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<document id="EC140AF879297F75F5F67CF427AF2CC9" ID-CLB-Dataset="7175" ID-DOI="10.5252/geodiversitas2021v43a9" ID-GBIF-Dataset="04426a8e-c179-4af0-8e23-d72908e8b4cc" ID-ISSN="1638-9395" ID-Zenodo-Dep="4720776" ID-ZooBank="urn:lsid:zoobank.org:pub:11D0D852-39D7-449C-9EB3-C3D804114556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1619445770811" checkinUser="felipe" docAuthor="Georgalis, Georgios L., Čerňanský, Andrej &amp; Klembara, Jozef" docDate="2021" docId="03A1633BFF97FFF531B2F90AF9A33FE1" docLanguage="en" docName="Geodiversitas.43.9.219-293.pdf" docOrigin="Geodiversitas 43 (9)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Saniwa sp." docType="treatment" docVersion="7" lastPageNumber="269" masterDocId="FF981B43FFA3FFC1303EFFB8FA5E380D" masterDocTitle="Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material" masterLastPageNumber="293" masterPageNumber="219" pageNumber="269" updateTime="1698937615995" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="7C5EA7C3BD68A0C47443431E78AA2FC4">
<mods:title id="30717516736312481E2DA527B186D99D">Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material</mods:title>
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<mods:name id="8B0E75756C33B65324FDEC6BB6BBFA33" type="personal">
<mods:role id="CD1A5540968BE3362F77E670A8826692">
<mods:roleTerm id="38BE125B8BE1FDCA438BE4A8FBA8BEBB">Author</mods:roleTerm>
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<mods:namePart id="19026DC49BDF4B233E14965B363C2880">Georgalis, Georgios L.</mods:namePart>
<mods:affiliation id="9043C5C832289E7DACC0322814A15CB2">Palaeontological Institute and Museum, University of Zurich, Karl Schmid-Strasse 4, 8006 Zurich (Switzerland) and Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia) and Department of Earth Sciences, University of Torino, Via Valperga Caluso 35, 10125 Turin (Italy)</mods:affiliation>
<mods:nameIdentifier id="BADEFD516DC970ABE1F319AEC9EDD712" type="email">georgios.georgalis@pim.uzh.ch</mods:nameIdentifier>
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<mods:name id="5284630F92FB869C6DEA1457B3D7D5C3" type="personal">
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<mods:namePart id="F4891C2BDDE8D94AA929ED9008A4D9F9">Čerňanský, Andrej</mods:namePart>
<mods:affiliation id="D42E49D1781A4D1DB0BA386214D42537">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia)</mods:affiliation>
</mods:name>
<mods:name id="9E8C0CD16D856A0AD35E0ABEE774B1CB" type="personal">
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<mods:roleTerm id="E6102F3FAE7689F289A30D3C86B1C9E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBCD09B935D5BDD92008EBA8B1610D05">Klembara, Jozef</mods:namePart>
<mods:affiliation id="CA0E7043E774B4BAC01DC8EC7D93A618">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia)</mods:affiliation>
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<mods:title id="6A1E15B8A90AB9C048A68125196A81FD">Geodiversitas</mods:title>
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<mods:date id="BA7CCD69AC3F8E0A9D27C531278ADC5E">2021</mods:date>
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<mods:number id="A01C1EA4E6EE0E2DD55F050D3753360E">2021-04-22</mods:number>
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<mods:number id="CAE54F37258DF11F656A511EFBBA4962">43</mods:number>
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<treatment id="03A1633BFF97FFF531B2F90AF9A33FE1" ID-DOI="http://doi.org/10.5281/zenodo.4721479" ID-GBIF-Taxon="181601110" ID-Zenodo-Dep="4721479" LSID="urn:lsid:plazi:treatment:03A1633BFF97FFF531B2F90AF9A33FE1" httpUri="http://treatment.plazi.org/id/03A1633BFF97FFF531B2F90AF9A33FE1" lastPageNumber="269" pageId="52" pageNumber="269">
<subSubSection id="C31281A6FF97FFF531B2F90AFBA13EC0" box="[396,511,1714,1741]" pageId="52" pageNumber="269" type="nomenclature">
<paragraph id="8BB7D22DFF97FFF531B2F90AFBA13EC0" blockId="52.[396,511,1714,1741]" box="[396,511,1714,1741]" pageId="52" pageNumber="269">
<heading id="D0FF6541FF97FFF531B2F90AFBA13EC0" box="[396,511,1714,1741]" centered="true" fontSize="11" level="2" pageId="52" pageNumber="269" reason="2">
<taxonomicName id="4C08A9AEFF97FFF531B2F90AFBA13EC0" box="[396,511,1714,1741]" class="Reptilia" family="Varanidae" genus="Saniwa" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="269" phylum="Chordata" rank="species" species="sp.">
<emphasis id="B97C0E3FFF97FFF531B2F90AFB843EC1" box="[396,474,1714,1740]" italics="true" pageId="52" pageNumber="269">Saniwa</emphasis>
sp.
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31281A6FF97FFF531BDF96AF8563EE1" box="[387,520,1746,1773]" pageId="52" pageNumber="269" type="description">
<paragraph id="8BB7D22DFF97FFF531BDF96AF8563EE1" blockId="52.[387,520,1746,1773]" box="[387,520,1746,1773]" pageId="52" pageNumber="269">
(
<figureCitation id="1333CEA8FF97FFF531B2F96AFBA13EE1" box="[396,511,1746,1773]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="64.[132,143,1985,2002]" captionStartId-1="65.[132,143,903,920]" captionStartId-2="66.[132,143,912,929]" captionTargetBox-0="[135,1448,218,1869]" captionTargetBox-1="[135,1452,217,862]" captionTargetBox-2="[137,1452,217,869]" captionTargetId-0="figure-15@64.[132,1455,215,1878]" captionTargetId-1="figure-617@65.[132,1455,234,865]" captionTargetId-2="figure-567@66.[132,1455,215,873]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionTargetPageId-2="66" captionText-0="FIG. 59. — Saniwa sp. Photographs of presacral vertebra NHMW 2019/0065/0001 in anterior (A), posterior (B), dorsal (C), ventral (D), and left lateral (E) views. Scale bar: 2 mm." captionText-1="FIG. 60. — Saniwa sp. A-E, photographs of presacral vertebra NHMW 2019/0065/0002 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views;F-J, photographs of presacral vertebra NHMW 2019/0065/0003 in anterior (F), posterior (G), dorsal (H), ventral (I), and left lateral (J) views.Scale bar: 2 mm." captionText-2="FIG. 61. — Anguimorpha indet. A-E, photographs of presacral vertebra NHMW 2019/0046/0005 in anterior (A), posterior (B), dorsal (C), ventral (D), and left lateral (E) views; F-J, photographs of presacral vertebra 2019/0046/0004 in anterior (F), posterior (G), dorsal (H), ventral (I), and right lateral (J) views. Scale bar: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720967" figureDoi-1="http://doi.org/10.5281/zenodo.4720969" figureDoi-2="http://doi.org/10.5281/zenodo.4720971" httpUri-0="https://zenodo.org/record/4720967/files/figure.png" httpUri-1="https://zenodo.org/record/4720969/files/figure.png" httpUri-2="https://zenodo.org/record/4720971/files/figure.png" pageId="52" pageNumber="269">Figs 59-61</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31281A6FF97FFF530BAF8B3F8133F30" pageId="52" pageNumber="269" type="materials_examined">
<paragraph id="8BB7D22DFF97FFF530BAF8B3F8133F30" blockId="52.[132,774,1803,1853]" pageId="52" pageNumber="269">
REFERRED SPECIMENS. —
<materialsCitation id="3B60D870FF97FFF531A8F8B3F8133F30" ID-GBIF-Occurrence="3091185323" pageId="52" pageNumber="269">Three presacral vertebrae (NHMW 2019/0065/0001-NHMW 2019/0065/0003).</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31281A6FF97FFF530BAF8CAFE923EE1" pageId="52" pageNumber="269" type="description">
<paragraph id="8BB7D22DFF97FFF530BAF8CAFB4B3F87" blockId="52.[132,775,1906,2028]" box="[132,277,1906,1932]" pageId="52" pageNumber="269">DESCRIPTION</paragraph>
<paragraph id="8BB7D22DFF97FFF530BAF829FB923FA1" blockId="52.[132,775,1906,2028]" box="[132,460,1937,1964]" pageId="52" pageNumber="269">
<emphasis id="B97C0E3FFF97FFF530BAF829FB923FA1" box="[132,460,1937,1964]" italics="true" pageId="52" pageNumber="269">
Presacral vertebrae (
<figureCitation id="1333CEA8FF97FFF5316DF82AFB9D3FA6" box="[339,451,1937,1963]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="64.[132,143,1985,2002]" captionStartId-1="65.[132,143,903,920]" captionStartId-2="66.[132,143,912,929]" captionTargetBox-0="[135,1448,218,1869]" captionTargetBox-1="[135,1452,217,862]" captionTargetBox-2="[137,1452,217,869]" captionTargetId-0="figure-15@64.[132,1455,215,1878]" captionTargetId-1="figure-617@65.[132,1455,234,865]" captionTargetId-2="figure-567@66.[132,1455,215,873]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionTargetPageId-2="66" captionText-0="FIG. 59. — Saniwa sp. Photographs of presacral vertebra NHMW 2019/0065/0001 in anterior (A), posterior (B), dorsal (C), ventral (D), and left lateral (E) views. Scale bar: 2 mm." captionText-1="FIG. 60. — Saniwa sp. A-E, photographs of presacral vertebra NHMW 2019/0065/0002 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views;F-J, photographs of presacral vertebra NHMW 2019/0065/0003 in anterior (F), posterior (G), dorsal (H), ventral (I), and left lateral (J) views.Scale bar: 2 mm." captionText-2="FIG. 61. — Anguimorpha indet. A-E, photographs of presacral vertebra NHMW 2019/0046/0005 in anterior (A), posterior (B), dorsal (C), ventral (D), and left lateral (E) views; F-J, photographs of presacral vertebra 2019/0046/0004 in anterior (F), posterior (G), dorsal (H), ventral (I), and right lateral (J) views. Scale bar: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720967" figureDoi-1="http://doi.org/10.5281/zenodo.4720969" figureDoi-2="http://doi.org/10.5281/zenodo.4720971" httpUri-0="https://zenodo.org/record/4720967/files/figure.png" httpUri-1="https://zenodo.org/record/4720969/files/figure.png" httpUri-2="https://zenodo.org/record/4720971/files/figure.png" pageId="52" pageNumber="269">Figs 59-61</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BB7D22DFF97FFF530BAF809FE923EE1" blockId="52.[132,775,1906,2028]" lastBlockId="52.[810,1457,1076,1773]" pageId="52" pageNumber="269">
The three vertebrae are large (
<figureCitation id="1333CEA8FF97FFF531E4F809F80D3FC6" box="[474,595,1969,1996]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="64.[132,143,1985,2002]" captionStartId-1="65.[132,143,903,920]" captionStartId-2="66.[132,143,912,929]" captionTargetBox-0="[135,1448,218,1869]" captionTargetBox-1="[135,1452,217,862]" captionTargetBox-2="[137,1452,217,869]" captionTargetId-0="figure-15@64.[132,1455,215,1878]" captionTargetId-1="figure-617@65.[132,1455,234,865]" captionTargetId-2="figure-567@66.[132,1455,215,873]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionTargetPageId-2="66" captionText-0="FIG. 59. — Saniwa sp. Photographs of presacral vertebra NHMW 2019/0065/0001 in anterior (A), posterior (B), dorsal (C), ventral (D), and left lateral (E) views. Scale bar: 2 mm." captionText-1="FIG. 60. — Saniwa sp. A-E, photographs of presacral vertebra NHMW 2019/0065/0002 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views;F-J, photographs of presacral vertebra NHMW 2019/0065/0003 in anterior (F), posterior (G), dorsal (H), ventral (I), and left lateral (J) views.Scale bar: 2 mm." captionText-2="FIG. 61. — Anguimorpha indet. A-E, photographs of presacral vertebra NHMW 2019/0046/0005 in anterior (A), posterior (B), dorsal (C), ventral (D), and left lateral (E) views; F-J, photographs of presacral vertebra 2019/0046/0004 in anterior (F), posterior (G), dorsal (H), ventral (I), and right lateral (J) views. Scale bar: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720967" figureDoi-1="http://doi.org/10.5281/zenodo.4720969" figureDoi-2="http://doi.org/10.5281/zenodo.4720971" httpUri-0="https://zenodo.org/record/4720967/files/figure.png" httpUri-1="https://zenodo.org/record/4720969/files/figure.png" httpUri-2="https://zenodo.org/record/4720971/files/figure.png" pageId="52" pageNumber="269">Figs 59-61</figureCitation>
), with centrum lengths ranging from about
<quantity id="4CF07FC8FF97FFF53191F869F8173FE1" box="[431,585,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" metricValueMax="9.6" metricValueMin="8.8" pageId="52" pageNumber="269" unit="mm" value="9.2" valueMax="9.6" valueMin="8.8">8.8 to 9.6 mm</quantity>
(seeAppendix 1). The centrum is almost triangular in ventral view and widens anteriorly (though not to that extent as in melanosaurine vertebrae described above). The subcentral ridges are straight in ventral view. The prezygapophyses are either much dorsally tilted (NHMW 2019/0065/0003) or only slightly so (in the other
<specimenCount id="9D0E19A4FF97FFF53352FB6CFE503CE2" box="[876,1038,1236,1263]" pageId="52" pageNumber="269" type="generic">two specimens</specimenCount>
). The prezygapophyseal articular facets are massive and broad in dorsal view. The postzygapophyseal articular facets are also massive. The neural spine develops in height mostly in the posterior half of the neural arch. The neural arch is vaulted in posterior view. There are slight signs of “pseudozygosphene” and “pseudozygantrum” (sensu
<bibRefCitation id="EF99AFDCFF97FFF53312FA2BF9BC3DA0" author="HOFFSTETTER R." box="[812,994,1427,1453]" pageId="52" pageNumber="269" pagination="1051 - 1064" refId="ref37022" refString="HOFFSTETTER R. 1969. - Presence de Varanidae (Reptilia, Sauria) dans le Miocene de Catalogne. Considerations sur l'histoire de la famille. Bulletin du Museum national d'Histoire naturelle, 2 eme serie, 40: 1051 - 1064. https: // www. biodiversitylibrary. org / page / 55344240" type="journal article" year="1969">Hoffstetter 1969</bibRefCitation>
). The cotyle and the condyle are strongly depressed. The centrum appear more convex in lateral view is NHMW 2019/0065/0001 and NHMW 2019/0065/0002, while it is more straight in NHMW 2019/0065/0003. In all specimens though, the dorsal level of the cotyle can be clearly visible in ventral view of the specimen. Precondylar constriction can be observed (even slightly though) in NHMW 2019/0065/0001, as the respective portion of the centrum is eroded in the other
<specimenCount id="9D0E19A4FF97FFF5341CF92BFE9A3EA1" box="[1058,1220,1682,1709]" pageId="52" pageNumber="269" type="generic">two specimens</specimenCount>
. Anocotylar foramina are present and are most prominent in the largest vertebra NHMW 2019/0065/0001 (
<figureCitation id="1333CEA8FF97FFF53461F96AFE9F3EE1" box="[1119,1217,1746,1773]" captionStart="FIG" captionStartId="64.[132,143,1985,2002]" captionTargetBox="[135,1448,218,1869]" captionTargetId="figure-15@64.[132,1455,215,1878]" captionTargetPageId="64" captionText="FIG. 59. — Saniwa sp. Photographs of presacral vertebra NHMW 2019/0065/0001 in anterior (A), posterior (B), dorsal (C), ventral (D), and left lateral (E) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720967" httpUri="https://zenodo.org/record/4720967/files/figure.png" pageId="52" pageNumber="269">Fig. 59A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31281A6FF97FFF53313F8AAF9A33FE1" pageId="52" pageNumber="269" type="discussion">
<paragraph id="8BB7D22DFF97FFF53313F8AAF9D13F27" blockId="52.[813,1456,1810,2028]" box="[813,911,1810,1836]" pageId="52" pageNumber="269">REMARKS</paragraph>
<paragraph id="8BB7D22DFF97FFF53313F889F9A33FE1" blockId="52.[813,1456,1810,2028]" pageId="52" pageNumber="269">
These three vertebrae can be referred to
<taxonomicName id="4C08A9AEFF97FFF534E5F88AFF773F41" authorityName="Leidy" authorityYear="1870" box="[1243,1321,1842,1868]" class="Reptilia" family="Varanidae" genus="Saniwa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="269" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF97FFF534E5F88AFF773F41" box="[1243,1321,1842,1868]" italics="true" pageId="52" pageNumber="269">Saniwa</emphasis>
</taxonomicName>
on the basis of their triangular centrum that widens anteriorly and the slight presence of “pseudozygosphene” and “pseudozygantrum” (
<bibRefCitation id="EF99AFDCFF97FFF533BCF829FE7D3FA6" author="GILMORE C. W." box="[898,1059,1937,1963]" pageId="52" pageNumber="269" pagination="1 - 28" refId="ref36312" refString="GILMORE C. W. 1922. - A new description of Saniwa ensidens Leidy, an extinct varanid lizard from Wyoming. Proceedings of the United States National Museum 60: 1 - 28. https: // doi. org / 10.5479 / si. 00963801.60 - 2418.1" type="journal article" year="1922">Gilmore 1922</bibRefCitation>
;
<bibRefCitation id="EF99AFDCFF97FFF5340AF82AFF4F3FA6" author="RAGE J. - C. &amp; AUGE M." box="[1076,1297,1937,1964]" pageId="52" pageNumber="269" pagination="103 - 116" refId="ref38912" refString="RAGE J. - C. &amp; AUGE M. 2003. - Amphibians and squamate reptiles from the lower Eocene of Silveirinha (Portugal). Ciencias da Terra (UNL) 15: 103 - 116. http: // hdl. handle. net / 10362 / 4731" type="journal article" year="2003">Rage &amp; Augé 2003</bibRefCitation>
;
<bibRefCitation id="EF99AFDCFF97FFF5351CF829FFC03FA1" author="AUGE M." box="[1314,1438,1937,1964]" pageId="52" pageNumber="269" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé 2005</bibRefCitation>
). See Discussion below for further information on European material of
<taxonomicName id="4C08A9AEFF97FFF53396F869F9A83FE6" authorityName="Leidy" authorityYear="1870" box="[936,1014,2001,2027]" class="Reptilia" family="Varanidae" genus="Saniwa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="269" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF97FFF53396F869F9A83FE6" box="[936,1014,2001,2027]" italics="true" pageId="52" pageNumber="269">Saniwa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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