="0AAE2D35FF81C20AFD6AFC18930FFC43" author="Sakai, T." box="[657,831,929,953]" page="500" pageId="1" pageNumber="385" year="1976">
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<bibRefCitation id="EF9E76D5FF81C20AFD6AFC18930FFC43" author="Sakai, T." box="[657,831,929,953]" pageId="1" pageNumber="385" refId="ref8631" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. 3 Vols. Kodansha Ltd., Tokyo, xxix + 773 pp. (English text), 461 pp. (Japanese text), 16 pp., 251 pls. (plates volume)." type="book" year="1976">Sakai 1976: 500</bibRefCitation>
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</treatmentCitation>
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.—
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<treatmentCitation id="0AAE2D35FF81C20AFCA6FC18949AFC43" author="Manning, R. B. & Holthuis, L. B." box="[861,1194,929,953]" page="151" pageId="1" pageNumber="385" year="1981">
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<bibRefCitation id="EF9E76D5FF81C20AFCA6FC18949AFC43" author="Manning, R. B. & Holthuis, L. B." box="[861,1194,929,953]" pageId="1" pageNumber="385" pagination="1 - 379" refId="ref7873" refString="Manning, R. B. & Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contribution of Zoology, 306, 1 - 379. https: // doi. org / 10.5479 / si. 00810282.306" type="journal article" year="1981">Manning & Holthuis 1981: 151</bibRefCitation>
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</treatmentCitation>
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.—
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<treatmentCitation id="0AAE2D35FF81C20AFB32FC18954CFC43" author="Davie, P. J. F." box="[1225,1404,929,953]" page="138" pageId="1" pageNumber="385" year="1989">
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<bibRefCitation id="EF9E76D5FF81C20AFB32FC18954CFC43" author="Davie, P. J. F." box="[1225,1404,929,953]" pageId="1" pageNumber="385" pagination="129 - 156" refId="ref6730" refString="Davie, P. J. F. (1989) A re-appraisal of Heteropanope Stimpson, and Pilumnopeus A. Milne Edwards (Crustacea: Decapoda: Pilumnidae) with descriptions of new species and new genera. Memoirs of the Queensland Museum, 27, 129 - 156." type="journal article" year="1989">Davie 1989: 138</bibRefCitation>
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</treatmentCitation>
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.—
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<treatmentCitation id="0AAE2D35FF81C20AFF3CFC799140FC2D" author="Wada, K." box="[199,368,959,984]" page="396" pageId="1" pageNumber="385" year="1995">
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<bibRefCitation id="EF9E76D5FF81C20AFF3CFC799140FC2D" author="Wada, K." box="[199,368,959,984]" pageId="1" pageNumber="385" pagination="379 - 418" refId="ref8968" refString="Wada, K. (1995) Brachyura. In: Guide to seashore animals of Japan with color pictures and keys. Hoikusha, Osaka, pp. 379 - 418." type="book chapter" year="1995">Wada 1995: 396</bibRefCitation>
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(key), 403.
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</paragraph>
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<treatmentCitationGroup id="AB1F2C0AFF81C20AFF6CFC669374FC0C" box="[151,836,990,1014]" pageId="1" pageNumber="385">
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<taxonomicName id="4C0F70A7FF81C20AFF6CFC669119FC0C" ID-CoL="7PK64" authorityName="Stimpson" authorityYear="1858" box="[151,297,991,1014]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
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<emphasis id="B97BD736FF81C20AFF6CFC669119FC0C" box="[151,297,991,1014]" italics="true" pageId="1" pageNumber="385">Heteropanope</emphasis>
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</taxonomicName>
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—
|
|
<treatmentCitation id="0AAE2D35FF81C20AFEBFFC6691C8FC0C" author="De Man, J. G." box="[324,504,990,1014]" page="52" pageId="1" pageNumber="385" year="1887">
|
|
<bibRefCitation id="EF9E76D5FF81C20AFEBFFC6691C8FC0C" author="De Man, J. G." box="[324,504,990,1014]" pageId="1" pageNumber="385" pagination="1 - 305" refId="ref6901" refString="De Man, J. G. (1887) Report on the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson F. R. S., Superintendent of the Museum. Journal of the Linnean Society, Zoology, 22, 1 - 305. https: // doi. org / 10.1111 / j. 1096 - 3642.1887. tb 00027. x" type="journal article" year="1887">De Man 1887: 52</bibRefCitation>
|
|
</treatmentCitation>
|
|
(part);
|
|
<treatmentCitation id="0AAE2D35FF81C20AFDBEFC6792CEFC0C" author="Alcock, A." box="[581,766,990,1014]" page="207" pageId="1" pageNumber="385" year="1898">
|
|
<bibRefCitation id="EF9E76D5FF81C20AFDBEFC6792CEFC0C" author="Alcock, A." box="[581,766,990,1014]" pageId="1" pageNumber="385" pagination="67 - 233" refId="ref6419" refString="Alcock, A. (1898) Materials for a carcinological fauna of India. No. 3. The Brachyura Cyclometopa. Part I. The family Xanthidae. Journal of the Asiatic Society of Bengal, 67, 67 - 233." type="journal article" year="1898">Alcock 1898: 207</bibRefCitation>
|
|
</treatmentCitation>
|
|
(part).
|
|
</treatmentCitationGroup>
|
|
</paragraph>
|
|
<paragraph id="8BB00B24FF81C20AFF6CFC4792D1FBEF" blockId="1.[151,1437,867,1045]" box="[151,737,1021,1045]" pageId="1" pageNumber="385">
|
|
<treatmentCitationGroup id="AB1F2C0AFF81C20AFF6CFC4792D1FBEF" box="[151,737,1021,1045]" pageId="1" pageNumber="385">
|
|
<taxonomicName id="4C0F70A7FF81C20AFF6CFC4791F6FBEF" authorityName="A. Milne-Edwards" authorityYear="1867" box="[151,454,1021,1045]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="subGenus" subGenus="Pilumnopeus">
|
|
<emphasis id="B97BD736FF81C20AFF6CFC479119FBEF" box="[151,297,1022,1045]" italics="true" pageId="1" pageNumber="385">Heteropanope</emphasis>
|
|
(
|
|
<emphasis id="B97BD736FF81C20AFEC2FC44918FFBEF" box="[313,447,1021,1045]" italics="true" pageId="1" pageNumber="385">Pilumnopeus</emphasis>
|
|
)
|
|
</taxonomicName>
|
|
—
|
|
<treatmentCitation id="0AAE2D35FF81C20AFE1AFC4492AAFBEF" author="Monod, Th." box="[481,666,1021,1045]" page="264" pageId="1" pageNumber="385" year="1956">
|
|
<bibRefCitation id="EF9E76D5FF81C20AFE1AFC4492AAFBEF" author="Monod, Th." box="[481,666,1021,1045]" pageId="1" pageNumber="385" pagination="1 - 674" refId="ref7988" refString="Monod, Th. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut Francais d'Afrique Noire (IFAN), 45, 1 - 674, figs. 1 - 884, tabs. 1 - 10." type="journal article" year="1956">Monod 1956: 264</bibRefCitation>
|
|
</treatmentCitation>
|
|
(part).
|
|
</treatmentCitationGroup>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C31558AFFF81C20AFF6CFBFA91C2F8AB" pageId="1" pageNumber="385" type="discussion">
|
|
<paragraph id="8BB00B24FF81C20AFF6CFBFA9552F9F7" blockId="1.[151,1437,1091,1873]" pageId="1" pageNumber="385">
|
|
<emphasis id="B97BD736FF81C20AFF6CFBFA913BFBA7" bold="true" box="[151,267,1091,1117]" pageId="1" pageNumber="385">Remarks.</emphasis>
|
|
<bibRefCitation id="EF9E76D5FF81C20AFEE8FBFA9199FBA7" author="Davie, P. J. F." box="[275,425,1091,1117]" pageId="1" pageNumber="385" pagination="129 - 156" refId="ref6730" refString="Davie, P. J. F. (1989) A re-appraisal of Heteropanope Stimpson, and Pilumnopeus A. Milne Edwards (Crustacea: Decapoda: Pilumnidae) with descriptions of new species and new genera. Memoirs of the Queensland Museum, 27, 129 - 156." type="journal article" year="1989">Davie (1989)</bibRefCitation>
|
|
diagnosed
|
|
<taxonomicName id="4C0F70A7FF81C20AFDD1FBFA928CFBA7" authorityName="A. Milne Edwards" authorityYear="1867" box="[554,700,1091,1117]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|
<emphasis id="B97BD736FF81C20AFDD1FBFA928CFBA7" box="[554,700,1091,1117]" italics="true" pageId="1" pageNumber="385">Pilumnopeus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A2486A4DFF81C20AFD3FFBFD9330FBA4" box="[708,768,1092,1118]" pageId="1" pageNumber="385" sensu="stricto">s. str.</taxonomicNameLabel>
|
|
with several key characters. The most significant is that in
|
|
<taxonomicName id="4C0F70A7FF81C20AFF6CFBDE91B3FB7B" box="[151,387,1127,1153]" pageId="1" pageNumber="385">
|
|
<emphasis id="B97BD736FF81C20AFF6CFBDE9119FB7B" box="[151,297,1127,1153]" italics="true" pageId="1" pageNumber="385">Pilumnopeus</emphasis>
|
|
species
|
|
</taxonomicName>
|
|
, the male thoracic sternites 3 and 4 are proportionately transversely narrower, with the distance between the suture between sternites 2 and 3, and that between sternites 3 and 4 subequal or slightly longer than the distance between the suture between sternites 3 and 4, and the tip of the sternopleonal cavity. In
|
|
<taxonomicName id="4C0F70A7FF81C20AFB01FB1690DBFB17" pageId="1" pageNumber="385">
|
|
<emphasis id="B97BD736FF81C20AFB01FB1695ACFB33" box="[1274,1436,1199,1225]" italics="true" pageId="1" pageNumber="385">Benthopanope</emphasis>
|
|
species
|
|
</taxonomicName>
|
|
, the male thoracic sternites 3 and 4 are proportionately transversely broader, with the distance between the suture of sternites 2 and 3, and that between sternites 3 and 4, is distinctly longer than the distance between the suture between sternites 3 and 4, and the tip of the sternopleonal cavity. Both genera can be separated from
|
|
<taxonomicName id="4C0F70A7FF81C20AFB06FAA595ACFACF" authorityName="Stimpson" authorityYear="1858" box="[1277,1436,1308,1333]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97BD736FF81C20AFB06FAA595ACFACF" box="[1277,1436,1308,1333]" italics="true" pageId="1" pageNumber="385">Heteropanope</emphasis>
|
|
</taxonomicName>
|
|
by their dorsal surface of the carapace possessing ridges and distinct regions (versus surface smooth and regions barely visible in
|
|
<taxonomicName id="4C0F70A7FF81C20AFEADFADD91C5FA87" authorityName="Stimpson" authorityYear="1858" box="[342,501,1380,1405]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97BD736FF81C20AFEADFADD91C5FA87" box="[342,501,1380,1405]" italics="true" pageId="1" pageNumber="385">Heteropanope</emphasis>
|
|
</taxonomicName>
|
|
), and male thoracic sternite 8 is visible when the pleon is closed (versus sternite 8 not visible in
|
|
<taxonomicName id="4C0F70A7FF81C20AFECCFA3191E6FA5B" authorityName="Stimpson" authorityYear="1858" box="[311,470,1416,1441]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97BD736FF81C20AFECCFA3191E6FA5B" box="[311,470,1416,1441]" italics="true" pageId="1" pageNumber="385">Heteropanope</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
<bibRefCitation id="EF9E76D5FF81C20AFE14FA3E92B6FA5B" author="Davie, P. J. F." box="[495,646,1415,1441]" pageId="1" pageNumber="385" pagination="129 - 156" refId="ref6730" refString="Davie, P. J. F. (1989) A re-appraisal of Heteropanope Stimpson, and Pilumnopeus A. Milne Edwards (Crustacea: Decapoda: Pilumnidae) with descriptions of new species and new genera. Memoirs of the Queensland Museum, 27, 129 - 156." type="journal article" year="1989">Davie (1989)</bibRefCitation>
|
|
noted that
|
|
<taxonomicName id="4C0F70A7FF81C20AFCF2FA3194F4FA5B" authority="A. Milne-Edwards, 1867" authorityName="A. Milne-Edwards" authorityYear="1867" box="[777,1220,1415,1441]" class="Malacostraca" family="Pilumnidae" genus="Eurycarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97BD736FF81C20AFCF2FA3193AFFA5B" box="[777,927,1416,1441]" italics="true" pageId="1" pageNumber="385">Eurycarcinus</emphasis>
|
|
A.
|
|
<bibRefCitation id="EF9E76D5FF81C20AFC30FA3E94F4FA5B" author="Milne-Edwards, A." box="[971,1220,1415,1441]" pageId="1" pageNumber="385" pagination="263 - 288" refId="ref7922" refString="Milne-Edwards, A. (1867) Descriptions de quelques especes nouvelles de Crustaces Brachyoures, Tribu des Oxyrhinques. Familie des Maiens. Annales de la Societe Entomologique de France, Series 4, 7, 263 - 288." type="journal article" year="1867">Milne-Edwards, 1867</bibRefCitation>
|
|
</taxonomicName>
|
|
, was closely allied to these three genera but refrained from further taxonomic treatment given the lack of material for examination.
|
|
<bibRefCitation id="EF9E76D5FF81C20AFF6CFA69917FFA13" author="Ng, P. K. L. & Abdelsalam, K. M. & Mona, M. H. & Nour Eldeen, M. F." box="[151,335,1487,1514]" pageId="1" pageNumber="385" pagination="471 - 487" refId="ref8028" refString="Ng, P. K. L., Abdelsalam, K. M., Mona, M. H. & Nour Eldeen, M. F. (2018) A synopsis of the genus Eurycarcinus A. Milne-Edwards, 1867 (Decapoda, Brachyura, Pilumnidae). Crustaceana, 91 (4), 471 - 487. https: // doi. org / 10.1163 / 15685403 - 00003775" type="journal article" year="2018">
|
|
Ng
|
|
<emphasis id="B97BD736FF81C20AFF39FA6990CCFA13" box="[194,252,1487,1513]" italics="true" pageId="1" pageNumber="385">et al.</emphasis>
|
|
(2018)
|
|
</bibRefCitation>
|
|
subsequently revised the genus and showed that
|
|
<taxonomicName id="4C0F70A7FF81C20AFC7AFA699440FA13" box="[897,1136,1487,1513]" pageId="1" pageNumber="385">
|
|
<emphasis id="B97BD736FF81C20AFC7AFA699427FA13" box="[897,1047,1488,1513]" italics="true" pageId="1" pageNumber="385">Eurycarcinus</emphasis>
|
|
species
|
|
</taxonomicName>
|
|
are morphologically quite different, with a large exposed male thoracic sternite 8 when the pleon is closed (see also
|
|
<bibRefCitation id="EF9E76D5FF81C20AFB7AFA4A9565F9F7" author="Trivedi, J. N. & Mitra, S. & Patel, P. & Maheta, N. & Patel, K. & Ng, P. K. L." box="[1153,1365,1523,1549]" pageId="1" pageNumber="385" pagination="1 - 18" refId="ref8799" refString="Trivedi, J. N., Mitra, S., Patel, P., Maheta, N., Patel, K. & Ng, P. K. L. (2021) On the Indian species of Eurycarcinus A. Milne-Edwards, 1867, Heteropanope Stimpson, 1858, and Pilumnopeus A. Milne-Edwards, 1867 (Decapoda: Brachyura: Pilumnidae). Nauplius, 29 (e 2021004), 1 - 18. https: // doi. org / 10.1590 / 2358 - 2936 e 2021004" type="journal article" year="2021">
|
|
Trivedi
|
|
<emphasis id="B97BD736FF81C20AFB21FA4D9522F9F7" box="[1242,1298,1523,1549]" italics="true" pageId="1" pageNumber="385">et al.</emphasis>
|
|
2021
|
|
</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BB00B24FF81C20AFF3CF9AE9560F93B" blockId="1.[151,1437,1091,1873]" pageId="1" pageNumber="385">
|
|
Although
|
|
<bibRefCitation id="EF9E76D5FF81C20AFEC3F9AE9236F9CB" author="Davie, P. J. F." box="[312,518,1559,1585]" pageId="1" pageNumber="385" pagination="129 - 156" refId="ref6730" refString="Davie, P. J. F. (1989) A re-appraisal of Heteropanope Stimpson, and Pilumnopeus A. Milne Edwards (Crustacea: Decapoda: Pilumnidae) with descriptions of new species and new genera. Memoirs of the Queensland Museum, 27, 129 - 156." type="journal article" year="1989">Davie (1989: 144)</bibRefCitation>
|
|
transferred
|
|
<taxonomicName id="4C0F70A7FF81C20AFD75F9A192CAF9CB" authorityName="Patel & Trivedi & Mitra & Ng" authorityYear="2024" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[654,762,1559,1585]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="indicus">
|
|
<emphasis id="B97BD736FF81C20AFD75F9A192CAF9CB" box="[654,762,1559,1585]" italics="true" pageId="1" pageNumber="385">P. indicus</emphasis>
|
|
</taxonomicName>
|
|
to
|
|
<taxonomicName id="4C0F70A7FF81C20AFCE4F9AE93F1F9CB" authorityName="Davie" authorityYear="1989" box="[799,961,1559,1585]" class="Malacostraca" family="Pilumnidae" genus="Benthopanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97BD736FF81C20AFCE4F9AE93F1F9CB" box="[799,961,1559,1585]" italics="true" pageId="1" pageNumber="385">Benthopanope</emphasis>
|
|
</taxonomicName>
|
|
, he did not examine specimens or provide an explanation. The present study shows that the
|
|
<taxonomicName id="4C0F70A7FF81C20AFD24F98594B7F9AF" authority="De Man, 1887" authorityName="De Man" authorityYear="1887" box="[735,1159,1595,1622]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="indica">
|
|
<emphasis id="B97BD736FF81C20AFD24F98593E0F9AF" box="[735,976,1595,1621]" italics="true" pageId="1" pageNumber="385">Heteropanope indica</emphasis>
|
|
<bibRefCitation id="EF9E76D5FF81C20AFC20F98594B7F9AF" author="De Man, J. G." box="[987,1159,1595,1622]" pageId="1" pageNumber="385" pagination="1 - 305" refId="ref6901" refString="De Man, J. G. (1887) Report on the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson F. R. S., Superintendent of the Museum. Journal of the Linnean Society, Zoology, 22, 1 - 305. https: // doi. org / 10.1111 / j. 1096 - 3642.1887. tb 00027. x" type="journal article" year="1887">De Man, 1887</bibRefCitation>
|
|
</taxonomicName>
|
|
, is not a
|
|
<taxonomicName id="4C0F70A7FF81C20AFB01F98295ACF9AF" authorityName="Davie" authorityYear="1989" box="[1274,1436,1595,1621]" class="Malacostraca" family="Pilumnidae" genus="Benthopanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97BD736FF81C20AFB01F98295ACF9AF" box="[1274,1436,1595,1621]" italics="true" pageId="1" pageNumber="385">Benthopanope</emphasis>
|
|
</taxonomicName>
|
|
as its male sternal condition is clearly that of a
|
|
<taxonomicName id="4C0F70A7FF81C20AFD39F9E69364F983" authorityName="A. Milne Edwards" authorityYear="1867" box="[706,852,1631,1657]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97BD736FF81C20AFD39F9E69364F983" box="[706,852,1631,1657]" italics="true" pageId="1" pageNumber="385">Pilumnopeus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
<taxonomicName id="4C0F70A7FF81C20AFC9EF9D99525F983" authority="Rathbun, 1932" authorityName="Rathbun" authorityYear="1932" box="[869,1301,1631,1657]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="pearsei">
|
|
<emphasis id="B97BD736FF81C20AFC9EF9D99450F983" box="[869,1120,1632,1657]" italics="true" pageId="1" pageNumber="385">Heteropanope pearsei</emphasis>
|
|
<bibRefCitation id="EF9E76D5FF81C20AFB91F9E69525F983" author="Rathbun, M. J." box="[1130,1301,1631,1657]" pageId="1" pageNumber="385" pagination="28 - 38" refId="ref8390" refString="Rathbun, M. J. (1932) Preliminary descriptions of new species of Japanese crabs. Proceedings of the biological Society of Washington, 45, 28 - 38." type="journal article" year="1932">Rathbun, 1932</bibRefCitation>
|
|
</taxonomicName>
|
|
, which was synonymised by T.
|
|
<bibRefCitation id="EF9E76D5FF81C20AFE8CF93A9239F964" author="Sakai, T." box="[375,521,1667,1694]" pageId="1" pageNumber="385" refId="ref8590" refString="Sakai, T. (1965) The crabs of Sagami Bay, collected by His Majesty the Emperor of Japan. Biological Laboratory, Imperial Household, Tokyo, 206 pp., 27 text-figs., 100 pls." type="book" year="1965">Sakai (1965)</bibRefCitation>
|
|
under
|
|
<taxonomicName id="4C0F70A7FF81C20AFDA0F93D92F9F967" authorityName="Patel & Trivedi & Mitra & Ng" authorityYear="2024" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[603,713,1667,1693]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="indicus">
|
|
<emphasis id="B97BD736FF81C20AFDA0F93D92F9F967" box="[603,713,1667,1693]" italics="true" pageId="1" pageNumber="385">P. indicus</emphasis>
|
|
</taxonomicName>
|
|
but treated as a separate species of
|
|
<taxonomicName id="4C0F70A7FF81C20AFB93F93A953AF967" authorityName="Davie" authorityYear="1989" box="[1128,1290,1667,1693]" class="Malacostraca" family="Pilumnidae" genus="Benthopanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97BD736FF81C20AFB93F93A953AF967" box="[1128,1290,1667,1693]" italics="true" pageId="1" pageNumber="385">Benthopanope</emphasis>
|
|
</taxonomicName>
|
|
by
|
|
<bibRefCitation id="EF9E76D5FF81C20AFACCF93D90EBF93B" author="Ng, P. K. L. & Wang, C. - H. & Ho, P. - H. & Shih, H. - T." pageId="1" pageNumber="385" pagination="1 - 86" refId="ref8322" refString="Ng, P. K. L., Wang, C. - H., Ho, P. - H. & Shih, H. - T. (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series, No. 11, 1 - 86, 8 colour pls." type="book chapter" year="2001">
|
|
Ng
|
|
<emphasis id="B97BD736FF81C20AFA99F93D95ADF967" box="[1378,1437,1667,1693]" italics="true" pageId="1" pageNumber="385">et al.</emphasis>
|
|
(2001
|
|
</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="EF9E76D5FF81C20AFF1CF91E9111F93B" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[231,289,1703,1729]" pageId="1" pageNumber="385" pagination="1 - 286" refId="ref8200" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">2008</bibRefCitation>
|
|
) is also here referred to
|
|
<taxonomicName id="4C0F70A7FF81C20AFDCBF91E92F2F93B" authorityName="A. Milne Edwards" authorityYear="1867" box="[560,706,1703,1729]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97BD736FF81C20AFDCBF91E92F2F93B" box="[560,706,1703,1729]" italics="true" pageId="1" pageNumber="385">Pilumnopeus</emphasis>
|
|
</taxonomicName>
|
|
. Its taxonomy is discussed together with
|
|
<taxonomicName id="4C0F70A7FF81C20AFB69F91194CEF93B" authorityName="Patel & Trivedi & Mitra & Ng" authorityYear="2024" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[1170,1278,1703,1729]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="indicus">
|
|
<emphasis id="B97BD736FF81C20AFB69F91194CEF93B" box="[1170,1278,1703,1729]" italics="true" pageId="1" pageNumber="385">P. indicus</emphasis>
|
|
</taxonomicName>
|
|
below.
|
|
</paragraph>
|
|
<paragraph id="8BB00B24FF81C20AFF3CF97291C2F8AB" blockId="1.[151,1437,1091,1873]" pageId="1" pageNumber="385">
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<taxonomicName id="4C0F70A7FF81C20AFF3CF9729169F91F" authorityName="A. Milne Edwards" authorityYear="1867" box="[199,345,1739,1765]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
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<emphasis id="B97BD736FF81C20AFF3CF9729169F91F" box="[199,345,1739,1765]" italics="true" pageId="1" pageNumber="385">Pilumnopeus</emphasis>
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</taxonomicName>
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|
now comprises 11 species:
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<taxonomicName id="4C0F70A7FF81C20AFD6CF97593EDF91F" authority="(De Man, 1902)" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[663,989,1739,1766]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="africanus">
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<emphasis id="B97BD736FF81C20AFD6CF975932DF91F" box="[663,797,1739,1765]" italics="true" pageId="1" pageNumber="385">P. africanus</emphasis>
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|
(De Man, 1902)
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|
</taxonomicName>
|
|
,
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|
<taxonomicName id="4C0F70A7FF81C20AFC11F975953CF91C" authority="(Monod, 1956)" baseAuthorityName="Monod" baseAuthorityYear="1956" box="[1002,1292,1739,1766]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="caparti">
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<emphasis id="B97BD736FF81C20AFC11F9759468F91F" box="[1002,1112,1740,1765]" italics="true" pageId="1" pageNumber="385">P. caparti</emphasis>
|
|
(
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|
<bibRefCitation id="EF9E76D5FF81C20AFB9CF9729535F91C" author="Monod, Th." box="[1127,1285,1739,1766]" pageId="1" pageNumber="385" pagination="1 - 674" refId="ref7988" refString="Monod, Th. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut Francais d'Afrique Noire (IFAN), 45, 1 - 674, figs. 1 - 884, tabs. 1 - 10." type="journal article" year="1956">Monod, 1956</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C0F70A7FF81C20AFAE2F9759158F8F3" authority="(Maccagno, 1936)" baseAuthorityName="Maccagno" baseAuthorityYear="1936" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="convexus">
|
|
<emphasis id="B97BD736FF81C20AFAE2F97595ADF91F" box="[1305,1437,1740,1765]" italics="true" pageId="1" pageNumber="385">P. convexus</emphasis>
|
|
(Maccagno, 1936)
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C0F70A7FF81C20AFE8FF94992A9F8F3" authority="Balss, 1933" authorityName="Balss" authorityYear="1933" box="[372,665,1775,1801]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="granulatus">
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|
<emphasis id="B97BD736FF81C20AFE8FF949923AF8F3" box="[372,522,1775,1801]" italics="true" pageId="1" pageNumber="385">P. granulatus</emphasis>
|
|
<bibRefCitation id="EF9E76D5FF81C20AFDE9F95692A9F8F3" author="Balss, H." box="[530,665,1775,1801]" pageId="1" pageNumber="385" pagination="1 - 47" refId="ref6577" refString="Balss, H. (1933) Beitrage zur Kenntnis der Gattung Pilumnus (Crustacea, Decapoda) und verwandter Gattungen. Capita Zoologica, 4, 1 - 47." type="journal article" year="1933">Balss, 1933</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C0F70A7FF81C20AFD5EF94993E2F8F3" authority="(De Man, 1887)" authorityName="Patel & Trivedi & Mitra & Ng" authorityYear="2024" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[677,978,1775,1802]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="indicus">
|
|
<emphasis id="B97BD736FF81C20AFD5EF9499323F8F3" box="[677,787,1775,1801]" italics="true" pageId="1" pageNumber="385">P. indicus</emphasis>
|
|
(
|
|
<bibRefCitation id="EF9E76D5FF81C20AFCD9F95693FBF8F3" author="De Man, J. G." box="[802,971,1775,1802]" pageId="1" pageNumber="385" pagination="1 - 305" refId="ref6901" refString="De Man, J. G. (1887) Report on the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson F. R. S., Superintendent of the Museum. Journal of the Linnean Society, Zoology, 22, 1 - 305. https: // doi. org / 10.1111 / j. 1096 - 3642.1887. tb 00027. x" type="journal article" year="1887">De Man, 1887</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C0F70A7FF81C20AFC24F9499538F8F3" authority="Cano, 1889" authorityName="Cano" authorityYear="1889" box="[991,1288,1775,1801]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="laevimanus">
|
|
<emphasis id="B97BD736FF81C20AFC24F949944CF8F3" box="[991,1148,1775,1801]" italics="true" pageId="1" pageNumber="385">P. laevimanus</emphasis>
|
|
Cano, 1889
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C0F70A7FF81C20AFAEFF9499161F8D7" authority="(Rathbun, 1931)" baseAuthorityName="Rathbun" baseAuthorityYear="1931" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="makianus">
|
|
<emphasis id="B97BD736FF81C20AFAEFF94995ACF8F3" box="[1300,1436,1775,1801]" italics="true" pageId="1" pageNumber="385">P. makianus</emphasis>
|
|
(Rathbun, 1931)
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C0F70A7FF81C20AFEA5F8AD92F5F8D4" authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[350,709,1811,1838]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="marginatus">
|
|
<emphasis id="B97BD736FF81C20AFEA5F8AD91C9F8D7" box="[350,505,1812,1837]" italics="true" pageId="1" pageNumber="385">P. marginatus</emphasis>
|
|
(Stimpson, 1858)
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C0F70A7FF81C20AFD2AF8AD9431F8D7" authority="(Rathbun, 1932)" baseAuthorityName="Rathbun" baseAuthorityYear="1932" box="[721,1025,1811,1837]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="pearsei">
|
|
<emphasis id="B97BD736FF81C20AFD2AF8AD930FF8D7" box="[721,831,1812,1837]" italics="true" pageId="1" pageNumber="385">P. pearsei</emphasis>
|
|
(
|
|
<bibRefCitation id="EF9E76D5FF81C20AFCB4F8AA93CAF8D7" author="Rathbun, M. J." box="[847,1018,1811,1837]" pageId="1" pageNumber="385" pagination="28 - 38" refId="ref8390" refString="Rathbun, M. J. (1932) Preliminary descriptions of new species of Japanese crabs. Proceedings of the biological Society of Washington, 45, 28 - 38." type="journal article" year="1932">Rathbun, 1932</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C0F70A7FF81C20AFBF5F8AD955CF8D4" authority="(Kinahan, 1856)" baseAuthorityName="Kinahan" baseAuthorityYear="1856" box="[1038,1388,1811,1838]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="serratifrons">
|
|
<emphasis id="B97BD736FF81C20AFBF5F8AD949DF8D7" box="[1038,1197,1811,1837]" italics="true" pageId="1" pageNumber="385">P. serratifrons</emphasis>
|
|
(
|
|
<bibRefCitation id="EF9E76D5FF81C20AFB47F8AA9554F8D4" author="Kinahan, J. R." box="[1212,1380,1811,1838]" pageId="1" pageNumber="385" pagination="111 - 134" refId="ref7752" refString="Kinahan, J. R. (1856) Remarks on the Habits and Distribution of Marine Crustacea on the eastern shores of Port Phillip, Victoria, Australia; with Descriptions of undescribed Species and Genera. Journal of the Royal Dublin Society, 1 (3), 111 - 134, pls. 3 + 4." type="journal article" year="1856">Kinahan, 1856</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C0F70A7FF81C20AFF6CF88191DEF8AB" authority="(Audouin, 1826)" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[151,494,1847,1873]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="vauquelini">
|
|
<emphasis id="B97BD736FF81C20AFF6CF8819119F8AB" box="[151,297,1847,1873]" italics="true" pageId="1" pageNumber="385">P. vauquelini</emphasis>
|
|
(Audouin, 1826)
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |