treatments-xml/data/03/A4/87/03A487F2FFFBFFB8751A6BE99751B0C8.xml

<document id="61E817CB64F9B535F89180F1B2B371D0" ID-CLB-Dataset="22375" ID-DOI="10.5852/ejt.2015.150" ID-GBIF-Dataset="f492ad9b-a83d-4d8e-bdf9-20ef759b75c1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3787588" ID-ZooBank="D546A676-35F8-4025-8660-F123A0BA67F7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588704123207" checkinUser="carolina" docAuthor="Enghoff, Henrik &amp; Frederiksen, Sara B." docDate="2015" docId="03A487F2FFFBFFB8751A6BE99751B0C8" docLanguage="en" docName="ejt-2015-150.pdf.imf" docOrigin="European Journal of Taxonomy 150" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aquattuor Frederiksen 2013" docType="treatment" docVersion="6" lastPageNumber="7" masterDocId="FF9DFF8AFFF9FFBE775F69219518B21B" masterDocTitle="A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="3" updateTime="1698838626763" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E702D3C8F99D208878D08C4149198516">A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="B2ACD916865C4ACBD012543D722F75A2">Enghoff, Henrik</mods:namePart>
<mods:affiliation id="4DA0CE30E4268A781FD0EC0FE39BE27F">Zoological Museum, Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100, Copenhagen Ø, Denmark &amp; Corresponding author: henghoff @ snm. ku. dk &amp; urn: lsid: zoobank. org: author: FB 09 A 817 - 000 D- 43 C 3 - BCC 4 - 2 BC 1 E 5373635</mods:affiliation>
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<mods:namePart id="72C17552A4C1C761BD3CC9769011ACFD">Frederiksen, Sara B.</mods:namePart>
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<mods:affiliation id="96CA1660C5A284E3F50AA99F9EAFEDA1">Zoological Museum, Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100, Copenhagen Ø, Denmark &amp; Department of Ecology, Animal Ecology, University of Marburg, Karl-von-Frisch-Strasse 8, D- 35032 Marburg, Germany &amp; Email: sara. frederiksen @ snm. ku. dk &amp; urn: lsid: zoobank. org: author: 6 DE 62 DD 5 - 0055 - 424 A-A 136 - 0 EF 290 AF 5 AFD</mods:affiliation>
<mods:nameIdentifier id="6273E36637032866F522CB3E899069B9" type="email">sara.frederiksen@snm.ku.dk</mods:nameIdentifier>
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<subSubSection id="C317656FFFFBFFBC751A6BE996F7B0F9" box="[581,1007,712,738]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BB236E4FFFBFFBC751A6BE996F7B0F9" blockId="2.[581,1007,712,738]" box="[581,1007,712,738]" pageId="2" pageNumber="3">
<heading id="D0FA8188FFFBFFBC751A6BE996F7B0F9" box="[581,1007,712,738]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
Genus 
<taxonomicName id="4C0D4D67FFFBFFBC75C86BE996F7B0F9" ID-CoL="7NQQQ" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[663,1007,712,738]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFBFFBC75C86BE9960BB0F9" bold="true" box="[663,787,712,738]" italics="true" pageId="2" pageNumber="3">Aquattuor</emphasis>
Frederiksen, 2013
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C317656FFFFBFFBC77E26A2B912DB15C" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BB236E4FFFBFFBC77E26A2B912DB15C" blockId="2.[189,1398,778,840]" pageId="2" pageNumber="3">
This genus was established by 
<bibRefCitation id="EF9C4B15FFFBFFBC75466A2B961DB13E" author="Frederiksen S. B." box="[537,773,778,805]" pageId="2" pageNumber="3" pagination="59 - 66" refId="ref8188" refString="Frederiksen S. B. 2013 b. East African odontopygid millipedes 3: Two new genera; Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3694: 59 - 66. http: // dx. doi. org / 10.11646 / zootaxa. 3694.1.4" type="journal article" year="2013">Frederiksen (2013b)</bibRefCitation>
for the 
<typeStatus id="54B68846FFFBFFBC74066A2B9694B13F" box="[857,908,778,804]" pageId="2" pageNumber="3">type</typeStatus>
and until now only species, 
<taxonomicName id="4C0D4D67FFFBFFBC73966A2B906EB13F" authorityName="Frederiksen" authorityYear="2013" box="[1225,1398,778,804]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B979EAF6FFFBFFBC73966A2B906EB13F" box="[1225,1398,778,804]" italics="true" pageId="2" pageNumber="3">A. denticulatus</emphasis>
</taxonomicName>
(East Usambara Mts, 
<collectingCountry id="F31A7674FFFBFFBC76E46A0C9732B153" box="[443,554,813,840]" name="Tanzania" pageId="2" pageNumber="3">Tanzania</collectingCountry>
). The name is regarded as a masculine noun.
</paragraph>
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<paragraph id="8BB236E4FFFBFFBC77E26A4F942EB193" blockId="2.[189,310,878,904]" box="[189,310,878,904]" pageId="2" pageNumber="3">
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<emphasis id="B979EAF6FFFBFFBC77E26A4F942EB193" bold="true" box="[189,310,878,904]" pageId="2" pageNumber="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFBFFBC77E26ABC97AAB63B" blockId="2.[189,1399,925,1056]" pageId="2" pageNumber="3">
A genus of 
<taxonomicName id="4C0D4D67FFFBFFBC76146ABC9718B1AC" authorityName="Attems" authorityYear="1909" box="[331,512,925,951]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Odontopygidae</taxonomicName>
<taxonomicName id="4C0D4D67FFFBFFBC75556ABC97A8B1AC" authorityName="Hoffman" authorityYear="1991" box="[522,688,925,951]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Prionopetalini">Prionopetalini</taxonomicName>
in which the gonopodal proplica is apically expanded into a distal ‘palette’, the metaplica has an oblique flange, the solenomere is simple, thin and whiplike, the telomere terminates in a long, curved gutter-like to tubular part, and the limbus of the body rings consists of large, rectangular, easily detached flaps.
</paragraph>
</subSubSection>
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<heading id="D0FA8188FFFBFFBC77E26D69945CB679" bold="true" box="[189,324,1096,1122]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B979EAF6FFFBFFBC77E26D69945CB679" bold="true" box="[189,324,1096,1122]" pageId="2" pageNumber="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFBFFBC77E26D579426B6A8" blockId="2.[189,1398,1142,1204]" pageId="2" pageNumber="3">The genus owes its name to the unique limbus flaps which resemble tiny sheets of paper in the standard A4 format.</paragraph>
</subSubSection>
<subSubSection id="C317656FFFFBFFBD77E26DFA963CB1CC" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
<paragraph id="8BB236E4FFFBFFBC77E26DFA9456B6EE" blockId="2.[189,334,1243,1269]" box="[189,334,1243,1269]" pageId="2" pageNumber="3">
<heading id="D0FA8188FFFBFFBC77E26DFA9456B6EE" bold="true" box="[189,334,1243,1269]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B979EAF6FFFBFFBC77E26DFA9456B6EE" bold="true" box="[189,334,1243,1269]" pageId="2" pageNumber="3">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFBFFBC77E26C2894B3B75C" blockId="2.[189,1398,1289,1351]" pageId="2" pageNumber="3">This description only applies to males and, as far as non-gonopodal characters are concerned, only includes a selection.</paragraph>
<paragraph id="8BB236E4FFFBFFBC77B56C4F972EB793" blockId="2.[189,1399,1390,1487]" box="[234,566,1390,1416]" pageId="2" pageNumber="3">
<emphasis id="B979EAF6FFFBFFBC77B56C4F972EB793" bold="true" box="[234,566,1390,1416]" pageId="2" pageNumber="3">Non-gonopodal characters</emphasis>
</paragraph>
<paragraph id="8BB236E4FFFBFFBC77E26CB39757B7D4" blockId="2.[189,1399,1390,1487]" pageId="2" pageNumber="3">
BODY LENGTH. 
<quantity id="4CF59B01FFFBFFBC76336CB094E9B7B7" box="[364,497,1425,1452]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.5" metricValueMin="1.9" pageId="2" pageNumber="3" unit="mm" value="27.0" valueMax="35.0" valueMin="19.0">19–35 mm</quantity>
. Midbody vertical diameter 
<quantity id="4CF59B01FFFBFFBC74156CB096C6B7B7" box="[842,990,1425,1452]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.1" metricValueMin="1.4" pageId="2" pageNumber="3" unit="mm" value="1.75" valueMax="2.1" valueMin="1.4">1.4–2.1 mm</quantity>
. 44–54 podous rings, no apodous rings in front of telson. See 
<figureCitation id="13362A61FFFBFFBC755B6C959757B7D4" box="[516,591,1460,1487]" captionStart="Fig" captionStartId="4.[189,232,1924,1950]" captionTargetBox="[187,1400,1040,1895]" captionTargetId="figure@4.[170,1431,1031,1907]" captionTargetPageId="4" captionText="Fig. 2. Body size (numbers of podous rings and midbody vertical diameter) in ♂♂ of species of Aquattuor Frederiksen, 2013. In cases of (almost) coinciding values, symbols have been slightly displaced horizontally." figureDoi="http://doi.org/10.5281/zenodo.3787592" httpUri="https://zenodo.org/record/3787592/files/figure.png" pageId="2" pageNumber="3">Fig. 2.</figureCitation>
</paragraph>
<paragraph id="8BB236E4FFFBFFBC77E26CD791E5B44D" blockId="2.[189,1399,1526,1623]" pageId="2" pageNumber="3">
COLOUR. Colours of specimens from Udzungwa and East Usambara Mountains all partly or completely faded, leaving only newly collected specimens from Mt. 
<collectingRegion id="49C9F806FFFBFFBC743B6F3896EAB428" box="[868,1010,1561,1587]" country="Tanzania" name="Kilimanjaro" pageId="2" pageNumber="3">Kilimanjaro</collectingRegion>
with fresh colour (
<figureCitation id="13362A61FFFBFFBC73896F389004B428" box="[1238,1308,1561,1587]" captionStart="Fig" captionStartId="3.[189,232,1912,1938]" captionTargetBox="[189,1397,1250,1881]" captionTargetId="figure@3.[173,1411,1244,1888]" captionTargetPageId="3" captionText="Fig. 1. Aquattuor claudiahempae sp. nov., two paratype ♂♂ after 18 months in alcohol. Scale bar = 1 mm (B). Photographs by S. Reboleira." figureDoi="http://doi.org/10.5281/zenodo.3787590" httpUri="https://zenodo.org/record/3787590/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
). Light mid-dorsal longitudinal stripe or traces thereof sometimes present, even on faded specimens.
</paragraph>
<paragraph id="8BB236E4FFFBFFBC77E26F5F944AB4A0" blockId="2.[189,1398,1662,1723]" pageId="2" pageNumber="3">
LIMBUS. Consisting of relatively large (
<emphasis id="B979EAF6FFFBFFBC75D46F5E9786B483" box="[651,670,1663,1688]" italics="true" pageId="2" pageNumber="3">c.</emphasis>
0.04 × 
<quantity id="4CF59B01FFFBFFBC75A36F5F9670B483" box="[764,872,1662,1688]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="3" unit="mm" value="0.06">0.06 mm</quantity>
) rectangular flaps which are easily detached (
<figureCitation id="13362A61FFFBFFBC77996F809412B4A0" box="[198,266,1697,1723]" captionStart="Fig" captionStartId="5.[189,232,1894,1920]" captionTargetBox="[236,1347,264,1860]" captionTargetId="figure@5.[233,1355,258,1863]" captionTargetPageId="5" captionText="Fig. 3. Aquattuor spp. A. A. stereosathe Enghoff sp. nov., paratype, front end. B–C. A. udzungwensis Enghoff sp. nov., paratype, telson, lateral and posterior view. D–E. A. udzungwensis Enghoff sp. nov., limbus. D. Specimen from West Kilombero Scarp FR. E. Paratype. Scale bars: A–D = 0.1 mm; E = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3787594" httpUri="https://zenodo.org/record/3787594/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
D–E).
</paragraph>
<paragraph id="8BB236E4FFFBFFBD77E26FC294EEB33E" blockId="2.[189,1399,1762,1895]" lastBlockId="3.[189,1399,267,470]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
TELSON (
<figureCitation id="13362A61FFFBFFBC76756FC3946CB4E6" box="[298,372,1762,1789]" captionStart="Fig" captionStartId="5.[189,232,1894,1920]" captionTargetBox="[236,1347,264,1860]" captionTargetId="figure@5.[233,1355,258,1863]" captionTargetPageId="5" captionText="Fig. 3. Aquattuor spp. A. A. stereosathe Enghoff sp. nov., paratype, front end. B–C. A. udzungwensis Enghoff sp. nov., paratype, telson, lateral and posterior view. D–E. A. udzungwensis Enghoff sp. nov., limbus. D. Specimen from West Kilombero Scarp FR. E. Paratype. Scale bars: A–D = 0.1 mm; E = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3787594" httpUri="https://zenodo.org/record/3787594/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
B–C). Preanal ring with wrinkled/coriaceous sculpture dorsally. Anal valves with wrinkled/coriaceous sculpture, with a pronounced dorsal denticle and a smaller ventral one, denticles well set off from rest of valve rather than just being “sharp corners”. Each valve with three setae. Free margin (“lip”) of anal valves raised and provided with three small tubercles on which the setae are borne. 
<emphasis id="B979EAF6FFFAFFBD77B5682A9471B33E" bold="true" box="[234,361,267,293]" pageId="3" pageNumber="4">Gonopods</emphasis>
(see 
<figureCitation id="13362A61FFFAFFBD76F8682A94F5B33E" box="[423,493,267,293]" captionStart="Fig" captionStartId="7.[189,232,1860,1886]" captionTargetBox="[228,1353,267,1835]" captionTargetId="figure@7.[227,1360,265,1835]" captionTargetPageId="7" captionText="Fig. 4. Aquattuor submajor Enghoff sp. nov., paratype. A. Gonopods in situ, right gonopod coloured. B. Left gonopod coxa and basal part of telopodite. C. Left gonopod telopodite and part of coxa, dorsal (basal) view.Articifical colours: orange = coxa; blue = basomere; yellow = solenomere; pink = telomere; red = gonopod sternum; light green = ventral part of eighth pleurotergite (right side); dark green = sternum of reduced ninth leg-pair. mp = metaplica; pa = palette; pp = proplica. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787596" httpUri="https://zenodo.org/record/3787596/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
)
</paragraph>
<paragraph id="8BB236E4FFFAFFBD77E2680E94E6B3CE" blockId="3.[189,1399,267,470]" pageId="3" pageNumber="4">
COXA. In anterior or posterior view 3½–4½ × as long as broad. Margins of proplica (
<emphasis id="B979EAF6FFFAFFBD7395680E91FFB353" box="[1226,1255,303,328]" italics="true" pageId="3" pageNumber="4">pp</emphasis>
) parallel or slightly diverging in basal 
<emphasis id="B979EAF6FFFAFFBD76A368729708B377" box="[508,528,339,364]" italics="true" pageId="3" pageNumber="4">c.</emphasis>
⅔, apical 
<emphasis id="B979EAF6FFFAFFBD75D0687297BBB377" box="[655,675,339,364]" italics="true" pageId="3" pageNumber="4">c.</emphasis>
⅓ set off by mesal (
<emphasis id="B979EAF6FFFAFFBD74C2687296A2B377" box="[925,954,339,364]" italics="true" pageId="3" pageNumber="4">mi</emphasis>
) and lateral (
<emphasis id="B979EAF6FFFAFFBD730268739175B377" box="[1117,1133,338,364]" italics="true" pageId="3" pageNumber="4">li</emphasis>
) incisions; apical part of proplica appearing like a rounded “palette” (
<emphasis id="B979EAF6FFFAFFBD75B368579611B394" box="[748,777,374,399]" italics="true" pageId="3" pageNumber="4">pa</emphasis>
). Proplical lobe (
<emphasis id="B979EAF6FFFAFFBD7487685496E2B394" box="[984,1018,373,399]" italics="true" pageId="3" pageNumber="4">prl</emphasis>
) protruding into mesal incision. Metaplica (
<emphasis id="B979EAF6FFFAFFBD761C68B8947FB3A9" box="[323,359,409,434]" italics="true" pageId="3" pageNumber="4">mp</emphasis>
) with irregular surface; at level of mesal and lateral incisions a stout ridge (
<emphasis id="B979EAF6FFFAFFBD738468B89012B3A9" box="[1243,1290,409,434]" italics="true" pageId="3" pageNumber="4">mpr</emphasis>
) running obliquely across metaplica.
</paragraph>
<paragraph id="8BB236E4FFFAFFBD77E268DC94C0B045" blockId="3.[189,1398,509,606]" pageId="3" pageNumber="4">
TELOPODITE. Basomere (
<emphasis id="B979EAF6FFFAFFBD768C68DC94E8B00C" box="[467,496,509,535]" italics="true" pageId="3" pageNumber="4">ba</emphasis>
) with small spine (
<emphasis id="B979EAF6FFFAFFBD759168DC97F0B00C" box="[718,744,509,535]" italics="true" pageId="3" pageNumber="4">bs</emphasis>
) just distal of torsotope (
<figureCitation id="13362A61FFFAFFBD735668DC918CB00C" box="[1033,1172,509,535]" captionStart="Fig" captionStartId="19.[189,232,1896,1922]" captionTargetBox="[188,1397,267,1871]" captionTargetId="figure@19.[181,1400,265,1871]" captionTargetPageId="19" captionText="Fig. 11. Aquattuor denticulatus Frederiksen, 2013, paratype. A. Gonopods, posterior view. B. Telomere, distal part, lateral view. C. Gonopods, anterior view. D. Telomere tip. li = lateral incision; bs = basomeral spine; pa = palette; mpl = mesal-posterior lamella of telomere. Scale bars: A, C = 0.1 mm; B = 0.02 mm; D = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3787610" httpUri="https://zenodo.org/record/3787610/files/figure.png" pageId="3" pageNumber="4">Fig. 11A, C</figureCitation>
), arculus 90°. Posttorsal narrowing (
<emphasis id="B979EAF6FFFAFFBD76CC6B0094A8B021" box="[403,432,545,570]" italics="true" pageId="3" pageNumber="4">pn</emphasis>
) pronounced (e.g., 
<figureCitation id="13362A61FFFAFFBD75C36B0197E6B020" box="[668,766,544,571]" captionStart="Fig" captionStartId="8.[189,232,1890,1916]" captionTargetBox="[198,1390,269,1859]" captionTargetId="figure@8.[194,1393,266,1862]" captionTargetPageId="8" captionText="Fig. 5. Aquattuor submajor Enghoff sp. nov., paratype, left gonopod. A. Anterior view. B. Close-up of mesal incision. C. Tip of telomere. D. Posterior view. btl = basal telomeral lamella; li = lateral incision; mi = mesal incision; mpl = mesal-posterior lamella of telomere; pa = palette; pn = posttorsal narrowing. Scale bars: A, D = 0.1 mm; B–C = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3787598" httpUri="https://zenodo.org/record/3787598/files/figure.png" pageId="3" pageNumber="4">Figs 5A</figureCitation>
, 
<figureCitation id="13362A61FFFAFFBD74526B01962DB020" box="[781,821,544,571]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[245,1346,267,1856]" captionTargetId="figure@10.[237,1351,265,1860]" captionTargetPageId="10" captionText="Fig. 6. Aquattuor major Enghoff sp. nov., paratype. A. Left gonopod, anterior view. B. Left gonopod, posterior view. C–D. Telomere tip, two different views. li = lateral incision; pa = palette; mpl = mesal posterior lamella of telomere; pn = posttorsal narrowing; tt = triangular tooth of telomere. Scale bars: A–B = 0.1 mm; C–D = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3787600" httpUri="https://zenodo.org/record/3787600/files/figure.png" pageId="3" pageNumber="4">6A</figureCitation>
). Telopodite beyond 
<emphasis id="B979EAF6FFFAFFBD736F6B009155B021" box="[1072,1101,545,570]" italics="true" pageId="3" pageNumber="4">pn</emphasis>
divided into solenomere (
<emphasis id="B979EAF6FFFAFFBD77986B6295E8B046" box="[199,240,579,605]" italics="true" pageId="3" pageNumber="4">slm</emphasis>
) and telomere (
<emphasis id="B979EAF6FFFAFFBD76F56B6594DFB046" box="[426,455,580,605]" italics="true" pageId="3" pageNumber="4">tm</emphasis>
).
</paragraph>
<paragraph id="8BB236E4FFFAFFBD77E26BA4918BB0D9" blockId="3.[189,1398,645,707]" pageId="3" pageNumber="4">SOLENOMERE. Long, thin, whiplike, often resting within telomeral gutter but on preserved specimens frequently free (probably an artefact), apical part with oblique-longitudinal flutings.</paragraph>
<paragraph id="8BB236E4FFFAFFBD77E26BCB963CB1CC" blockId="3.[189,1399,745,983]" pageId="3" pageNumber="4">
TELOMERE. At base expanded into a complicated basal telomeral lamella (
<emphasis id="B979EAF6FFFAFFBD73536BC89133B118" box="[1036,1067,745,771]" italics="true" pageId="3" pageNumber="4">btl</emphasis>
) obscuring origin of solenomere (
<figureCitation id="13362A61FFFAFFBD765B6A2D9478B13C" box="[260,352,780,807]" captionStart="Fig" captionStartId="8.[189,232,1890,1916]" captionTargetBox="[198,1390,269,1859]" captionTargetId="figure@8.[194,1393,266,1862]" captionTargetPageId="8" captionText="Fig. 5. Aquattuor submajor Enghoff sp. nov., paratype, left gonopod. A. Anterior view. B. Close-up of mesal incision. C. Tip of telomere. D. Posterior view. btl = basal telomeral lamella; li = lateral incision; mi = mesal incision; mpl = mesal-posterior lamella of telomere; pa = palette; pn = posttorsal narrowing. Scale bars: A, D = 0.1 mm; B–C = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3787598" httpUri="https://zenodo.org/record/3787598/files/figure.png" pageId="3" pageNumber="4">Fig. 5D</figureCitation>
). Main part of telomere consisting of a long, narrow lamella folded into a more or less narrow gutter, sometimes even tube-like in distalmost part; gutter/tube describing a 90°–360° curve, usually in almost one plane, but sometimes (
<taxonomicName id="4C0D4D67FFFAFFBD75176A72964BB176" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[584,851,851,877]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="stereosathe" status="sp. nov.">
<emphasis id="B979EAF6FFFAFFBD75176A72977AB176" box="[584,610,851,877]" italics="true" pageId="3" pageNumber="4">A.</emphasis>
<emphasis id="B979EAF6FFFAFFBD75376A7297F5B176" box="[616,749,851,877]" italics="true" pageId="3" pageNumber="4">stereosathe</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFAFFBD74076A7296ABB176" box="[856,947,851,877]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
) clearly in three dimensions (note: this applies to preserved specimens). Posterior margin of gutter expanded into mesal-posterior lamella (
<emphasis id="B979EAF6FFFAFFBD721E6A579075B18B" box="[1345,1389,886,912]" italics="true" pageId="3" pageNumber="4">mpl</emphasis>
) (e.g., 
<figureCitation id="13362A61FFFAFFBD765C6AB8947BB1AF" box="[259,355,921,948]" captionStart="Fig" captionStartId="8.[189,232,1890,1916]" captionTargetBox="[198,1390,269,1859]" captionTargetId="figure@8.[194,1393,266,1862]" captionTargetPageId="8" captionText="Fig. 5. Aquattuor submajor Enghoff sp. nov., paratype, left gonopod. A. Anterior view. B. Close-up of mesal incision. C. Tip of telomere. D. Posterior view. btl = basal telomeral lamella; li = lateral incision; mi = mesal incision; mpl = mesal-posterior lamella of telomere; pa = palette; pn = posttorsal narrowing. Scale bars: A, D = 0.1 mm; B–C = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3787598" httpUri="https://zenodo.org/record/3787598/files/figure.png" pageId="3" pageNumber="4">Fig. 5D</figureCitation>
). Tip of telomere variable: margins smooth or denticulate-laciniate, internal surface of gutter more or less microspiculate (e.g., 
<figureCitation id="13362A61FFFAFFBD75C96A9D97C2B1CC" box="[662,730,956,983]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[245,1346,267,1856]" captionTargetId="figure@10.[237,1351,265,1860]" captionTargetPageId="10" captionText="Fig. 6. Aquattuor major Enghoff sp. nov., paratype. A. Left gonopod, anterior view. B. Left gonopod, posterior view. C–D. Telomere tip, two different views. li = lateral incision; pa = palette; mpl = mesal posterior lamella of telomere; pn = posttorsal narrowing; tt = triangular tooth of telomere. Scale bars: A–B = 0.1 mm; C–D = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3787600" httpUri="https://zenodo.org/record/3787600/files/figure.png" pageId="3" pageNumber="4">Fig. 6</figureCitation>
C–D).
</paragraph>
</subSubSection>
<subSubSection id="C317656FFFFAFFBD77E26ADF9617B6AB" pageId="3" pageNumber="4" type="distribution">
<paragraph id="8BB236E4FFFAFFBD77E26ADF944EB603" blockId="3.[189,342,1022,1048]" box="[189,342,1022,1048]" pageId="3" pageNumber="4">
<heading id="D0FA8188FFFAFFBD77E26ADF944EB603" bold="true" box="[189,342,1022,1048]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B979EAF6FFFAFFBD77E26ADF944EB603" bold="true" box="[189,342,1022,1048]" pageId="3" pageNumber="4">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFAFFBD77E26D0D9617B6AB" blockId="3.[189,1399,1068,1201]" pageId="3" pageNumber="4">
Species of 
<taxonomicName id="4C0D4D67FFFAFFBD76606D0C94AEB65C" authorityName="Frederiksen" authorityYear="2013" box="[319,438,1069,1095]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFAFFBD76606D0C94AEB65C" box="[319,438,1069,1095]" italics="true" pageId="3" pageNumber="4">Aquattuor</emphasis>
</taxonomicName>
are so far known only from 
<collectingCountry id="F31A7674FFFAFFBD74586D0D966CB65D" box="[775,884,1068,1094]" name="Tanzania" pageId="3" pageNumber="4">Tanzania</collectingCountry>
. They have been found in several mountain blocks belonging to the Eastern Arc Mts (
<bibRefCitation id="EF9C4B15FFFAFFBD75BF6D7196FAB671" author="Burgess N. &amp; Butinsky T. M. &amp; Cordeiro N. J. &amp; Doggart N. H. &amp; Fjeldsa J. &amp; Howell K. M. &amp; Kilahama F. B. &amp; Loader S. P. &amp; Lovett J. C. &amp; Mbilinyi B. &amp; Menegon M. &amp; Moyer D. C. &amp; Nashanda E. &amp; Perkin A. &amp; Rovero F. &amp; Stanley W. T. &amp; Stuart S. N." box="[736,994,1104,1130]" pageId="3" pageNumber="4" pagination="209 - 231" refId="ref7941" refString="Burgess N., Butinsky T. M., Cordeiro N. J., Doggart N. H., Fjeldsa J., Howell K. M., Kilahama F. B., Loader S. P., Lovett J. C., Mbilinyi B., Menegon M., Moyer D. C., Nashanda E., Perkin A., Rovero F., Stanley W. T. &amp; Stuart S. N. 2007. The biological importance of the Eastern Arc Mountains of Tanzania and Kenya. Biological Conservation 134: 209 - 231. http: // dx. doi. org / 10.1016 / j. biocon. 2006.08.015" type="journal article" year="2007">
Burgess 
<emphasis id="B979EAF6FFFAFFBD740F6D71968BB671" box="[848,915,1104,1130]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
<date id="FFB31024FFFAFFBD74FD6D7196FAB671" box="[930,994,1104,1130]" pageId="3" pageNumber="4" value="2007">2007</date>
</bibRefCitation>
): Udzungwa (5 species), Nguru (1 species), Rubeho (1 species) and East Usambara (1 species). One further species has been found on Mt. 
<collectingRegion id="49C9F806FFFAFFBD77B16DB79498B6AB" box="[238,384,1174,1200]" country="Tanzania" name="Kilimanjaro" pageId="3" pageNumber="4">Kilimanjaro</collectingRegion>
. For an overview, see 
<figureCitation id="13362A61FFFAFFBD75DB6DB79613B6AB" box="[644,779,1174,1200]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="21.[189,232,1915,1941]" captionStartId-1="22.[189,232,1908,1934]" captionTargetBox-0="[187,1400,780,1886]" captionTargetBox-1="[190,1402,936,1878]" captionTargetId-0="figure@21.[183,1402,775,1888]" captionTargetId-1="figure@22.[178,1408,927,1882]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Fig. 12. Map of part of Tanzania showing the mountains where Aquattuor species have been found. Based on Burgess et al. (2007: fig. 1)." captionText-1="Fig. 13. Map of the Udzungwa Mts showing the forest reserves where Aquattuor species have been found, and the epithets of the species found. Based on Marshall et al. (2010: fig. 1)." figureDoi-0="http://doi.org/10.5281/zenodo.3787612" figureDoi-1="http://doi.org/10.5281/zenodo.3787614" httpUri-0="https://zenodo.org/record/3787612/files/figure.png" httpUri-1="https://zenodo.org/record/3787614/files/figure.png" pageId="3" pageNumber="4">Figs 12–13</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="DF72666CFFFAFFBD77E26E59978DB5AD" ID-DOI="http://doi.org/10.5281/zenodo.3787590" ID-Zenodo-Dep="3787590" httpUri="https://zenodo.org/record/3787590/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1912,1938]" targetBox="[189,1397,1250,1881]" targetPageId="3">
<paragraph id="8BB236E4FFFAFFBD77E26E59978DB5AD" blockId="3.[189,1399,1912,1974]" pageId="3" pageNumber="4">
<emphasis id="B979EAF6FFFAFFBD77E26E599415B588" bold="true" box="[189,269,1912,1939]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
<taxonomicName id="4C0D4D67FFFAFFBD76486E589753B588" authority="Enghoff &amp; Frederiksen, 2015" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[279,587,1913,1939]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="claudiahempae" status="sp. nov.">
<emphasis id="B979EAF6FFFAFFBD76486E589753B588" box="[279,587,1913,1939]" italics="true" pageId="3" pageNumber="4">Aquattuor claudiahempae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFAFFBD750A6E5897ADB588" box="[597,693,1913,1939]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
, two paratype ♂♂ after 18 months in alcohol. Scale bar = 1 mm (B). Photographs by S. Reboleira.
</paragraph>
</caption>
<subSubSection id="C317656FFFFDFFBA77E2682A91E8B3A6" pageId="4" pageNumber="5" type="description">
<paragraph id="8BB236E4FFFDFFBA77E2682A9745B33E" blockId="4.[189,605,267,293]" box="[189,605,267,293]" pageId="4" pageNumber="5">
<heading id="D0FA8188FFFDFFBA77E2682A9745B33E" bold="true" box="[189,605,267,293]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B979EAF6FFFDFFBA77E2682A9745B33E" bold="true" box="[189,605,267,293]" pageId="4" pageNumber="5">Intercalary cuticular microscutes</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFDFFBA77E2681B91E8B3A6" blockId="4.[189,1399,313,446]" pageId="4" pageNumber="5">
In 
<taxonomicName id="4C0D4D67FFFDFFBA7784681B94D6B348" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[219,462,313,340]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="submajor" status="sp. nov.">
<emphasis id="B979EAF6FFFDFFBA7784681B9471B348" box="[219,361,313,340]" italics="true" pageId="4" pageNumber="5">A. submajor</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFDFFBA768B681B9735B34F" box="[468,557,314,340]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C0D4D67FFFDFFBA753D681B9691B348" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[610,905,313,340]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="udzungwensis" status="sp. nov.">
<emphasis id="B979EAF6FFFDFFBA753D681B963CB348" box="[610,804,313,340]" italics="true" pageId="4" pageNumber="5">A. udzungwensis</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFDFFBA74D1681B96F2B34F" box="[910,1002,314,340]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, intercalary cuticular microscutes, similar to those first reported by 
<bibRefCitation id="EF9C4B15FFFDFFBA756A687D9645B36C" author="Akkari N. &amp; Enghoff H." box="[565,861,348,375]" pageId="4" pageNumber="5" pagination="1 - 24" refId="ref7853" refString="Akkari N. &amp; Enghoff H. 2011. On some surface structures of potential taxonomic importance in families of the suborders Polydesmidea and Dalodesmidea (Polydesmida, Diplopoda). Zookeys 156: 1 - 24. http: // dx. doi. org / 10.3897 / zookeys. 156.2134" type="journal article" year="2011">
Akkari &amp; Enghoff (
<date id="FFB31024FFFDFFBA7447687C964CB36C" box="[792,852,349,375]" pageId="4" pageNumber="5" value="2011">2011</date>
)
</bibRefCitation>
in polydesmidan millipedes and subsequently found, 
<emphasis id="B979EAF6FFFDFFBA764768A09494B381" box="[280,396,384,410]" italics="true" pageId="4" pageNumber="5">inter alia</emphasis>
, in the odontopygid genus 
<taxonomicName id="4C0D4D67FFFDFFBA75AD68A19012B381" authority="Attems, 1914" authorityName="Attems" authorityYear="1914" box="[754,1290,384,410]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFDFFBA75AD68A19693B381" box="[754,907,384,410]" italics="true" pageId="4" pageNumber="5">Chaleponcus</emphasis>
Attems, 1914 (
<bibRefCitation id="EF9C4B15FFFDFFBA730A68A1901AB381" author="Enghoff H." box="[1109,1282,384,410]" pageId="4" pageNumber="5" pagination="1 - 75" refId="ref8067" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff 2014</bibRefCitation>
)
</taxonomicName>
and the callipodidan genus 
<taxonomicName id="4C0D4D67FFFDFFBA76FE688597C3B3A6" authority="Mauries, 1978" authorityName="Mauries" authorityYear="1978" box="[417,731,419,445]" class="Diplopoda" family="Dorypetalidae" genus="Lusitanipus" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFDFFBA76FE68859732B3A6" box="[417,554,420,445]" italics="true" pageId="4" pageNumber="5">Lusitanipus</emphasis>
Mauriès, 1978
</taxonomicName>
(
<bibRefCitation id="EF9C4B15FFFDFFBA75B568829131B3A5" author="Reboleira A. S. P. S. &amp; Enghoff H." box="[746,1065,419,446]" pageId="4" pageNumber="5" pagination="567 - 576" refId="ref8576" refString="Reboleira A. S. P. S. &amp; Enghoff H. 2015. Redescription of Lusitanipus alternans (Verhoeff, 1893) (Diplopoda, Callipoda, Dorypetalidae) and ecological data on its Laboulbeniales ectoparasites in caves. Zootaxa 3957: 567 - 576. http: // dx. doi. org / 10.11646 / zootaxa. 3957.5.5" type="journal article" year="2015">
Reboleira &amp; Enghoff 
<date id="FFB31024FFFDFFBA74B568829131B3A5" box="[1002,1065,419,446]" pageId="4" pageNumber="5" value="2015">2015</date>
</bibRefCitation>
), were observed.
</paragraph>
</subSubSection>
<subSubSection id="C317656FFFFDFFB877E268C49751B0C8" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BB236E4FFFDFFBA77E268C494CEB3E5" blockId="4.[189,470,484,511]" box="[189,470,484,511]" pageId="4" pageNumber="5">
<heading id="D0FA8188FFFDFFBA77E268C494CEB3E5" bold="true" box="[189,470,484,511]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B979EAF6FFFDFFBA77E268C494CEB3E5" bold="true" box="[189,470,484,511]" pageId="4" pageNumber="5">Notes on identification</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFDFFBA77E26B329465B0C6" blockId="4.[189,1399,531,733]" pageId="4" pageNumber="5">
The species of 
<taxonomicName id="4C0D4D67FFFDFFBA762C6B3294F2B036" authorityName="Frederiksen" authorityYear="2013" box="[371,490,531,557]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFDFFBA762C6B3294F2B036" box="[371,490,531,557]" italics="true" pageId="4" pageNumber="5">Aquattuor</emphasis>
</taxonomicName>
are all quite similar, even in gonopod structure. In this respect, they form a contrast to the previously treated (much larger) species-swarm centred in the Udzungwa Mts., i.e., the 
<taxonomicName id="4C0D4D67FFFDFFBA77E26B7894EBB068" authorityName="Kraus" authorityYear="1958" box="[189,499,601,627]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="B979EAF6FFFDFFBA77E26B7894EBB068" box="[189,499,601,627]" italics="true" pageId="4" pageNumber="5">Chaleponcus dabagaensis</emphasis>
</taxonomicName>
-group (
<bibRefCitation id="EF9C4B15FFFDFFBA750C6B7897E6B068" author="Enghoff H." box="[595,766,601,627]" pageId="4" pageNumber="5" pagination="1 - 75" refId="ref8067" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">
Enghoff 
<date id="FFB31024FFFDFFBA75E06B7897E6B068" box="[703,766,601,627]" pageId="4" pageNumber="5" value="2014">2014</date>
</bibRefCitation>
), in which gonopods are highly diverse. Differences between 
<taxonomicName id="4C0D4D67FFFDFFBA76776B5C9487B08C" authorityName="Frederiksen" authorityYear="2013" box="[296,415,637,663]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFDFFBA76776B5C9487B08C" box="[296,415,637,663]" italics="true" pageId="4" pageNumber="5">Aquattuor</emphasis>
</taxonomicName>
species concern body size (
<figureCitation id="13362A61FFFDFFBA75B16B5D962FB08C" box="[750,823,636,663]" captionStart="Fig" captionStartId="4.[189,232,1924,1950]" captionTargetBox="[187,1400,1040,1895]" captionTargetId="figure@4.[170,1431,1031,1907]" captionTargetPageId="4" captionText="Fig. 2. Body size (numbers of podous rings and midbody vertical diameter) in ♂♂ of species of Aquattuor Frederiksen, 2013. In cases of (almost) coinciding values, symbols have been slightly displaced horizontally." figureDoi="http://doi.org/10.5281/zenodo.3787592" httpUri="https://zenodo.org/record/3787592/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
), details of the outline of the gonopod coxa, the curvature of the telomere, the development of the posterior margin of the telomere, and the armature of the telomere tip.
</paragraph>
<paragraph id="8BB236E4FFFDFFBA77E26A25946EB104" blockId="4.[189,374,772,799]" box="[189,374,772,799]" pageId="4" pageNumber="5">
<heading id="D0FA8188FFFDFFBA77E26A25946EB104" bold="true" box="[189,374,772,799]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B979EAF6FFFDFFBA77E26A25946EB104" bold="true" box="[189,374,772,799]" pageId="4" pageNumber="5">Similar genera</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFDFFB877E26A1291FCB394" blockId="4.[189,1399,819,986]" lastBlockId="6.[189,1399,267,399]" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
By far the most diagnostic character of 
<taxonomicName id="4C0D4D67FFFDFFBA75F26A12963CB156" authorityName="Frederiksen" authorityYear="2013" box="[685,804,819,845]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFDFFBA75F26A12963CB156" box="[685,804,819,845]" italics="true" pageId="4" pageNumber="5">Aquattuor</emphasis>
</taxonomicName>
is the large (sub)rectangular-lobed limbus. Subrectangular limbus lobes do occur in certain other odontopygid genera, e.g., 
<taxonomicName id="4C0D4D67FFFDFFBA73686A77906FB16B" authority="Attems, 1912" authorityName="Attems" authorityYear="1912" box="[1079,1399,854,880]" class="Diplopoda" family="Odontopygidae" genus="Allantogonus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFDFFBA73686A7791CBB16B" box="[1079,1235,854,880]" italics="true" pageId="4" pageNumber="5">Allantogonus</emphasis>
Attems, 1912
</taxonomicName>
(
<bibRefCitation id="EF9C4B15FFFDFFBA779A6A58944AB188" author="Kraus O." box="[197,338,889,915]" pageId="4" pageNumber="5" pagination="1 - 207" refId="ref8365" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">
Kraus 
<date id="FFB31024FFFDFFBA764C6A58944AB188" box="[275,338,889,915]" pageId="4" pageNumber="5" value="1960">1960</date>
</bibRefCitation>
: fig. 5) and 
<taxonomicName id="4C0D4D67FFFDFFBA76876A58962DB188" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" box="[472,821,889,916]" class="Diplopoda" family="Odontopygidae" genus="Syndesmogenus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFDFFBA76876A58978AB188" box="[472,658,889,915]" italics="true" pageId="4" pageNumber="5">Syndesmogenus</emphasis>
<bibRefCitation id="EF9C4B15FFFDFFBA75C76A5B962DB188" author="Attems C." box="[664,821,889,916]" pageId="4" pageNumber="5" pagination="1 - 64" refId="ref7905" refString="Attems C. 1909. Myriopoda. Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906 3 (19): 1 - 64." type="journal article" year="1909">Attems, 1909</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF9C4B15FFFDFFBA741C6A5896C8B188" author="Kraus O." box="[835,976,889,915]" pageId="4" pageNumber="5" pagination="1 - 143" refId="ref8396" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." type="journal article" year="1966">
Kraus 
<date id="FFB31024FFFDFFBA74CE6A5896C8B188" box="[913,976,889,915]" pageId="4" pageNumber="5" value="1966">1966</date>
</bibRefCitation>
: figs 97–99), but in these genera, the lobes are much smaller and are not prone to detachment. The gonopods of 
<taxonomicName id="4C0D4D67FFFDFFBA73666ABC91A8B1AC" authorityName="Frederiksen" authorityYear="2013" box="[1081,1200,925,951]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFDFFBA73666ABC91A8B1AC" box="[1081,1200,925,951]" italics="true" pageId="4" pageNumber="5">Aquattuor</emphasis>
</taxonomicName>
, with their long, whip-like solenomere and slender telomere, superficially resemble those of 
<taxonomicName id="4C0D4D67FFFDFFBA73676AE191CCB1C1" authorityName="Attems" authorityYear="1912" box="[1080,1236,960,986]" class="Diplopoda" family="Odontopygidae" genus="Allantogonus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFDFFBA73676AE191CCB1C1" box="[1080,1236,960,986]" italics="true" pageId="4" pageNumber="5">Allantogonus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9C4B15FFFDFFBA73BB6AE1906BB1C1" author="Kraus O." box="[1252,1395,960,986]" pageId="4" pageNumber="5" pagination="1 - 207" refId="ref8365" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">
Kraus 
<date id="FFB31024FFFDFFBA726B6AE1906BB1C1" box="[1332,1395,960,986]" pageId="4" pageNumber="5" value="1960">1960</date>
</bibRefCitation>
: figs 7, 11, 17) and 
<taxonomicName id="4C0D4D67FFFFFFB876C1682A963DB33E" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[414,805,267,293]" class="Diplopoda" family="Odontopygidae" genus="Lamelloramus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFFFFB876C1682A975FB33E" box="[414,583,267,293]" italics="true" pageId="6" pageNumber="7">Lamelloramus</emphasis>
Frederiksen, 2013
</taxonomicName>
(Frederiksen 
<date id="FFB31024FFFFFFB87495682A9111B33E" box="[970,1033,267,293]" pageId="6" pageNumber="7" value="2013">2013</date>
: figs 5, 9). However, the coxal apex in 
<taxonomicName id="4C0D4D67FFFFFFB87642680E94A1B352" authorityName="Attems" authorityYear="1912" box="[285,441,303,329]" class="Diplopoda" family="Odontopygidae" genus="Allantogonus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFFFFB87642680E94A1B352" box="[285,441,303,329]" italics="true" pageId="6" pageNumber="7">Allantogonus</emphasis>
</taxonomicName>
is folded basad over the anterior side; the metaplical flange of 
<taxonomicName id="4C0D4D67FFFFFFB873EF680E9041B352" authorityName="Frederiksen" authorityYear="2013" box="[1200,1369,303,329]" class="Diplopoda" family="Odontopygidae" genus="Lamelloramus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFFFFB873EF680E9041B352" box="[1200,1369,303,329]" italics="true" pageId="6" pageNumber="7">Lamelloramus</emphasis>
</taxonomicName>
is situated completely differently on the median side, and the telomere is either strongly curling up or looping in these genera. The limbus is completely different from that of 
<taxonomicName id="4C0D4D67FFFFFFB873536854919BB394" authorityName="Frederiksen" authorityYear="2013" box="[1036,1155,373,399]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFFFFB873536854919BB394" box="[1036,1155,373,399]" italics="true" pageId="6" pageNumber="7">Aquattuor</emphasis>
</taxonomicName>
in both.
</paragraph>
<caption id="DF72666CFFFDFFBA77E26EA594DFB5FE" ID-DOI="http://doi.org/10.5281/zenodo.3787592" ID-Zenodo-Dep="3787592" httpUri="https://zenodo.org/record/3787592/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1924,1950]" targetBox="[187,1400,1040,1895]" targetPageId="4">
<paragraph id="8BB236E4FFFDFFBA77E26EA594DFB5FE" blockId="4.[189,1398,1924,2021]" pageId="4" pageNumber="5">
<emphasis id="B979EAF6FFFDFFBA77E26EA59408B584" bold="true" box="[189,272,1924,1951]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
Body size (numbers of podous rings and midbody vertical diameter) in ♂♂ of species of 
<taxonomicName id="4C0D4D67FFFDFFBA77E26E899738B5D9" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[189,544,1960,1986]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFDFFBA77E26E89942CB5D9" box="[189,308,1960,1986]" italics="true" pageId="4" pageNumber="5">Aquattuor</emphasis>
Frederiksen, 2013
</taxonomicName>
. In cases of (almost) coinciding values, symbols have been slightly displaced horizontally.
</paragraph>
</caption>
<caption id="DF72666CFFFCFFBB77E26E4795EAB5F0" ID-DOI="http://doi.org/10.5281/zenodo.3787594" ID-Zenodo-Dep="3787594" httpUri="https://zenodo.org/record/3787594/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1894,1920]" targetBox="[236,1347,264,1860]" targetPageId="5">
<paragraph id="8BB236E4FFFCFFBB77E26E4795EAB5F0" blockId="5.[189,1399,1894,2027]" pageId="5" pageNumber="6">
<emphasis id="B979EAF6FFFCFFBB77E26E479414B59B" bold="true" box="[189,268,1894,1920]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="4C0D4D67FFFCFFBB764A6E469494B59A" authorityName="Frederiksen" authorityYear="2013" box="[277,396,1895,1921]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAF6FFFCFFBB764A6E469494B59A" box="[277,396,1895,1921]" italics="true" pageId="5" pageNumber="6">Aquattuor</emphasis>
</taxonomicName>
spp. 
<emphasis id="B979EAF6FFFCFFBB76916E4794FBB59B" bold="true" box="[462,483,1894,1920]" pageId="5" pageNumber="6">A</emphasis>
. 
<taxonomicName id="4C0D4D67FFFCFFBB76AC6E46961BB59B" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[499,771,1894,1921]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="stereosathe" status="sp. nov.">
<emphasis id="B979EAF6FFFCFFBB76AC6E469783B59B" box="[499,667,1894,1921]" italics="true" pageId="5" pageNumber="6">A. stereosathe</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFCFFBB74536E469672B59A" box="[780,874,1895,1921]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, front end. 
<emphasis id="B979EAF6FFFCFFBB73356E4791B9B59B" bold="true" box="[1130,1185,1894,1920]" pageId="5" pageNumber="6">B–C</emphasis>
. 
<taxonomicName id="4C0D4D67FFFCFFBB73EE6E469405B5BF" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="udzungwensis" status="sp. nov.">
<emphasis id="B979EAF6FFFCFFBB73EE6E46906EB59B" box="[1201,1398,1894,1921]" italics="true" pageId="5" pageNumber="6">A. udzungwensis</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFCFFBB767A6EAB949BB5BF" box="[293,387,1930,1956]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, telson, lateral and posterior view. 
<emphasis id="B979EAF6FFFCFFBB74C86EAB96D6B5BF" bold="true" box="[919,974,1930,1956]" pageId="5" pageNumber="6">D–E</emphasis>
. 
<taxonomicName id="4C0D4D67FFFCFFBB74816EAB9014B5BF" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[990,1292,1930,1956]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="udzungwensis" status="sp. nov.">
<emphasis id="B979EAF6FFFCFFBB74816EAB91BBB5BF" box="[990,1187,1930,1956]" italics="true" pageId="5" pageNumber="6">A. udzungwensis</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFCFFBB724B6EAB906AB5BF" box="[1300,1394,1930,1956]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, limbus. D. Specimen from West Kilombero Scarp FR. E. Paratype. Scale bars: A–D = 0.1 mm; E = 0.01 mm.
</paragraph>
</caption>
<paragraph id="8BB236E4FFFFFFB877E268969751B3CA" blockId="6.[189,585,438,465]" box="[189,585,438,465]" pageId="6" pageNumber="7">
<emphasis id="B979EAF6FFFFFFB877E268969491B3CB" bold="true" box="[189,393,438,465]" pageId="6" pageNumber="7">Included species</emphasis>
(alphabetically)
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E268C4967FB3E4" blockId="6.[189,871,485,723]" box="[189,871,485,511]" pageId="6" pageNumber="7">
<taxonomicName id="4C0D4D67FFFFFFB877E268C4961DB3E4" authority="Enghoff &amp; Frederiksen" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[189,773,485,511]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="claudiahempae" status="sp. nov.">
<emphasis id="B979EAF6FFFFFFB877E268C494F6B3E4" box="[189,494,485,511]" italics="true" pageId="6" pageNumber="7">Aquattuor claudiahempae</emphasis>
Enghoff &amp; Frederiksen
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFFFFB8745268C4967FB3E4" box="[781,871,485,511]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26B2897EAB038" blockId="6.[189,871,485,723]" box="[189,754,520,547]" pageId="6" pageNumber="7">
<taxonomicName id="4C0D4D67FFFFFFB877E26B289750B039" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[189,584,520,547]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B979EAF6FFFFFFB877E26B289475B039" box="[189,365,520,547]" italics="true" pageId="6" pageNumber="7">A. denticulatus</emphasis>
Frederiksen, 2013
</taxonomicName>
(
<typeStatus id="54B68846FFFFFFB875086B299793B039" box="[599,651,520,546]" pageId="6" pageNumber="7">type</typeStatus>
species)
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26B0D970FB05D" blockId="6.[189,871,485,723]" box="[189,535,555,582]" pageId="6" pageNumber="7">
<taxonomicName id="4C0D4D67FFFFFFB877E26B0D94ADB05E" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[189,437,555,582]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longipala" status="sp. nov.">
<emphasis id="B979EAF6FFFFFFB877E26B0D9456B05E" box="[189,334,555,582]" italics="true" pageId="6" pageNumber="7">A. longipala</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFFFFB876E36B0D970FB05D" box="[444,535,556,582]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26B6E94F5B072" blockId="6.[189,871,485,723]" box="[189,493,590,617]" pageId="6" pageNumber="7">
<taxonomicName id="4C0D4D67FFFFFFB877E26B6E9493B073" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[189,395,590,617]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="major" status="sp. nov.">
<emphasis id="B979EAF6FFFFFFB877E26B6E943CB073" box="[189,292,591,617]" italics="true" pageId="6" pageNumber="7">A. major</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFFFFB876CD6B6E94F5B072" box="[402,493,591,617]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26B539733B097" blockId="6.[189,871,485,723]" box="[189,555,626,652]" pageId="6" pageNumber="7">
<taxonomicName id="4C0D4D67FFFFFFB877E26B5394D1B097" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[189,457,626,652]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="stereosathe" status="sp. nov.">
<emphasis id="B979EAF6FFFFFFB877E26B53947BB097" box="[189,355,626,652]" italics="true" pageId="6" pageNumber="7">A. stereosathe</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFFFFB8768F6B539733B097" box="[464,555,626,652]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26BB4970DB0B4" blockId="6.[189,871,485,723]" box="[189,533,661,687]" pageId="6" pageNumber="7">
<taxonomicName id="4C0D4D67FFFFFFB877E26BB494ACB0B4" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[189,436,661,687]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="submajor" status="sp. nov.">
<emphasis id="B979EAF6FFFFFFB877E26BB49455B0B4" box="[189,333,661,687]" italics="true" pageId="6" pageNumber="7">A. submajor</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFFFFB876E46BB4970DB0B4" box="[443,533,661,687]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26B989751B0C8" blockId="6.[189,871,485,723]" box="[189,585,696,723]" pageId="6" pageNumber="7">
<taxonomicName id="4C0D4D67FFFFFFB877E26B9894F0B0C9" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[189,488,696,723]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="udzungwensis" status="sp. nov.">
<emphasis id="B979EAF6FFFFFFB877E26B989499B0C9" box="[189,385,696,723]" italics="true" pageId="6" pageNumber="7">A. udzungwensis</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFFFFB876B06B989751B0C8" box="[495,585,697,723]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>