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<document id="35E02B29E50CCE2A6907CEFC43FE9DB1" ID-DOI="10.11646/zootaxa.5043.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5531958" ID-ZooBank="51F1E65D-E5CF-4D2F-93DE-DC64507F8603" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632752098661" checkinUser="plazi" docAuthor="Johnston, Nikolas P., Wallman, James F., Szpila, Krzysztof &amp; Pape, Thomas" docDate="2021" docId="03A1CC39AC20FFE25EC5FA911AD4FC4D" docLanguage="en" docName="zootaxa.5043.1.1.pdf" docOrigin="Zootaxa 5043 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5043.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Protomiltogramma nigrisensa Johnston &amp; Wallman &amp; Szpila &amp; Pape 2021, sp. n." docType="treatment" docVersion="3" lastPageNumber="22" masterDocId="FF98B441AC33FFF75E52FFAB182EFF84" masterDocTitle="Integrative taxonomy reveals remarkable diversity in Australian Protomiltogramma (Diptera: Sarcophagidae)" masterLastPageNumber="104" masterPageNumber="1" pageNumber="20" updateTime="1699415761683" updateUser="plazi">
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<mods:title id="353DEFAB67D08592904257DC6CD02D04">Integrative taxonomy reveals remarkable diversity in Australian Protomiltogramma (Diptera: Sarcophagidae)</mods:title>
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<mods:namePart id="3945CD09DE467547021BC10C51C8F61C">Johnston, Nikolas P.</mods:namePart>
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<mods:namePart id="737FFCA61984FC61470748597C2DCE6F">Wallman, James F.</mods:namePart>
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<mods:namePart id="0CBC01D4C2E26295499DC72FBF6D632C">Szpila, Krzysztof</mods:namePart>
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<mods:namePart id="6E39EC306ADD1E0AC8B287BD1009D299">Pape, Thomas</mods:namePart>
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<mods:nameIdentifier id="C1FB8FA034A6D6A6BE7F137BBE960A75" type="email">tpape@snm.ku.dk</mods:nameIdentifier>
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<paragraph id="8BB77D2FAC20FFE45EC5FA911A65FAD1" blockId="19.[151,587,1338,1401]" box="[151,587,1338,1365]" pageId="19" pageNumber="20">
<heading id="D0FFCA43AC20FFE45EC5FA911A65FAD1" bold="true" box="[151,587,1338,1365]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="4">
<emphasis id="B97CA13DAC20FFE45EC5FA911A65FAD1" bold="true" box="[151,587,1338,1365]" pageId="19" pageNumber="20">
<taxonomicName id="4C0806ACAC20FFE45EC5FA911A2CFAD1" authority="Johnston &amp; Wallman &amp; Szpila &amp; Pape, 2021" authorityName="Johnston &amp; Wallman &amp; Szpila &amp; Pape" authorityYear="2021" box="[151,514,1338,1365]" class="Insecta" family="Sarcophagidae" genus="Protomiltogramma" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrisensa" status="sp. nov.">
<emphasis id="B97CA13DAC20FFE45EC5FA911A2CFAD1" bold="true" box="[151,514,1338,1365]" italics="true" pageId="19" pageNumber="20">Protomiltogramma nigrisensa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F1C46AC20FFE45C5BFA901A65FAD1" box="[521,587,1339,1365]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3122EA4AC20FFE45EC5FAF4196CFAFD" box="[151,322,1374,1401]" pageId="19" pageNumber="20" type="description">
<paragraph id="8BB77D2FAC20FFE45EC5FAF4196CFAFD" blockId="19.[151,587,1338,1401]" box="[151,322,1374,1401]" pageId="19" pageNumber="20">
(
<figureCitation id="133361AAAC20FFE45ECDFAF418C9FAFD" box="[159,231,1375,1401]" captionStart="FIGURE 8" captionStartId="20.[151,250,1944,1968]" captionTargetBox="[179,1407,181,1920]" captionTargetId="figure-19@20.[179,1407,181,1920]" captionTargetPageId="20" captionText="FIGURE 8. Protomiltogramma nigrisensa sp. n., paratype, male, Watheroo National Park, NT, 12.xii.2015, N.P. Johnston, J.F. Wallman and K. Szpila (ANIC). A. Habitus, lateral view. B. Head, lateral view. C. Habitus, dorsal view. D. Head, dorsal view. E. Abdomen, dorsal view. F. Head anterior view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531984" httpUri="https://zenodo.org/record/5531984/files/figure.png" pageId="19" pageNumber="20">Figs 8</figureCitation>
,
<figureCitation id="133361AAAC20FFE45EA6FAF51914FAFD" box="[244,314,1374,1401]" captionStart="FIGURE 15" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[193,1405,182,1919]" captionTargetId="figure-17@31.[181,1406,181,1919]" captionTargetPageId="31" captionText="FIGURE 15. Protomiltogramma spp., male terminalia. AD. Protomiltogramma kapnos sp. n. EH. Protomiltogramma laticeps Malloch. IL. Protomiltogramma nigrisensa sp. n. MP. Protomiltogramma nigriseta (Baranov). Abbreviations: c = cercus, s = surstylus and p = acrophallus. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5532012" httpUri="https://zenodo.org/record/5532012/files/figure.png" pageId="19" pageNumber="20">15IL</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3122EA4AC20FFE45EC5FA0D19ABF98D" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BB77D2FAC20FFE45EC5FA0D19ABF98D" blockId="19.[151,1437,1446,2013]" pageId="19" pageNumber="20">
<emphasis id="B97CA13DAC20FFE45EC5FA0D193CFA44" bold="true" box="[151,274,1446,1472]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Distinguished from other Australian
<taxonomicName id="4C0806ACAC20FFE45CE4FA0C1BA2FA45" box="[694,908,1447,1473]" class="Insecta" family="Sarcophagidae" genus="Protomiltogramma" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B97CA13DAC20FFE45CE4FA0C1BA2FA45" box="[694,908,1447,1473]" italics="true" pageId="19" pageNumber="20">Protomiltogramma</emphasis>
</taxonomicName>
through the combination of: dark brown/black ground colour of 1st flagellomere (particularly laterally) and pale yellow/white microtomentum on fronto-orbital and parafacial plates.
</paragraph>
</subSubSection>
<subSubSection id="C3122EA4AC20FFE45E95F9B81A3FF91D" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BB77D2FAC20FFE45E95F9B81A03F9D5" blockId="19.[151,1437,1446,2013]" pageId="19" pageNumber="20">
<emphasis id="B97CA13DAC20FFE45E95F9B81959F9A8" bold="true" box="[199,375,1554,1581]" pageId="19" pageNumber="20">Type material.</emphasis>
<materialsCitation id="3B607772AC20FFE45F2DF9B81A03F9D5" collectingDate="2015-12-12" collectionCode="ANIC" collectorName="Mount Lindesay National Park, WA &amp; N. P. Johnston &amp; J. F. Wallman &amp; K. Szpila" location="Mitchell River" municipality="Mount Lindesay National Park" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54B3C38DAC20FFE45F2DF9B819C9F9A9" box="[383,487,1555,1581]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0EB6A6AC20FFE45FA2F9B81A26F9A9" box="[496,520,1555,1581]" pageId="19" pageNumber="20" type="male"></specimenCount>
:
<location id="8ED72BF4AC20FFE45C44F9B81A93F9A9" LSID="urn:lsid:plazi:treatment:03A1CC39AC20FFE25EC5FA911AD4FC4D:8ED72BF4AC20FFE45C44F9B81A93F9A9" box="[534,701,1555,1581]" municipality="Mount Lindesay National Park" name="Mitchell River" pageId="19" pageNumber="20">Mitchell River</location>
,
<collectorName id="26FD18F9AC20FFE45C98F9B81C4CF9A9" box="[714,1122,1555,1581]" pageId="19" pageNumber="20">
<collectingMunicipality id="6BD3E755AC20FFE45C98F9B81C02F9A9" box="[714,1068,1555,1581]" pageId="19" pageNumber="20">Mount Lindesay National Park</collectingMunicipality>
, WA
</collectorName>
<date id="FFB65BEFAC20FFE45A39F9B91CC3F9A9" box="[1131,1261,1554,1581]" pageId="19" pageNumber="20" value="2015-12-12">
<collectingDate id="EFF2A207AC20FFE45A39F9B91CC3F9A9" box="[1131,1261,1554,1581]" pageId="19" pageNumber="20" value="2015-12-12">12.xii.2015</collectingDate>
</date>
,
<collectorName id="26FD18F9AC20FFE45AA8F9B81DB6F9A9" box="[1274,1432,1555,1581]" pageId="19" pageNumber="20">N.P. Johnston</collectorName>
,
<collectorName id="26FD18F9AC20FFE45EC5F99C1906F9D5" box="[151,296,1591,1617]" pageId="19" pageNumber="20">J.F. Wallman</collectorName>
and
<collectorName id="26FD18F9AC20FFE45F0DF99C19E9F9D5" box="[351,455,1591,1617]" pageId="19" pageNumber="20">K. Szpila</collectorName>
(
<collectionCode id="ED19E5EAAC20FFE45F84F99C1A0FF9D5" box="[470,545,1591,1617]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="19" pageNumber="20">ANIC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB77D2FAC20FFE45E95F9F01A46F9F1" blockId="19.[151,1437,1446,2013]" box="[199,616,1627,1653]" pageId="19" pageNumber="20">
<materialsCitation id="3B607772AC20FFE45E95F9F01A4BF9F1" box="[199,613,1627,1653]" collectionCode="ANIC" pageId="19" pageNumber="20" specimenCount="7" specimenCount-male="7" typeStatus="paratype">
<typeStatus id="54B3C38DAC20FFE45E95F9F01916F9F1" box="[199,312,1627,1653]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D0EB6A6AC20FFE45F11F9F01943F9F1" box="[323,365,1627,1653]" pageId="19" pageNumber="20" type="male">7 ♂</specimenCount>
(
<collectionCode id="ED19E5EAAC20FFE45F29F9F019EDF9F1" box="[379,451,1627,1653]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="19" pageNumber="20">ANIC</collectionCode>
) (see
<tableCitation id="C68A4894AC20FFE45C57F9F01A75F9F1" box="[517,603,1627,1653]" captionStart="TABLE 2" captionStartId="34.[151,239,465,491]" captionText="TABLE 2. List of Protomiltogramma material examined. Holotypes and paratypes are shaded in grey. All specimens are from Australia unless otherwise indicated. Detailed specimen label data is presented. Specimens are in alphabetical order of species, then in alphabetical order of depository, then in ascending order of collection date. Label data are given verbatim, except for Australian state names, which have been given standardised acronyms throughout, and location coordinates, which have been placed inside parentheses. Label data include locality (with type of habitat, flower visited or similar information in brackets), GPS coordinates of location (when available), date of collection and collector(s). Specimen depository is indicated by the following acronyms under the code header: AM = Australian Museum, Sydney; ANIC = Australian National Insect Collection, Canberra; NHMD = Natural History Museum of Denmark, Copenhagen; NMHUK = Natural History Museum, London, UK; NMNH = Smithsonian National Museum of Natural History, Washington, D.C., USA; QM = Queensland Museum, Brisbane. Specimens collected during this revision were also given a unique code, which is not a depository: FLC = University of Wollongong “Fly Library Code”. “*” indicates that the sex was not identified." pageId="19" pageNumber="20">Table 2</tableCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB77D2FAC20FFE45E95F9D51A3FF91D" blockId="19.[151,1437,1446,2013]" box="[199,529,1662,1689]" pageId="19" pageNumber="20">
<emphasis id="B97CA13DAC20FFE45E95F9D519AFF91C" bold="true" box="[199,385,1662,1688]" pageId="19" pageNumber="20">Other material.</emphasis>
See
<tableCitation id="C68A4894AC20FFE45FE5F9D41A23F91D" box="[439,525,1663,1689]" captionStart="TABLE 2" captionStartId="34.[151,239,465,491]" captionText="TABLE 2. List of Protomiltogramma material examined. Holotypes and paratypes are shaded in grey. All specimens are from Australia unless otherwise indicated. Detailed specimen label data is presented. Specimens are in alphabetical order of species, then in alphabetical order of depository, then in ascending order of collection date. Label data are given verbatim, except for Australian state names, which have been given standardised acronyms throughout, and location coordinates, which have been placed inside parentheses. Label data include locality (with type of habitat, flower visited or similar information in brackets), GPS coordinates of location (when available), date of collection and collector(s). Specimen depository is indicated by the following acronyms under the code header: AM = Australian Museum, Sydney; ANIC = Australian National Insect Collection, Canberra; NHMD = Natural History Museum of Denmark, Copenhagen; NMHUK = Natural History Museum, London, UK; NMNH = Smithsonian National Museum of Natural History, Washington, D.C., USA; QM = Queensland Museum, Brisbane. Specimens collected during this revision were also given a unique code, which is not a depository: FLC = University of Wollongong “Fly Library Code”. “*” indicates that the sex was not identified." pageId="19" pageNumber="20">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3122EA4AC20FFE25E95F90919A2FD75" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="description">
<paragraph id="8BB77D2FAC20FFE45E95F9091A9CF939" blockId="19.[151,1437,1446,2013]" box="[199,690,1698,1725]" pageId="19" pageNumber="20">
<emphasis id="B97CA13DAC20FFE45E95F9091977F938" bold="true" box="[199,345,1698,1724]" pageId="19" pageNumber="20">Description.</emphasis>
Body length:
<quantity id="4CF0D0CAAC20FFE45FAAF9091A7BF939" box="[504,597,1698,1725]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="19" pageNumber="20" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">58 mm</quantity>
(n
<emphasis id="B97CA13DAC20FFE45C28F9081AA3F938" box="[634,653,1699,1724]" italics="true" pageId="19" pageNumber="20">=</emphasis>
4).
</paragraph>
<paragraph id="8BB77D2FAC20FFE45E95F96C1D69F859" blockId="19.[151,1437,1446,2013]" pageId="19" pageNumber="20">
<emphasis id="B97CA13DAC20FFE45E95F96C1926F965" box="[199,264,1735,1761]" italics="true" pageId="19" pageNumber="20">Male.</emphasis>
Head (
<figureCitation id="133361AAAC20FFE45F0AF96C19C3F965" box="[344,493,1735,1761]" captionStart="FIGURE 8" captionStartId="20.[151,250,1944,1968]" captionTargetBox="[179,1407,181,1920]" captionTargetId="figure-19@20.[179,1407,181,1920]" captionTargetPageId="20" captionText="FIGURE 8. Protomiltogramma nigrisensa sp. n., paratype, male, Watheroo National Park, NT, 12.xii.2015, N.P. Johnston, J.F. Wallman and K. Szpila (ANIC). A. Habitus, lateral view. B. Head, lateral view. C. Habitus, dorsal view. D. Head, dorsal view. E. Abdomen, dorsal view. F. Head anterior view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531984" httpUri="https://zenodo.org/record/5531984/files/figure.png" pageId="19" pageNumber="20">Fig. 8B, D, F</figureCitation>
). Frontal vitta orange-brown, 0.18 of head width at height of anterior ocellus; frontoorbital plate with gold microtomentum; 2 proclinate FO setae, a patch of unordered reclinate FO setae, slightly weaker than proclinate FO setae; ocellar setae weaker than reclinate FO setae; 1012 frontal setae; 3rd aristomere black, thickened on basal 0.75, fine and hairlike on distal 0.25, equal in length to 1st flagellomere; 1st flagellomere black and 2.5x of pedicel length; pedicel setose, with a single strong seta and multiple weak setulae; scape without setae; parafacial plate with white microtomentum, as wide as 0.75 of distance between antennal insertion and eye margin; gena and postgena grey with black setae; genal groove and facial ridge slightly darker than parafacial plate; vibrissa located midway between tip of 1st flagellomere and lower facial margin; 12 supra-vibrissal setae.
</paragraph>
<caption id="DF772DA7AC27FFE35EC5F8331B61F873" ID-DOI="http://doi.org/10.5281/zenodo.5531984" ID-Zenodo-Dep="5531984" httpUri="https://zenodo.org/record/5531984/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1944,1968]" targetBox="[179,1407,181,1920]" targetPageId="20">
<paragraph id="8BB77D2FAC27FFE35EC5F8331B61F873" blockId="20.[151,1437,1944,2040]" pageId="20" pageNumber="21">
<emphasis id="B97CA13DAC27FFE35EC5F833193DF834" bold="true" box="[151,275,1944,1968]" pageId="20" pageNumber="21">FIGURE 8.</emphasis>
<taxonomicName id="4C0806ACAC27FFE35F4BF8331A64F834" authority="Johnston &amp; Wallman &amp; Szpila &amp; Pape, 2021" authorityName="Johnston &amp; Wallman &amp; Szpila &amp; Pape" authorityYear="2021" box="[281,586,1944,1968]" class="Insecta" family="Sarcophagidae" genus="Protomiltogramma" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nigrisensa" status="sp. nov.">
<emphasis id="B97CA13DAC27FFE35F4BF8331A64F834" box="[281,586,1944,1968]" italics="true" pageId="20" pageNumber="21">Protomiltogramma nigrisensa</emphasis>
</taxonomicName>
<emphasis id="B97CA13DAC27FFE35C02F8331AA7F834" bold="true" box="[592,649,1944,1968]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A24F1C46AC27FFE35C02F8331AA7F834" box="[592,649,1944,1968]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratype, male, Watheroo National Park, NT, 12.xii.2015, N.P. Johnston, J.F. Wallman and K. Szpila (ANIC).
<emphasis id="B97CA13DAC27FFE35FB7F81719D3F850" bold="true" box="[485,509,1980,2004]" pageId="20" pageNumber="21">A.</emphasis>
Habitus, lateral view.
<emphasis id="B97CA13DAC27FFE35CB0F8171AD7F850" bold="true" box="[738,761,1980,2004]" pageId="20" pageNumber="21">B.</emphasis>
Head, lateral view.
<emphasis id="B97CA13DAC27FFE35D96F8171BF3F850" bold="true" box="[964,989,1980,2004]" pageId="20" pageNumber="21">C.</emphasis>
Habitus, dorsal view.
<emphasis id="B97CA13DAC27FFE35A92F8171CF7F850" bold="true" box="[1216,1241,1980,2004]" pageId="20" pageNumber="21">D.</emphasis>
Head, dorsal view.
<emphasis id="B97CA13DAC27FFE35EC5F84B1880F87C" bold="true" box="[151,174,2016,2040]" pageId="20" pageNumber="21">E.</emphasis>
Abdomen, dorsal view.
<emphasis id="B97CA13DAC27FFE35FF6F84B1999F87C" bold="true" box="[420,439,2016,2040]" pageId="20" pageNumber="21">F.</emphasis>
Head anterior view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BB77D2FAC26FFE25E95FF3C1BA9FE29" blockId="21.[151,1437,151,971]" pageId="21" pageNumber="22">
Thorax (
<figureCitation id="133361AAAC26FFE25F75FF3C198FFF35" box="[295,417,151,177]" captionStart="FIGURE 8" captionStartId="20.[151,250,1944,1968]" captionTargetBox="[179,1407,181,1920]" captionTargetId="figure-19@20.[179,1407,181,1920]" captionTargetPageId="20" captionText="FIGURE 8. Protomiltogramma nigrisensa sp. n., paratype, male, Watheroo National Park, NT, 12.xii.2015, N.P. Johnston, J.F. Wallman and K. Szpila (ANIC). A. Habitus, lateral view. B. Head, lateral view. C. Habitus, dorsal view. D. Head, dorsal view. E. Abdomen, dorsal view. F. Head anterior view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531984" httpUri="https://zenodo.org/record/5531984/files/figure.png" pageId="21" pageNumber="22">Fig. 8A, C</figureCitation>
). Dorsum dark grey with silver microtomentum; a major median, two major lateral and two minor lateral stripes (mediad to major lateral stripes and approximately half their width), major lateral stripes ending just before anterior margin of scutellum, minor lateral stripes ending just posterior to suture; scutellum with less pronounced microtomentum and some darkening on lateral margins; notopleuron with two strong setae; proepisternum bare; katepisternal setae 1+23 (with posterior katepisternal setae as an unordered patch), numerous weaker setae covering entire katepisternum.
<emphasis id="B97CA13DAC26FFE25FA7FEE01A01FEE0" box="[501,559,331,356]" italics="true" pageId="21" pageNumber="22">Wing</emphasis>
: some specimens with weak fumosity on basal 0.25; dorsal surface of vein R
<subScript id="178C7F6AAC26FFE25BC6FEF61DB2FEEF" attach="left" box="[1428,1436,349,363]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
bare, R
<subScript id="178C7F6AAC26FFE25EBAFE2A192CFE0B" attach="left" box="[232,258,385,399]" fontSize="6" pageId="21" pageNumber="22">4+5</subScript>
with a patch of setae basally.
<emphasis id="B97CA13DAC26FFE25C1CFEC41AADFE0C" box="[590,643,367,392]" italics="true" pageId="21" pageNumber="22">Legs</emphasis>
: black; fore-tarsal claws slightly shorter than tarsomere 5; pulvilli 0.75 of tarsal claw length; mid tibia with 1 AD, 1 PD, 2 P and 1 V setae.
</paragraph>
<paragraph id="8BB77D2FAC26FFE25E95FE1C1AB3FD49" blockId="21.[151,1437,151,971]" pageId="21" pageNumber="22">
Abdomen (
<figureCitation id="133361AAAC26FFE25F17FE1C19CEFE55" box="[325,480,439,465]" captionStart="FIGURE 8" captionStartId="20.[151,250,1944,1968]" captionTargetBox="[179,1407,181,1920]" captionTargetId="figure-19@20.[179,1407,181,1920]" captionTargetPageId="20" captionText="FIGURE 8. Protomiltogramma nigrisensa sp. n., paratype, male, Watheroo National Park, NT, 12.xii.2015, N.P. Johnston, J.F. Wallman and K. Szpila (ANIC). A. Habitus, lateral view. B. Head, lateral view. C. Habitus, dorsal view. D. Head, dorsal view. E. Abdomen, dorsal view. F. Head anterior view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531984" httpUri="https://zenodo.org/record/5531984/files/figure.png" pageId="21" pageNumber="22">Fig. 8A, C, E</figureCitation>
). Black with silver microtomentum on anterior margins of T35 (occupying approximately 0.25 of each tergite); median stripe absent; microtomentum variable depending on viewing angle, when viewed dorso-laterally distinct median spots are visible on T3 and T4; setae on entire posterior margin of T3, T4 and T5; backward and up-curved setae on T5 absent.
<emphasis id="B97CA13DAC26FFE25CA7FD881B41FDB9" box="[757,879,547,573]" italics="true" pageId="21" pageNumber="22">Terminalia</emphasis>
(
<figureCitation id="133361AAAC26FFE25D2DFD881BD3FDB9" box="[895,1021,546,573]" captionStart="FIGURE 15" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[193,1405,182,1919]" captionTargetId="figure-17@31.[181,1406,181,1919]" captionTargetPageId="31" captionText="FIGURE 15. Protomiltogramma spp., male terminalia. AD. Protomiltogramma kapnos sp. n. EH. Protomiltogramma laticeps Malloch. IL. Protomiltogramma nigrisensa sp. n. MP. Protomiltogramma nigriseta (Baranov). Abbreviations: c = cercus, s = surstylus and p = acrophallus. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5532012" httpUri="https://zenodo.org/record/5532012/files/figure.png" pageId="21" pageNumber="22">Fig. 15IL</figureCitation>
): cercus (lateral view) broad at base, narrowing slightly distally, distal 0.25 weakly hook-shaped, dorsal surface with only few dispersed setae; cerci (posterior view) elongated triangle-shaped, fused along basal 0.6 and separate on distal 0.4; surstylus (lateral view) 0.75 of cercus length, tapered to a single point; surstyli (posterior view) curved inwards towards cerci; acrophallus thin and needle-like, equal in length to cercus.
</paragraph>
<paragraph id="8BB77D2FAC26FFE25E95FD7C19A2FD75" blockId="21.[151,1437,151,971]" box="[199,396,727,753]" pageId="21" pageNumber="22">
<emphasis id="B97CA13DAC26FFE25E95FD7C1935FD75" box="[199,283,727,753]" italics="true" pageId="21" pageNumber="22">Female</emphasis>
. As male.
</paragraph>
</subSubSection>
<subSubSection id="C3122EA4AC26FFE25E95FD501986FCD9" pageId="21" pageNumber="22" type="etymology">
<paragraph id="8BB77D2FAC26FFE25E95FD501986FCD9" blockId="21.[151,1437,151,971]" pageId="21" pageNumber="22">
<emphasis id="B97CA13DAC26FFE25E95FD501960FC91" bold="true" box="[199,334,763,789]" pageId="21" pageNumber="22">Etymology.</emphasis>
The species epithet “
<taxonomicName id="4C0806ACAC26FFE25C69FD501A9EFC90" authorityName="Johnston &amp; Wallman &amp; Szpila &amp; Pape" authorityYear="2021" box="[571,688,763,788]" class="Insecta" family="Sarcophagidae" genus="Protomiltogramma" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="nigrisensa">
<emphasis id="B97CA13DAC26FFE25C69FD501A9EFC90" box="[571,688,763,788]" italics="true" pageId="21" pageNumber="22">nigrisensa</emphasis>
</taxonomicName>
” [from the Latin
<emphasis id="B97CA13DAC26FFE25D21FD501B87FC90" box="[883,937,763,788]" italics="true" pageId="21" pageNumber="22">nigri</emphasis>
meaning black and
<emphasis id="B97CA13DAC26FFE25ACAFD501D25FC90" box="[1176,1291,763,788]" italics="true" pageId="21" pageNumber="22">sensorium</emphasis>
meaning an organ of sensation], which should be treated as a noun in apposition, refers to the characteristic dark brown/black antennae of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3122EA4AC26FFE25E95FCCD1C4AFC21" pageId="21" pageNumber="22" type="biology_ecology">
<paragraph id="8BB77D2FAC26FFE25E95FCCD1C4AFC21" blockId="21.[151,1437,151,971]" pageId="21" pageNumber="22">
<emphasis id="B97CA13DAC26FFE25E95FCCD1909FC04" bold="true" box="[199,295,870,896]" pageId="21" pageNumber="22">Biology.</emphasis>
Females were observed close to the nests of unknown species of
<taxonomicName id="4C0806ACAC26FFE25A5EFCCC1C4EFC05" authorityName="Fabricius" authorityYear="1775" box="[1036,1120,871,897]" class="Insecta" family="Crabronidae" genus="Bembix" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B97CA13DAC26FFE25A5EFCCC1C4EFC05" box="[1036,1120,871,897]" italics="true" pageId="21" pageNumber="22">Bembix</emphasis>
</taxonomicName>
(
<taxonomicName id="4C0806ACAC26FFE25A3DFCCC1D20FC05" box="[1135,1294,871,897]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<taxonomicName id="4C0806ACAC26FFE25B4BFCCC1DBFFC06" authorityName="Latreille" authorityYear="1802" box="[1305,1425,871,898]" class="Insecta" family="Sphecidae" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">Sphecidae</taxonomicName>
), whereas males were collected while perching on sticks and rocks on loose sandy trails.
</paragraph>
</subSubSection>
<subSubSection id="C3122EA4AC26FFE25E95FC051AD4FC4D" box="[199,762,942,971]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8BB77D2FAC26FFE25E95FC051AD4FC4D" blockId="21.[151,1437,151,971]" box="[199,762,942,971]" pageId="21" pageNumber="22">
<emphasis id="B97CA13DAC26FFE25E95FC05194FFC4C" bold="true" box="[199,353,942,968]" pageId="21" pageNumber="22">Distribution.</emphasis>
Australasia—
<collectingCountry id="F31F3DBFAC26FFE25C52FC041A49FC4D" box="[512,615,943,969]" name="Australia" pageId="21" pageNumber="22">Australia</collectingCountry>
(QLD, WA).
</paragraph>
</subSubSection>
</treatment>
</document>