344 lines
38 KiB
XML
344 lines
38 KiB
XML
<document id="B75895F78505CCCB8D249FAB39DD051F" ID-DOI="10.11646/zootaxa.5514.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="13849728" ID-ZooBank="B20A1F98-0B55-48DF-AD7F-EA4339591E32" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727441401877" checkinUser="plazi" docAuthor="Szadziewski, Ryszard, Dominiak, Patrycja, Rumišek, Mario, Neinhuis, Christoph, Wanke, Stefan & Rupp, Thomas" docDate="2024" docId="03A4879AFFE3DA64FF43FBBAFD7A333E" docLanguage="en" docName="zootaxa.5514.2.5.pdf" docOrigin="Zootaxa 5514 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5514.2.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Schizohelea algeriana Szadziewski, Dominiak, Rumisek et Rupp 2024, sp. nov." docType="treatment" docUuid="ECFD6277-C1D3-4E49-962E-114036E7748F" docUuidSource="ZooBank" docVersion="2" lastPageNumber="179" masterDocId="FF9DFFE2FFEBDA6EFFD4FFD8FFFD3223" masterDocTitle="Predatory midges of the genus Schizohelea Kieffer, 1917 in Europe and North Africa (Diptera: Ceratopogonidae)" masterLastPageNumber="187" masterPageNumber="169" pageNumber="177" updateTime="1727443309841" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6D7C467AD34E4271EE892925404C4A48">Predatory midges of the genus Schizohelea Kieffer, 1917 in Europe and North Africa (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="D5F31F5751194EFBDCE9947D9D2E9542">Szadziewski, Ryszard</mods:namePart>
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<mods:affiliation id="F4E4E01AAB398EC4DE59577F400071DE">Department of Invertebrate Zoology and Parasitology, Faculty of Biology, University of Gdańsk, Wita Stwosza 59, 80 - 308 Gdańsk, Poland</mods:affiliation>
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<mods:namePart id="8BA188E8707D6C98FB4F859A0324E880">Dominiak, Patrycja</mods:namePart>
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<mods:affiliation id="F2BE6966A54A34DFA3B81A94E263C18B">TromsØ University Museum, UiT - The Arctic University of Norway, NO- 9037 TromsØ, Norway</mods:affiliation>
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<mods:namePart id="A5B3A7E7E38FC7D505141126315BB36F">Rumišek, Mario</mods:namePart>
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<mods:affiliation id="B2F1A9E374143A11B59D7DA7518CD182">Division of Zoology, Faculty of Science, HR- 10000 Zagreb, Croatia</mods:affiliation>
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<mods:namePart id="728423DE2355CA7F563CB012E1AFD40B">Neinhuis, Christoph</mods:namePart>
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<mods:affiliation id="E783597B02D4DA7E0C722B419F67EB82">Chair of Botany, Faculty of Biology, Technische Universität Dresden, 01062 Dresden, Germany</mods:affiliation>
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<mods:namePart id="FC2F8F033633115261C26A6D09229C9C">Wanke, Stefan</mods:namePart>
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<mods:affiliation id="F4A343ED9F8AC3445DC5BBCEA118417B">Chair of Botany, Faculty of Biology, Technische Universität Dresden, 01062 Dresden, Germany & Institut für Ökologie, Evolution und Diversität, Goethe-Universität Frankfurt, 60438 Frankfurt am Main, Germany & Abteilung Botanik und molekulare Evolutionsforschung, Senckenberg Gesellschaft für Naturforschung, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:namePart id="90F6D16BED81135B91B6B65A3EE15DC5">Rupp, Thomas</mods:namePart>
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<mods:affiliation id="C311A488D4480599FE80C4144460FE29">Department of Environment and Biodiversity, Paris-Lodron University of Salzburg, Hellbrunnerstr. 34, 5020 Salzburg, Austria</mods:affiliation>
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<mods:title id="77054103D292510BBB6369BA5A22DBB6">Zootaxa</mods:title>
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<mods:date id="6350848ED27AF479C76BD5EC760B6EC4">2024</mods:date>
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<mods:number id="348086F390050D490DD8EBF7006377A6">2024-09-26</mods:number>
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<mods:number id="6850F4CA1E6032A8D2102742EB2C69FD">5514</mods:number>
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<mods:url id="3CFA5130B4D979ECA974EE4D3498D686">http://dx.doi.org/10.11646/zootaxa.5514.2.5</mods:url>
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<treatment id="03A4879AFFE3DA64FF43FBBAFD7A333E" ID-DOI="http://doi.org/10.5281/zenodo.13849814" ID-Zenodo-Dep="13849814" LSID="urn:lsid:zoobank.org:act:ECFD6277-C1D3-4E49-962E-114036E7748F" httpUri="http://treatment.plazi.org/id/03A4879AFFE3DA64FF43FBBAFD7A333E" lastPageId="10" lastPageNumber="179" pageId="8" pageNumber="177">
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<subSubSection id="C3176507FFE3DA66FF43FBBAFBDF365E" box="[151,1058,1122,1149]" pageId="8" pageNumber="177" type="nomenclature">
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<paragraph id="8BB2368CFFE3DA66FF43FBBAFBDF365E" blockId="8.[151,1058,1122,1221]" box="[151,1058,1122,1149]" pageId="8" pageNumber="177">
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<heading id="D0FA81E0FFE3DA66FF43FBBAFBDF365E" bold="true" box="[151,1058,1122,1149]" fontSize="11" level="1" pageId="8" pageNumber="177" reason="1">
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<emphasis id="B979EA9EFFE3DA66FF43FBBAFBDF365E" bold="true" box="[151,1058,1122,1149]" pageId="8" pageNumber="177">
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<taxonomicName id="4C0D4D0FFFE3DA66FF43FBBAFC45365E" authority="Szadziewski, Dominiak, Rumisek et Rupp" authorityName="Szadziewski, Dominiak, Rumisek et Rupp" authorityYear="2024" box="[151,952,1122,1149]" class="Insecta" family="Ceratopogonidae" genus="Schizohelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="177" phylum="Arthropoda" rank="species" species="algeriana" status="sp. nov.">
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<emphasis id="B979EA9EFFE3DA66FF43FBBAFE59365E" bold="true" box="[151,420,1122,1149]" italics="true" pageId="8" pageNumber="177">Schizohelea algeriana</emphasis>
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Szadziewski, Dominiak, Rumišek et Rupp
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24A57E5FFE3DA66FC10FBBBFBDF365E" box="[964,1058,1123,1149]" pageId="8" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3176507FFE3DA66FF43FB5FFC4436E6" pageId="8" pageNumber="177" type="description">
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<paragraph id="8BB2368CFFE3DA66FF43FB5FFEC63682" blockId="8.[151,1058,1122,1221]" box="[151,315,1159,1185]" pageId="8" pageNumber="177">
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(
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<figureCitation id="13362A09FFE3DA66FF4BFB5FFEE53682" box="[159,280,1159,1185]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[203,1383,181,1919]" captionTargetId="figure-29@6.[203,1383,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Schizohelea incerta (Clastrier, 1963) (a–d) and S. algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov. (e–h): a—male maxillary palp; b—male wing; c—female wing; d—male thorax and legs; e—male wing; f—female wing; g—female genitalia; h—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.13849736" httpUri="https://zenodo.org/record/13849736/files/figure.png" pageId="8" pageNumber="177">Figs. 4e–h</figureCitation>
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,
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<figureCitation id="13362A09FFE3DA66FEF7FB5FFECF3682" box="[291,306,1159,1185]" captionStart="FIGURE 6" captionStartId="9.[152,255,1288,1312]" captionTargetBox="[200,1387,194,1251]" captionTargetId="figure-25@9.[187,1399,181,1264]" captionTargetPageId="9" captionText="FIGURE 6. Schizohelea algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov., male genitalia: a—ventral view; b— gonostylus; c—gonocoxal expansions; d—parameres; e—aedeagus." figureDoi="http://doi.org/10.5281/zenodo.13849742" httpUri="https://zenodo.org/record/13849742/files/figure.png" pageId="8" pageNumber="177">6</figureCitation>
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)
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</paragraph>
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<paragraph id="8BB2368CFFE3DA66FF43FB73FC4436E6" blockId="8.[151,1058,1122,1221]" box="[151,953,1195,1221]" pageId="8" pageNumber="177">
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<subSubSection id="C3176507FFE3DA66FF43FB2AFAC737BE" pageId="8" pageNumber="177" type="diagnosis">
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<paragraph id="8BB2368CFFE3DA66FF43FB2AFAC737BE" blockId="8.[151,1437,1266,1761]" pageId="8" pageNumber="177">
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<emphasis id="B979EA9EFFE3DA66FF43FB2AFEF7372F" bold="true" box="[151,266,1266,1292]" pageId="8" pageNumber="177">Diagnosis</emphasis>
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. The new species is unique in having wings with a long second radial cell which in both sexes is 1.7–2.5 times longer than first one. Macrotrichia on wing membrane absent. The male is distinct in having markedly curved gonostylus, simple aedeagus with rounded apicomedian plate armed with strong teeth, stout and curved parameres, each with enlarged bulbous apex, and strong and slightly 3-lobed apex of lateroventral tubercle of gonocoxite. Female with unarmed legs, and arms of sternite 9 broad and blunt, with dark recurved apical margin each.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3176507FFE3DA67FF13FA7EFB7A3447" lastPageId="9" lastPageNumber="178" pageId="8" pageNumber="177" type="description">
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<paragraph id="8BB2368CFFE3DA66FF13FA7EFEAC37E3" blockId="8.[151,1437,1266,1761]" box="[199,337,1446,1472]" pageId="8" pageNumber="177">
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<emphasis id="B979EA9EFFE3DA66FF13FA7EFEAC37E3" bold="true" box="[199,337,1446,1472]" pageId="8" pageNumber="177">Description</emphasis>
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</paragraph>
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<paragraph id="8BB2368CFFE3DA66FF13FA13FE5634C2" blockId="8.[151,1437,1266,1761]" pageId="8" pageNumber="177">
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<emphasis id="B979EA9EFFE3DA66FF13FA13FEF937C6" bold="true" box="[199,260,1483,1509]" pageId="8" pageNumber="177">Male</emphasis>
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. Flagellum barely visible. Third palpal segment
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<quantity id="4CF59B69FFE3DA66FCFAFA12FC5C37C6" box="[814,929,1482,1509]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="177" unit="mm" value="0.075">0.075 mm</quantity>
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long. Wing length
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<quantity id="4CF59B69FFE3DA66FBA9FA12FB1837C6" box="[1149,1253,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="177" unit="mm" value="1.5">1.50 mm</quantity>
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, costal ratio CR 0.63; second radial cell about 2.1 times longer than first one (
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<figureCitation id="13362A09FFE3DA66FCA5FA37FC3A342A" box="[881,967,1519,1545]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[203,1383,181,1919]" captionTargetId="figure-29@6.[203,1383,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Schizohelea incerta (Clastrier, 1963) (a–d) and S. algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov. (e–h): a—male maxillary palp; b—male wing; c—female wing; d—male thorax and legs; e—male wing; f—female wing; g—female genitalia; h—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.13849736" httpUri="https://zenodo.org/record/13849736/files/figure.png" pageId="8" pageNumber="177">Fig. 4e</figureCitation>
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). Wing membrane without macrotrichia. Scutellum with 4 marginal bristles and some short setae. Legs including hind femur slender. Hind basitarsus with subbasal spine and a single row of palisade setae, tibial comb with 9 spines, hind spur short. Tarsal ratio TR(1) 1.5, TR(2) 1.6, TR(3) 2.1. Male genitalia (
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<figureCitation id="13362A09FFE3DA66FDEBF983FD7E3456" box="[575,643,1627,1653]" captionStart="FIGURE 6" captionStartId="9.[152,255,1288,1312]" captionTargetBox="[200,1387,194,1251]" captionTargetId="figure-25@9.[187,1399,181,1264]" captionTargetPageId="9" captionText="FIGURE 6. Schizohelea algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov., male genitalia: a—ventral view; b— gonostylus; c—gonocoxal expansions; d—parameres; e—aedeagus." figureDoi="http://doi.org/10.5281/zenodo.13849742" httpUri="https://zenodo.org/record/13849742/files/figure.png" pageId="8" pageNumber="177">Fig. 6</figureCitation>
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). Gonocoxite with strong lateroventral expansion with slightly 3-lobed apex (
|
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<figureCitation id="13362A09FFE3DA66FF0EF9A7FED634BA" box="[218,299,1663,1689]" captionStart="FIGURE 6" captionStartId="9.[152,255,1288,1312]" captionTargetBox="[200,1387,194,1251]" captionTargetId="figure-25@9.[187,1399,181,1264]" captionTargetPageId="9" captionText="FIGURE 6. Schizohelea algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov., male genitalia: a—ventral view; b— gonostylus; c—gonocoxal expansions; d—parameres; e—aedeagus." figureDoi="http://doi.org/10.5281/zenodo.13849742" httpUri="https://zenodo.org/record/13849742/files/figure.png" pageId="8" pageNumber="177">Fig. 6c</figureCitation>
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). Gonostylus slender, distinctly C-curved, 1.3 times longer than gonocoxite (
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<figureCitation id="13362A09FFE3DA66FBAAF9A7FB2C34BA" box="[1150,1233,1663,1689]" captionStart="FIGURE 6" captionStartId="9.[152,255,1288,1312]" captionTargetBox="[200,1387,194,1251]" captionTargetId="figure-25@9.[187,1399,181,1264]" captionTargetPageId="9" captionText="FIGURE 6. Schizohelea algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov., male genitalia: a—ventral view; b— gonostylus; c—gonocoxal expansions; d—parameres; e—aedeagus." figureDoi="http://doi.org/10.5281/zenodo.13849742" httpUri="https://zenodo.org/record/13849742/files/figure.png" pageId="8" pageNumber="177">Fig. 6b</figureCitation>
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). Parameres stout, curved with enlarged bulbous apex (
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<figureCitation id="13362A09FFE3DA66FDF8F97BFD82349E" box="[556,639,1699,1725]" captionStart="FIGURE 6" captionStartId="9.[152,255,1288,1312]" captionTargetBox="[200,1387,194,1251]" captionTargetId="figure-25@9.[187,1399,181,1264]" captionTargetPageId="9" captionText="FIGURE 6. Schizohelea algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov., male genitalia: a—ventral view; b— gonostylus; c—gonocoxal expansions; d—parameres; e—aedeagus." figureDoi="http://doi.org/10.5281/zenodo.13849742" httpUri="https://zenodo.org/record/13849742/files/figure.png" pageId="8" pageNumber="177">Fig. 6d</figureCitation>
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). Aedeagus arch-shaped with broad, dentate apicomedian plate, median tooth strongest (
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<figureCitation id="13362A09FFE3DA66FE98F91FFE6334C2" box="[332,414,1735,1761]" captionStart="FIGURE 6" captionStartId="9.[152,255,1288,1312]" captionTargetBox="[200,1387,194,1251]" captionTargetId="figure-25@9.[187,1399,181,1264]" captionTargetPageId="9" captionText="FIGURE 6. Schizohelea algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov., male genitalia: a—ventral view; b— gonostylus; c—gonocoxal expansions; d—parameres; e—aedeagus." figureDoi="http://doi.org/10.5281/zenodo.13849742" httpUri="https://zenodo.org/record/13849742/files/figure.png" pageId="8" pageNumber="177">Fig. 6e</figureCitation>
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).
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</paragraph>
|
||
<caption id="DF726604FFE2DA67FF4CFAD0FCB43767" ID-DOI="http://doi.org/10.5281/zenodo.13849742" ID-Zenodo-Dep="13849742" httpUri="https://zenodo.org/record/13849742/files/figure.png" pageId="9" pageNumber="178" startId="9.[152,255,1288,1312]" targetBox="[200,1387,194,1251]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8BB2368CFFE2DA67FF4CFAD0FCB43767" blockId="9.[151,1436,1288,1348]" pageId="9" pageNumber="178">
|
||
<emphasis id="B979EA9EFFE2DA67FF4CFAD0FEE43703" bold="true" box="[152,281,1288,1312]" pageId="9" pageNumber="178">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C0D4D0FFFE2DA67FEF5FAD0FC43373C" authority="Szadziewski, Dominiak, Rumisek et Rupp" authorityName="Szadziewski, Dominiak, Rumisek et Rupp" authorityYear="2024" box="[289,958,1288,1312]" class="Insecta" family="Ceratopogonidae" genus="Schizohelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="algeriana" status="sp. nov.">
|
||
<emphasis id="B979EA9EFFE2DA67FEF5FAD0FDF93703" box="[289,516,1288,1312]" italics="true" pageId="9" pageNumber="178">Schizohelea algeriana</emphasis>
|
||
Szadziewski, Dominiak, Rumišek et Rupp
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A24A57E5FFE2DA67FC1EFAD0FBE1373C" box="[970,1052,1288,1311]" pageId="9" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male genitalia: a—ventral view; b— gonostylus; c—gonocoxal expansions; d—parameres; e—aedeagus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB2368CFFE2DA67FF13FA4EFB7A3447" blockId="9.[151,1437,1430,2033]" pageId="9" pageNumber="178">
|
||
<emphasis id="B979EA9EFFE2DA67FF13FA4EFEE33793" bold="true" box="[199,286,1430,1456]" pageId="9" pageNumber="178">Female</emphasis>
|
||
. Flagellum length
|
||
<quantity id="4CF59B69FFE2DA67FE25FA4EFD6B3792" box="[497,662,1430,1457]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="6.0" metricValueMin="5.1" pageId="9" pageNumber="178" unit="mm" value="0.5549999999999999" valueMax="0.6" valueMin="0.51">0.51–0.60 mm</quantity>
|
||
, antennal ratio AR 0.91–1.14. Third palpal segment
|
||
<quantity id="4CF59B69FFE2DA67FB0AFA4EFA603792" box="[1246,1437,1430,1457]" metricMagnitude="-5" metricUnit="m" metricValue="5.8999999999999995" metricValueMax="6.5" metricValueMin="5.3" pageId="9" pageNumber="178" unit="mm" value="0.059" valueMax="0.065" valueMin="0.053">
|
||
<geoCoordinate id="EE39504BFFE2DA67FB0AFA4EFAE03792" box="[1246,1309,1430,1457]" degrees="0.053" direction="north" orientation="latitude" pageId="9" pageNumber="178" precision="55" value="0.053">0.053</geoCoordinate>
|
||
<geoCoordinate id="EE39504BFFE2DA67FAC9FA4EFA973792" box="[1309,1386,1430,1457]" degrees="0.065" direction="west" orientation="longitude" pageId="9" pageNumber="178" precision="55" value="-0.065">–0.065</geoCoordinate>
|
||
mm
|
||
</quantity>
|
||
long. Mandible armed with 9–12 teeth. Wing length
|
||
<quantity id="4CF59B69FFE2DA67FD3AFA62FC6837F6" box="[750,917,1466,1493]" metricMagnitude="-3" metricUnit="m" metricValue="1.345" metricValueMax="1.45" metricValueMin="1.24" pageId="9" pageNumber="178" unit="mm" value="1.345" valueMax="1.45" valueMin="1.24">1.24–1.45 mm</quantity>
|
||
, costal ratio CR 0.69–0.70; second radial cell 1.7–2.5 times longer than first radial cell (
|
||
<figureCitation id="13362A09FFE2DA67FDAEFA06FD3637DB" box="[634,715,1502,1528]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[203,1383,181,1919]" captionTargetId="figure-29@6.[203,1383,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Schizohelea incerta (Clastrier, 1963) (a–d) and S. algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov. (e–h): a—male maxillary palp; b—male wing; c—female wing; d—male thorax and legs; e—male wing; f—female wing; g—female genitalia; h—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.13849736" httpUri="https://zenodo.org/record/13849736/files/figure.png" pageId="9" pageNumber="178">Fig. 4f</figureCitation>
|
||
). Wing membrane without macrotrichia. Scutellum with 4 long and 4–5 short marginal setae. Tarsal ratio TR(1) 1.6–1.8, TR(2) 1.9–2.0, TR(3) 2.1–2.4. Two subequal spermathecae present, both slightly ovoid with short neck, length
|
||
<quantity id="4CF59B69FFE2DA67FD37F9FEFC583462" box="[739,933,1574,1601]" metricMagnitude="-5" metricUnit="m" metricValue="6.35" metricValueMax="7.199999999999999" metricValueMin="5.5" pageId="9" pageNumber="178" unit="mm" value="0.0635" valueMax="0.072" valueMin="0.055">
|
||
<geoCoordinate id="EE39504BFFE2DA67FD37F9FEFCDC3462" box="[739,801,1574,1601]" degrees="0.055" direction="north" orientation="latitude" pageId="9" pageNumber="178" precision="55" value="0.055">0.055</geoCoordinate>
|
||
<geoCoordinate id="EE39504BFFE2DA67FCF5F9FEFC933462" box="[801,878,1574,1601]" degrees="0.072" direction="west" orientation="longitude" pageId="9" pageNumber="178" precision="55" value="-0.072">–0.072</geoCoordinate>
|
||
mm
|
||
</quantity>
|
||
, rudimentary spermatheca distinct (
|
||
<figureCitation id="13362A09FFE2DA67FAEEF9FEFA723463" box="[1338,1423,1574,1600]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[203,1383,181,1919]" captionTargetId="figure-29@6.[203,1383,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Schizohelea incerta (Clastrier, 1963) (a–d) and S. algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov. (e–h): a—male maxillary palp; b—male wing; c—female wing; d—male thorax and legs; e—male wing; f—female wing; g—female genitalia; h—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.13849736" httpUri="https://zenodo.org/record/13849736/files/figure.png" pageId="9" pageNumber="178">Fig. 4h</figureCitation>
|
||
). Abdominal sternite 9 with blunt singular arms; blunt apex with recurved margin (
|
||
<figureCitation id="13362A09FFE2DA67FBF3F992FB873447" box="[1063,1146,1610,1636]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[203,1383,181,1919]" captionTargetId="figure-29@6.[203,1383,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Schizohelea incerta (Clastrier, 1963) (a–d) and S. algeriana Szadziewski, Dominiak, Rumišek et Rupp, sp. nov. (e–h): a—male maxillary palp; b—male wing; c—female wing; d—male thorax and legs; e—male wing; f—female wing; g—female genitalia; h—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.13849736" httpUri="https://zenodo.org/record/13849736/files/figure.png" pageId="9" pageNumber="178">
|
||
Fig.
|
||
<quantity id="4CF59B69FFE2DA67FB8FF992FB873447" box="[1115,1146,1610,1636]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="9" pageNumber="178" unit="g" value="4.0">4g</quantity>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176507FFE2DA67FF13F9B6FD2F353B" pageId="9" pageNumber="178" type="materials_examined">
|
||
<paragraph id="8BB2368CFFE2DA67FF13F9B6FF0E348F" blockId="9.[151,1437,1430,2033]" pageId="9" pageNumber="178">
|
||
<emphasis id="B979EA9EFFE2DA67FF13F9B6FE9334AB" bold="true" box="[199,366,1646,1672]" pageId="9" pageNumber="178">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B653CD1FFE2DA67FEA8F9B6FF0E348F" collectingDate="1981-04-24" collectionCode="R, CEIG" collectorName="Szadziewski" country="Algeria" location="Ras Isly" pageId="9" pageNumber="178" specimenCount="1" stateProvince="Setif" typeStatus="holotype">
|
||
<typeStatus id="54B6882EFFE2DA67FEA8F9B6FE1934AB" box="[380,484,1646,1672]" pageId="9" pageNumber="178" type="holotype">Holotype</typeStatus>
|
||
male,
|
||
<emphasis id="B979EA9EFFE2DA67FDE4F9B6FD7434AB" bold="true" box="[560,649,1646,1672]" pageId="9" pageNumber="178">
|
||
<collectingCountry id="F31A761CFFE2DA67FDE4F9B6FD7434AB" box="[560,649,1646,1672]" name="Algeria" pageId="9" pageNumber="178">Algeria</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49C9F86EFFE2DA67FD43F9B6FCC534AB" box="[663,824,1646,1672]" country="Algeria" name="Setif" pageId="9" pageNumber="178">Sétif province</collectingRegion>
|
||
,
|
||
<location id="8ED26057FFE2DA67FC97F9B7FC6234AB" LSID="urn:lsid:plazi:treatment:03A4879AFFE3DA64FF43FBBAFD7A333E:8ED26057FFE2DA67FC97F9B7FC6234AB" box="[835,927,1646,1673]" country="Algeria" name="Ras Isly" pageId="9" pageNumber="178" stateProvince="Setif">Ras Isly</location>
|
||
near
|
||
<collectingRegion id="49C9F86EFFE2DA67FC09F9B6FBDA34AB" box="[989,1063,1646,1672]" country="Algeria" name="Bejaia" pageId="9" pageNumber="178">Béjaia</collectingRegion>
|
||
,
|
||
<date id="FFB3104CFFE2DA67FBE7F9B6FB4E34AB" box="[1075,1203,1646,1672]" pageId="9" pageNumber="178" value="1981-04-24">
|
||
<collectingDate id="EFF7E9A4FFE2DA67FBE7F9B6FB4E34AB" box="[1075,1203,1646,1672]" pageId="9" pageNumber="178" value="1981-04-24">24.04.1981</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED1CAE49FFE2DA67FB6BF9B7FB2834AA" box="[1215,1237,1647,1673]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="178">R</collectionCode>
|
||
.
|
||
<collectorName id="26F8535AFFE2DA67FB34F9B6FA9134AB" box="[1248,1388,1646,1672]" pageId="9" pageNumber="178">Szadziewski</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED1CAE49FFE2DA67FF4BF94AFF1B348F" box="[159,230,1682,1708]" pageId="9" pageNumber="178">CEIG</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB2368CFFE2DA67FF13F96EFD2F353B" blockId="9.[151,1437,1430,2033]" pageId="9" pageNumber="178">
|
||
Other specimens:
|
||
<materialsCitation id="3B653CD1FFE2DA67FE45F96EFCCE34F2" box="[401,819,1718,1745]" collectorName="Ras Isly" country="Algeria" location="Setif province" pageId="9" pageNumber="178" specimenCount="1" stateProvince="Setif">
|
||
<emphasis id="B979EA9EFFE2DA67FE45F96EFE1734F3" bold="true" box="[401,490,1718,1744]" pageId="9" pageNumber="178">
|
||
<collectingCountry id="F31A761CFFE2DA67FE45F96EFE1734F3" box="[401,490,1718,1744]" name="Algeria" pageId="9" pageNumber="178">Algeria</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49C9F86EFFE2DA67FE23F96EFD6534F3" box="[503,664,1718,1744]" country="Algeria" name="Setif" pageId="9" pageNumber="178">Sétif province</collectingRegion>
|
||
,
|
||
<collectorName id="26F8535AFFE2DA67FD76F96FFD0034F3" box="[674,765,1718,1745]" pageId="9" pageNumber="178">Ras Isly</collectorName>
|
||
near
|
||
</materialsCitation>
|
||
<materialsCitation id="3B653CD1FFE2DA67FCEEF96EFF3E34D7" collectingDate="1981-05-04" collectionCode="R" collectorName="Szadziewski" country="Algeria" location="Bejaia" pageId="9" pageNumber="178" specimenCount="1" specimenCount-female="1" stateProvince="Bejaia">
|
||
<collectingRegion id="49C9F86EFFE2DA67FCEEF96EFC7934F3" box="[826,900,1718,1744]" country="Algeria" name="Bejaia" pageId="9" pageNumber="178">Béjaia</collectingRegion>
|
||
,
|
||
<taxonomicName id="4C0D4D0FFFE2DA67FC5AF96EFC0434F3" box="[910,1017,1718,1744]" class="Magnoliopsida" family="Apiaceae" kingdom="Plantae" order="Apiales" pageId="9" pageNumber="178" phylum="Tracheophyta" rank="family">Apiaceae</taxonomicName>
|
||
,
|
||
<date id="FFB3104CFFE2DA67FBD0F96EFB8B34F2" box="[1028,1142,1718,1745]" pageId="9" pageNumber="178" value="1981-05-04">
|
||
<collectingDate id="EFF7E9A4FFE2DA67FBD0F96EFB8B34F2" box="[1028,1142,1718,1745]" pageId="9" pageNumber="178" value="1981-05-04">4.05.1981</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D0BFD05FFE2DA67FB55F96EFB1834F3" box="[1153,1253,1718,1744]" count="1" pageId="9" pageNumber="178" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="ED1CAE49FFE2DA67FB24F96FFAFB34F2" box="[1264,1286,1719,1745]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="178">R</collectionCode>
|
||
.
|
||
<collectorName id="26F8535AFFE2DA67FAC4F96EFA6134F3" box="[1296,1436,1718,1744]" pageId="9" pageNumber="178">Szadziewski</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B653CD1FFE2DA67FF1AF902FE0134D6" box="[206,508,1754,1781]" collectorName="Tichi" country="Algeria" location="Bejaia Province" pageId="9" pageNumber="178" specimenCount="1" stateProvince="Bejaia">
|
||
<collectingRegion id="49C9F86EFFE2DA67FF1AF902FE7F34D7" box="[206,386,1754,1780]" country="Algeria" name="Bejaia" pageId="9" pageNumber="178">Béjaia Province</collectingRegion>
|
||
,
|
||
<collectorName id="26F8535AFFE2DA67FE58F902FE3B34D7" box="[396,454,1754,1780]" pageId="9" pageNumber="178">Tichi</collectorName>
|
||
near
|
||
</materialsCitation>
|
||
<materialsCitation id="3B653CD1FFE2DA67FDD6F902FBDD34D7" box="[514,1056,1754,1781]" collectingDate="1981-05-08" collectionCode="R" collectorName="Szadziewski" country="Algeria" location="Bejaia" pageId="9" pageNumber="178" specimenCount="1" specimenCount-female="1" stateProvince="Bejaia">
|
||
<collectingRegion id="49C9F86EFFE2DA67FDD6F902FDB134D7" box="[514,588,1754,1780]" country="Algeria" name="Bejaia" pageId="9" pageNumber="178">Béjaia</collectingRegion>
|
||
,
|
||
<date id="FFB3104CFFE2DA67FD83F902FD3734D6" box="[599,714,1754,1781]" pageId="9" pageNumber="178" value="1981-05-08">
|
||
<collectingDate id="EFF7E9A4FFE2DA67FD83F902FD3734D6" box="[599,714,1754,1781]" pageId="9" pageNumber="178" value="1981-05-08">8.05.1981</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D0BFD05FFE2DA67FD00F902FCCA34D7" box="[724,823,1754,1780]" count="1" pageId="9" pageNumber="178" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="ED1CAE49FFE2DA67FC96F903FCA534D6" box="[834,856,1755,1781]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="178">R</collectionCode>
|
||
.
|
||
<collectorName id="26F8535AFFE2DA67FCB5F902FC1034D7" box="[865,1005,1754,1780]" pageId="9" pageNumber="178">Szadziewski</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B653CD1FFE2DA67FBFEF902FD33353B" collectingDate="1981-04-16" collectionCode="R, CEIG" collectorName="Szadziewski" country="Algeria" location="Ziama Mansouria" pageId="9" pageNumber="178" specimenCount="1" specimenCount-female="1" stateProvince="Jijel">
|
||
<collectingRegion id="49C9F86EFFE2DA67FBFEF902FB3834D7" box="[1066,1221,1754,1780]" country="Algeria" name="Jijel" pageId="9" pageNumber="178">Jijel Province</collectingRegion>
|
||
,
|
||
<location id="8ED26057FFE2DA67FB04F902FA6534D7" LSID="urn:lsid:plazi:treatment:03A4879AFFE3DA64FF43FBBAFD7A333E:8ED26057FFE2DA67FB04F902FA6534D7" box="[1232,1432,1754,1780]" country="Algeria" name="Ziama Mansouria" pageId="9" pageNumber="178" stateProvince="Jijel">Ziama Mansouria</location>
|
||
,
|
||
<date id="FFB3104CFFE2DA67FF43F926FEEA353B" box="[151,279,1790,1816]" pageId="9" pageNumber="178" value="1981-04-16">
|
||
<collectingDate id="EFF7E9A4FFE2DA67FF43F926FEEA353B" box="[151,279,1790,1816]" pageId="9" pageNumber="178" value="1981-04-16">16.04.1981</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D0BFD05FFE2DA67FEF7F926FE7A353B" box="[291,391,1790,1816]" count="1" pageId="9" pageNumber="178" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="ED1CAE49FFE2DA67FE46F927FE55353A" box="[402,424,1791,1817]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="178">R</collectionCode>
|
||
.
|
||
<collectorName id="26F8535AFFE2DA67FE67F926FDC2353B" box="[435,575,1790,1816]" pageId="9" pageNumber="178">Szadziewski</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED1CAE49FFE2DA67FDAAF926FD38353B" box="[638,709,1790,1816]" pageId="9" pageNumber="178">CEIG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176507FFE2DA67FF13F8FAFBB1351F" box="[199,1100,1826,1853]" pageId="9" pageNumber="178" type="etymology">
|
||
<paragraph id="8BB2368CFFE2DA67FF13F8FAFBB1351F" blockId="9.[151,1437,1430,2033]" box="[199,1100,1826,1853]" pageId="9" pageNumber="178">
|
||
<emphasis id="B979EA9EFFE2DA67FF13F8FAFEB5351F" bold="true" box="[199,328,1826,1852]" pageId="9" pageNumber="178">Etymology</emphasis>
|
||
. The specific name refers to
|
||
<collectingCountry id="F31A761CFFE2DA67FD58F8FAFD1E351F" box="[652,739,1826,1852]" name="Algeria" pageId="9" pageNumber="178">Algeria</collectingCountry>
|
||
, the country of the
|
||
<typeStatus id="54B6882EFFE2DA67FC6FF8FBFC16351E" box="[955,1003,1827,1853]" pageId="9" pageNumber="178">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176507FFE2DA67FF13F89EFE3F3543" box="[199,450,1862,1889]" pageId="9" pageNumber="178" type="distribution">
|
||
<paragraph id="8BB2368CFFE2DA67FF13F89EFE3F3543" blockId="9.[151,1437,1430,2033]" box="[199,450,1862,1889]" pageId="9" pageNumber="178">
|
||
<emphasis id="B979EA9EFFE2DA67FF13F89EFEA43543" bold="true" box="[199,345,1862,1888]" pageId="9" pageNumber="178">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F31A761CFFE2DA67FEB2F89EFE433543" box="[358,446,1862,1888]" name="Algeria" pageId="9" pageNumber="178">Algeria</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176507FFE2DA64FF13F8B2FD7A333E" lastPageId="10" lastPageNumber="179" pageId="9" pageNumber="178" type="discussion">
|
||
<paragraph id="8BB2368CFFE2DA64FF13F8B2FCD432F6" blockId="9.[151,1437,1430,2033]" lastBlockId="10.[151,1437,151,285]" lastPageId="10" lastPageNumber="179" pageId="9" pageNumber="178">
|
||
<emphasis id="B979EA9EFFE2DA67FF13F8B2FEB535A7" bold="true" box="[199,328,1898,1924]" pageId="9" pageNumber="178">Comments</emphasis>
|
||
. Similar to
|
||
<taxonomicName id="4C0D4D0FFFE2DA67FE18F8B2FCEA35A7" authority="(Remm, 1967)" baseAuthorityName="Remm" baseAuthorityYear="1967" box="[460,791,1898,1925]" class="Insecta" family="Ceratopogonidae" genus="Schizohelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="lampropeza">
|
||
<emphasis id="B979EA9EFFE2DA67FE18F8B2FE2735A7" box="[460,474,1898,1924]" italics="true" pageId="9" pageNumber="178">S</emphasis>
|
||
.
|
||
<emphasis id="B979EA9EFFE2DA67FE3CF8B2FD9635A7" box="[488,619,1898,1924]" italics="true" pageId="9" pageNumber="178">lampropeza</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9C4B7DFFE2DA67FDAEF8B2FCED35A7" author="Remm, H." box="[634,784,1898,1924]" pageId="9" pageNumber="178" pagination="3 - 37" refId="ref8684" refString="Remm, H. (1967) On the fauna of Ceratopogonidae (Diptera) in the Caucasus [in Russian]. Tartu Riikliku Ulikooli Toimetised, 194, 3 - 37." type="journal article" year="1967">Remm, 1967</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0D4D0FFFE2DA67FCF7F8B2FBD735A7" authority="(Remm, 1980)" baseAuthorityName="Remm" baseAuthorityYear="1980" box="[803,1066,1898,1925]" class="Insecta" family="Ceratopogonidae" genus="Schizohelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="pekae">
|
||
<emphasis id="B979EA9EFFE2DA67FCF7F8B2FCCC35A7" box="[803,817,1898,1924]" italics="true" pageId="9" pageNumber="178">S</emphasis>
|
||
.
|
||
<emphasis id="B979EA9EFFE2DA67FCEBF8B2FC7D35A7" box="[831,896,1898,1924]" italics="true" pageId="9" pageNumber="178">pekae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9C4B7DFFE2DA67FC5BF8B2FBDF35A7" author="Remm, H." box="[911,1058,1898,1924]" pageId="9" pageNumber="178" pagination="85 - 128" refId="ref8718" refString="Remm, H. (1980) New species of the family Ceratopogonidae (Diptera) from the Middle Asia [in Russian, English summary]. Tartu Riikliku Ulikooli Toimetised, 516, 85 - 128." type="journal article" year="1980">Remm, 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0D4D0FFFE2DA67FBB4F8B2FA6135A7" authority="(Remm, 1993)" baseAuthorityName="Remm" baseAuthorityYear="1993" box="[1120,1436,1898,1925]" class="Insecta" family="Ceratopogonidae" genus="Schizohelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="spathulata">
|
||
<emphasis id="B979EA9EFFE2DA67FBB4F8B2FB9335A7" box="[1120,1134,1898,1924]" italics="true" pageId="9" pageNumber="178">S</emphasis>
|
||
.
|
||
<emphasis id="B979EA9EFFE2DA67FBA8F8B2FB0F35A7" box="[1148,1266,1898,1924]" italics="true" pageId="9" pageNumber="178">spathulata</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9C4B7DFFE2DA67FAD5F8B2FA6935A7" author="Remm, H." box="[1281,1428,1898,1924]" pageId="9" pageNumber="178" pagination="180 - 200" refId="ref8757" refString="Remm, H. (1993) New species of Ceratopogonidae (Diptera) from the CIS [in Russian, Estonian and English summaries]. Eesti Teaduste Akadeemia Toimetised, Bioloogia, 42, 180 - 200." type="journal article" year="1993">Remm, 1993</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from eastern Eurasia, the newly described species has simple, arch-shaped aedeagus. It differs from all the above species in the shape of the aedeagal apicomedian plate, which is armed with strong teeth. Additionally, parameres in
|
||
<taxonomicName id="4C0D4D0FFFE2DA67FF63F80EFEB935D3" authorityName="Szadziewski, Dominiak, Rumisek et Rupp" authorityYear="2024" box="[183,324,2006,2033]" class="Insecta" family="Ceratopogonidae" genus="Schizohelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="algeriana">
|
||
<emphasis id="B979EA9EFFE2DA67FF63F80EFF3835D3" box="[183,197,2006,2032]" italics="true" pageId="9" pageNumber="178">S</emphasis>
|
||
.
|
||
<emphasis id="B979EA9EFFE2DA67FF03F80EFEB935D3" box="[215,324,2006,2032]" italics="true" pageId="9" pageNumber="178">algeriana</emphasis>
|
||
</taxonomicName>
|
||
are stout, curved and have enlarged bulbous apices. The males of the above-mentioned species described by Remm have a smooth apicomedian plate of the aedeagus and slender parameres. The new species is also unique in having wings with a long second radial cell.
|
||
</paragraph>
|
||
<paragraph id="8BB2368CFFE1DA64FF13FF07FD7A333E" blockId="10.[151,1437,151,285]" pageId="10" pageNumber="179">
|
||
We are not entirely sure if the three examined females were correctly associated with the
|
||
<typeStatus id="54B6882EFFE1DA64FB5AFF07FB0D32DA" box="[1166,1264,223,249]" pageId="10" pageNumber="179" type="holotype">holotype</typeStatus>
|
||
male, therefore we are not including them in the type series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |