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<document id="C91F1089E68DA17007E4A71EE99FD77F" ID-DOI="10.11646/palaeoentomology.6.2.11" ID-ISSN="2624-2834" ID-Zenodo-Dep="7928885" ID-ZooBank="1E80FE4B-F846-4243-ABB9-15CF7F8341F4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683887594507" checkinUser="plazi" docAuthor="GOROCHOV, ANDREI V. &amp; CORAM, ROBERT A." docDate="2023" docId="03A487E6D645D331FC987AD5FD65FA06" docLanguage="en" docName="palaeoentomology.6.2.11.pdf" docOrigin="Palaeoentomology 6 (2)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.2.11" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Daniilacheta Gorochov &amp; Coram 2023, gen. nov." docType="treatment" docVersion="2" lastPageNumber="200" masterDocId="FF9DFF9ED644D333FFB27E7DFFD7FF86" masterDocTitle="New and little known taxa of the order Orthoptera (Insecta) from the Upper Triassic and Lower Jurassic of England" masterLastPageNumber="204" masterPageNumber="198" pageNumber="199" updateTime="1683888079122" updateUser="ExternalLinkService">
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<mods:title id="6062CE87644C347A19D58D3AF75398E6">New and little known taxa of the order Orthoptera (Insecta) from the Upper Triassic and Lower Jurassic of England</mods:title>
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<mods:namePart id="6F99A3991B5690BD1FC9AA8B3B522E3E">GOROCHOV, ANDREI V.</mods:namePart>
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<mods:namePart id="26F20084FB41569A7C1807C34F7CBC25">CORAM, ROBERT A.</mods:namePart>
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<mods:title id="3702E49E9D0DC45209FEEA471E1D9CE5">Palaeoentomology</mods:title>
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<mods:date id="F726745EDADB2BBC7399BC15DCD780AD">2023</mods:date>
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<mods:number id="645A5B6EFDDB9F0A8F604CAAF0D6B1FE">2023-04-28</mods:number>
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<subSubSection id="C317657BD645D332FC987AD5FAA8FB45" box="[810,1407,1192,1219]" pageId="1" pageNumber="199" type="nomenclature">
<paragraph id="8BB236F0D645D332FC987AD5FAA8FB45" blockId="1.[810,1407,1192,1219]" box="[810,1407,1192,1219]" pageId="1" pageNumber="199">
<heading id="D0FA819CD645D332FC987AD5FAA8FB45" box="[810,1407,1192,1219]" fontSize="10" level="2" pageId="1" pageNumber="199" reason="2">
<emphasis id="B979EAE2D645D332FC987AD5FAA8FB45" bold="true" box="[810,1407,1192,1219]" pageId="1" pageNumber="199">
Genus
<taxonomicName id="4C0D4D73D645D332FCCF7AD5FAD8FB44" authority="Gorochov &amp; Coram" authorityName="Gorochov &amp; Coram" authorityYear="2023" box="[893,1295,1192,1219]" class="Insecta" family="Protogryllidae" genus="Daniilacheta" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B979EAE2D645D332FCCF7AD5FBC5FB45" bold="true" box="[893,1042,1192,1219]" italics="true" pageId="1" pageNumber="199">Daniilacheta</emphasis>
Gorochov &amp; Coram
</taxonomicName>
<taxonomicNameLabel id="A24A5799D645D332FAA47AD4FAA8FB45" box="[1302,1407,1193,1219]" pageId="1" pageNumber="199" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C317657BD645D332FC987A8FFB03FAD2" pageId="1" pageNumber="199" type="type_taxon">
<paragraph id="8BB236F0D645D332FC987A8FFBDDFAB6" blockId="1.[810,1452,1265,2093]" pageId="1" pageNumber="199">
<emphasis id="B979EAE2D645D332FC987A8FFC10FA8D" bold="true" box="[810,967,1265,1292]" pageId="1" pageNumber="199">
<typeStatus id="54B68852D645D332FC987A8FFCB2FA8A" box="[810,869,1266,1292]" pageId="1" pageNumber="199">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C0D4D73D645D332FC627A8CFB15FA8D" authority="GOROCHOV &amp; CORAM, 2023" authorityName="GOROCHOV &amp; CORAM" authorityYear="2023" box="[976,1218,1265,1291]" class="Insecta" family="Protogryllidae" genus="Daniilacheta" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="species" species="aristovi" status="gen. et sp. nov.">
<emphasis id="B979EAE2D645D332FC627A8CFB15FA8D" box="[976,1218,1265,1291]" italics="true" pageId="1" pageNumber="199">Daniilacheta aristovi</emphasis>
</taxonomicName>
<emphasis id="B979EAE2D645D332FB787A8FFA50FA8A" bold="true" box="[1226,1415,1266,1292]" pageId="1" pageNumber="199">
<taxonomicNameLabel id="A24A5799D645D332FB787A8FFA56FA8A" box="[1226,1409,1266,1292]" pageId="1" pageNumber="199" rank="species">gen. et sp. nov.</taxonomicNameLabel>
,
</emphasis>
by present designation.
</paragraph>
<paragraph id="8BB236F0D645D332FCED7B47FB03FAD2" blockId="1.[810,1452,1265,2093]" box="[863,1236,1338,1365]" pageId="1" pageNumber="199">
<emphasis id="B979EAE2D645D332FCED7B47FC28FAD2" bold="true" box="[863,1023,1338,1364]" pageId="1" pageNumber="199">Composition.</emphasis>
Only
<typeStatus id="54B68852D645D332FBF77B46FBA2FAD3" box="[1093,1141,1339,1365]" pageId="1" pageNumber="199">type</typeStatus>
species.
</paragraph>
</subSubSection>
<subSubSection id="C317657BD645D332FCED7B22FB5AFA44" pageId="1" pageNumber="199" type="etymology">
<paragraph id="8BB236F0D645D332FCED7B22FB5AFA44" blockId="1.[810,1452,1265,2093]" pageId="1" pageNumber="199">
<emphasis id="B979EAE2D645D332FCED7B22FC31FAFF" bold="true" box="[863,998,1375,1401]" pageId="1" pageNumber="199">Etymology.</emphasis>
After the first name of the Russian palaeoentomologist Daniil [=Danila] S. Aristov (in his memory) and the genus
<taxonomicName id="4C0D4D73D645D332FB8A7BDAFB51FA47" box="[1080,1158,1447,1473]" class="Insecta" family="Gryllidae" genus="Acheta" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FB8A7BDAFB51FA47" box="[1080,1158,1447,1473]" italics="true" pageId="1" pageNumber="199">Acheta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C317657BD645D332FCED7BB1FBB8F865" pageId="1" pageNumber="199" type="diagnosis">
<paragraph id="8BB236F0D645D332FCED7BB1FBB8F865" blockId="1.[810,1452,1265,2093]" pageId="1" pageNumber="199">
<emphasis id="B979EAE2D645D332FCED7BB1FC0EFA60" bold="true" box="[863,985,1484,1510]" pageId="1" pageNumber="199">Diagnosis.</emphasis>
Male tegmen (
<figureCitation id="13362A75D645D332FB127BB1FAD7FA60" box="[1184,1280,1484,1510]" captionStart="FIGURE 1" captionStartId="3.[136,235,1357,1381]" captionTargetBox="[249,1337,167,1332]" captionTargetId="figure-213@3.[248,1340,164,1334]" captionTargetPageId="3" captionText="FIGURE 1. Daniilacheta aristovi gen. et sp. nov., male tegmen, holotype LD/Orth 19, Dorset, Lower Jurassic. A, Reconstruction of spread venation. B, Photograph of imprint (part). C, Photograph of imprint (counterpart). Abbreviations: c area with chords (distal parts of CuP, 1A and 2A); d diagonal vein (oblique crossvein crossing CuA2 and having two branches near stridulatory vein); h harp area with harp veins or “oblique veins” (crossveins between CuA1 and proximal part of CuP); lancet lancet-like area between R and MA. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.7928895" httpUri="https://zenodo.org/record/7928895/files/figure.png" pageId="1" pageNumber="199">Fig. 1A</figureCitation>
) with a very long apical area and distinctly developed stridulatory apparatus having the following characters: lancet-like area also distinctly developed, probably moderately wide and with two or three crossveins in its distal half; distal parts of Sc and R+1MA1 strongly protruding beyond this area; harp area with strongly arcuate crossveins (“oblique veins”); diagonal vein and proximal part of CuA2 (before it) clearly developed; rest of CuA2 (behind diagonal vein) and majority of branches of MP+CuA1 almost straight to very slightly arcuate, obliquely longitudinal, with areas between them rather narrow and more or less equal in shape but gradually narrowing to tegminal apex; stridulatory vein obliquely transverse, rather long; chords strongly curved but not long.
</paragraph>
</subSubSection>
<subSubSection id="C317657BD645D331FCED7993FD65FA06" lastPageId="2" lastPageNumber="200" pageId="1" pageNumber="199" type="discussion">
<paragraph id="8BB236F0D645D331FCED7993FE9DFD7E" blockId="1.[810,1452,1265,2093]" lastBlockId="2.[136,778,158,1409]" lastPageId="2" lastPageNumber="200" pageId="1" pageNumber="199">
<emphasis id="B979EAE2D645D332FCED7993FC07F78E" bold="true" box="[863,976,2030,2056]" pageId="1" pageNumber="199">Remarks.</emphasis>
The new genus is similar to
<taxonomicName id="4C0D4D73D645D332FAB17993FA7CF78E" authorityName="Gorochov" authorityYear="1984" box="[1283,1451,2030,2056]" class="Insecta" family="Protogryllidae" genus="Bacharogryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FAB17993FA7CF78E" box="[1283,1451,2030,2056]" italics="true" pageId="1" pageNumber="199">Bacharogryllus</emphasis>
</taxonomicName>
in the presence of two or more crossveins in the lancet-like area, but is distinguished from it by this area having more numerous crossveins in its distal half (
<taxonomicName id="4C0D4D73D646D331FDEF7EBFFCDDFF5A" authorityName="Gorochov" authorityYear="1984" box="[605,778,194,220]" class="Insecta" family="Protogryllidae" genus="Bacharogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D646D331FDEF7EBFFCDDFF5A" box="[605,778,194,220]" italics="true" pageId="2" pageNumber="200">Bacharogryllus</emphasis>
</taxonomicName>
has one crossvein, whereas
<taxonomicName id="4C0D4D73D646D331FE7D7E9BFDB6FE86" authority="GOROCHOV &amp; CORAM, 2023" authorityName="Gorochov &amp; Coram" authorityYear="2023" box="[463,609,230,256]" class="Insecta" family="Protogryllidae" genus="Daniilacheta" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B979EAE2D646D331FE7D7E9BFDB6FE86" box="[463,609,230,256]" italics="true" pageId="2" pageNumber="200">Daniilacheta</emphasis>
</taxonomicName>
<emphasis id="B979EAE2D646D331FDDE7E9BFD0EFE86" bold="true" box="[620,729,230,256]" pageId="2" pageNumber="200">
<taxonomicNameLabel id="A24A5799D646D331FDDE7E9BFD0EFE86" box="[620,729,230,256]" pageId="2" pageNumber="200" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
has two or more) and by a shorter harp area with significantly more arcuate crossveins. The new genus differs from
<taxonomicName id="4C0D4D73D646D331FF3A7F2FFE97FEEA" authorityName="Haughton" authorityYear="1924" box="[136,320,338,364]" class="Insecta" family="Protogryllidae" genus="Archaegryllodes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D646D331FF3A7F2FFE97FEEA" box="[136,320,338,364]" italics="true" pageId="2" pageNumber="200">Archaegryllodes</emphasis>
</taxonomicName>
in having more numerous crossveins in the lancet-like area (
<taxonomicName id="4C0D4D73D646D331FE157F0BFD88FE16" authorityName="Haughton" authorityYear="1924" box="[423,607,374,400]" class="Insecta" family="Protogryllidae" genus="Archaegryllodes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D646D331FE157F0BFD88FE16" box="[423,607,374,400]" italics="true" pageId="2" pageNumber="200">Archaegryllodes</emphasis>
</taxonomicName>
has only one crossvein) as well as having a shorter harp area and a longer stridulatory vein. Differences from
<taxonomicName id="4C0D4D73D646D331FDCF7FC3FCDDFE5E" authorityName="Handlirsch" authorityYear="1906" box="[637,778,446,472]" class="Insecta" family="Protogryllidae" genus="Protogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D646D331FDCF7FC3FCDDFE5E" box="[637,778,446,472]" italics="true" pageId="2" pageNumber="200">Protogryllus</emphasis>
</taxonomicName>
are problematic with respect to the above discussion, but some specimens known from male tegmina and included in this genus by different authors are distinguished from
<taxonomicName id="4C0D4D73D646D331FF3A7C33FECDFDEE" authority="GOROCHOV &amp; CORAM, 2023" authorityName="Gorochov &amp; Coram" authorityYear="2023" box="[136,282,590,616]" class="Insecta" family="Protogryllidae" genus="Daniilacheta" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B979EAE2D646D331FF3A7C33FECDFDEE" box="[136,282,590,616]" italics="true" pageId="2" pageNumber="200">Daniilacheta</emphasis>
</taxonomicName>
<emphasis id="B979EAE2D646D331FE957C33FE42FDEE" bold="true" box="[295,405,590,616]" pageId="2" pageNumber="200">
<taxonomicNameLabel id="A24A5799D646D331FE957C33FE42FDEE" box="[295,405,590,616]" pageId="2" pageNumber="200" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
by the lancet-like area lacking crossveins, the harp area with almost straight crossveins, the apical area and distal parts of Sc and R+1MA1 distinctly shorter, and some of the MP+CuA1 branches clearly S-shaped.
</paragraph>
<paragraph id="8BB236F0D646D331FF0F7D7EFD65FA06" blockId="2.[136,778,158,1409]" pageId="2" pageNumber="200">
It is necessary also to mention that
<taxonomicName id="4C0D4D73D646D331FDEA7D7FFE94FCC6" authority="Gorochov, 1992" authorityName="Gorochov" authorityYear="1992" class="Insecta" family="Protogryllidae" genus="Aenigmagryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D646D331FDEA7D7FFCDEFC9A" box="[600,777,770,796]" italics="true" pageId="2" pageNumber="200">Aenigmagryllus</emphasis>
Gorochov, 1992
</taxonomicName>
(Upper Jurassic or Lower Cretaceous of Mongolia), which has an unknown lancet-like area, is included in
<taxonomicName id="4C0D4D73D646D331FE817D13FE04FC0E" authorityName="Zeuner" authorityYear="1937" box="[307,467,878,904]" class="Insecta" family="Protogryllidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="family">Protogryllidae</taxonomicName>
without attribution to any subfamily and differs from the new genus in having less arcuate crossveins of the harp area, a strongly S-shaped portion of CuA2 located behind the diagonal vein, and a shorter costal (longest) chord. The two other subfamilies of
<taxonomicName id="4C0D4D73D646D331FF1B7A5FFE9EFBBA" authorityName="Zeuner" authorityYear="1937" box="[169,329,1058,1084]" class="Insecta" family="Protogryllidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="family">Protogryllidae</taxonomicName>
(Karataogryllinae Gorochov, 1985 and Falsispeculinae Gorochov, 1985) can be respectively distinguished from the new genus by a much longer costal chord of the male tegmen and by the absence of crossveins in the distal part of the lancet-like area mentioned above (Falsispeculinae and
<taxonomicName id="4C0D4D73D646D331FDD87AABFCDDFB76" authorityName="Zeuner" authorityYear="1937" box="[618,778,1238,1264]" class="Insecta" family="Protogryllidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="subFamily" subFamily="Protogryllinae">Protogryllinae</taxonomicName>
differ from each other mainly in the structure of the base of this lancet-like area, unknown in the new genus, and in this connection, inclusion of this genus in
<taxonomicName id="4C0D4D73D646D331FDD87B3FFCDDFADA" authorityName="Zeuner" authorityYear="1937" box="[618,778,1346,1372]" class="Insecta" family="Protogryllidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="subFamily" subFamily="Protogryllinae">Protogryllinae</taxonomicName>
but not to Falsispeculinae is somewhat tentative).
</paragraph>
</subSubSection>
</treatment>
</document>