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<document id="0D88C2DDA7462E05914FC342724691BD" ID-DOI="10.5281/zenodo.12586111" ID-ISSN="2064-2474" ID-Zenodo-Dep="12586111" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="felipe" checkinTime="1719595482395" checkinUser="felipe" docAuthor="Papp, L., Merz, B. &amp; Földvári, M." docDate="2006" docId="03A487CEFF8BF507E86EFAF3D3E8FC67" docLanguage="en" docName="ActaZoolAcadSciH.52.2.97-269.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 52 (2), No. 2" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Thaumalea nigronitida L. PAPP 2006, sp. n." docType="treatment" docVersion="3" lastPageNumber="124" masterDocId="FF9DFFB6FF92F51CEA41FF99D277FF96" masterDocTitle="DIPTERA OF THAILAND A summary of the families and genera with references to the species representations" masterLastPageNumber="269" masterPageNumber="97" pageNumber="122" updateTime="1719712550900" updateUser="admin" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="CD1CA26A703B7E044FA33B9F41055A72">DIPTERA OF THAILAND A summary of the families and genera with references to the species representations</mods:title>
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<mods:namePart id="D740174DF4A1A0A06894865A4CA693BD">Papp, L.</mods:namePart>
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<mods:namePart id="045BBBCEEA7AF739DD8D65ABB1CE14A7">Merz, B.</mods:namePart>
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<mods:namePart id="9BC1C9BDCE22B8EC71763DB67051F95E">Földvári, M.</mods:namePart>
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<mods:typeOfResource id="E7F4F3B7FDB395321F99532F103563D2">text</mods:typeOfResource>
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<mods:title id="E576F7FFFF84C31F5113F8ED1526EA1A">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
</mods:titleInfo>
<mods:part id="6EE4B9861B5F254CC0FED12155994381">
<mods:date id="895C6489A9AE86B3C02FD61F12B5CA07">2006</mods:date>
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<mods:number id="5B238A32F61556F428922FD0F409FDAD">2006-07-31</mods:number>
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<mods:number id="E8E8FD756ACD0FBCCA75EFE14E2D9407">52</mods:number>
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<mods:identifier id="ED3A97D08409BCE33B16D458155B0DE4" type="DOI">10.5281/zenodo.12586111</mods:identifier>
<mods:identifier id="1C04FF9953E8D5C8DF63212DDD5A5AA8" type="ISSN">2064-2474</mods:identifier>
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<treatment id="03A487CEFF8BF507E86EFAF3D3E8FC67" LSID="urn:lsid:plazi:treatment:03A487CEFF8BF507E86EFAF3D3E8FC67" httpUri="http://treatment.plazi.org/id/03A487CEFF8BF507E86EFAF3D3E8FC67" lastPageId="27" lastPageNumber="124" pageId="25" pageNumber="122">
<subSubSection id="C3176553FF8BF505E86EFAF3D674FA13" box="[559,1027,1386,1413]" pageId="25" pageNumber="122" type="nomenclature">
<paragraph id="8BB236D8FF8BF505E86EFAF3D674FA13" blockId="25.[559,1027,1386,1413]" box="[559,1027,1386,1413]" pageId="25" pageNumber="122">
<heading id="D0FA81B4FF8BF505E86EFAF3D674FA13" box="[559,1027,1386,1413]" centered="true" fontSize="10" level="2" pageId="25" pageNumber="122" reason="6">
<taxonomicName id="4C0D4D5BFF8BF505E86EFAF3D1C1FA12" ID-CoL="564RT" authority="L. PAPP" authorityName="L. PAPP" authorityYear="2006" box="[559,950,1386,1413]" class="Insecta" family="Thaumaleidae" genus="Thaumalea" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="122" phylum="Arthropoda" rank="species" species="nigronitida" status="sp. nov.">
<emphasis id="B979EACAFF8BF505E86EFAF3D126FA12" bold="true" box="[559,849,1386,1412]" pageId="25" pageNumber="122">Thaumalea nigronitida</emphasis>
L.
<smallCapsWord id="8D54A004FF8BF505E938FAF2D1C1FA12" baselines="1406,1406" box="[889,950,1387,1413]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Papp" pageId="25" pageNumber="122">PAPP</smallCapsWord>
</taxonomicName>
,
<taxonomicNameLabel id="A24A57B1FF8BF505E985FAF2D674FA13" box="[964,1027,1387,1413]" pageId="25" pageNumber="122" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3176553FF8BF507E88FFA17D3E8FC67" lastPageId="27" lastPageNumber="124" pageId="25" pageNumber="122" type="description">
<paragraph id="8BB236D8FF8BF505E88FFA17D112FA3E" blockId="25.[718,869,1422,1448]" box="[718,869,1422,1448]" pageId="25" pageNumber="122">
<heading id="D0FA81B4FF8BF505E88FFA17D112FA3E" box="[718,869,1422,1448]" centered="true" fontSize="11" level="3" pageId="25" pageNumber="122" reason="3">
(
<figureCitation id="13362A5DFF8BF505E897FA17D12BFA3E" box="[726,860,1422,1448]" captionStart="Figs 13–16" captionStartId="26.[317,360,1744,1765]" captionTargetBox="[337,1250,840,1712]" captionTargetId="figure-193@26.[332,1254,835,1716]" captionTargetPageId="26" captionText="Figs 13–16. Thaumalea nigronitida sp. n., holotype female.13 = apex of abdomen, lateral view, 14 = same, ventral view, 15 = genital sclerites, ventral view, 16 = egg. Scales: 0.2 mm for Figs 13–14, 0.1" figureDoi="http://doi.org/10.5281/zenodo.12586119" httpUri="https://zenodo.org/record/12586119/files/figure.png" pageId="25" pageNumber="122">Figs 13–16</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BB236D8FF8BF505EB3FFA48D3B1F9B6" blockId="25.[317,1270,1489,1568]" pageId="25" pageNumber="122">
<materialsCitation id="3B653C85FF8BF505EB3FFA48D3B1F9B6" collectingDate="2004-10-30" collectionCode="HNHM, PAPP, FOLDVARI" collectorName="Doi Inthanon N. P." country="Thailand" location="THAILAND" pageId="25" pageNumber="122" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B6887AFF8BF505EB3FFA48D3A0FA70" box="[382,471,1489,1510]" pageId="25" pageNumber="122" type="holotype">Holotype</typeStatus>
female (
<collectionCode id="ED1CAE1DFF8BF505E86DFA48D00BFA70" box="[556,636,1489,1510]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="25" pageNumber="122">HNHM</collectionCode>
):
<collectingCountry id="F31A7648FF8BF505E8CAFA48D17CFA70" box="[651,779,1489,1510]" name="Thailand" pageId="25" pageNumber="122">THAILAND</collectingCountry>
:
<collectorName id="26F8530EFF8BF505E955FA48D1BBFA70" box="[788,972,1489,1510]" pageId="25" pageNumber="122">Doi Inthanon N. P.</collectorName>
, over a small rocky brook,
<date id="FFB31018FF8BF505EE92FA48D3E4F995" pageId="25" pageNumber="122" value="2004-10-30">
<collectingDate id="EFF7E9F0FF8BF505EE92FA48D3E4F995" pageId="25" pageNumber="122" value="2004-10-30">Oct 30, 2004</collectingDate>
</date>
, No. 9, L.
<smallCapsWord id="8D54A004FF8BF505EBB8FA77D05FF994" baselines="1534,1534" box="[505,552,1518,1539]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Papp" pageId="25" pageNumber="122">
<collectionCode id="ED1CAE1DFF8BF505EBB8FA77D05FF994" box="[505,552,1518,1539]" pageId="25" pageNumber="122">PAPP</collectionCode>
</smallCapsWord>
&amp; M.
<smallCapsWord id="8D54A004FF8BF505E829FA77D0BDF994" baselines="1534,1534" box="[616,714,1518,1539]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Foldvari" pageId="25" pageNumber="122">
<collectionCode id="ED1CAE1DFF8BF505E829FA77D0BDF994" box="[616,714,1518,1539]" pageId="25" pageNumber="122">FÖLDVÁRI</collectionCode>
</smallCapsWord>
(abdomen with genitalia in a microvial with glycerol, left hind leg lost).
</materialsCitation>
</paragraph>
<paragraph id="8BB236D8FF8BF505EB3FF9DDD091F9E0" blockId="25.[317,1270,1604,1827]" pageId="25" pageNumber="122">Measurements in mm: body length 1.98 to 2.00 mm (measured along the downcurved abdomen), wing length 2.47, wing breadth 1.18.</paragraph>
<paragraph id="8BB236D8FF8BF505EB3FF9E7D667F905" blockId="25.[317,1270,1604,1827]" box="[382,1040,1662,1683]" pageId="25" pageNumber="122">Body black, scutum shiny black, coxae black, legs blackish brown.</paragraph>
<paragraph id="8BB236D8FF8BF505EB3FF902D632F95B" blockId="25.[317,1270,1604,1827]" pageId="25" pageNumber="122">
Apical flagellomere with teo
<quantity id="4CF59B3DFF8BF505E8DEF902D14FF926" box="[671,824,1691,1712]" metricMagnitude="-5" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="25" pageNumber="122" unit="mm" value="0.0325" valueMax="0.035" valueMin="0.03">0.03–0.035 mm</quantity>
long apical setae. Five visible flagellomeres, first flagellomere thickened. Second and third palpomeres bulbously thickened.
</paragraph>
<paragraph id="8BB236D8FF8BF506EB3FF94DD3EEFDB6" blockId="25.[317,1270,1604,1827]" lastBlockId="26.[317,1271,436,774]" lastPageId="26" lastPageNumber="123" pageId="25" pageNumber="122">
Wing fumose, c, sc, r1 and r2+3 cells black. Subcostal vein short (less than
<quantity id="4CF59B3DFF8BF505EE12F94DD6ECF97F" box="[1107,1179,1748,1769]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="122" unit="mm" value="0.4">0.4 mm</quantity>
from H). Costal sections (from H): 33: 69: 108, i.e. 0.36: 0.75:
<quantity id="4CF59B3DFF8BF505E97FF968D1E1F890" box="[830,918,1777,1798]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="25" pageNumber="122" unit="mm" value="1.17">1.17 mm</quantity>
. Veins R1 and R4+5 setose dorsally, crossvein M-M concave, M-Cu
<quantity id="4CF59B3DFF8BF505E831F897D0A6F8B5" box="[624,721,1806,1827]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="25" pageNumber="122" unit="mm" value="0.075">0.075 mm</quantity>
long. Anterior medial vein almost straight, posterior medial vein (M3 or M3+4, “CuA1”, see remark below) downcurved, Cu1 S-formed, distal third almost straight. Cu2 runs close to Cu1, archely curved, faint and thin apically, but overruns crossvein Cu-M remarkably. Vein A1 present only as a straight basal vein. A2 continued to wing margin as a colourless vein.
</paragraph>
<paragraph id="8BB236D8FF88F506EB3FFDB1D61CFDAB" blockId="26.[317,1271,436,774]" box="[382,1131,552,573]" pageId="26" pageNumber="123">
Haltere broad lamelliform, sides curled,
<quantity id="4CF59B3DFF88F506E94BFDB1D116FDAB" box="[778,865,552,573]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="26" pageNumber="123" unit="mm" value="0.58">0.58 mm</quantity>
long, black on dorsal side.
</paragraph>
<paragraph id="8BB236D8FF88F506EB3FFDDDD1CAFD05" blockId="26.[317,1271,436,774]" pageId="26" pageNumber="123">
Abdomen rather thick, dark brown. Several hundreds of
<quantity id="4CF59B3DFF88F506E9DDFDDDD650FDCF" box="[924,1063,580,601]" metricMagnitude="-4" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="26" pageNumber="123" unit="mm" value="0.165" valueMax="0.17" valueMin="0.16">0.16–0.17 mm</quantity>
long ripen eggs (
<figureCitation id="13362A5DFF88F506EE8EFDDDD321FDE0" captionStart="Figs 13–16" captionStartId="26.[317,360,1744,1765]" captionTargetBox="[337,1250,840,1712]" captionTargetId="figure-193@26.[332,1254,835,1716]" captionTargetPageId="26" captionText="Figs 13–16. Thaumalea nigronitida sp. n., holotype female.13 = apex of abdomen, lateral view, 14 = same, ventral view, 15 = genital sclerites, ventral view, 16 = egg. Scales: 0.2 mm for Figs 13–14, 0.1" figureDoi="http://doi.org/10.5281/zenodo.12586119" httpUri="https://zenodo.org/record/12586119/files/figure.png" pageId="26" pageNumber="123">Fig. 16</figureCitation>
) were in holotype’s abdomen, some of them are preserved in the microvial with glycerol. Egg bullet-shaped, with longitudinal ridges and a small round micropyle.
</paragraph>
<paragraph id="8BB236D8FF88F506EB3FFD02D101F889" blockId="26.[317,1271,436,774]" lastBlockId="26.[317,1270,1744,1823]" pageId="26" pageNumber="123">
Tergite 8 and sternite 8 completely fused laterally with an emargination at place of fusion (
<figureCitation id="13362A5DFF88F506EE8FFD02D320FD5B" captionStart="Figs 13–16" captionStartId="26.[317,360,1744,1765]" captionTargetBox="[337,1250,840,1712]" captionTargetId="figure-193@26.[332,1254,835,1716]" captionTargetPageId="26" captionText="Figs 13–16. Thaumalea nigronitida sp. n., holotype female.13 = apex of abdomen, lateral view, 14 = same, ventral view, 15 = genital sclerites, ventral view, 16 = egg. Scales: 0.2 mm for Figs 13–14, 0.1" figureDoi="http://doi.org/10.5281/zenodo.12586119" httpUri="https://zenodo.org/record/12586119/files/figure.png" pageId="26" pageNumber="123">Fig. 13</figureCitation>
). Tergite 9 simple, without any projection on ventral caudal edge (that is definitely rounded). Cerci subquadratic in profile with broadly rounded apex. Sternite 8 and 9 partly fused, lateral lobes rather long rounded apically (
<figureCitation id="13362A5DFF88F506E815FD68D0EDFC90" box="[596,666,753,774]" captionStart="Figs 13–16" captionStartId="26.[317,360,1744,1765]" captionTargetBox="[337,1250,840,1712]" captionTargetId="figure-193@26.[332,1254,835,1716]" captionTargetPageId="26" captionText="Figs 13–16. Thaumalea nigronitida sp. n., holotype female.13 = apex of abdomen, lateral view, 14 = same, ventral view, 15 = genital sclerites, ventral view, 16 = egg. Scales: 0.2 mm for Figs 13–14, 0.1" figureDoi="http://doi.org/10.5281/zenodo.12586119" httpUri="https://zenodo.org/record/12586119/files/figure.png" pageId="26" pageNumber="123">Fig. 14</figureCitation>
). Inner genital sclerites (
<figureCitation id="13362A5DFF88F506E9C3FD68D1B2FC90" box="[898,965,753,774]" captionStart="Figs 13–16" captionStartId="26.[317,360,1744,1765]" captionTargetBox="[337,1250,840,1712]" captionTargetId="figure-193@26.[332,1254,835,1716]" captionTargetPageId="26" captionText="Figs 13–16. Thaumalea nigronitida sp. n., holotype female.13 = apex of abdomen, lateral view, 14 = same, ventral view, 15 = genital sclerites, ventral view, 16 = egg. Scales: 0.2 mm for Figs 13–14, 0.1" figureDoi="http://doi.org/10.5281/zenodo.12586119" httpUri="https://zenodo.org/record/12586119/files/figure.png" pageId="26" pageNumber="123">Fig. 15</figureCitation>
) of an intricate form with an anmm for
<figureCitation id="13362A5DFF88F506E94BF893D101F889" box="[778,886,1802,1823]" captionStart="Figs 13–16" captionStartId="26.[317,360,1744,1765]" captionTargetBox="[337,1250,840,1712]" captionTargetId="figure-193@26.[332,1254,835,1716]" captionTargetPageId="26" captionText="Figs 13–16. Thaumalea nigronitida sp. n., holotype female.13 = apex of abdomen, lateral view, 14 = same, ventral view, 15 = genital sclerites, ventral view, 16 = egg. Scales: 0.2 mm for Figs 13–14, 0.1" figureDoi="http://doi.org/10.5281/zenodo.12586119" httpUri="https://zenodo.org/record/12586119/files/figure.png" pageId="26" pageNumber="123">Figs 15–16</figureCitation>
</paragraph>
<caption id="DF726650FF88F506EB7CF949D682F895" ID-DOI="http://doi.org/10.5281/zenodo.12586119" ID-Zenodo-Dep="12586119" httpUri="https://zenodo.org/record/12586119/files/figure.png" pageId="26" pageNumber="123" startId="26.[317,360,1744,1765]" targetBox="[337,1250,840,1712]" targetPageId="26" targetType="figure">
<paragraph id="8BB236D8FF88F506EB7CF949D682F895" blockId="26.[317,1270,1744,1823]" pageId="26" pageNumber="123">
<emphasis id="B979EACAFF88F506EB7CF949D3DEF973" bold="true" box="[317,425,1744,1765]" pageId="26" pageNumber="123">Figs 13–16</emphasis>
.
<taxonomicName id="4C0D4D5BFF88F506EBF2F948D0E7F970" authorityName="Papp &amp; Merz &amp; Földvári" authorityYear="2006" box="[435,656,1745,1766]" class="Insecta" family="Thaumaleidae" genus="Thaumalea" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="123" phylum="Arthropoda" rank="species" species="nigronitida" status="sp. nov.">
<emphasis id="B979EACAFF88F506EBF2F948D0E7F970" box="[435,656,1745,1766]" italics="true" pageId="26" pageNumber="123">Thaumalea nigronitida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A57B1FF88F506E8D5F948D0BFF970" box="[660,712,1745,1766]" pageId="26" pageNumber="123" rank="species">sp. n.</taxonomicNameLabel>
, holotype female. 13 = apex of abdomen, lateral view, 14 = same, ventral view, 15 = genital sclerites, ventral view, 16 = egg. Scales: 0.2 mm for Figs 13–14, 0.1
</paragraph>
</caption>
<paragraph id="8BB236D8FF89F507EB7CFE2DD676FE70" blockId="27.[317,1269,436,486]" pageId="27" pageNumber="124">
terior cylindrical apodeme and a shovel-shaped medial part. Genital sclerites join to the latero-caudal edge of the sternal
<taxonomicName id="4C0D4D5BFF89F507EBBBFE48D03AFE70" authorityName="Yang, Saigusa &amp; Masunaga" authorityYear="2002" box="[506,589,465,486]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="124" phylum="Arthropoda" rank="species" species="complex">complex</taxonomicName>
through a pigmented sclerotized thickening.
</paragraph>
<paragraph id="8BB236D8FF89F507EB3FFD97D0CBFC88" blockId="27.[317,1270,526,1009]" pageId="27" pageNumber="124">
The other Oriental
<taxonomicName id="4C0D4D5BFF89F507E820FD97D16AFDBE" box="[609,797,526,552]" class="Insecta" family="Thaumaleidae" genus="Thaumalea" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="124" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B979EACAFF89F507E820FD97D092FDBE" box="[609,741,526,552]" italics="true" pageId="27" pageNumber="124">Thaumalea</emphasis>
spp.
</taxonomicName>
were described from
<collectingCountry id="F31A7648FF89F507EE61FD97D6F4FDBE" box="[1056,1155,526,552]" name="Pakistan" pageId="27" pageNumber="124">Pakistan</collectingCountry>
(
<bibRefCitation id="EF9C4B29FF89F507EED4FD97D30EFDDD" author="SCHMID, F." pageId="27" pageNumber="124" pagination="1 - 23" refId="ref93024" refString="SCHMID, F. (1958) Quelques Dipteres Nematoceres nouveaux ou interessants (Thaumaleides et Limnobiides). Institut royal des Sciences naturelles de Belgique Bulletin 34 (10): 1 - 23." type="journal article" year="1958">
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1958
</bibRefCitation>
) and from
<taxonomicName id="4C0D4D5BFF89F507EBB6FDA8D179FDDD" authority="(SCHMID 1970)" baseAuthorityName="SCHMID" baseAuthorityYear="1970" box="[503,782,561,587]" class="Insecta" family="Chironomidae" genus="Nilothauma" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="124" phylum="Arthropoda" rank="species" species="india">
N India (
<bibRefCitation id="EF9C4B29FF89F507E81FFDA8D173FDDD" author="SCHMID, F." box="[606,772,561,587]" pageId="27" pageNumber="124" pagination="499 - 510" refId="ref93060" refString="SCHMID, F. (1970) Quelques thaumaleides nouvelles ou peu connues (Diptera, Thaumaleidae). Le Naturaliste Canadien 97: 499 - 510." type="journal article" year="1970">
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1970
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)
</taxonomicName>
. All were described based on male genitalia. One may regard a description of a species based on a female as questionable. However, I think this black-bodied and black-winged small sp. is characteristic enough to represent the first record of the family from SE Asia. The new species is related to
<taxonomicName id="4C0D4D5BFF89F507EBF2FD27D09AFD40" authority="SCHMID" authorityName="SCHMID" box="[435,749,702,728]" class="Insecta" family="Thaumaleidae" genus="Thaumalea" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="124" phylum="Arthropoda" rank="species" species="karakoramica">
<emphasis id="B979EACAFF89F507EBF2FD27D0F2FD4E" box="[435,645,702,728]" italics="true" pageId="27" pageNumber="124">Th. karakoramica</emphasis>
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by its body colour. I have to note that in the HNHM there is a
<taxonomicName id="4C0D4D5BFF89F507E846FD78D0FCFD6D" box="[519,651,737,763]" class="Insecta" family="Thaumaleidae" genus="Thaumalea" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B979EACAFF89F507E846FD78D0FCFD6D" box="[519,651,737,763]" italics="true" pageId="27" pageNumber="124">Thaumalea</emphasis>
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female also from
<collectingCountry id="F31A7648FF89F507E916FD78D1CBFD6D" box="[855,956,737,763]" name="Vietnam" pageId="27" pageNumber="124">Vietnam</collectingCountry>
(but that is a “regular” species with its large reddish body).
</paragraph>
<paragraph id="8BB236D8FF89F507EB3FFCBED3E8FC67" blockId="27.[317,1270,526,1009]" pageId="27" pageNumber="124">
Remark. I question the hitherto prevailing nomenclature of the wing veins in thaumaleid taxonomy. I am convinced that not one but two medial veins present also in thaumaleids. In
<taxonomicName id="4C0D4D5BFF89F507E81CFCF7D18FFC10" authority="KOVALEV" authorityName="KOVALEV" box="[605,1016,878,904]" class="Insecta" family="Thaumaleidae" genus="Mesothaumalea" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="124" phylum="Arthropoda" rank="species" species="fossilis">
<emphasis id="B979EACAFF89F507E81CFCF7D102FC1E" box="[605,885,878,904]" italics="true" pageId="27" pageNumber="124">Mesothaumalea fossilis</emphasis>
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</taxonomicName>
(Upper Jurassic, see
<bibRefCitation id="EF9C4B29FF89F507EB7CFC08D0C1FC3D" author="KRZEMINSKI, W. &amp; EVENHUIS, N. L." box="[317,694,913,939]" pageId="27" pageNumber="124" pagination="535 - 564" refId="ref82866" refString="KRZEMINSKI, W. and EVENHUIS, N. L. (2000) 1.14. Review of the Diptera palaeontological records. Pp. 535 - 564. In: PAPP, L. and DARVAS, B. (eds): Contribution to a Manual of the Palaearctic Diptera, Vol. 1. Science Herald, Budapest, 978 pp." type="book chapter" year="2000">
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&amp;
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2000
</bibRefCitation>
) there are clearly two medial veins. The presence of the characteristic Cu2 (its basal part running close to Cu1) corroborates this opinion.
</paragraph>
</subSubSection>
</treatment>
</document>
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