416 lines
52 KiB
XML
416 lines
52 KiB
XML
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<mods:title id="704B88E294E90627A167E784D8124C4B">First Systematic Study using the Variability of the Residual Colour Patterns: The Case of the Paleogene Seraphsidae (Mollusca, Gastropoda, Stromboidea)</mods:title>
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<mods:namePart id="95A8C319DFDC48578609EAA061D37682">Merle, Didier</mods:namePart>
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<mods:namePart id="4BDF61C8218D84D94E0F89079FE3B36C">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="27A05D42B01621269E39198689AF362B">Saint Martin, Jean-Paul</mods:namePart>
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<mods:title id="269E477E432BBFDB596F85115323F3DC">Geodiversitas</mods:title>
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<mods:date id="283F1D2A276563F9749A30ECC2508EC3">2010</mods:date>
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<taxonomicName id="4C0F52DDFF8B1E43FF1EFF2AFE00FF26" authority="Caze & Merle & Pacaud & Saint Martin, 2010" authorityName="Caze & Merle & Pacaud & Saint Martin" authorityYear="2010" box="[248,447,215,242]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="445" phylum="Mollusca" rank="species" species="peterjungi" status="sp. nov.">
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<emphasis id="B97BF54CFF8B1E43FF1EFF2AFE00FF26" bold="true" box="[248,447,215,242]" italics="true" pageId="28" pageNumber="445">Seraphs peterjungi</emphasis>
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J-N; 19)
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, Lutetian (Middle Eocene).
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</materialsCitation>
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;
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,
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(leg.
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)
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</materialsCitation>
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;
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<materialsCitation id="3B672303FF8B1E43FE42FDB3FF4DFD55" accessionNumber="A28559" collectionCode="MNHN" collectorName="Ledon" pageId="28" pageNumber="445" specimenCount="1" typeStatus="paratype">
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,
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<collectionCode id="ED1EB19BFF8B1E43FE3BFDB3FD8FFDB2" box="[477,560,590,614]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="445">MNHN</collectionCode>
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<accessionNumber id="945CB4BDFF8B1E43FDD0FDB3FD3AFDB2" box="[566,645,590,614]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A28559" pageId="28" pageNumber="445" type="EnaNcbi">A28559</accessionNumber>
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(
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<collectorName id="26FA4C88FF8B1E43FF9EFD94FF07FD55" box="[120,184,617,641]" pageId="28" pageNumber="445">Ledon</collectorName>
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coll.)
|
||
</materialsCitation>
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;
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||
</paragraph>
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||
<paragraph id="8BB0295EFF8B1E43FF97FD7EFDB8FD4F" blockId="28.[112,648,379,932]" box="[113,519,643,667]" pageId="28" pageNumber="445">
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<materialsCitation id="3B672303FF8B1E43FF97FD7EFDBCFD4F" accessionNumber="A28789" box="[113,515,643,667]" collectionCode="MNHN" pageId="28" pageNumber="445" specimenCount="1">
|
||
Beynes, Yvelines,
|
||
<specimenCount id="9D09E2D7FF8B1E43FEFBFD7EFEF1FD4F" box="[285,334,643,667]" pageId="28" pageNumber="445" type="generic">1 ex.</specimenCount>
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,
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<collectionCode id="ED1EB19BFF8B1E43FEBEFD7EFE14FD4F" box="[344,427,643,667]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="445">MNHN</collectionCode>
|
||
<accessionNumber id="945CB4BDFF8B1E43FE57FD7EFDBCFD4F" box="[433,515,643,667]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A28789" pageId="28" pageNumber="445" type="EnaNcbi">A28789</accessionNumber>
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</materialsCitation>
|
||
;
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||
</paragraph>
|
||
<paragraph id="8BB0295EFF8B1E43FF97FD63FE64FCD1" blockId="28.[112,648,379,932]" pageId="28" pageNumber="445">
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<materialsCitation id="3B672303FF8B1E43FF97FD63FDCAFD62" accessionNumber="A27634" box="[113,629,670,694]" collectionCode="MNHN" collectorName="Pacaud" pageId="28" pageNumber="445" specimenCount="5">
|
||
Fercourt, Oise,
|
||
<specimenCount id="9D09E2D7FF8B1E43FEECFD63FE83FD62" box="[266,316,670,694]" pageId="28" pageNumber="445" type="generic">5 ex.</specimenCount>
|
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,
|
||
<collectionCode id="ED1EB19BFF8B1E43FEA3FD63FE27FD62" box="[325,408,670,694]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="445">MNHN</collectionCode>
|
||
A27634 (leg.
|
||
<collectorName id="26FA4C88FF8B1E43FDC3FD63FDD0FD62" box="[549,623,670,694]" pageId="28" pageNumber="445">Pacaud</collectorName>
|
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)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B672303FF8B1E43FF97FD45FE9FFD3F" accessionNumber="A27632" collectionCode="MNHN" collectorName="Galerie" pageId="28" pageNumber="445" specimenCount="1">
|
||
Grignon, Yvelines,
|
||
<specimenCount id="9D09E2D7FF8B1E43FEC8FD45FEDFFD04" box="[302,352,696,720]" pageId="28" pageNumber="445" type="generic">1 ex.</specimenCount>
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,
|
||
<collectionCode id="ED1EB19BFF8B1E43FE8DFD45FE01FD04" box="[363,446,696,720]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="445">MNHN</collectionCode>
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<accessionNumber id="945CB4BDFF8B1E43FE23FD45FDAAFD04" box="[453,533,696,720]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A27632" pageId="28" pageNumber="445" type="EnaNcbi">A27632</accessionNumber>
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(ex
|
||
<collectorName id="26FA4C88FF8B1E43FDA6FD45FD39FD04" box="[576,646,696,720]" pageId="28" pageNumber="445">Galerie</collectorName>
|
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de Zoologie coll.)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B672303FF8B1E43FECAFD2EFD3CFD3F" accessionNumber="A28512" box="[300,643,723,747]" collectionCode="MNHN" collectorName="Caze" isTailEnumeration="true" pageId="28" pageNumber="445" specimenCount="1">
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<specimenCount id="9D09E2D7FF8B1E43FECAFD2EFEE2FD3F" box="[300,349,723,747]" pageId="28" pageNumber="445" type="generic">1 ex.</specimenCount>
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,
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<collectionCode id="ED1EB19BFF8B1E43FE8EFD2EFE04FD3F" box="[360,443,723,747]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="445">MNHN</collectionCode>
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<accessionNumber id="945CB4BDFF8B1E43FE24FD2EFDAEFD3F" box="[450,529,723,747]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A28512" pageId="28" pageNumber="445" type="EnaNcbi">A28512</accessionNumber>
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(leg.
|
||
<collectorName id="26FA4C88FF8B1E43FDAFFD2EFDC3FD3F" box="[585,636,723,747]" pageId="28" pageNumber="445">Caze</collectorName>
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)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B672303FF8B1E43FF97FD10FE68FCD1" accessionNumber="A28530" box="[113,471,749,773]" collectionCode="MNHN" collectorName="Ballot" pageId="28" pageNumber="445" specimenCount="1">
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,
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<collectionCode id="ED1EB19BFF8B1E43FF4AFD10FF40FCD1" box="[172,255,749,773]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="445">MNHN</collectionCode>
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<accessionNumber id="945CB4BDFF8B1E43FEE0FD10FEEAFCD1" box="[262,341,749,773]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A28530" pageId="28" pageNumber="445" type="EnaNcbi">A28530</accessionNumber>
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(
|
||
<collectorName id="26FA4C88FF8B1E43FE84FD10FE22FCD1" box="[354,413,749,773]" pageId="28" pageNumber="445">Ballot</collectorName>
|
||
coll.)
|
||
</materialsCitation>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8B1E43FF97FCFAFE5FFCEE" blockId="28.[112,648,379,932]" pageId="28" pageNumber="445">
|
||
<materialsCitation id="3B672303FF8B1E43FF97FCFAFD3DFCCB" accessionNumber="A11186" box="[113,642,775,799]" collectionCode="MNHN" collectorName="d'Orbigny" pageId="28" pageNumber="445" specimenCount="1">
|
||
Parnes, Oise,
|
||
<specimenCount id="9D09E2D7FF8B1E43FF13FCFAFE9AFCCB" box="[245,293,775,799]" pageId="28" pageNumber="445" type="generic">1 ex.</specimenCount>
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||
,
|
||
<collectionCode id="ED1EB19BFF8B1E43FEC8FCFAFE3EFCCB" box="[302,385,775,799]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="445">MNHN</collectionCode>
|
||
A11186 (
|
||
<collectorName id="26FA4C88FF8B1E43FE04FCFAFDF6FCCB" box="[482,585,775,799]" pageId="28" pageNumber="445">d’Orbigny</collectorName>
|
||
coll.)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B672303FF8B1E43FF97FCDFFE64FCEE" accessionNumber="A28511" box="[113,475,802,826]" collectionCode="MNHN" collectorName="Roissy" isTailEnumeration="true" pageId="28" pageNumber="445" specimenCount="1">
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<specimenCount id="9D09E2D7FF8B1E43FF97FCDFFF1CFCEE" box="[113,163,802,826]" pageId="28" pageNumber="445" type="generic">1 ex.</specimenCount>
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,
|
||
<collectionCode id="ED1EB19BFF8B1E43FF4AFCDFFF40FCEE" box="[172,255,802,826]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="445">MNHN</collectionCode>
|
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<accessionNumber id="945CB4BDFF8B1E43FEE0FCDFFEEAFCEE" box="[262,341,802,826]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A28511" pageId="28" pageNumber="445" type="EnaNcbi">A28511</accessionNumber>
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(
|
||
<collectorName id="26FA4C88FF8B1E43FE84FCDFFE1DFCEE" box="[354,418,802,826]" pageId="28" pageNumber="445">Roissy</collectorName>
|
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coll.)
|
||
</materialsCitation>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8B1E43FF97FCC1FF4BFCBB" blockId="28.[112,648,379,932]" pageId="28" pageNumber="445">
|
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<materialsCitation id="3B672303FF8B1E43FF97FCC1FF4EFCBB" accessionNumber="A28521" collectionCode="MNHN" collectorName="Pacaud" pageId="28" pageNumber="445" specimenCount="1">
|
||
Parnes (Grande Cronière), Oise,
|
||
<specimenCount id="9D09E2D7FF8B1E43FE4FFCC1FE66FC80" box="[425,473,828,852]" pageId="28" pageNumber="445" type="generic">1 ex.</specimenCount>
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,
|
||
<collectionCode id="ED1EB19BFF8B1E43FE06FCC1FD8CFC80" box="[480,563,828,852]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="445">MNHN</collectionCode>
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<accessionNumber id="945CB4BDFF8B1E43FDDEFCC1FD38FC80" box="[568,647,828,852]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A28521" pageId="28" pageNumber="445" type="EnaNcbi">A28521</accessionNumber>
|
||
(leg.
|
||
<collectorName id="26FA4C88FF8B1E43FF46FCAAFF55FCBB" box="[160,234,855,879]" pageId="28" pageNumber="445">Pacaud</collectorName>
|
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)
|
||
</materialsCitation>
|
||
;
|
||
</paragraph>
|
||
<caption id="DF7079D6FF8B1E43FD43FCBCFBFFFCBD" ID-DOI="http://doi.org/10.5281/zenodo.5381188" ID-Zenodo-Dep="5381188" httpUri="https://zenodo.org/record/5381188/files/figure.png" pageId="28" pageNumber="445" startId="28.[677,688,833,850]" targetBox="[682,1206,216,790]" targetPageId="28">
|
||
<paragraph id="8BB0295EFF8B1E43FD43FCBCFBFFFCBD" blockId="28.[675,1210,833,873]" pageId="28" pageNumber="445">
|
||
FIG. 18.– Distribution of the pattern types with their intermediaries (
|
||
<emphasis id="B97BF54CFF8B1E43FD4EFCA5FD69FCBD" bold="true" box="[680,726,856,873]" pageId="28" pageNumber="445">Inter.</emphasis>
|
||
) in
|
||
<taxonomicName id="4C0F52DDFF8B1E43FD14FCA5FB82FCBD" authority="(Cossmann, 1889)" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[754,1085,856,873]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="445" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8B1E43FD14FCA5FC22FCBD" box="[754,925,856,873]" italics="true" pageId="28" pageNumber="445">Seraphs chilophorus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9E54AFFF8B1E43FC41FCA5FB87FCBD" author="COSSMANN M." box="[935,1080,856,873]" pageId="28" pageNumber="445" pagination="3 - 385" refId="ref23409" refString="COSSMANN M. 1889. - Catalogue illustre des coquilles fossiles de l'Eocene des environs de Paris (4 e fascicule). Annales de la Societe royale malacologique de Belgique 24: 3 - 385." type="journal article" year="1889">Cossmann, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB0295EFF8B1E43FF97FC8CFF75FC70" blockId="28.[112,648,379,932]" pageId="28" pageNumber="445">
|
||
Saulx-Marchaix, Yvelines,
|
||
<specimenCount id="9D09E2D7FF8B1E43FE81FC8CFE29FC5D" box="[359,406,881,905]" pageId="28" pageNumber="445" type="generic">1 ex.</specimenCount>
|
||
,
|
||
<collectionCode id="ED1EB19BFF8B1E43FE7BFC8CFE50FC5D" box="[413,495,881,905]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="445">MNHN</collectionCode>
|
||
<accessionNumber id="945CB4BDFF8B1E43FE15FC8CFDFFFC5D" box="[499,576,881,905]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A28520" pageId="28" pageNumber="445" type="EnaNcbi">A28520</accessionNumber>
|
||
(1965- 11 coll.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3157AD5FF8B1E43FF97FC41FE8EFC3A" pageId="28" pageNumber="445" type="etymology">
|
||
<paragraph id="8BB0295EFF8B1E43FF97FC41FE8EFC3A" blockId="28.[113,647,955,1006]" pageId="28" pageNumber="445">
|
||
ETYMOLOGY. — Dedicated to Peter Jung for his work on the
|
||
<taxonomicName id="4C0F52DDFF8B1E43FF51FC2BFE92FC3A" authorityName="Jung" authorityYear="1974" box="[183,301,982,1006]" class="Gastropoda" family="Seraphsidae" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="445" phylum="Mollusca" rank="family">Seraphsidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3157AD5FF8B1E42FF97FBEDFE5BFE24" lastPageId="29" lastPageNumber="446" pageId="28" pageNumber="445" type="description">
|
||
<paragraph id="8BB0295EFF8B1E43FF97FBEDFE71FBFC" blockId="28.[111,649,1040,1478]" box="[113,462,1040,1066]" pageId="28" pageNumber="445">
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||
<materialsCitation id="3B672303FF8B1E43FF97FBEDFE71FBFC" box="[113,462,1040,1066]" collectionCode="DESCRIPTION, OF, THE" pageId="28" pageNumber="445" specimenCount="1" typeStatus="holotype">
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<collectionCode id="ED1EB19BFF8B1E43FF97FBEDFEE7FBFC" box="[113,344,1040,1066]" pageId="28" pageNumber="445">DESCRIPTION OF THE</collectionCode>
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<typeStatus id="54B497FCFF8B1E43FEB9FBE8FE71FBFC" box="[351,462,1045,1064]" pageId="28" pageNumber="445" type="holotype">HOLOTYPE</typeStatus>
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</materialsCitation>
|
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</paragraph>
|
||
<paragraph id="8BB0295EFF8B1E43FF97FBD2FE63FA12" blockId="28.[111,649,1040,1478]" pageId="28" pageNumber="445">
|
||
Shell
|
||
<quantity id="4CF784BBFF8B1E43FF57FBD2FEBBFB9E" box="[177,260,1071,1098]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="28" pageNumber="445" unit="mm" value="23.0">23 mm</quantity>
|
||
in height and
|
||
<quantity id="4CF784BBFF8B1E43FE57FBD2FE4AFB9E" box="[433,501,1071,1098]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="28" pageNumber="445" unit="mm" value="7.0">7 mm</quantity>
|
||
in diameter, slender with acute apical part. Aperture long and narrow. Callus of the inner lip well developed and delineated, continuing adapically on the spire and forming a narrow band parallel to the outer lip. Columella bent backwards near the base (
|
||
<figureCitation id="133435DBFF8B1E43FDFBFB33FDC2FB3D" box="[541,637,1230,1257]" captionStart="FIG" captionStartId="24.[113,124,1544,1561]" captionTargetBox="[126,1202,227,1464]" captionTargetId="figure-192@24.[113,1210,214,1500]" captionTargetPageId="24" captionText="FIG. 15. — Seraphs Montfort, 1810 shells in normal light; A-I, S. chilophorus (Cossmann, 1889); A-E, MNHN A28779 (Ledon coll.), Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, labral view; C, detailed view of the apical part; D, dorsal view;E, detailed view of the apical part; F-H, MNHN A28780 (Ledon coll.), Chaussy (Les Garennes), Lutetian;F, ventral view;G, dorsal view; H, detailed view of the apical part; I, MNHN A27783, detailed view the oblique grooves (leg. Pacaud), Saint-Lubin-de-la-Haye, Eure-et-Loir, France, Lutetian; J-N, S. peterjungi n. sp., holotype MNHN A28578 (Pacaud leg.), Chaussy (Les Garennes), Lutetian; J, ventral view; K, labral view; L, detailed view of the apical part; M, detailed view the oblique grooves; N, dorsal view. Scale bars: A-B, D, F-G, J-K, N, 10 mm; C, I, 5 mm; E, H, L, M, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381182" httpUri="https://zenodo.org/record/5381182/files/figure.png" pageId="28" pageNumber="445">Fig. 15K</figureCitation>
|
||
). Outer lip slightly thickened and, in labral view, sinuous near the adapical end of the aperture. Outer lip extending to the apex and continuing on the opposite side for a short distance (
|
||
<figureCitation id="133435DBFF8B1E43FE3DFAB1FDC4FAB3" box="[475,635,1356,1383]" captionStart="FIG" captionStartId="24.[113,124,1544,1561]" captionTargetBox="[126,1202,227,1464]" captionTargetId="figure-192@24.[113,1210,214,1500]" captionTargetPageId="24" captionText="FIG. 15. — Seraphs Montfort, 1810 shells in normal light; A-I, S. chilophorus (Cossmann, 1889); A-E, MNHN A28779 (Ledon coll.), Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, labral view; C, detailed view of the apical part; D, dorsal view;E, detailed view of the apical part; F-H, MNHN A28780 (Ledon coll.), Chaussy (Les Garennes), Lutetian;F, ventral view;G, dorsal view; H, detailed view of the apical part; I, MNHN A27783, detailed view the oblique grooves (leg. Pacaud), Saint-Lubin-de-la-Haye, Eure-et-Loir, France, Lutetian; J-N, S. peterjungi n. sp., holotype MNHN A28578 (Pacaud leg.), Chaussy (Les Garennes), Lutetian; J, ventral view; K, labral view; L, detailed view of the apical part; M, detailed view the oblique grooves; N, dorsal view. Scale bars: A-B, D, F-G, J-K, N, 10 mm; C, I, 5 mm; E, H, L, M, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381182" httpUri="https://zenodo.org/record/5381182/files/figure.png" pageId="28" pageNumber="445">Fig. 15K, L, N</figureCitation>
|
||
). Siphonal notch moderately deep on the dorsal side of the shell. Surface sculpture of narrow oblique grooves near the base (
|
||
<figureCitation id="133435DBFF8B1E43FE83FA56FE70FA12" box="[357,463,1451,1478]" captionStart="FIG" captionStartId="24.[113,124,1544,1561]" captionTargetBox="[126,1202,227,1464]" captionTargetId="figure-192@24.[113,1210,214,1500]" captionTargetPageId="24" captionText="FIG. 15. — Seraphs Montfort, 1810 shells in normal light; A-I, S. chilophorus (Cossmann, 1889); A-E, MNHN A28779 (Ledon coll.), Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, labral view; C, detailed view of the apical part; D, dorsal view;E, detailed view of the apical part; F-H, MNHN A28780 (Ledon coll.), Chaussy (Les Garennes), Lutetian;F, ventral view;G, dorsal view; H, detailed view of the apical part; I, MNHN A27783, detailed view the oblique grooves (leg. Pacaud), Saint-Lubin-de-la-Haye, Eure-et-Loir, France, Lutetian; J-N, S. peterjungi n. sp., holotype MNHN A28578 (Pacaud leg.), Chaussy (Les Garennes), Lutetian; J, ventral view; K, labral view; L, detailed view of the apical part; M, detailed view the oblique grooves; N, dorsal view. Scale bars: A-B, D, F-G, J-K, N, 10 mm; C, I, 5 mm; E, H, L, M, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381182" httpUri="https://zenodo.org/record/5381182/files/figure.png" pageId="28" pageNumber="445">Fig. 15M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8B1E43FF97FA11FE7AF9D7" blockId="28.[112,647,1516,1700]" box="[113,453,1516,1542]" pageId="28" pageNumber="445">COLOUR PATTERN DESCRIPTION</paragraph>
|
||
<paragraph id="8BB0295EFF8B1E43FF97F9F7FB2EFA9C" blockId="28.[112,647,1516,1700]" lastBlockId="28.[674,1210,944,1352]" pageId="28" pageNumber="445">
|
||
The colour pattern shows two different and superimposed levels of residual pigmentation: a dark one and a white and fluorescent one. The background displays an intermediate colour between these two levels. Th e darker elements correspond to several thick spiral rows of patches. Th e lighter fluorescent elements, overlaid on the darker, consist of narrow spiral lines. Th e morphology of the dark elements is similar to the pattern of
|
||
<taxonomicName id="4C0F52DDFF8B1E43FC79FBEDFB95FBFD" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[927,1066,1039,1066]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="445" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8B1E43FC79FBEDFB95FBFD" box="[927,1066,1039,1066]" italics="true" pageId="28" pageNumber="445">S. chilophorus</emphasis>
|
||
</taxonomicName>
|
||
: several spiral rows of dark triangles of which the apex is directed towards the outer lip. Th eir width, depending on the size of the patches, is variable (
|
||
<figureCitation id="133435DBFF8B1E43FBF6FB92FB30FB5E" box="[1040,1167,1135,1162]" captionStart="FIG" captionStartId="30.[113,124,1589,1606]" captionTargetBox="[113,1210,215,1520]" captionTargetId="figure-133@30.[113,1210,234,1520]" captionTargetPageId="30" captionText="FIG. 19. — Seraphs peterjungi n. sp. under UV light: A, B, holotype MNHN A28578,Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, dorsal view; C, D, paratype MNHN A28537 (Faullummel coll.), Chaussy (Les Garennes), Lutetian; C, ventral view; D, dorsal view; E, F, paratype MNHN A27273 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; E, ventral view; F, dorsal view; G, H, paratype MNHN A28931 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; G, detailed view of the dots; H, dorsal view. Scale bars: A-F, H, 10 mm; G, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381190" httpUri="https://zenodo.org/record/5381190/files/figure.png" pageId="28" pageNumber="445">Fig. 19A, B</figureCitation>
|
||
). In addition, some tiny and isolated triangles, orientated towards the outer lip, are visible between the rows (
|
||
<figureCitation id="133435DBFF8B1E43FD4AFB33FC87FB3D" box="[684,824,1230,1257]" captionStart="FIG" captionStartId="30.[113,124,1589,1606]" captionTargetBox="[113,1210,215,1520]" captionTargetId="figure-133@30.[113,1210,234,1520]" captionTargetPageId="30" captionText="FIG. 19. — Seraphs peterjungi n. sp. under UV light: A, B, holotype MNHN A28578,Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, dorsal view; C, D, paratype MNHN A28537 (Faullummel coll.), Chaussy (Les Garennes), Lutetian; C, ventral view; D, dorsal view; E, F, paratype MNHN A27273 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; E, ventral view; F, dorsal view; G, H, paratype MNHN A28931 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; G, detailed view of the dots; H, dorsal view. Scale bars: A-F, H, 10 mm; G, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381190" httpUri="https://zenodo.org/record/5381190/files/figure.png" pageId="28" pageNumber="445">Fig. 19G, H</figureCitation>
|
||
). Th eir size, distribution and shape are heterogenous and irregular (
|
||
<figureCitation id="133435DBFF8B1E43FC1EFB10FB3EFADC" box="[1016,1153,1261,1288]" captionStart="FIG" captionStartId="30.[113,124,1589,1606]" captionTargetBox="[113,1210,215,1520]" captionTargetId="figure-133@30.[113,1210,234,1520]" captionTargetPageId="30" captionText="FIG. 19. — Seraphs peterjungi n. sp. under UV light: A, B, holotype MNHN A28578,Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, dorsal view; C, D, paratype MNHN A28537 (Faullummel coll.), Chaussy (Les Garennes), Lutetian; C, ventral view; D, dorsal view; E, F, paratype MNHN A27273 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; E, ventral view; F, dorsal view; G, H, paratype MNHN A28931 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; G, detailed view of the dots; H, dorsal view. Scale bars: A-F, H, 10 mm; G, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381190" httpUri="https://zenodo.org/record/5381190/files/figure.png" pageId="28" pageNumber="445">Fig. 19G, H</figureCitation>
|
||
). On the apical part of the shell, the elements are more dense and coalescent, tending to be shapeless.
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8B1E43FD43FA91FC99FA51" blockId="28.[674,1210,1388,1700]" box="[677,806,1388,1414]" pageId="28" pageNumber="445">VARIABILITY</paragraph>
|
||
<paragraph id="8BB0295EFF8B1E42FD43FA71FDF7FF24" blockId="28.[674,1210,1388,1700]" lastBlockId="29.[106,647,217,496]" lastPageId="29" lastPageNumber="446" pageId="28" pageNumber="445">
|
||
The variability is only observable on the darker elements of the residual pattern. Th e rows can be more or less distant, from widely spaced to contiguous (
|
||
<figureCitation id="133435DBFF8B1E43FD4AFA16FCF8F9D2" box="[684,839,1515,1542]" captionStart="FIG" captionStartId="30.[113,124,1589,1606]" captionTargetBox="[113,1210,215,1520]" captionTargetId="figure-133@30.[113,1210,234,1520]" captionTargetPageId="30" captionText="FIG. 19. — Seraphs peterjungi n. sp. under UV light: A, B, holotype MNHN A28578,Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, dorsal view; C, D, paratype MNHN A28537 (Faullummel coll.), Chaussy (Les Garennes), Lutetian; C, ventral view; D, dorsal view; E, F, paratype MNHN A27273 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; E, ventral view; F, dorsal view; G, H, paratype MNHN A28931 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; G, detailed view of the dots; H, dorsal view. Scale bars: A-F, H, 10 mm; G, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381190" httpUri="https://zenodo.org/record/5381190/files/figure.png" pageId="28" pageNumber="445">Fig. 19A, E, F</figureCitation>
|
||
). Dark patches can show considerable shape variation from triangular to completely distorted (
|
||
<figureCitation id="133435DBFF8B1E43FCEAF9D7FC3EF991" box="[780,897,1578,1605]" captionStart="FIG" captionStartId="30.[113,124,1589,1606]" captionTargetBox="[113,1210,215,1520]" captionTargetId="figure-133@30.[113,1210,234,1520]" captionTargetPageId="30" captionText="FIG. 19. — Seraphs peterjungi n. sp. under UV light: A, B, holotype MNHN A28578,Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, dorsal view; C, D, paratype MNHN A28537 (Faullummel coll.), Chaussy (Les Garennes), Lutetian; C, ventral view; D, dorsal view; E, F, paratype MNHN A27273 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; E, ventral view; F, dorsal view; G, H, paratype MNHN A28931 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; G, detailed view of the dots; H, dorsal view. Scale bars: A-F, H, 10 mm; G, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381190" httpUri="https://zenodo.org/record/5381190/files/figure.png" pageId="28" pageNumber="445">Fig. 19E, F</figureCitation>
|
||
). Th ey can be well separated or connected. The fluorescent elements, narrow spiral lines, do not show perceptible variation except in their number: between 16 and 20. SHELL AND COLOUR PATTERN COMPARISONS
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8A1E42FF97FF05FEB4FEA6" blockId="29.[106,647,217,496]" pageId="29" pageNumber="446">
|
||
The shell of
|
||
<taxonomicName id="4C0F52DDFF8A1E42FF0AFF0AFE1FFEC5" authority="Caze & Merle & Pacaud & Saint Martin, 2010" authorityName="Caze & Merle & Pacaud & Saint Martin" authorityYear="2010" box="[236,416,247,273]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="446" phylum="Mollusca" rank="species" species="peterjungi" status="sp. nov.">
|
||
<emphasis id="B97BF54CFF8A1E42FF0AFF0AFE1FFEC5" box="[236,416,247,273]" italics="true" pageId="29" pageNumber="446">Seraphs peterjungi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2484837FF8A1E42FE43FF05FE63FEC7" box="[421,476,248,275]" pageId="29" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
is indistinguishable from that of
|
||
<taxonomicName id="4C0F52DDFF8A1E42FECDFEEAFE4CFEE5" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[299,499,279,305]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="446" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8A1E42FECDFEEAFE4CFEE5" box="[299,499,279,305]" italics="true" pageId="29" pageNumber="446">Seraphs chilophorus</emphasis>
|
||
</taxonomicName>
|
||
. Accordingly, for shell comparison with the other species, see
|
||
<taxonomicName id="4C0F52DDFF8A1E42FF95FEAAFEBBFEA4" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[115,260,342,369]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="446" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8A1E42FF95FEAAFEBBFEA4" box="[115,260,342,369]" italics="true" pageId="29" pageNumber="446">S. chilophorus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8A1E42FF6EFE8BFE5BFE24" blockId="29.[106,647,217,496]" pageId="29" pageNumber="446">
|
||
The dark underlying elements of
|
||
<taxonomicName id="4C0F52DDFF8A1E42FE19FE8AFD38FE44" authority="Caze & Merle & Pacaud & Saint Martin, 2010" authorityName="Caze & Merle & Pacaud & Saint Martin" authorityYear="2010" box="[511,647,375,401]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="446" phylum="Mollusca" rank="species" species="peterjungi" status="sp. nov.">
|
||
<emphasis id="B97BF54CFF8A1E42FE19FE8AFD38FE44" box="[511,647,375,401]" italics="true" pageId="29" pageNumber="446">S. peterjungi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2484837FF8A1E42FF8CFE6BFF1CFE65" box="[106,163,406,433]" pageId="29" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
resemble the colour pattern of
|
||
<taxonomicName id="4C0F52DDFF8A1E42FE17FE6BFD3FFE64" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[497,640,406,432]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="446" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8A1E42FE17FE6BFD3FFE64" box="[497,640,406,432]" italics="true" pageId="29" pageNumber="446">S. chilophorus</emphasis>
|
||
</taxonomicName>
|
||
, but it appears to be less variable.
|
||
<taxonomicName id="4C0F52DDFF8A1E42FE24FE48FD39FE1B" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[450,646,437,463]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="446" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8A1E42FE24FE48FD39FE1B" box="[450,646,437,463]" italics="true" pageId="29" pageNumber="446">Seraphs chilophorus</emphasis>
|
||
</taxonomicName>
|
||
never has fluorescent narrow lines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3157AD5FF8A1E42FF97FDE8FE02FB9E" pageId="29" pageNumber="446" type="discussion">
|
||
<paragraph id="8BB0295EFF8A1E42FF97FDE8FF4CFDF9" blockId="29.[111,648,533,1098]" box="[113,243,533,559]" pageId="29" pageNumber="446">DISCUSSION</paragraph>
|
||
<paragraph id="8BB0295EFF8A1E42FF97FDC9FE02FB9E" blockId="29.[111,648,533,1098]" pageId="29" pageNumber="446">
|
||
The comparisons between
|
||
<taxonomicName id="4C0F52DDFF8A1E42FE77FDC8FDABFD9A" authority="Caze & Merle & Pacaud & Saint Martin, 2010" authorityName="Caze & Merle & Pacaud & Saint Martin" authorityYear="2010" box="[401,532,565,591]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="446" phylum="Mollusca" rank="species" species="peterjungi" status="sp. nov.">
|
||
<emphasis id="B97BF54CFF8A1E42FE77FDC8FDABFD9A" box="[401,532,565,591]" italics="true" pageId="29" pageNumber="446">S. peterjungi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2484837FF8A1E42FDFDFDC9FDE8FD9B" box="[539,599,564,591]" pageId="29" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C0F52DDFF8A1E42FF94FDA9FEBBFDBA" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[114,260,596,622]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="446" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8A1E42FF94FDA9FEBBFDBA" box="[114,260,596,622]" italics="true" pageId="29" pageNumber="446">S. chilophorus</emphasis>
|
||
</taxonomicName>
|
||
are respectively based on
|
||
<specimenCount id="9D09E2D7FF8A1E42FDC2FDA9FF15FD5B" pageId="29" pageNumber="446" type="generic">24 specimens</specimenCount>
|
||
and
|
||
<specimenCount id="9D09E2D7FF8A1E42FF3BFD89FEC6FD5B" box="[221,377,628,655]" pageId="29" pageNumber="446" type="generic">163 specimens</specimenCount>
|
||
(
|
||
<tableCitation id="C68D1CE5FF8A1E42FE63FD89FE6BFD5B" box="[389,468,628,655]" captionStart="TABLE" captionStartId="9.[285,294,219,236]" captionText="TABLE 1. — Comparison between positive and" pageId="29" pageNumber="446">Table 1</tableCitation>
|
||
). Using the shell shape, no distinction can be made between the two populations. Using the residual pattern, we can observe a strong similarity in the dark component, suggesting a close relationship. However,
|
||
<taxonomicName id="4C0F52DDFF8A1E42FDD6FD0EFF19FCF8" authority="Caze & Merle & Pacaud & Saint Martin, 2010" authorityName="Caze & Merle & Pacaud & Saint Martin" authorityYear="2010" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="446" phylum="Mollusca" rank="species" species="peterjungi" status="sp. nov.">
|
||
<emphasis id="B97BF54CFF8A1E42FDD6FD0EFF19FCF8" italics="true" pageId="29" pageNumber="446">S. peterjungi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2484837FF8A1E42FF56FCEFFF56FCF9" box="[176,233,786,813]" pageId="29" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
shows an additional element with some light and fluorescent spiral lines (
|
||
<figureCitation id="133435DBFF8A1E42FE35FCCFFD9DFC99" box="[467,546,818,845]" captionStart="FIG" captionStartId="30.[113,124,1589,1606]" captionTargetBox="[113,1210,215,1520]" captionTargetId="figure-133@30.[113,1210,234,1520]" captionTargetPageId="30" captionText="FIG. 19. — Seraphs peterjungi n. sp. under UV light: A, B, holotype MNHN A28578,Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, dorsal view; C, D, paratype MNHN A28537 (Faullummel coll.), Chaussy (Les Garennes), Lutetian; C, ventral view; D, dorsal view; E, F, paratype MNHN A27273 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; E, ventral view; F, dorsal view; G, H, paratype MNHN A28931 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; G, detailed view of the dots; H, dorsal view. Scale bars: A-F, H, 10 mm; G, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381190" httpUri="https://zenodo.org/record/5381190/files/figure.png" pageId="29" pageNumber="446">Fig. 19</figureCitation>
|
||
), lacking in all examined
|
||
<taxonomicName id="4C0F52DDFF8A1E42FEC5FCAFFE09FCBF" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[291,438,849,876]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="446" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8A1E42FEC5FCAFFE09FCBF" box="[291,438,849,876]" italics="true" pageId="29" pageNumber="446">S. chilophorus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133435DBFF8A1E42FE21FCACFDA8FCB8" box="[455,535,849,876]" captionStart="FIG" captionStartId="26.[113,124,1589,1606]" captionTargetBox="[114,1210,215,1525]" captionTargetId="figure-144@26.[113,1210,240,1526]" captionTargetPageId="26" captionText="FIG. 16. — Seraphs chilophorus (Cossmann, 1889) under UV light: A, B, MNHN A28927 (Roissy coll.), Parnes, Oise, France, Lutetian; A, ventral view; B, dorsal view; C-E, neotype MNHN A25009, Fontenay-en-Vexin, Eure, France, Lutetian; C, ventral view; D, dorsal view; E, detailed view of the dots; F-H, MNHN A28924 (leg. Pacaud), Fercourt, Oise, France, Lutetian; F, dorsal view; G, H, detailed views of the dots; I-K, MNHN A28928 (leg. Pacaud), Parnes, Oise, France, Lutetian; I, ventral view; J, K, dorsal views (K negative view). Scale bars: A-D, F, I-K, 10 mm; E, G, H, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381184" httpUri="https://zenodo.org/record/5381184/files/figure.png" pageId="29" pageNumber="446">Fig. 16</figureCitation>
|
||
) and suggesting a morphological discontinuity. Considering the number of specimens studied and the lack of intermediaries between the two colour patterns, we cannot regard both populations as a single species. In this case, the residual pattern does seem to be a unique discriminating criterion enabling the differentiation of fossil species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |