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<document ID-DOI="http://doi.org/10.5281/zenodo.4758931" ID-GBIF-Dataset="93917b8d-1ce7-425e-912a-5892214c81b4" ID-GBIF-Taxon="182073759" ID-ISSN="1854-0392" ID-Zenodo-Dep="4758931" approvalRequired="4" approvalRequired_for_textStreams="4" checkinTime="1620936844672" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Sheldon, Andrew L." docDate="2009" docId="03A621374904FF90FE9BE80CFB05357E" docLanguage="en" docName="Illiesia.5.2.11-19.pdf" docOrigin="Illiesia 5 (2)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Phanoperla belalong Stark &amp; Sheldon 2009, sp. n." docType="treatment" docVersion="6" lastPageNumber="18" masterDocId="FF9F594F4903FF97FFD4E926FFE6341B" masterDocTitle="Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species" masterLastPageNumber="19" masterPageNumber="11" pageId="7" pageNumber="18" updateTime="1632022693049" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>and Andrew L. Sheldon &amp; Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sheldon, Andrew L.</mods:namePart>
<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu &amp; Division of Biological Sciences, University of Montana, Missoula, Montana, U. S. A. 59812 E-mail: andylsheldon @ comcast. net &amp; and Andrew L. Sheldon &amp; Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-02-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>11</mods:start>
<mods:end>19</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4759022</mods:identifier>
<mods:identifier type="GBIF-Dataset">93917b8d-1ce7-425e-912a-5892214c81b4</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4759022</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4758931" ID-GBIF-Taxon="182073759" ID-Zenodo-Dep="4758931" LSID="urn:lsid:plazi:treatment:03A621374904FF90FE9BE80CFB05357E" httpUri="http://treatment.plazi.org/id/03A621374904FF90FE9BE80CFB05357E" lastPageNumber="18" pageId="7" pageNumber="18">
<subSubSection box="[335,651,298,322]" pageId="7" pageNumber="18" type="nomenclature">
<paragraph blockId="7.[335,651,298,322]" box="[335,651,298,322]" pageId="7" pageNumber="18">
<heading bold="true" box="[335,651,298,322]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="18" reason="2">
<emphasis bold="true" box="[335,651,298,322]" pageId="7" pageNumber="18">
<taxonomicName authority="Stark &amp; Sheldon, 2009" authorityName="Stark &amp; Sheldon" authorityYear="2009" box="[335,580,298,322]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="belalong" status="sp. n.">
<emphasis bold="true" box="[335,580,298,322]" italics="true" pageId="7" pageNumber="18">Phanoperla belalong</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[587,651,298,322]" pageId="7" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[422,564,334,358]" pageId="7" pageNumber="18" type="description">
<paragraph blockId="7.[422,564,334,358]" box="[422,564,334,358]" pageId="7" pageNumber="18">
(
<figureCitation box="[431,555,334,358]" captionStart="Figs" captionStartId="6.[189,240,1434,1458]" captionTargetBox="[324,1344,217,1354]" captionTargetId="figure-254@6.[324,1307,217,1354]" captionTargetPageId="6" captionText="Figs. 11‐14. Phanoperla astrospinata (11‐12) and P. belalong (13‐14). 11. Male terminalia, 12. Aedeagus, lateral aspect, 13. Male terminalia, 14. Aedeagus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4759032" httpUri="https://zenodo.org/record/4759032/files/figure.png" pageId="7" pageNumber="18">Figs. 13‐14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="18" type="materials_examined">
<paragraph blockId="7.[189,798,404,1927]" pageId="7" pageNumber="18">
<emphasis bold="true" box="[189,440,405,429]" pageId="7" pageNumber="18">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111531310" collectingDate="2008-07-03" collectionCode="ALS, USNM" collectorName="A. L. Sheldon" country="Brunei" latitude="4.54822" location="Kuala Belalong Field Studies Centre" longLatPrecision="1" longitude="115.15823" municipality="Sungai Belalong" pageId="7" pageNumber="18" specimenCode="B39-08" specimenCount="1" specimenCount-male="1" stateProvince="Temburong" typeStatus="holotype">
<typeStatus box="[462,573,405,429]" pageId="7" pageNumber="18">Holotype</typeStatus>
<specimenCount box="[596,616,404,429]" pageId="7" pageNumber="18" type="male">♂</specimenCount>
from
<collectingCountry name="Brunei" pageId="7" pageNumber="18">Brunei Darussalam</collectingCountry>
,
<collectingRegion box="[346,583,441,465]" country="Brunei" name="Temburong" pageId="7" pageNumber="18">Temburong District</collectingRegion>
,
<collectingMunicipality box="[597,793,441,465]" pageId="7" pageNumber="18">Sungai Belalong</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03A621374904FF90FE9BE80CFB05357E:8ED0C6FA4904FF90FF69E8F8FD9935ED" box="[189,639,478,502]" country="Brunei" latitude="4.54822" longLatPrecision="1" longitude="115.15823" municipality="Sungai Belalong" name="Kuala Belalong Field Studies Centre" pageId="7" pageNumber="18" stateProvince="Temburong">Kuala Belalong Field Studies Centre</location>
,
<geoCoordinate box="[656,792,473,503]" degrees="04.54822" direction="north" orientation="latitude" pageId="7" pageNumber="18" precision="1" value="4.54822">
04.54822
<emphasis bold="true" box="[756,769,473,503]" pageId="7" pageNumber="18">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate box="[189,333,511,541]" degrees="115.15823" direction="east" orientation="longitude" pageId="7" pageNumber="18" precision="1" value="115.15823">
115.15823
<emphasis bold="true" box="[302,315,511,541]" pageId="7" pageNumber="18">°</emphasis>
E
</geoCoordinate>
,
<collectionCode box="[346,397,516,540]" pageId="7" pageNumber="18">ALS</collectionCode>
<specimenCode box="[405,485,516,540]" collectionCode="ALS" pageId="7" pageNumber="18">B39‐08</specimenCode>
,
<date box="[498,631,515,540]" pageId="7" pageNumber="18" value="2008-07-03">
<collectingDate box="[498,631,515,540]" pageId="7" pageNumber="18" value="2008-07-03">3 July 2008</collectingDate>
</date>
,
<collectorName box="[643,797,515,540]" pageId="7" pageNumber="18">A.L. Sheldon</collectorName>
(
<collectionCode box="[197,284,551,575]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="18" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111531302" collectingDate="2008-06-18" collectionCode="ALS" collectorName="Same" country="Brunei" pageId="7" pageNumber="18" specimenCode="B24-08" specimenCount="1" stateProvince="Temburong" typeStatus="paratype">
<typeStatus box="[306,426,551,575]" pageId="7" pageNumber="18">Paratypes</typeStatus>
.
<collectorName box="[440,503,551,575]" pageId="7" pageNumber="18">Same</collectorName>
site but
<collectionCode box="[614,665,551,575]" pageId="7" pageNumber="18">ALS</collectionCode>
<specimenCode box="[675,756,551,575]" collectionCode="ALS" pageId="7" pageNumber="18">B24‐08</specimenCode>
,
<date pageId="7" pageNumber="18" value="2008-06-18">
<collectingDate pageId="7" pageNumber="18" value="2008-06-18">18 June 2008</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3111531330" collectingDate="2008-06-27" collectionCode="BPS, ALS" collectorName="A. L. Sheldon" country="Brunei" location="Same" pageId="7" pageNumber="18" specimenCode="B36-08" specimenCount="1" specimenCount-male="1" stateProvince="Temburong" typeStatus="paratype">
<collectorName box="[312,464,586,611]" pageId="7" pageNumber="18">A.L. Sheldon</collectorName>
,
<specimenCount box="[473,511,586,611]" pageId="7" pageNumber="18" type="male">1 ♂</specimenCount>
(
<collectionCode box="[524,575,587,611]" collectionName="BPS" country="USA" lsid="urn:lsid:biocol.org:col:14922" name="California Department of Food and Agriculture" pageId="7" pageNumber="18" type="Herbarium">BPS</collectionCode>
).
<location LSID="urn:lsid:plazi:treatment:03A621374904FF90FE9BE80CFB05357E:8ED0C6FA4904FF90FD9BEB6DFD683678" box="[591,654,587,611]" country="Brunei" name="Same" pageId="7" pageNumber="18" stateProvince="Temburong">Same</location>
site but
<collectionCode box="[747,797,587,611]" pageId="7" pageNumber="18">ALS</collectionCode>
<specimenCode box="[189,270,622,646]" collectionCode="ALS" pageId="7" pageNumber="18">B36‐08</specimenCode>
,
<date box="[281,430,622,646]" pageId="7" pageNumber="18" value="2008-06-27">
<collectingDate box="[281,430,622,646]" pageId="7" pageNumber="18" value="2008-06-27">27 June 2008</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3111531303" box="[440,721,621,646]" collectionCode="BPS" collectorName="A. L. Sheldon" country="Brunei" pageId="7" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Temburong" typeStatus="paratype">
<collectorName box="[440,596,622,646]" pageId="7" pageNumber="18">A.L. Sheldon</collectorName>
,
<specimenCount box="[607,647,621,646]" pageId="7" pageNumber="18" type="male">1 ♂</specimenCount>
(
<collectionCode box="[661,712,622,646]" collectionName="BPS" country="USA" lsid="urn:lsid:biocol.org:col:14922" name="California Department of Food and Agriculture" pageId="7" pageNumber="18" type="Herbarium">BPS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="18" type="description">
<paragraph blockId="7.[189,798,404,1927]" pageId="7" pageNumber="18">
<emphasis bold="true" box="[189,364,657,681]" pageId="7" pageNumber="18">Adult habitus.</emphasis>
Body color pale yellow brown. Head yellow brown without pattern; ocelli almost touch. Basal 4‐5 antennal segments pale, rest of flagellum brown. Pronotum pale with darker rugosities and dark brown median and marginal sutures. Femora pale, tibiae brown.Wings pale, veins amber.
</paragraph>
<paragraph blockId="7.[189,798,404,1927]" pageId="7" pageNumber="18">
<emphasis bold="true" box="[189,258,871,895]" pageId="7" pageNumber="18">Male.</emphasis>
Forewing length
<quantity box="[524,634,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="18" unit="mm" value="7.5">7.5 mm</quantity>
. Tergum 8 unmodified, tergum 9 with lateral and mesal sensilla basiconica patches united, or almost so (
<figureCitation box="[698,784,942,966]" captionStart="Figs" captionStartId="6.[189,240,1434,1458]" captionTargetBox="[324,1344,217,1354]" captionTargetId="figure-254@6.[324,1307,217,1354]" captionTargetPageId="6" captionText="Figs. 11‐14. Phanoperla astrospinata (11‐12) and P. belalong (13‐14). 11. Male terminalia, 12. Aedeagus, lateral aspect, 13. Male terminalia, 14. Aedeagus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4759032" httpUri="https://zenodo.org/record/4759032/files/figure.png" pageId="7" pageNumber="18">Fig. 13</figureCitation>
). Hemiterga with attenuated tips; basal callus offset from apical section of hemiterga by broad, shallow emargination. Aedeagal tube short and plump with a few fine setal spines on bulb; apex of tube bearing a pair of ventromedian lobes and a pair of dorsolateral lobes (
<figureCitation box="[268,352,1155,1180]" captionStart="Figs" captionStartId="6.[189,240,1434,1458]" captionTargetBox="[324,1344,217,1354]" captionTargetId="figure-254@6.[324,1307,217,1354]" captionTargetPageId="6" captionText="Figs. 11‐14. Phanoperla astrospinata (11‐12) and P. belalong (13‐14). 11. Male terminalia, 12. Aedeagus, lateral aspect, 13. Male terminalia, 14. Aedeagus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4759032" httpUri="https://zenodo.org/record/4759032/files/figure.png" pageId="7" pageNumber="18">Fig. 14</figureCitation>
); apical half of tube, including lobes armed with fine triangular spines. Aedeagal sac bearing lateral groups of 6‐7 large black spines in fan shaped clusters on either side of a cylindrical lobe densely armed with black, scale‐ like spines; dorsad to large fan shaped clusters, a small membranous lobe, armed with brown, triangular spines occurs on each side of sac. Dorsal sclerite spear shaped but apex curved dorsad over base of sac.
</paragraph>
<paragraph blockId="7.[189,798,404,1927]" box="[189,415,1475,1499]" pageId="7" pageNumber="18">
<emphasis bold="true" box="[189,283,1475,1499]" pageId="7" pageNumber="18">Female.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="7.[189,798,404,1927]" box="[189,396,1511,1535]" pageId="7" pageNumber="18">
<emphasis bold="true" box="[189,264,1511,1535]" pageId="7" pageNumber="18">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="18" type="etymology">
<paragraph blockId="7.[189,798,404,1927]" pageId="7" pageNumber="18">
<emphasis bold="true" box="[189,327,1547,1571]" pageId="7" pageNumber="18">Etymology.</emphasis>
The species name, used as a noun in apposition, is based on the
<typeStatus box="[525,577,1583,1607]" pageId="7" pageNumber="18">type</typeStatus>
locality of Sungai Belalong and the Kuala Belalong Field Studies Centre.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="18" type="diagnosis">
<paragraph blockId="7.[189,798,404,1927]" lastBlockId="7.[835,1443,227,357]" pageId="7" pageNumber="18">
<emphasis bold="true" box="[189,318,1689,1713]" pageId="7" pageNumber="18">Diagnosis.</emphasis>
The aedeagal features of this species are similar to both
<taxonomicName authority="Stark &amp; Sivec" authorityName="Stark &amp; Sivec" authorityYear="2007" box="[376,677,1725,1749]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="flabellare">
<emphasis box="[376,506,1725,1749]" italics="true" pageId="7" pageNumber="18">P. flabellare</emphasis>
Stark &amp; Sivec
</taxonomicName>
and to
<taxonomicName authority="(Banks)" baseAuthorityName="Banks" baseAuthorityYear="1939" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="anomala">
<emphasis italics="true" pageId="7" pageNumber="18">P. anomala</emphasis>
(Banks)
</taxonomicName>
(
<bibRefCitation author="Stark, B. P. &amp; I. Sivec" box="[390,612,1761,1785]" pageId="7" pageNumber="18" pagination="53 - 64" refId="ref4750" refString="Stark, B. P. &amp; I. Sivec. 2007. Studies of Indonesian Perlidae (Plecoptera), with descriptions of three new species. Illiesia, 3: 53 - 64." type="journal article" year="2007">Stark &amp; Sivec 2007</bibRefCitation>
;
<bibRefCitation author="Zwick, P." box="[623,761,1761,1785]" pageId="7" pageNumber="18" pagination="87 - 126" refId="ref4786" refString="Zwick, P. 1982. A revision of the Oriental stonefly genus Phanoperla (Plecoptera: Perlidae). Systematic Entomology, 7: 87 - 126." type="journal article" year="1982">Zwick 1982</bibRefCitation>
). It differs from the former in bearing two pairs of prominent, membranous, spine‐ covered lobes ventroapically and dorsolaterally on the tube. In addition, a small spiny lobe occurs dorsad to the large fan shaped clusters of spines on the sac. The latter species shares the ventroapical spiny lobes on the aedeagal tube but the sac is armed with an almost complete ring of large black spines.
</paragraph>
</subSubSection>
</treatment>
</document>
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