treatments-xml/data/03/A4/87/03A487DBFFD9FFE158B15898FE8FDB43.xml

<document ID-DOI="10.11646/zootaxa.5239.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7635170" ID-ZooBank="5F1C330D-6344-4A7C-AD67-87F880D832F1" approvalRequired="5" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="4" checkinTime="1676283582796" checkinUser="plazi" docAuthor="Lencioni, F. A. A." docDate="2023" docId="03A487DBFFD9FFE158B15898FE8FDB43" docLanguage="en" docName="zootaxa.5239.4.3.pdf" docOrigin="Zootaxa 5239 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5239.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helveciagrion Machado 1980" docType="treatment" docVersion="2" lastPageNumber="510" masterDocId="FF9DFFA3FFDFFFEB58265E5EFFC5DF76" masterDocTitle="Analysis of Telebasis Selys, 1865 - Part I. (Odonata: Coenagrionidae)" masterLastPageNumber="520" masterPageNumber="500" pageNumber="506" updateTime="1676284197801" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Analysis of Telebasis Selys, 1865 - Part I. (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart>Lencioni, F. A. A.</mods:namePart>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<subSubSection box="[151,521,1734,1761]" pageId="6" pageNumber="506" type="nomenclature">
<paragraph blockId="6.[151,1436,1734,1833]" box="[151,521,1734,1761]" pageId="6" pageNumber="506">
<heading bold="true" box="[151,521,1734,1761]" fontSize="11" level="1" pageId="6" pageNumber="506" reason="1">
<taxonomicName authority="Machado, 1980" authorityName="Machado" authorityYear="1980" box="[151,521,1734,1761]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[151,521,1734,1761]" pageId="6" pageNumber="506">
<emphasis bold="true" box="[151,322,1734,1761]" italics="true" pageId="6" pageNumber="506">Helveciagrion</emphasis>
<bibRefCitation author="Machado, A. B. M." box="[330,521,1734,1761]" pageId="6" pageNumber="506" pagination="59 - 87" refId="ref9803" refString="Machado, A. B. M. (1980) Helveciagrion n. g., com descricao de uma nova especie do Parque Estadual do Rio Doce, Minas Gerais (Odonata: Coenagrionidae). Lundiana, 1, 59 - 87." type="journal article" year="1980">Machado, 1980</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="506" type="description">
<paragraph blockId="6.[151,1436,1734,1833]" pageId="6" pageNumber="506">
<figureCitation box="[151,216,1770,1797]" captionStart="FIGURE 5" captionStartId="7.[152,255,1078,1102]" captionTargetBox="[284,1303,181,1055]" captionTargetId="figure-20@7.[284,1303,181,1055]" captionTargetPageId="7" captionText="FIGURE 5. Helveciagrion obsoletum—male appendages in posterior view, showing a pair of heavily sclerotized tubercles." figureDoi="http://doi.org/10.5281/zenodo.7635184" httpUri="https://zenodo.org/record/7635184/files/figure.png" pageId="6" pageNumber="506">Fig. 5</figureCitation>
(♁ app., posterior view). 
<figureCitation box="[503,568,1771,1797]" captionStart="FIGURE 6" captionStartId="7.[152,255,1997,2021]" captionTargetBox="[266,1186,1162,1914]" captionTargetId="figure-44@7.[266,1320,1127,1974]" captionTargetPageId="7" captionText="FIGURE 6. Helveciagrion obsoletum—male appendages in lateral view, showing a pair of heavily sclerotized tubercles." figureDoi="http://doi.org/10.5281/zenodo.7635186" httpUri="https://zenodo.org/record/7635186/files/figure.png" pageId="6" pageNumber="506">Fig. 6</figureCitation>
(♁ app., lateral view). 
<figureCitation box="[825,890,1771,1797]" captionStart="FIGURE 7" captionStartId="8.[152,255,1102,1126]" captionTargetBox="[176,1418,181,1078]" captionTargetId="figure-15@8.[151,1436,181,1078]" captionTargetPageId="8" captionText="FIGURE 7. Helveciagrion obsoletum—female thorax in dorsal view, showing the mesepisternal fossae where male tubercles fit." figureDoi="http://doi.org/10.5281/zenodo.7635188" httpUri="https://zenodo.org/record/7635188/files/figure.png" pageId="6" pageNumber="506">Fig. 7</figureCitation>
(
<specimenCount box="[907,926,1771,1797]" pageId="6" pageNumber="506" type="female">♀</specimenCount>
prothorax, dorsal view). 
<figureCitation box="[1207,1272,1771,1797]" captionStart="FIGURE 8" captionStartId="8.[152,255,1913,1937]" captionTargetBox="[151,1436,1151,1890]" captionTargetId="figure-39@8.[151,1436,1151,1890]" captionTargetPageId="8" captionText="FIGURE 8. Tandem pair of Helveciagrion obsoletum." figureDoi="http://doi.org/10.5281/zenodo.7635190" httpUri="https://zenodo.org/record/7635190/files/figure.png" pageId="6" pageNumber="506">Fig. 8</figureCitation>
(tandem pair), 
<figureCitation box="[151,217,1807,1833]" captionStart="FIGURE 9" captionStartId="9.[152,255,1156,1180]" captionTargetBox="[251,1336,181,1132]" captionTargetId="figure-20@9.[251,1336,181,1132]" captionTargetPageId="9" captionText="FIGURE 9. Detail of the contact between cercus and mesepisternal fossae." figureDoi="http://doi.org/10.5281/zenodo.7635192" httpUri="https://zenodo.org/record/7635192/files/figure.png" pageId="6" pageNumber="506">Fig. 9</figureCitation>
(detail of the contact between cercus and mesepisternal fossae), 
<figureCitation box="[942,1021,1807,1833]" captionStart="FIGURE 10" captionStartId="9.[152,255,1985,2009]" captionTargetBox="[226,1362,1204,1961]" captionTargetId="figure-38@9.[226,1362,1204,1961]" captionTargetPageId="9" captionText="FIGURE 10. Helveciagrion obsoletum male, Brotas—S.P., 17.i.2012 (FAAL)." figureDoi="http://doi.org/10.5281/zenodo.7635194" httpUri="https://zenodo.org/record/7635194/files/figure.png" pageId="6" pageNumber="506">Fig. 10</figureCitation>
(live specimen),
</paragraph>
<paragraph blockId="6.[151,1141,1870,1894]" box="[151,1141,1870,1894]" pageId="6" pageNumber="506">
<typeStatus box="[151,201,1870,1894]" pageId="6" pageNumber="506">Type</typeStatus>
species. 
<taxonomicName authority="Machado, 1980" authorityName="Machado" authorityYear="1980" box="[294,715,1870,1894]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="506" phylum="Arthropoda" rank="species" species="vulcanoae">
<emphasis box="[294,549,1870,1894]" italics="true" pageId="6" pageNumber="506">Helveciagrion vulcanoae</emphasis>
<bibRefCitation author="Machado, A. B. M." box="[555,715,1870,1894]" pageId="6" pageNumber="506" pagination="59 - 87" refId="ref9803" refString="Machado, A. B. M. (1980) Helveciagrion n. g., com descricao de uma nova especie do Parque Estadual do Rio Doce, Minas Gerais (Odonata: Coenagrionidae). Lundiana, 1, 59 - 87." type="journal article" year="1980">Machado, 1980</bibRefCitation>
</taxonomicName>
, by original designation (
<bibRefCitation author="Machado, A. B. M." box="[972,1130,1870,1894]" pageId="6" pageNumber="506" pagination="59 - 87" refId="ref9803" refString="Machado, A. B. M. (1980) Helveciagrion n. g., com descricao de uma nova especie do Parque Estadual do Rio Doce, Minas Gerais (Odonata: Coenagrionidae). Lundiana, 1, 59 - 87." type="journal article" year="1980">Machado 1980</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="510" pageId="6" pageNumber="506" type="etymology">
<paragraph blockId="6.[151,1436,1934,2033]" pageId="6" pageNumber="506">
<emphasis bold="true" box="[151,280,1935,1961]" pageId="6" pageNumber="506">Etymology</emphasis>
: 
<taxonomicName authority="Helvecio" authorityName="Machado" authorityYear="1980" box="[302,462,1935,1961]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis box="[302,462,1935,1961]" italics="true" pageId="6" pageNumber="506">Helveciagrion</emphasis>
</taxonomicName>
= 
<emphasis bold="true" box="[505,606,1934,1961]" italics="true" pageId="6" pageNumber="506">Helvecio</emphasis>
, after Dom Helvecio Gomes de Oliveira (b. 1876, d. 1960), Former Archbishop of Mariana–MG + 
<emphasis bold="true" box="[500,576,1971,1997]" italics="true" pageId="6" pageNumber="506">agrion</emphasis>
, neuter form of agrios (ἄγριος) meaning “wild”, which in damselflies is now used as an indicator that a genus belongs to the 
<taxonomicName authority="(Fliedner 2006)" baseAuthorityName="Fliedner" baseAuthorityYear="2006" box="[685,1047,2007,2033]" class="Insecta" family="Coenagrionidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="506" phylum="Arthropoda" rank="family">
Coenagrionidae (
<bibRefCitation author="Fliedner, H." box="[876,1040,2007,2033]" pageId="6" pageNumber="506" pagination="5 - 23" refId="ref9336" refString="Fliedner, H. (2006) Die wissenschaftlichen Namen der Libellen in Burmeisters &quot; Handbuch der Entomologie &quot;. Virgo, 9, 5 - 23." type="journal article" year="2006">Fliedner 2006</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635184" ID-Zenodo-Dep="7635184" box="[152,1409,1078,1103]" httpUri="https://zenodo.org/record/7635184/files/figure.png" pageId="7" pageNumber="507" startId="7.[152,255,1078,1102]" targetBox="[284,1303,181,1055]" targetPageId="7">
<paragraph blockId="7.[152,1409,1078,1103]" box="[152,1409,1078,1103]" pageId="7" pageNumber="507">
<emphasis bold="true" box="[152,274,1078,1103]" pageId="7" pageNumber="507">FIGURE 5</emphasis>
. 
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1876" box="[287,542,1079,1103]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="507" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis box="[287,542,1079,1103]" italics="true" pageId="7" pageNumber="507">Helveciagrion obsoletum</emphasis>
</taxonomicName>
—male appendages in posterior view, showing a pair of heavily sclerotized tubercles.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635186" ID-Zenodo-Dep="7635186" box="[152,1382,1997,2022]" httpUri="https://zenodo.org/record/7635186/files/figure.png" pageId="7" pageNumber="507" startId="7.[152,255,1997,2021]" targetBox="[266,1186,1162,1914]" targetPageId="7">
<paragraph blockId="7.[152,1382,1997,2022]" box="[152,1382,1997,2022]" pageId="7" pageNumber="507">
<emphasis bold="true" box="[152,280,1997,2022]" pageId="7" pageNumber="507">FIGURE 6.</emphasis>
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1876" box="[287,542,1998,2022]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="507" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis box="[287,542,1998,2022]" italics="true" pageId="7" pageNumber="507">Helveciagrion obsoletum</emphasis>
</taxonomicName>
—male appendages in lateral view, showing a pair of heavily sclerotized tubercles.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635188" ID-Zenodo-Dep="7635188" box="[152,1437,1102,1126]" httpUri="https://zenodo.org/record/7635188/files/figure.png" pageId="8" pageNumber="508" startId="8.[152,255,1102,1126]" targetBox="[176,1418,181,1078]" targetPageId="8">
<paragraph blockId="8.[152,1437,1102,1126]" box="[152,1437,1102,1126]" pageId="8" pageNumber="508">
<emphasis bold="true" box="[152,278,1102,1126]" pageId="8" pageNumber="508">FIGURE 7.</emphasis>
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1876" box="[283,537,1102,1126]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="508" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis box="[283,537,1102,1126]" italics="true" pageId="8" pageNumber="508">Helveciagrion obsoletum</emphasis>
</taxonomicName>
—female thorax in dorsal view, showing the mesepisternal fossae where male tubercles fit.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635190" ID-Zenodo-Dep="7635190" box="[152,709,1913,1937]" httpUri="https://zenodo.org/record/7635190/files/figure.png" pageId="8" pageNumber="508" startId="8.[152,255,1913,1937]" targetBox="[151,1436,1151,1890]" targetPageId="8">
<paragraph blockId="8.[152,709,1913,1937]" box="[152,709,1913,1937]" pageId="8" pageNumber="508">
<emphasis bold="true" box="[152,280,1913,1937]" pageId="8" pageNumber="508">FIGURE 8.</emphasis>
Tandem pair of 
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1876" box="[447,703,1913,1937]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="508" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis box="[447,703,1913,1937]" italics="true" pageId="8" pageNumber="508">Helveciagrion obsoletum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635192" ID-Zenodo-Dep="7635192" box="[152,920,1156,1180]" httpUri="https://zenodo.org/record/7635192/files/figure.png" pageId="9" pageNumber="509" startId="9.[152,255,1156,1180]" targetBox="[251,1336,181,1132]" targetPageId="9">
<paragraph blockId="9.[152,920,1156,1180]" box="[152,920,1156,1180]" pageId="9" pageNumber="509">
<emphasis bold="true" box="[152,280,1156,1180]" pageId="9" pageNumber="509">FIGURE 9.</emphasis>
Detail of the contact between cercus and mesepisternal fossae.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635194" ID-Zenodo-Dep="7635194" box="[152,956,1985,2009]" httpUri="https://zenodo.org/record/7635194/files/figure.png" pageId="9" pageNumber="509" startId="9.[152,255,1985,2009]" targetBox="[226,1362,1204,1961]" targetPageId="9">
<paragraph blockId="9.[152,956,1985,2009]" box="[152,956,1985,2009]" pageId="9" pageNumber="509">
<emphasis bold="true" box="[152,286,1985,2009]" pageId="9" pageNumber="509">FIGURE 10</emphasis>
. 
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1876" box="[299,555,1985,2009]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="509" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis box="[299,555,1985,2009]" italics="true" pageId="9" pageNumber="509">Helveciagrion obsoletum</emphasis>
</taxonomicName>
male, Brotas—S.P., 17.i.2012 (FAAL).
</paragraph>
</caption>
<paragraph blockId="10.[151,1437,150,1077]" pageId="10" pageNumber="510">
<emphasis bold="true" box="[199,514,150,176]" pageId="10" pageNumber="510">Generic characterization.</emphasis>
Medium sized 
<taxonomicName baseAuthorityName="Fliedner" baseAuthorityYear="2006" box="[718,895,151,177]" class="Insecta" family="Coenagrionidae" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="family">Coenagrionidae</taxonomicName>
(
<quantity box="[920,1057,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.8" metricValueMin="3.0" pageId="10" pageNumber="510" unit="mm" value="34.0" valueMax="38.0" valueMin="30.0">30–38 mm</quantity>
), angulated frons, male cercus approximate at the base, with a pair of heavily sclerotized tubercles of different sizes (
<figureCitation box="[1126,1222,186,213]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="7.[152,255,1078,1102]" captionStartId-1="7.[152,255,1997,2021]" captionTargetBox-0="[284,1303,181,1055]" captionTargetBox-1="[266,1186,1162,1914]" captionTargetId-0="figure-20@7.[284,1303,181,1055]" captionTargetId-1="figure-44@7.[266,1320,1127,1974]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="FIGURE 5. Helveciagrion obsoletum—male appendages in posterior view, showing a pair of heavily sclerotized tubercles." captionText-1="FIGURE 6. Helveciagrion obsoletum—male appendages in lateral view, showing a pair of heavily sclerotized tubercles." figureDoi-0="http://doi.org/10.5281/zenodo.7635184" figureDoi-1="http://doi.org/10.5281/zenodo.7635186" httpUri-0="https://zenodo.org/record/7635184/files/figure.png" httpUri-1="https://zenodo.org/record/7635186/files/figure.png" pageId="10" pageNumber="510">Fig. 5–6</figureCitation>
) that fit the pair of mesepisternal fossae in the pterothorax of the female (
<figureCitation box="[747,814,223,249]" captionStart="FIGURE 7" captionStartId="8.[152,255,1102,1126]" captionTargetBox="[176,1418,181,1078]" captionTargetId="figure-15@8.[151,1436,181,1078]" captionTargetPageId="8" captionText="FIGURE 7. Helveciagrion obsoletum—female thorax in dorsal view, showing the mesepisternal fossae where male tubercles fit." figureDoi="http://doi.org/10.5281/zenodo.7635188" httpUri="https://zenodo.org/record/7635188/files/figure.png" pageId="10" pageNumber="510">Fig. 7</figureCitation>
, 
<figureCitation box="[825,841,223,249]" captionStart="FIGURE 9" captionStartId="9.[152,255,1156,1180]" captionTargetBox="[251,1336,181,1132]" captionTargetId="figure-20@9.[251,1336,181,1132]" captionTargetPageId="9" captionText="FIGURE 9. Detail of the contact between cercus and mesepisternal fossae." figureDoi="http://doi.org/10.5281/zenodo.7635192" httpUri="https://zenodo.org/record/7635192/files/figure.png" pageId="10" pageNumber="510">9</figureCitation>
), paraprocts subequal to or longer than cercus, female lacking flap–like structures on anterior margin of prothorax and lacking mesanepisternal groove. Anterior part of female pterothorax with a pair of mesepisternal fossae (
<figureCitation box="[775,844,295,321]" captionStart="FIGURE 7" captionStartId="8.[152,255,1102,1126]" captionTargetBox="[176,1418,181,1078]" captionTargetId="figure-15@8.[151,1436,181,1078]" captionTargetPageId="8" captionText="FIGURE 7. Helveciagrion obsoletum—female thorax in dorsal view, showing the mesepisternal fossae where male tubercles fit." figureDoi="http://doi.org/10.5281/zenodo.7635188" httpUri="https://zenodo.org/record/7635188/files/figure.png" pageId="10" pageNumber="510">Fig. 7</figureCitation>
). Penis long and narrow, with inner fold small, S 
<quantity box="[1394,1436,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="510" unit="in" value="3.0">3 in</quantity>
lateral view expanded laterally, rounded, or triangular–shaped and lacking accessory lobes or processes, ectal region of S3 without lateral fold.
</paragraph>
<paragraph blockId="10.[151,1437,150,1077]" pageId="10" pageNumber="510">
<emphasis bold="true" box="[199,396,402,428]" pageId="10" pageNumber="510">Species included</emphasis>
: 
<taxonomicName authority="(Garrison, 1994)" authorityName="Lencioni" authorityYear="2023" baseAuthorityName="Garrison" baseAuthorityYear="1994" box="[411,892,403,429]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="boomsmae" status="comb. nov.">
<emphasis box="[411,699,403,429]" italics="true" pageId="10" pageNumber="510">Helveciagrion boomsmae</emphasis>
(
<bibRefCitation author="Garrison, R. W." box="[713,884,403,429]" pageId="10" pageNumber="510" pagination="211 - 282" refId="ref9443" refString="Garrison, R. W. (1994) Telebasis boomsmae spec, nov., a new damselfly from Belize (Zygoptera: Coenagrionidae). Odonatologica, 23, 211 - 282." type="journal article" year="1994">Garrison, 1994</bibRefCitation>
)
</taxonomicName>
<emphasis bold="true" box="[900,1028,403,429]" pageId="10" pageNumber="510">
<taxonomicNameLabel box="[900,1028,403,429]" pageId="10" pageNumber="510" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
; 
<taxonomicName authority="(Calvert, 1902)" authorityName="Lencioni" authorityYear="2023" baseAuthorityName="Calvert" baseAuthorityYear="1902" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="collopistes" status="comb. nov.">
<emphasis box="[1043,1331,403,429]" italics="true" pageId="10" pageNumber="510">Helveciagrion collopistes</emphasis>
(
<bibRefCitation author="Calvert, P. P." pageId="10" pageNumber="510" pagination="73 - 128" refId="ref9040" refString="Calvert, P. P. (1902) Odonata. In: Godman, F. D. &amp; Salvin, O. (Eds.) Biologia Centrali Americana - Insecta Neuroptera. R. H. Porter and Dulau &amp; Co., London, pp. 73 - 128." type="book chapter" year="1902">Calvert, 1902</bibRefCitation>
)
</taxonomicName>
<emphasis bold="true" box="[227,359,439,465]" pageId="10" pageNumber="510">
<taxonomicNameLabel box="[227,359,439,465]" pageId="10" pageNumber="510" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
; 
<taxonomicName authority="(Bick &amp; Bick, 1995)" authorityName="Lencioni" authorityYear="2023" baseAuthorityName="Bick &amp; Bick" baseAuthorityYear="1995" box="[377,903,438,465]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="garrisoni" status="comb. nov.">
<emphasis box="[377,652,439,465]" italics="true" pageId="10" pageNumber="510">Helveciagrion garrisoni</emphasis>
(
<bibRefCitation author="Bick, G. H. &amp; Bick, J. C." box="[670,894,438,465]" pageId="10" pageNumber="510" pagination="1 - 15" refId="ref8883" refString="Bick, G. H. &amp; Bick, J. C. (1995) A review of the genus Telebasis with descriptions of eight new species (Zygoptera: Coenagrionidae). Odonatologica, 24, 1 - 15." type="journal article" year="1995">Bick &amp; Bick, 1995</bibRefCitation>
)
</taxonomicName>
<emphasis bold="true" box="[913,1045,439,465]" pageId="10" pageNumber="510">
<taxonomicNameLabel box="[913,1045,439,465]" pageId="10" pageNumber="510" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
; 
<taxonomicName authority="(Selys, 1876)" baseAuthorityName="Selys" baseAuthorityYear="1876" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis box="[1063,1349,439,465]" italics="true" pageId="10" pageNumber="510">Helveciagrion obsoletum</emphasis>
(Selys, 1876)
</taxonomicName>
; 
<taxonomicName authority="(Haber, 2020)" authorityName="Lencioni" authorityYear="2023" baseAuthorityName="Haber" baseAuthorityYear="2020" box="[228,655,475,501]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="rojinegra" status="comb. nov.">
<emphasis box="[228,498,475,501]" italics="true" pageId="10" pageNumber="510">Helveciagrion rojinegra</emphasis>
(
<bibRefCitation author="Haber, W. A." box="[509,647,475,501]" pageId="10" pageNumber="510" pagination="129 - 138" refId="ref9571" refString="Haber, W. A. (2020) Telebasis rojinegra (Odonata: Zygoptera: Coenagrionidae) sp. nov. from Costa Rica. Zootaxa, 4577 (1), 129 - 138. https: // doi. org / 10.11646 / zootaxa. 4755.1.5" type="journal article" year="2020">Haber, 2020</bibRefCitation>
)
</taxonomicName>
<emphasis bold="true" box="[658,784,475,501]" pageId="10" pageNumber="510">
<taxonomicNameLabel box="[658,784,475,501]" pageId="10" pageNumber="510" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
; 
<taxonomicName authority="(Calvert, 1909)" baseAuthorityName="Calvert" baseAuthorityYear="1909" box="[796,1267,475,501]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="simulacrum">
<emphasis box="[796,1092,475,501]" italics="true" pageId="10" pageNumber="510">Helveciagrion simulacrum</emphasis>
(
<bibRefCitation author="Calvert, P. P." box="[1104,1259,475,501]" pageId="10" pageNumber="510" pagination="73 - 280" refId="ref9094" refString="Calvert, P. P. (1909) Contributions to a knowledge of the Odonata of the Neotropical region, exclusive of Mexico and Central America. Annals of the Carnegie Museum, 6, 73 - 280. https: // doi. org / 10.5962 / p. 202300" type="journal article" year="1909">Calvert, 1909</bibRefCitation>
)
</taxonomicName>
; 
<taxonomicName authority="Machado, 1980" authorityName="Machado" authorityYear="1980" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="vulcanoae">
<emphasis italics="true" pageId="10" pageNumber="510">Helveciagrion vulcanoae</emphasis>
<bibRefCitation author="Machado, A. B. M." box="[273,450,511,537]" pageId="10" pageNumber="510" pagination="59 - 87" refId="ref9803" refString="Machado, A. B. M. (1980) Helveciagrion n. g., com descricao de uma nova especie do Parque Estadual do Rio Doce, Minas Gerais (Odonata: Coenagrionidae). Lundiana, 1, 59 - 87." type="journal article" year="1980">Machado, 1980</bibRefCitation>
</taxonomicName>
; 
<taxonomicName authority="(Garrison, 2009)" authorityName="Lencioni" authorityYear="2023" baseAuthorityName="Garrison" baseAuthorityYear="2009" box="[462,954,511,537]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="williamsoni" status="comb. nov.">
<emphasis box="[462,760,511,537]" italics="true" pageId="10" pageNumber="510">Helveciagrion williamsoni</emphasis>
(
<bibRefCitation author="Garrison, R. W." box="[775,946,511,537]" pageId="10" pageNumber="510" pagination="1 - 121" refId="ref9526" refString="Garrison, R. W. (2009) A synopsis of the genus Telebasis (Odonata: Coenagrionidae). International Journal of Odonatology, 12 (1), 1 - 121. https: // doi. org / 10.1080 / 13887890.2009.9748331" type="journal article" year="2009">Garrison, 2009</bibRefCitation>
)
</taxonomicName>
<emphasis bold="true" box="[961,1089,511,537]" pageId="10" pageNumber="510">
<taxonomicNameLabel box="[961,1089,511,537]" pageId="10" pageNumber="510" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="510" type="distribution">
<paragraph blockId="10.[151,1437,150,1077]" box="[199,665,546,573]" pageId="10" pageNumber="510">
<emphasis bold="true" box="[199,345,546,572]" pageId="10" pageNumber="510">Distribution</emphasis>
. From Mexico to 
<collectingCountry box="[547,658,547,573]" name="Argentina" pageId="10" pageNumber="510">Argentina</collectingCountry>
.
</paragraph>
<paragraph blockId="10.[151,1437,150,1077]" box="[199,905,582,609]" pageId="10" pageNumber="510">
<emphasis bold="true" box="[199,507,582,609]" pageId="10" pageNumber="510">For species identification:</emphasis>
<bibRefCitation author="Garrison, R. W." box="[514,693,583,609]" pageId="10" pageNumber="510" pagination="1 - 121" refId="ref9526" refString="Garrison, R. W. (2009) A synopsis of the genus Telebasis (Odonata: Coenagrionidae). International Journal of Odonatology, 12 (1), 1 - 121. https: // doi. org / 10.1080 / 13887890.2009.9748331" type="journal article" year="2009">Garrison (2009)</bibRefCitation>
and 
<bibRefCitation author="Haber, W. A." box="[748,901,583,609]" pageId="10" pageNumber="510" pagination="129 - 138" refId="ref9571" refString="Haber, W. A. (2020) Telebasis rojinegra (Odonata: Zygoptera: Coenagrionidae) sp. nov. from Costa Rica. Zootaxa, 4577 (1), 129 - 138. https: // doi. org / 10.11646 / zootaxa. 4755.1.5" type="journal article" year="2020">Haber (2020)</bibRefCitation>
.
</paragraph>
<paragraph blockId="10.[151,1437,150,1077]" pageId="10" pageNumber="510">
In addition to the original description, I examined specimens of the following species: 
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1876" box="[1165,1318,618,645]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis bold="true" box="[1165,1318,618,645]" italics="true" pageId="10" pageNumber="510">H. obsoletum</emphasis>
</taxonomicName>
1♁—(NC 16)— 
<collectingCountry box="[215,285,655,681]" name="Brazil" pageId="10" pageNumber="510">Brazil</collectingCountry>
: Miranda (Salobra)—MS, 
<geoCoordinate box="[592,740,655,683]" degrees="20" direction="south" minutes="12" orientation="latitude" pageId="10" pageNumber="510" precision="15" seconds="10" value="-20.202778">20° 12′ 10′′ S</geoCoordinate>
&amp; 
<geoCoordinate box="[773,931,654,683]" degrees="56" direction="west" minutes="32" orientation="longitude" pageId="10" pageNumber="510" precision="15" seconds="53" value="-56.548054">56° 32′ 53′′ W</geoCoordinate>
, 1–2/1941, Leg. C.I.O. Cruz (exchange—Ex- MNRJ collection (nº 18640)); 
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1876" box="[500,656,690,717]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis bold="true" box="[500,656,690,717]" italics="true" pageId="10" pageNumber="510">H. obsoletum</emphasis>
</taxonomicName>
1♁, 
<specimenCount box="[715,739,691,717]" pageId="10" pageNumber="510" type="female">1♀</specimenCount>
—(NC 3346)— 
<collectingCountry box="[929,1012,691,717]" name="Bolivia" pageId="10" pageNumber="510">Bolivia</collectingCountry>
: 
<collectingRegion box="[1037,1162,691,717]" country="Bolivia" name="Santa Cruz" pageId="10" pageNumber="510">Santa Cruz</collectingRegion>
Dept., San Ramon, Rio San Julian, 
<geoCoordinate box="[286,440,727,755]" degrees="16" direction="south" minutes="38" orientation="latitude" pageId="10" pageNumber="510" precision="15" seconds="43" value="-16.645277">16º 38′ 43″ S</geoCoordinate>
&amp; 
<geoCoordinate box="[480,645,727,755]" degrees="62" direction="west" minutes="30" orientation="longitude" pageId="10" pageNumber="510" precision="15" seconds="32" value="-62.50889">62º 30′ 32″ W</geoCoordinate>
, 
<quantity box="[658,733,726,753]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="10" pageNumber="510" unit="m" value="215.0">215 m</quantity>
, 
<date box="[747,886,727,753]" pageId="10" pageNumber="510" value="2003-08-12">12.viii.2003</date>
, Leg. F.A.A. Lencioni; 
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1876" box="[1159,1315,726,753]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis bold="true" box="[1159,1315,726,753]" italics="true" pageId="10" pageNumber="510">H. obsoletum</emphasis>
</taxonomicName>
1♁—(NC 3367)— 
<collectingCountry box="[244,327,763,789]" name="Bolivia" pageId="10" pageNumber="510">Bolivia</collectingCountry>
: 
<collectingRegion box="[347,469,763,789]" country="Bolivia" name="Santa Cruz" pageId="10" pageNumber="510">Santa Cruz</collectingRegion>
Dept., San Ramon, Rio San Julian, 
<geoCoordinate box="[867,1012,763,791]" degrees="16" direction="south" minutes="38" orientation="latitude" pageId="10" pageNumber="510" precision="15" seconds="43" value="-16.645277">16º 38′ 43″ S</geoCoordinate>
&amp; 
<geoCoordinate box="[1046,1201,763,791]" degrees="62" direction="west" minutes="30" orientation="longitude" pageId="10" pageNumber="510" precision="15" seconds="32" value="-62.50889">62º 30′ 32″ W</geoCoordinate>
, 
<quantity box="[1211,1283,762,789]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="10" pageNumber="510" unit="m" value="215.0">215 m</quantity>
, 
<date box="[1294,1432,763,789]" pageId="10" pageNumber="510" value="2003-08-12">12.viii.2003</date>
, Leg. F.A.A. Lencioni; 
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1876" box="[403,556,798,825]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis bold="true" box="[403,556,798,825]" italics="true" pageId="10" pageNumber="510">H. obsoletum</emphasis>
</taxonomicName>
<specimenCount box="[563,586,799,825]" pageId="10" pageNumber="510" type="female">1♀</specimenCount>
—(NC 3372)— 
<collectingCountry box="[774,856,799,825]" name="Bolivia" pageId="10" pageNumber="510">Bolivia</collectingCountry>
: 
<collectingRegion box="[877,1000,799,825]" country="Bolivia" name="Santa Cruz" pageId="10" pageNumber="510">Santa Cruz</collectingRegion>
Dept., San Ramon, Rio San Julian, 
<geoCoordinate degrees="16" direction="south" minutes="38" orientation="latitude" pageId="10" pageNumber="510" precision="15" seconds="43" value="-16.645277">16º 38′ 43″ S</geoCoordinate>
&amp; 
<geoCoordinate box="[298,462,835,863]" degrees="62" direction="west" minutes="30" orientation="longitude" pageId="10" pageNumber="510" precision="15" seconds="32" value="-62.50889">62º 30′ 32″ W</geoCoordinate>
, 
<quantity box="[475,550,834,861]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="10" pageNumber="510" unit="m" value="215.0">215 m</quantity>
, 
<date box="[564,702,835,861]" pageId="10" pageNumber="510" value="2003-08-12">12.viii.2003</date>
, Leg. F.A.A. Lencioni; 
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1876" box="[975,1131,834,861]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="obsoletum">
<emphasis bold="true" box="[975,1131,834,861]" italics="true" pageId="10" pageNumber="510">H. obsoletum</emphasis>
</taxonomicName>
<specimenCount box="[1140,1164,835,861]" pageId="10" pageNumber="510" type="female">1♀</specimenCount>
—(NC 3698)— 
<collectingCountry box="[1353,1436,835,861]" name="Bolivia" pageId="10" pageNumber="510">Bolivia</collectingCountry>
: 
<collectingRegion box="[169,222,871,897]" country="Bolivia" name="Beni" pageId="10" pageNumber="510">Beni</collectingRegion>
Dept., 
<collectingCountry box="[309,407,871,897]" name="Trinidad and Tobago" pageId="10" pageNumber="510">Trinidad</collectingCountry>
, forest N of town, 
<geoCoordinate box="[631,797,871,899]" degrees="14" direction="south" minutes="45" orientation="latitude" pageId="10" pageNumber="510" precision="1" seconds="1.5" value="-14.750417">14° 45′ 1.5″ S</geoCoordinate>
&amp; 
<geoCoordinate box="[838,1030,871,899]" degrees="64" direction="west" minutes="57" orientation="longitude" pageId="10" pageNumber="510" precision="1" seconds="20.7" value="-64.95575">64° 57′ 20.7″ W</geoCoordinate>
, 
<date box="[1035,1169,871,897]" pageId="10" pageNumber="510" value="2003-08-21">21.viii.2003</date>
, Leg. F.A.A. Lencioni; 
<taxonomicName authorityName="Machado" authorityYear="1980" box="[151,308,906,933]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="vulcanoae">
<emphasis bold="true" box="[151,308,906,933]" italics="true" pageId="10" pageNumber="510">H. vulcanoae</emphasis>
</taxonomicName>
1♁—(NC 26)— 
<collectingCountry box="[504,573,907,933]" name="Brazil" pageId="10" pageNumber="510">Brazil</collectingCountry>
: Lagoa Santa—MG, 
<date box="[832,954,907,933]" pageId="10" pageNumber="510" value="1966-11-02">02.xi.1966</date>
, Leg. N. Santos, J. Machado e J. Martins (exchange—MNRJ—Exc. 303—col. 10); 
<taxonomicName authorityName="Machado" authorityYear="1980" box="[631,787,942,969]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="vulcanoae">
<emphasis bold="true" box="[631,787,942,969]" italics="true" pageId="10" pageNumber="510">H. vulcanoae</emphasis>
</taxonomicName>
1♁—(NC 79)— 
<collectingCountry box="[982,1051,943,969]" name="Brazil" pageId="10" pageNumber="510">Brazil</collectingCountry>
: Marliéria—MG, Dom Helvécio lake, 
<geoCoordinate box="[213,354,979,1007]" degrees="19" direction="south" minutes="47" orientation="latitude" pageId="10" pageNumber="510" precision="15" seconds="5" value="-19.784721">19° 47′ 5′′ S</geoCoordinate>
&amp; 
<geoCoordinate box="[392,557,979,1007]" degrees="42" direction="west" minutes="35" orientation="longitude" pageId="10" pageNumber="510" precision="15" seconds="15" value="-42.587498">42° 35′ 15′′ W</geoCoordinate>
, 
<date box="[569,724,979,1005]" pageId="10" pageNumber="510" value="1980-05-18" valueMax="1980-05-22" valueMin="1980-05-18">18–22.v.1980</date>
, Leg. Angelo Machado (donation— 
<emphasis bold="true" box="[1133,1240,979,1005]" pageId="10" pageNumber="510">
<typeStatus box="[1133,1240,979,1005]" pageId="10" pageNumber="510">paratype</typeStatus>
</emphasis>
); 
<taxonomicName authorityName="Lencioni" authorityYear="2023" baseAuthorityName="Garrison" baseAuthorityYear="2009" box="[1265,1436,978,1005]" class="Insecta" family="Coenagrionidae" genus="Helveciagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="510" phylum="Arthropoda" rank="species" species="williamsoni">
<emphasis bold="true" box="[1265,1436,978,1005]" italics="true" pageId="10" pageNumber="510">H. williamsoni</emphasis>
</taxonomicName>
2♁♁—(NC 8)— 
<collectingCountry box="[336,455,1015,1041]" name="Venezuela" pageId="10" pageNumber="510">Venezuela</collectingCountry>
: 
<collectingRegion box="[473,597,1015,1041]" country="Venezuela" name="Zulia" pageId="10" pageNumber="510">Zulia State</collectingRegion>
, Encontrados, 9° 3′ 40 ‘′ N &amp; 
<geoCoordinate box="[920,1062,1015,1043]" degrees="72" direction="west" minutes="14" orientation="longitude" pageId="10" pageNumber="510" precision="15" seconds="8" value="-72.23555">72° 14′ 8′′ W</geoCoordinate>
, 
<date box="[1071,1191,1014,1041]" pageId="10" pageNumber="510" value="1920-04-25">25.iv.1920</date>
, Leg. J.H. Williamson et al (donation).
</paragraph>
</subSubSection>
</treatment>
</document>