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<document id="77C064A55DEBBEA6B51023A552CE67C5" ID-DOI="10.1649/0010-065X-77.3.432" ID-ISSN="1938-4394" ID-Zenodo-Dep="13243269" ID-ZooBank="urn:lsid:zoobank.org:pub:0C6B0BEF-48DF-4F5A-93C9-F8E20E7F8948" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722968213433" checkinUser="felipe" docAuthor="Caterino, Michael S. &amp; Yamamoto, Shûhei" docDate="2023" docId="03A487FEFF88FF9B05B1FEC57F6FFDC8" docLanguage="en" docName="TheColeopBull.77.3.432-438.pdf" docOrigin="The Coleopterists Bulletin 77 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-77.3.432" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Phasmister hkamticus Caterino and Yamamoto 2023, new species" docType="treatment" docVersion="2" lastPageNumber="435" masterDocId="FF9DFF86FF8AFF9F0775FFF37B47FE45" masterDocTitle="New Onthophiline Fossil Species (Coleoptera: Histeridae: Onthophilinae) from Mid-Cretaceous Burmese Amber" masterLastPageNumber="438" masterPageNumber="432" pageNumber="433" updateTime="1722971481944" updateUser="ExternalLinkService">
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<mods:title id="35F6BE36E1348E922D47004568E2D6B5">New Onthophiline Fossil Species (Coleoptera: Histeridae: Onthophilinae) from Mid-Cretaceous Burmese Amber</mods:title>
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<mods:namePart id="8A1E0F555205DF01AEBC35CED4514260">Caterino, Michael S.</mods:namePart>
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<mods:namePart id="E8ABB939E145BF3E3D7C89F1ECF559F9">Yamamoto, Shûhei</mods:namePart>
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<mods:title id="1C89A5685A14AF19470B349CDE10CA4E">The Coleopterists Bulletin</mods:title>
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<mods:date id="6B3EE219E3B948B415E7AD3DD1746D86">2023</mods:date>
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<treatment id="03A487FEFF88FF9B05B1FEC57F6FFDC8" LSID="urn:lsid:plazi:treatment:03A487FEFF88FF9B05B1FEC57F6FFDC8" httpUri="http://treatment.plazi.org/id/03A487FEFF88FF9B05B1FEC57F6FFDC8" lastPageId="4" lastPageNumber="435" pageId="2" pageNumber="433">
<subSubSection id="C3176563FF88FF9D05B1FEC578B5FFFD" pageId="2" pageNumber="433" type="nomenclature">
<paragraph id="8BB236E8FF88FF9D05B1FEC578B5FFFD" blockId="2.[708,1190,310,441]" pageId="2" pageNumber="433">
<heading id="D0FA8184FF88FF9D05B1FEC578B6FF2C" centered="true" fontSize="9" level="2" pageId="2" pageNumber="433" reason="2">
<emphasis id="B979EAFAFF88FF9D05B1FEC578B6FF2C" bold="true" pageId="2" pageNumber="433">
<taxonomicName id="4C0D4D6BFF88FF9D05B1FEC57FE5FF0E" authority="Caterino and Yamamoto" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[708,1186,310,332]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="hkamticus" status="sp. nov.">
<emphasis id="B979EAFAFF88FF9D05B1FEC578E7FF09" bold="true" box="[708,928,310,332]" italics="true" pageId="2" pageNumber="433">Phasmister hkamticus</emphasis>
Caterino and Yamamoto
</taxonomicName>
,
<taxonomicNameLabel id="A24A5781FF88FF9D040FFEA078B6FF2C" box="[890,1009,339,361]" pageId="2" pageNumber="433" rank="species">new species</taxonomicNameLabel>
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(
<figureCitation id="13362A6DFF88FF9D04F5FE507885FFFD" box="[896,962,419,441]" captionStart="Fig" captionStartId="2.[197,228,1700,1719]" captionTargetBox="[173,1193,659,1683]" captionTargetId="figure-194@2.[173,1193,659,1683]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Phasmister hkamticus, new species. A) Dorsal, B) Dorsolateral, C) Lateral, D) Ventral, E) Anterodorsal, F) Posterodorsal. All scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13243271" httpUri="https://zenodo.org/record/13243271/files/figure.png" pageId="2" pageNumber="433">Figs. 1</figureCitation>
,
<figureCitation id="13362A6DFF88FF9D04BBFE5078ACFFFD" box="[974,1003,419,440]" captionStart="Fig" captionStartId="4.[197,228,1192,1211]" captionTargetBox="[173,659,193,1175]" captionTargetId="figure-511@4.[173,659,193,1175]" captionTargetPageId="4" captionText="Fig. 3. Comparison of pronota of Phasmister spp. A) P. cristatus, B) P. hkamticus, new species, C) P. parallellus, new species." figureDoi="http://doi.org/10.5281/zenodo.13243277" httpUri="https://zenodo.org/record/13243277/files/figure.png" pageId="2" pageNumber="433">3B</figureCitation>
)
</paragraph>
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<subSubSection id="C3176563FF88FF9B05AEFE3878C9FF3D" lastPageId="4" lastPageNumber="435" pageId="2" pageNumber="433" type="materials_examined">
<paragraph id="8BB236E8FF88FF9D05AEFE3878D4FC23" blockId="2.[707,1193,459,614]" pageId="2" pageNumber="433">
<emphasis id="B979EAFAFF88FF9D05AEFE387837FFA4" bold="true" box="[731,880,459,481]" pageId="2" pageNumber="433">Type Material.</emphasis>
<materialsCitation id="3B653CB5FF88FF9D0400FE3878D4FC23" collectionCode="HUM" country="Myanmar" location="Hkamti District" municipality="Northern" pageId="2" pageNumber="433" specimenCount="2" stateProvince="Sagaing" typeStatus="holotype">
<typeStatus id="54B6884AFF88FF9D0400FE387889FFA4" box="[885,974,459,481]" pageId="2" pageNumber="433" type="holotype">Holotype</typeStatus>
specimen, of unknown sex (SEHU-0000121202); deposited in
<collectionCode id="ED1CAE2DFF88FF9D0340FE157F28FFBE" box="[1077,1135,486,507]" country="Germany" httpUri="http://grbio.org/cool/kch5-hv72" name="Humboldt University Zoologisches Museum" pageId="2" pageNumber="433">HUM</collectionCode>
.
<emphasis id="B979EAFAFF88FF9D0303FE157857FC53" bold="true" pageId="2" pageNumber="433">Type locality:</emphasis>
<collectingMunicipality id="6BD6AC92FF88FF9D0461FDF27821FC52" box="[788,870,513,535]" pageId="2" pageNumber="433">Northern</collectingMunicipality>
<collectingCountry id="F31A7678FF88FF9D041CFDF27882FC53" box="[873,965,513,534]" name="Myanmar" pageId="2" pageNumber="433">Myanmar</collectingCountry>
:
<collectingRegion id="49C9F80AFF88FF9D04B9FDF27F1AFC52" box="[972,1117,513,535]" country="Myanmar" name="Sagaing" pageId="2" pageNumber="433">Sagaing Region</collectingRegion>
,
<location id="8ED26033FF88FF9D0311FDF27855FC74" LSID="urn:lsid:plazi:treatment:03A487FEFF88FF9B05B1FEC57F6FFDC8:8ED26033FF88FF9D0311FDF27855FC74" country="Myanmar" municipality="Northern" name="Hkamti District" pageId="2" pageNumber="433" stateProvince="Sagaing">Hkamti District</location>
. The
<typeStatus id="54B6884AFF88FF9D043BFDE878E2FC74" box="[846,933,539,561]" pageId="2" pageNumber="433" type="holotype">holotype</typeStatus>
is embedded in a small, elongate-oval, yellowish transparent amber piece (9.8 × 4.9 ×
<quantity id="4CF59B0DFF88FF9D044FFDA278C0FC23" box="[826,903,593,614]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="433" unit="mm" value="1.7">1.7 mm</quantity>
).
</materialsCitation>
</paragraph>
<caption id="DF726660FF88FF9D07B0F95779D0F895" ID-DOI="http://doi.org/10.5281/zenodo.13243271" ID-Zenodo-Dep="13243271" httpUri="https://zenodo.org/record/13243271/files/figure.png" pageId="2" pageNumber="433" startId="2.[197,228,1700,1719]" targetBox="[173,1193,659,1683]" targetPageId="2" targetType="figure">
<paragraph id="8BB236E8FF88FF9D07B0F9577FEEF8F2" blockId="2.[173,1193,1700,1744]" box="[197,1193,1700,1720]" pageId="2" pageNumber="433">
<emphasis id="B979EAFAFF88FF9D07B0F9577BB8F8F2" bold="true" box="[197,255,1700,1719]" pageId="2" pageNumber="433">Fig. 1.</emphasis>
Habitus images of
<taxonomicName id="4C0D4D6BFF88FF9D06C2F9577933F8F2" authorityName="Caterino &amp; Yamamoto" authorityYear="2023" box="[439,628,1700,1719]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="hkamticus" status="sp. nov.">
<emphasis id="B979EAFAFF88FF9D06C2F9577933F8F2" box="[439,628,1700,1719]" italics="true" pageId="2" pageNumber="433">Phasmister hkamticus</emphasis>
</taxonomicName>
,
<emphasis id="B979EAFAFF88FF9D050AF95779AEF8F2" bold="true" box="[639,745,1700,1719]" pageId="2" pageNumber="433">
<taxonomicNameLabel id="A24A5781FF88FF9D050AF95779AEF8F2" box="[639,745,1700,1719]" pageId="2" pageNumber="433" rank="species">new species</taxonomicNameLabel>
</emphasis>
. A) Dorsal, B) Dorsolateral, C) Lateral, D) Ventral,
</paragraph>
<paragraph id="8BB236E8FF88FF9D07D8F94F79D0F895" blockId="2.[173,1193,1700,1744]" box="[173,663,1724,1744]" pageId="2" pageNumber="433">E) Anterodorsal, F) Posterodorsal. All scale bars 0.5 mm.</paragraph>
</caption>
<paragraph id="8BB236E8FF89FF9B07EDFF8D781EFF48" blockId="3.[152,184,126,145]" lastBlockId="4.[705,1193,194,909]" lastPageId="4" lastPageNumber="435" pageId="3" pageNumber="434">
434
<emphasis id="B979EAFAFF89FF9C07C5FF327AD4FE92" bold="true" box="[176,403,193,215]" pageId="3" pageNumber="434">Diagnostic Description.</emphasis>
Full body length,
<emphasis id="B979EAFAFF89FF9C054BFF317931FE92" box="[574,630,194,215]" italics="true" pageId="3" pageNumber="434">in situ</emphasis>
: with posterior and lateral margins raised, antennal
<quantity id="4CF59B0DFF89FF9C07EDFF2F7BADFEB7" box="[152,234,220,242]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="3" pageNumber="434" unit="mm" value="1.21">1.21 mm</quantity>
(apex of head to posterior end of abdomen); insertions open anteriorly; antennal club with disbody (pronotum + elytra) length along midline: tinct transverse annuli; prosternum with lateral slots
<quantity id="4CF59B0DFF89FF9C07EDFEE17BB2FF62" box="[152,245,274,295]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="434" unit="mm" value="1.02">1.02 mm</quantity>
; maximum (humeral) width:
<quantity id="4CF59B0DFF89FF9C0568FEE1793DFF62" box="[541,634,274,295]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="3" pageNumber="434" unit="mm" value="0.71">0.71 mm</quantity>
; for passage of the antennal funicle; hypomeral head width:
<quantity id="4CF59B0DFF89FF9C0661FEDF7A29FF07" box="[276,366,300,322]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="434" unit="mm" value="0.26">0.26 mm</quantity>
; pronotal length:
<quantity id="4CF59B0DFF89FF9C056AFEDF793EFF07" box="[543,633,300,322]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="434" unit="mm" value="0.34">0.34 mm</quantity>
; antennal depression open beneath.
<taxonomicName id="4C0D4D6BFF89FF9C0352FEDF7FD4FF04" authorityName="Caterino" authorityYear="2021" box="[1063,1171,300,321]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAFAFF89FF9C0352FEDF7FD4FF04" box="[1063,1171,300,321]" italics="true" pageId="3" pageNumber="434">Phasmister</emphasis>
</taxonomicName>
pronotal width:
<quantity id="4CF59B0DFF89FF9C065AFEB47AC1FF19" box="[303,390,327,349]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="434" unit="mm" value="0.56">0.56 mm</quantity>
. This species shares many
<taxonomicName id="4C0D4D6BFF89FF9C05D8FEB47849FF19" authorityName="Caterino &amp; Yamamoto" authorityYear="2023" box="[685,782,327,348]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="hkamticus">
<emphasis id="B979EAFAFF89FF9C05D8FEB47849FF19" box="[685,782,327,348]" italics="true" pageId="3" pageNumber="434">hkamticus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0D4D6BFF89FF9C04F9FEBB78B1FF18" authorityName="Caterino" authorityYear="2021" box="[908,1014,328,349]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B979EAFAFF89FF9C04F9FEBB78B1FF18" box="[908,1014,328,349]" italics="true" pageId="3" pageNumber="434">P. cristatus</emphasis>
</taxonomicName>
in the following of the generic level characters described for specific characters: prothorax less strongly trans-
<taxonomicName id="4C0D4D6BFF89FF9C07EDFE8F7AFFFFD4" authority="(Caterino 2021)" baseAuthorityName="Caterino" baseAuthorityYear="2021" box="[152,440,380,402]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAFAFF89FF9C07EDFE8F7A4EFFD4" box="[152,265,380,401]" italics="true" pageId="3" pageNumber="434">Phasmister</emphasis>
(Caterino 2021)
</taxonomicName>
: body subquadrate, verse, length/width ratio slightly greater (see
<figureCitation id="13362A6DFF89FF9C033AFE8F7FC1FFD4" box="[1103,1158,380,402]" captionStart="Fig" captionStartId="4.[197,228,1192,1211]" captionTargetBox="[173,659,193,1175]" captionTargetId="figure-511@4.[173,659,193,1175]" captionTargetPageId="4" captionText="Fig. 3. Comparison of pronota of Phasmister spp. A) P. cristatus, B) P. hkamticus, new species, C) P. parallellus, new species." figureDoi="http://doi.org/10.5281/zenodo.13243277" httpUri="https://zenodo.org/record/13243277/files/figure.png" pageId="3" pageNumber="434">Fig. 3</figureCitation>
); weakly compressed; elytra with every second inter- anterior corners of pronotum not as produced, less stria strongly raised to form a carina; humeral elytral deeply explanate; anterior margin of pronotum angles expanded; pronotum with raised lateral and rather narrow; median pronotal carina less dorsally anterior margins, depressed in anterior corners, with projecting (as seen in lateral view), little higher than various carinae on disc; pronotal disc strongly car- elytral carinae, more obtusely divergent near apex, inate; posterior pronotal margin sinuate, produced then angulate to anterior margin, enclosing a disposteriad in front of small scutellar shield; frons tinctly (irregularly) pentagonal space; sublateral elongate; propygidium deeply concave in basal half, punctate apically; pygidium with apex directed nearly anteriad.
</paragraph>
<caption id="DF726660FF89FF9C07C5F9647976F887" ID-DOI="http://doi.org/10.5281/zenodo.13243273" ID-Zenodo-Dep="13243273" httpUri="https://zenodo.org/record/13243273/files/figure.png" pageId="3" pageNumber="434" startId="3.[176,207,1687,1706]" targetBox="[151,1171,643,1669]" targetPageId="3" targetType="figure">
<paragraph id="8BB236E8FF89FF9C07C5F9647FD3F8EF" blockId="3.[152,1172,1687,1730]" box="[176,1172,1687,1706]" pageId="3" pageNumber="434">
<emphasis id="B979EAFAFF89FF9C07C5F9647BAEF8EF" bold="true" box="[176,233,1687,1706]" pageId="3" pageNumber="434">Fig. 2.</emphasis>
Habitus images of
<taxonomicName id="4C0D4D6BFF89FF9C06D7F9647918F8EF" authorityName="Caterino &amp; Yamamoto" authorityYear="2023" box="[418,607,1687,1706]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="parallelus" status="sp. nov.">
<emphasis id="B979EAFAFF89FF9C06D7F9647918F8EF" box="[418,607,1687,1706]" italics="true" pageId="3" pageNumber="434">Phasmister parallelus</emphasis>
</taxonomicName>
,
<emphasis id="B979EAFAFF89FF9C051FF9647993F8EF" bold="true" box="[618,724,1687,1706]" pageId="3" pageNumber="434">
<taxonomicNameLabel id="A24A5781FF89FF9C051FF9647993F8EF" box="[618,724,1687,1706]" pageId="3" pageNumber="434" rank="species">new species</taxonomicNameLabel>
</emphasis>
. A) Dorsal, B) Ventral, C) Lateral, D) Dorsolateral,
</paragraph>
<paragraph id="8BB236E8FF89FF9C07EDF95C7976F887" blockId="3.[152,1172,1687,1730]" box="[152,561,1711,1730]" pageId="3" pageNumber="434">E) Anterior, F) Posterior. All scale bars 0.5 mm.</paragraph>
</caption>
<paragraph id="8BB236E8FF8EFF9B05AEFEE278C9FF3D" blockId="4.[705,1193,194,909]" pageId="4" pageNumber="435">
<emphasis id="B979EAFAFF8EFF9B05AEFEE27F47FF62" bold="true" box="[731,1024,273,295]" pageId="4" pageNumber="435">Derivation of Specific Name.</emphasis>
The name
<taxonomicName id="4C0D4D6BFF8EFF9B0319FEE179B7FF07" authorityName="Caterino &amp; Yamamoto" authorityYear="2023" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="hkamticus">
<emphasis id="B979EAFAFF8EFF9B0319FEE179B7FF07" italics="true" pageId="4" pageNumber="435">hkamticus</emphasis>
</taxonomicName>
is an adjective referring to the novel source of amber (the Hkamti amber deposits) in which the specimen was found.
</paragraph>
</subSubSection>
<subSubSection id="C3176563FF8EFF9B05AEFE8F7F6FFDC8" pageId="4" pageNumber="435" type="discussion">
<paragraph id="8BB236E8FF8EFF9B05AEFE8F78B8FD67" blockId="4.[705,1193,194,909]" pageId="4" pageNumber="435">
<emphasis id="B979EAFAFF8EFF9B05AEFE8F7879FFD4" bold="true" box="[731,830,380,401]" pageId="4" pageNumber="435">Remarks.</emphasis>
The differences between this species and
<taxonomicName id="4C0D4D6BFF8EFF9B0586FE6B7822FFE8" authorityName="Caterino" authorityYear="2021" box="[755,869,408,429]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B979EAFAFF8EFF9B0586FE6B7822FFE8" box="[755,869,408,429]" italics="true" pageId="4" pageNumber="435">P. cristatus</emphasis>
</taxonomicName>
are well-marked, despite their clearly close relationship. The distinct conformation of pronotal carinae separates them most clearly, with the broader, pentagonal area between the median carinae (
<figureCitation id="13362A6DFF8EFF9B0460FDF17818FC52" box="[789,863,514,536]" captionStart="Fig" captionStartId="4.[197,228,1192,1211]" captionTargetBox="[173,659,193,1175]" captionTargetId="figure-511@4.[173,659,193,1175]" captionTargetPageId="4" captionText="Fig. 3. Comparison of pronota of Phasmister spp. A) P. cristatus, B) P. hkamticus, new species, C) P. parallellus, new species." figureDoi="http://doi.org/10.5281/zenodo.13243277" httpUri="https://zenodo.org/record/13243277/files/figure.png" pageId="4" pageNumber="435">Fig. 3B</figureCitation>
) of the new species quite distinct. The apparently lower overall elevation of the median pronotal carina also appears distinctive. The differences of both of these species from
<taxonomicName id="4C0D4D6BFF8EFF9B034BFDA17863FCC4" authorityName="Caterino &amp; Yamamoto" authorityYear="2023" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="parallelus" status="sp. nov.">
<emphasis id="B979EAFAFF8EFF9B034BFDA17863FCC4" italics="true" pageId="4" pageNumber="435">Phasmister parallelus</emphasis>
</taxonomicName>
,
<emphasis id="B979EAFAFF8EFF9B045AFD9F78E3FCC7" bold="true" box="[815,932,620,642]" pageId="4" pageNumber="435">
<taxonomicNameLabel id="A24A5781FF8EFF9B045AFD9F78E3FCC7" box="[815,932,620,642]" pageId="4" pageNumber="435" rank="species">new species</taxonomicNameLabel>
</emphasis>
are more noteworthy. Most distinct are the differences in tibial shape, particularly in the broad, outwardly angulate protibia of
<taxonomicName id="4C0D4D6BFF8EFF9B05B6FD4E7879FC94" authorityName="Caterino &amp; Yamamoto" authorityYear="2023" box="[707,830,700,722]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B979EAFAFF8EFF9B05B6FD4E7879FC94" box="[707,830,700,722]" italics="true" pageId="4" pageNumber="435">P. parallelus</emphasis>
</taxonomicName>
, with a medial rather than marginal groove.
<taxonomicName id="4C0D4D6BFF8EFF9B0466FD2478A0FCA9" authorityName="Caterino &amp; Yamamoto" authorityYear="2023" box="[787,999,727,748]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="hkamticus">
<emphasis id="B979EAFAFF8EFF9B0466FD2478A0FCA9" box="[787,999,727,748]" italics="true" pageId="4" pageNumber="435">Phasmister hkamticus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0D4D6BFF8EFF9B036DFD2B7FC3FCA8" authorityName="Caterino" authorityYear="2021" box="[1048,1156,728,749]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B979EAFAFF8EFF9B036DFD2B7FC3FCA8" box="[1048,1156,728,749]" italics="true" pageId="4" pageNumber="435">P. cristatus</emphasis>
</taxonomicName>
are both distinctly flattened dorsally, where the species described below is more convex.
</paragraph>
<paragraph id="8BB236E8FF8EFF9B05AEFCD47F6FFDC8" blockId="4.[705,1193,194,909]" pageId="4" pageNumber="435">
<emphasis id="B979EAFAFF8EFF9B05AEFCD4782AFD78" bold="true" box="[731,877,807,829]" pageId="4" pageNumber="435">Syninclusions.</emphasis>
No arthropod inclusions are included, but there is some small organic debris and a partial remnant of a boring of
<taxonomicName id="4C0D4D6BFF8EFF9B0481FCAF7F18FD37" authorityName="Lamarck" authorityYear="1809" box="[1012,1119,860,882]" class="Bivalvia" family="Pholadidae" kingdom="Animalia" order="Myida" pageId="4" pageNumber="435" phylum="Mollusca" rank="family">Pholadidae</taxonomicName>
(
<taxonomicName id="4C0D4D6BFF8EFF9B031EFCAF79A3FDC8" authorityName="Linnaeus" authorityYear="1758" class="Bivalvia" kingdom="Animalia" pageId="4" pageNumber="435" phylum="Mollusca" rank="class">Bivalvia</taxonomicName>
: Myoida) within the amber piece.
</paragraph>
</subSubSection>
</treatment>
</document>