309 lines
36 KiB
XML
309 lines
36 KiB
XML
<document id="E81EF255B388958A1A819DC772C4AC3F" ID-DOI="10.5852/ejt.2018.445" ID-GBIF-Dataset="5dfd4ff4-6c8b-47f4-9aa7-0f6b1b2af967" ID-Zenodo-Dep="1489598" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1545431021202" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2018" docId="03A10B47FFBFFFCFFDE7FE81FD12FCD8" docLanguage="en" docName="ejt-445_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 445" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Eviulisoma biquintum Enghoff 2018, sp. nov." docType="treatment" docUuid="0B213B4D-8455-4489-B253-13B7769E1611" docUuidSource="ZooBank" docVersion="8" lastPageNumber="74" masterDocId="FF98733FFFF8FF86FF85FF8AFFF2FFB8" masterDocTitle="A mountain of millipedes VII: The genus Eviulisoma Silvestri, 1910, in the Udzungwa Mountains, Tanzania, and related species from other Eastern Arc Mountains. With notes on Eoseviulisoma Brolemann, 1920, and Suohelisoma Hoffman, 1963 (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="90" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1545034857558" originalUpdateUser="plazi" pageNumber="72" updateTime="1698553157531" updateUser="plazi">
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<mods:titleInfo id="3756AB076E28149AC569611D4CDB41DA">
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<mods:title id="1B53FFB1878D7628F2320E2A028DFC03">A mountain of millipedes VII: The genus Eviulisoma Silvestri, 1910, in the Udzungwa Mountains, Tanzania, and related species from other Eastern Arc Mountains. With notes on Eoseviulisoma Brolemann, 1920, and Suohelisoma Hoffman, 1963 (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:roleTerm id="9470F734A2E1D5BA9F2043343D7B9C92">Author</mods:roleTerm>
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<mods:namePart id="92BDE83B66EC309E1C264C213018D9E2">Enghoff, Henrik</mods:namePart>
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<mods:typeOfResource id="DA27A229CD80DC2A1313DF807F31FC5E">text</mods:typeOfResource>
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<mods:title id="27C0F1E02A1FE3667457C04E4830D00C">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="657B425805903723C8E75442AFEEB52A">
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<mods:date id="6A597C9DF01A0AE49516302BAF194C27">2018</mods:date>
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<mods:number id="0CF5EB60BEE63C60218325BF95DAB55E">2018-06-19</mods:number>
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<mods:number id="7EDECCB717E4A71168E273DD421A0DE2">445</mods:number>
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<mods:classification id="5F2F68D05927053449866055A86EBC40">journal article</mods:classification>
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<mods:identifier id="70242B0AF1DEA2295ECA14A423541CD8" type="DOI">10.5852/ejt.2018.445</mods:identifier>
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<mods:identifier id="E608FEFBA9EA8774040AB6E67D26CE98" type="Zenodo-Dep">1489598</mods:identifier>
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<mods:identifier id="F3F8F7D8A3DE1D525E7B45B49E01D5DE" type="ZooBank">852A3F68-B728-413A-B12E-56F306D56C35</mods:identifier>
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<treatment id="03A10B47FFBFFFCFFDE7FE81FD12FCD8" ID-DOI="http://doi.org/10.5281/zenodo.5681630" ID-GBIF-Taxon="149668663" ID-Zenodo-Dep="5681630" LSID="urn:lsid:zoobank.org:act:0B213B4D-8455-4489-B253-13B7769E1611" httpUri="http://treatment.plazi.org/id/03A10B47FFBFFFCFFDE7FE81FD12FCD8" lastPageId="73" lastPageNumber="74" pageId="71" pageNumber="72">
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<subSubSection id="C312E9DAFFBFFFC1FDE7FE81FB4BFEF1" pageId="71" pageNumber="72" type="nomenclature">
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<paragraph id="8BB7BA51FFBFFFC1FDE7FE81FC20FE9E" blockId="71.[379,1209,267,365]" box="[610,978,267,294]" pageId="71" pageNumber="72">
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<heading id="D0FF0D3DFFBFFFC1FDE7FE81FC20FE9E" box="[610,978,267,294]" centered="true" fontSize="11" level="2" pageId="71" pageNumber="72" reason="2">
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<taxonomicName id="4C08C1D2FFBFFFC1FDE7FE81FC82FE9E" ID-CoL="3DHSK" authority="Enghoff, 2018" authorityName="Enghoff" authorityYear="2018" box="[610,880,267,294]" class="Diplopoda" family="Paradoxosomatidae" genus="Eviulisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="biquintum" status="sp. nov.">
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<emphasis id="B97C6643FFBFFFC1FDE7FE81FC82FE9E" bold="true" box="[610,880,267,294]" italics="true" pageId="71" pageNumber="72">Eviulisoma biquintum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24FDB38FFBFFFC1FCF2FE86FC20FE9E" box="[887,978,268,294]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
<paragraph id="8BB7BA51FFBFFFC1FEFEFEA4FB4BFEF1" blockId="71.[379,1209,267,365]" box="[379,1209,302,329]" pageId="71" pageNumber="72">urn:lsid:zoobank.org:act:0B213B4D-8455-4489-B253-13B7769E1611</paragraph>
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</subSubSection>
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<subSubSection id="C312E9DAFFBFFFC1FD33FED8FC8FFED4" box="[694,893,338,365]" pageId="71" pageNumber="72" type="description">
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<paragraph id="8BB7BA51FFBFFFC1FD33FED8FC8FFED4" blockId="71.[379,1209,267,365]" box="[694,893,338,365]" pageId="71" pageNumber="72">
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<figureCitation id="1333A6D4FFBFFFC1FD33FED8FCE6FED4" box="[694,788,338,364]" captionStart="Fig. 3" captionStartId="11.[189,232,1860,1886]" captionTargetBox="[190,1399,670,1825]" captionTargetId="figure@11.[170,1424,661,1834]" captionTargetPageId="11" captionText="Fig. 3. Eviulisoma spp., examples of colour patterns. A. E. akkariae sp. nov., paratype, showing a banded pattern (pale prozonites, dark metazonites) similar to that shown in Fig. 1. B. E. coxale sp. nov., tentatively referred female from type locality, showing dark lateral spots on a pale background. C. E. biquintum sp. nov., paratype from Mito Mitatu, showing a black body contrasting with white legs. Not to scale. Photographs by F. Vad and A. Illum." httpUri="https://zenodo.org/record/1489604/files/figure.png" pageId="71" pageNumber="72">Figs 3C</figureCitation>
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,
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<figureCitation id="1333A6D4FFBFFFC1FCA4FED8FCB3FED5" box="[801,833,338,365]" captionStart="Fig. 35" captionStartId="72.[189,232,1892,1918]" captionTargetBox="[190,1402,265,1863]" captionTargetId="figure@72.[180,1427,249,1874]" captionTargetPageId="72" captionText="Fig. 35. Eviulisoma biquintum sp. nov., holotype. A–D. Right gonopod. A. Mesal view. B. Ventral view. C. Lateral view. D. Acropodite, subapical view. E. Sternum 5. Abbreviations: cxl = coxal lobe; map = mesal acropodital process; prf = prefemoral part; slm = solenomere; sph = solenophore. Scale bars: A–C, E = 0.1 mm; D = 0.05 mm." httpUri="https://zenodo.org/record/1489668/files/figure.png" pageId="71" pageNumber="72">35</figureCitation>
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,
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<figureCitation id="1333A6D4FFBFFFC1FCC8FED8FC8FFED4" box="[845,893,338,364]" captionStart="Fig. 39" captionStartId="82.[189,232,1860,1886]" captionTargetBox="[233,1355,267,1830]" captionTargetId="figure@82.[220,1368,251,1847]" captionTargetPageId="82" captionText="Fig. 39. Suohelisoma ulugurense Hoffman, 1964, specimen from Uluguru Mts, Lupanga, M. Stoltze and N. Scharff leg. A–C. Right gonopod. A. Mesal view. B. Ventral view. C. Lateral view. D. Body ring 6 showing excavated sternum, ventral view. E. Anterior leg, showing scopulae. Abbreviations: cxl = coxallobe; map/ fet = mesal acropodital process / ʻfemoriteʼ; prf =prefemoral part; slm =solenomere; sph = solenophore. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1489676/files/figure.png" pageId="71" pageNumber="72">39C</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C312E9DAFFBFFFC1FF38FE19FF20FD9A" pageId="71" pageNumber="72" type="diagnosis">
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||
<paragraph id="8BB7BA51FFBFFFC1FF38FE19FEC4FE15" blockId="71.[189,310,403,429]" box="[189,310,403,429]" pageId="71" pageNumber="72">
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||
<heading id="D0FF0D3DFFBFFFC1FF38FE19FEC4FE15" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="71" pageNumber="72" reason="3">
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||
<emphasis id="B97C6643FFBFFFC1FF38FE19FEC4FE15" bold="true" box="[189,310,403,429]" pageId="71" pageNumber="72">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB7BA51FFBFFFC1FF38FE48FA85FE64" blockId="71.[189,1399,450,546]" box="[189,1399,449,476]" pageId="71" pageNumber="72">
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Differs from all other species of
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<taxonomicName id="4C08C1D2FFBFFFC1FDC6FE48FD35FE64" authorityName="Silvestri" authorityYear="1910" box="[579,711,450,476]" class="Diplopoda" family="Paradoxosomatidae" genus="Eviulisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
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<emphasis id="B97C6643FFBFFFC1FDC6FE48FD35FE64" box="[579,711,450,476]" italics="true" pageId="71" pageNumber="72">Eviulisoma</emphasis>
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</taxonomicName>
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by having two knobs on sternum 5, one between coxae 4
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</paragraph>
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<paragraph id="8BB7BA51FFBFFFC1FF38FE6FFF20FD9A" blockId="71.[189,1399,450,546]" pageId="71" pageNumber="72">
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and one between coxae 6. Differs from most species of
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<taxonomicName id="4C08C1D2FFBFFFC1FCC1FE6FFC3AFE47" authorityName="Silvestri" authorityYear="1910" box="[836,968,485,511]" class="Diplopoda" family="Paradoxosomatidae" genus="Eviulisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B97C6643FFBFFFC1FCC1FE6FFC3AFE47" box="[836,968,485,511]" italics="true" pageId="71" pageNumber="72">Eviulisoma</emphasis>
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||
</taxonomicName>
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by lacking an excavation of sternum 6.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C312E9DAFFBFFFC1FF38FDC0FE2DFD0E" pageId="71" pageNumber="72" type="etymology">
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||
<paragraph id="8BB7BA51FFBFFFC1FF38FDC0FEB6FDDC" blockId="71.[189,1399,586,694]" box="[189,324,586,612]" pageId="71" pageNumber="72">
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||
<heading id="D0FF0D3DFFBFFFC1FF38FDC0FEB6FDDC" bold="true" box="[189,324,586,612]" fontSize="11" level="3" pageId="71" pageNumber="72" reason="3">
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||
<emphasis id="B97C6643FFBFFFC1FF38FDC0FEB6FDDC" bold="true" box="[189,324,586,612]" pageId="71" pageNumber="72">Etymology</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BB7BA51FFBFFFC1FF38FDF2FE2DFD0E" blockId="71.[189,1399,586,694]" pageId="71" pageNumber="72">
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||
The name is composed of the Latin
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||
<emphasis id="B97C6643FFBFFFC1FDEEFDF2FD70FD2A" box="[619,642,632,658]" italics="true" pageId="71" pageNumber="72">bi</emphasis>
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||
, meaning ʻtwoʼ, and
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<emphasis id="B97C6643FFBFFFC1FC07FDF3FC10FD2A" box="[898,994,633,658]" italics="true" pageId="71" pageNumber="72">quintum</emphasis>
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, meaning ʻfifthʼ, and refers to the two knobs on sternum 5.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C312E9DAFFBFFFC1FF38FD57FC27FB82" pageId="71" pageNumber="72" type="materials_examined">
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||
<paragraph id="8BB7BA51FFBFFFC1FF38FD57FDDFFD4F" blockId="71.[189,557,733,759]" box="[189,557,733,759]" pageId="71" pageNumber="72">
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||
<emphasis id="B97C6643FFBFFFC1FF38FD57FE7EFD4F" bold="true" box="[189,396,733,759]" pageId="71" pageNumber="72">Material studied</emphasis>
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(total: 3 ♂♂)
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||
</paragraph>
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||
<paragraph id="8BB7BA51FFBFFFC1FF6FFC81FEAEFC9D" blockId="71.[189,1398,779,876]" box="[234,348,779,805]" pageId="71" pageNumber="72">
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||
<emphasis id="B97C6643FFBFFFC1FF6FFC81FEAEFC9D" bold="true" box="[234,348,779,805]" pageId="71" pageNumber="72">Holotype</emphasis>
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</paragraph>
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<paragraph id="8BB7BA51FFBFFFC1FF38FCA5FC03FCD4" blockId="71.[189,1398,779,876]" pageId="71" pageNumber="72">
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<materialsCitation id="3B60B00CFFBFFFC1FF38FCA5FC03FCD4" ID-GBIF-Occurrence="1937470265" collectingDate="1994-07" collectingDateMax="1994-11-31" collectingDateMin="1994-07" collectionCode="ZMUC" collectorName="David Moyer" country="Tanzania" county="Iringa District" latitude="-7.713611" location="Udekwa Village" longLatPrecision="21" longitude="36.420834" municipality="Nyambanike Mts" pageId="71" pageNumber="72" specimenCount="1" stateProvince="Iringa Region">
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<collectingCountry id="F31FFAC1FFBFFFC1FF38FCA5FEA5FCF1" box="[189,343,815,841]" name="Tanzania" pageId="71" pageNumber="72">TANZANIA</collectingCountry>
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:
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<specimenCount id="9D0E71D8FFBFFFC1FEE2FCA5FE8CFCF1" box="[359,382,815,841]" pageId="71" pageNumber="72" type="male">♂</specimenCount>
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,
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||
<collectingRegion id="49CC74B3FFBFFFC1FE17FCA4FDCFFCF0" box="[402,573,814,840]" country="Tanzania" name="Iringa" pageId="71" pageNumber="72">Iringa Region</collectingRegion>
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,
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||
<collectingCounty id="62D6C2DDFFBFFFC1FDCBFCA4FD0EFCF0" box="[590,764,814,840]" pageId="71" pageNumber="72">Iringa District</collectingCounty>
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,
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||
<collectingMunicipality id="6BD3202BFFBFFFC1FC88FCA4FC12FCF1" box="[781,992,814,841]" pageId="71" pageNumber="72">Nyambanike Mts</collectingMunicipality>
|
||
,
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||
<location id="8ED7EC8AFFBFFFC1FC77FCA4FB4AFCF0" LSID="urn:lsid:plazi:treatment:03A10B47FFBFFFCFFDE7FE81FD12FCD8:8ED7EC8AFFBFFFC1FC77FCA4FB4AFCF0" box="[1010,1208,814,840]" country="Tanzania" county="Iringa District" latitude="-7.713611" longLatPrecision="21" longitude="36.420834" municipality="Nyambanike Mts" name="Udekwa Village" pageId="71" pageNumber="72" stateProvince="Iringa Region">Udekwa Village</location>
|
||
,
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||
<location id="8ED7EC8AFFBFFFC1FB4CFCA4FA80FCF1" LSID="urn:lsid:plazi:treatment:03A10B47FFBFFFCFFDE7FE81FD12FCD8:8ED7EC8AFFBFFFC1FB4CFCA4FA80FCF1" box="[1225,1394,814,841]" country="Tanzania" county="Iringa District" latitude="-7.713611" longLatPrecision="21" longitude="36.420834" municipality="Nyambanike Mts" name="Ukami Forest" pageId="71" pageNumber="72" stateProvince="Iringa Region">Ukami Forest</location>
|
||
,
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||
<geoCoordinate id="EE3CDC96FFBFFFC1FF38FCDAFECCFCD4" box="[189,318,848,876]" direction="south" orientation="latitude" pageId="71" pageNumber="72" precision="15" value="-7.713611">7°42′49″ S</geoCoordinate>
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,
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<geoCoordinate id="EE3CDC96FFBFFFC1FECEFCDAFE2FFCD4" box="[331,477,848,876]" direction="east" orientation="longitude" pageId="71" pageNumber="72" precision="15" value="36.420834">36°25′15″ E</geoCoordinate>
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||
,
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||
<collectingDate id="EFF26579FFBFFFC1FE6CFCD8FD6CFCD4" box="[489,670,850,876]" pageId="71" pageNumber="72" value="1994-07" valueMax="1994-11-31" valueMin="1994-07-00">
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||
<date id="FFB69C91FFBFFFC1FE6CFCD8FD6CFCD4" box="[489,670,850,876]" pageId="71" pageNumber="72" value="1994-07" valueMax="1994-11-31" valueMin="1994-07-00">Jul.–Nov. 1994</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26FDDF87FFBFFFC1FD2FFCD8FCB5FCD4" box="[682,839,850,876]" pageId="71" pageNumber="72">David Moyer</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED192294FFBFFFC1FC0CFCD8FC17FCD4" box="[905,997,850,876]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="71" pageNumber="72">ZMUC</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF6FFC19FE95FC15" blockId="71.[189,1399,915,1082]" box="[234,359,915,941]" pageId="71" pageNumber="72">
|
||
<emphasis id="B97C6643FFBFFFC1FF6FFC19FE95FC15" bold="true" box="[234,359,915,941]" pageId="71" pageNumber="72">Paratypes</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38FC3CFC27FB82" blockId="71.[189,1399,915,1082]" pageId="71" pageNumber="72">
|
||
<materialsCitation id="3B60B00CFFBFFFC1FF38FC3CFBA4FC4C" ID-GBIF-Occurrence="1937470254" collectingDate="1979-11-23" collectingDateMax="2013-03-22" collectingDateMin="1979-11-23" collectionCode="VMNH, ZMUC" collectorName="Mwanihana FR & W. A. Rodgers & K. H. Bulstrode" country="Tanzania" county="Iringa District" elevation="914" latitude="-7.837861" location="Sanje River valley near Sanje Falls" longLatPrecision="1" longitude="36.845722" municipality="Uzungwa Mts" pageId="71" pageNumber="72" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region">
|
||
<collectingCountry id="F31FFAC1FFBFFFC1FF38FC3CFEA5FC68" box="[189,343,950,976]" name="Tanzania" pageId="71" pageNumber="72">TANZANIA</collectingCountry>
|
||
:
|
||
<specimenCount id="9D0E71D8FFBFFFC1FEE7FC3CFE62FC69" box="[354,400,950,977]" pageId="71" pageNumber="72" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingCounty id="62D6C2DDFFBFFFC1FE1AFC3CFDBAFC68" box="[415,584,950,976]" pageId="71" pageNumber="72">Iringa District</collectingCounty>
|
||
,
|
||
<collectorName id="26FDDF87FFBFFFC1FDD1FC3CFCFCFC69" box="[596,782,950,977]" pageId="71" pageNumber="72">Mwanihana FR</collectorName>
|
||
,
|
||
<collectingMunicipality id="6BD3202BFFBFFFC1FC9FFC3DFC31FC69" box="[794,963,951,977]" pageId="71" pageNumber="72">Uzungwa Mts</collectingMunicipality>
|
||
,
|
||
<location id="8ED7EC8AFFBFFFC1FC55FC3CFA80FC68" LSID="urn:lsid:plazi:treatment:03A10B47FFBFFFCFFDE7FE81FD12FCD8:8ED7EC8AFFBFFFC1FC55FC3CFA80FC68" box="[976,1394,950,977]" country="Tanzania" county="Iringa District" latitude="-7.837861" longLatPrecision="1" longitude="36.845722" municipality="Uzungwa Mts" name="Sanje River valley near Sanje Falls" pageId="71" pageNumber="72" stateProvince="Morogoro Region">Sanje River valley near Sanje Falls</location>
|
||
,
|
||
<elevation id="00255D62FFBFFFC1FF38FC50FEA0FC4B" box="[189,338,985,1012]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="71" pageNumber="72" unit="ft" value="3000.0">
|
||
<quantity id="4CF017B4FFBFFFC1FF38FC50FEE0FC4B" box="[189,274,985,1012]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="71" pageNumber="72" unit="ft" value="3000.0">3000 ft</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFF26579FFBFFFC1FEE5FC50FDF1FC4C" box="[352,515,986,1012]" pageId="71" pageNumber="72" value="1979-11-23">
|
||
<date id="FFB69C91FFBFFFC1FEE5FC50FDF1FC4C" box="[352,515,986,1012]" pageId="71" pageNumber="72" value="1979-11-23">23 Nov. 1979</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26FDDF87FFBFFFC1FD95FC50FD45FC4B" box="[528,695,985,1012]" pageId="71" pageNumber="72">W.A. Rodgers</collectorName>
|
||
and
|
||
<collectorName id="26FDDF87FFBFFFC1FD77FC50FC57FC4B" box="[754,933,985,1012]" pageId="71" pageNumber="72">K.H. Bulstrode</collectorName>
|
||
leg. (VMNH);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B60B00CFFBFFFC1FBDAFC50FC27FB82" ID-GBIF-Occurrence="1937470273" collectingDate="2013-03-22" collectingDateMax="2013-03-22" collectingDateMin="1979-11-23" collectingMethod="hand collected" collectionCode="VMNH, ZMUC" collectorName="Plot MM & T. Pape & N. Scharff" country="Tanzania" county="Iringa District" elevation="800" latitude="-7.837861" location="Udzungwa Mts National Park" longLatPrecision="1" longitude="36.845722" municipality="Uzungwa Mts" pageId="71" pageNumber="72" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region">
|
||
<specimenCount id="9D0E71D8FFBFFFC1FBDAFC50FB7EFC4C" box="[1119,1164,986,1012]" pageId="71" pageNumber="72" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingRegion id="49CC74B3FFBFFFC1FB1EFC50FA80FC4B" box="[1179,1394,985,1012]" country="Tanzania" name="Morogoro" pageId="71" pageNumber="72">Morogoro Region</collectingRegion>
|
||
,
|
||
<location id="8ED7EC8AFFBFFFC1FF38FC77FDDAFBAF" LSID="urn:lsid:plazi:treatment:03A10B47FFBFFFCFFDE7FE81FD12FCD8:8ED7EC8AFFBFFFC1FF38FC77FDDAFBAF" box="[189,552,1021,1047]" country="Tanzania" county="Iringa District" latitude="-7.837861" longLatPrecision="1" longitude="36.845722" municipality="Uzungwa Mts" name="Udzungwa Mts National Park" pageId="71" pageNumber="72" stateProvince="Morogoro Region">Udzungwa Mts National Park</location>
|
||
,
|
||
<location id="8ED7EC8AFFBFFFC1FDB2FC77FD3EFBAF" LSID="urn:lsid:plazi:treatment:03A10B47FFBFFFCFFDE7FE81FD12FCD8:8ED7EC8AFFBFFFC1FDB2FC77FD3EFBAF" box="[567,716,1021,1047]" country="Tanzania" county="Iringa District" latitude="-7.837861" longLatPrecision="1" longitude="36.845722" municipality="Uzungwa Mts" name="Mito Mitatu" pageId="71" pageNumber="72" stateProvince="Morogoro Region">Mito Mitatu</location>
|
||
,
|
||
<collectorName id="26FDDF87FFBFFFC1FD59FC77FCA0FBAF" box="[732,850,1021,1047]" pageId="71" pageNumber="72">Plot MM</collectorName>
|
||
1,
|
||
<geoCoordinate id="EE3CDC96FFBFFFC1FCEEFC71FBE6FBAF" box="[875,1044,1019,1047]" direction="south" orientation="latitude" pageId="71" pageNumber="72" precision="1" value="-7.837861">07°50′16.3″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3CDC96FFBFFFC1FBA0FC71FB22FBAF" box="[1061,1232,1019,1047]" direction="east" orientation="longitude" pageId="71" pageNumber="72" precision="1" value="36.845722">36°50′44.6″ E</geoCoordinate>
|
||
,
|
||
<elevation id="00255D62FFBFFFC1FB65FC77FA82FBAF" box="[1248,1392,1021,1047]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="71" pageNumber="72" unit="m" value="800.0">
|
||
<quantity id="4CF017B4FFBFFFC1FB65FC77FADCFBAF" box="[1248,1326,1021,1047]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="71" pageNumber="72" unit="m" value="800.0">800 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFF26579FFBFFFC1FF38FBAAFEAFFB82" box="[189,349,1056,1082]" pageId="71" pageNumber="72" value="2013-03-22">
|
||
<date id="FFB69C91FFBFFFC1FF38FBAAFEAFFB82" box="[189,349,1056,1082]" pageId="71" pageNumber="72" value="2013-03-22">22 Mar. 2013</date>
|
||
</collectingDate>
|
||
,
|
||
<collectingMethod id="5249C246FFBFFFC1FEECFBAAFDE4FB82" box="[361,534,1056,1082]" pageId="71" pageNumber="72">hand collected</collectingMethod>
|
||
,
|
||
<collectorName id="26FDDF87FFBFFFC1FDA4FBAAFD8BFB82" box="[545,633,1056,1082]" pageId="71" pageNumber="72">T. Pape</collectorName>
|
||
and
|
||
<collectorName id="26FDDF87FFBFFFC1FD37FBAAFCDEFB82" box="[690,812,1056,1082]" pageId="71" pageNumber="72">N. Scharff</collectorName>
|
||
leg. (ZMUC).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312E9DAFFBFFFCFFF38FBEBFB74FEF1" lastPageId="73" lastPageNumber="74" pageId="71" pageNumber="72" type="description">
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38FBEBFE50FBC4" blockId="71.[189,771,1121,1194]" box="[189,418,1121,1148]" pageId="71" pageNumber="72">
|
||
<emphasis id="B97C6643FFBFFFC1FF38FBEBFEBCFBC3" bold="true" box="[189,334,1121,1147]" pageId="71" pageNumber="72">Description</emphasis>
|
||
(male)
|
||
</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38FB1AFCF1FB12" blockId="71.[189,771,1121,1194]" box="[189,771,1168,1194]" pageId="71" pageNumber="72">
|
||
SIZE. Length
|
||
<quantity id="4CF017B4FFBFFFC1FED3FB1AFE2AFB12" box="[342,472,1168,1194]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="71" pageNumber="72" unit="mm" value="21.5" valueMax="24.0" valueMin="19.0">19–24 mm</quantity>
|
||
, max. width 2.0–
|
||
<quantity id="4CF017B4FFBFFFC1FD27FB1AFD0DFB12" box="[674,767,1168,1194]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="71" pageNumber="72" unit="mm" value="2.4">2.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38FB5BFCB6FAC0" blockId="71.[189,1399,1233,1401]" pageId="71" pageNumber="72">
|
||
COLOUR. Specimen from Mito Mitatu after 4 years in alcohol with entire body deep black, except for amber-coloured sternal regions and contrastingly white legs (
|
||
<figureCitation id="1333A6D4FFBFFFC1FC09FB7EFC1AFAB7" box="[908,1000,1268,1295]" captionStart="Fig. 3" captionStartId="11.[189,232,1860,1886]" captionTargetBox="[190,1399,670,1825]" captionTargetId="figure@11.[170,1424,661,1834]" captionTargetPageId="11" captionText="Fig. 3. Eviulisoma spp., examples of colour patterns. A. E. akkariae sp. nov., paratype, showing a banded pattern (pale prozonites, dark metazonites) similar to that shown in Fig. 1. B. E. coxale sp. nov., tentatively referred female from type locality, showing dark lateral spots on a pale background. C. E. biquintum sp. nov., paratype from Mito Mitatu, showing a black body contrasting with white legs. Not to scale. Photographs by F. Vad and A. Illum." httpUri="https://zenodo.org/record/1489604/files/figure.png" pageId="71" pageNumber="72">Fig. 3C</figureCitation>
|
||
). Specimen from Udekwa Village (
|
||
<typeStatus id="54B304F3FFBFFFC1FF40FA92FEC1FA8B" box="[197,307,1304,1331]" pageId="71" pageNumber="72" type="holotype">holotype</typeStatus>
|
||
): after 23 years in alcohol head reddish brown; antennae, collum, metazonites and telson dark brown; prozonites and legs yellowish. Specimen from Mwanihana FR overall pallid after 38 years in alcohol, only metazonites dorsally very light brownish.
|
||
</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38FA2AFD65FA02" blockId="71.[189,663,1439,1466]" box="[189,663,1439,1466]" pageId="71" pageNumber="72">ANTENNAE. Reaching back to ring 4 or 5.</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38FA6BFE24F9FA" blockId="71.[189,1398,1505,1602]" pageId="71" pageNumber="72">
|
||
BODY RINGS. Paranota represented by very indistinct keels on body ring 2 (as
|
||
<figureCitation id="1333A6D4FFBFFFC1FB0CFA6BFB1BFA43" box="[1161,1257,1505,1531]" captionStart="Fig. 4" captionStartId="12.[189,232,1931,1957]" captionTargetBox="[191,1410,510,1870]" captionTargetId="figure@12.[180,1417,500,1882]" captionTargetPageId="12" captionText="Fig. 4. Eviulisoma spp. A–B. E. paradisiacum sp. nov., ♂. A. Head and rings 1–8, lateral view. B. Head and collum, frontal view. C. E. Zebra sp. nov., ♂, head and rings 1–3, lateral view. Abbreviation: pn = paranotum. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/1489606/files/figure.png" pageId="71" pageNumber="72">Fig. 4C</figureCitation>
|
||
), otherwise completely absent. Stricture between pro- and metazonite smooth. No setae seen on post-collar metazonites (abraded?).
|
||
</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38F9E3FD07F93B" blockId="71.[189,757,1641,1667]" box="[189,757,1641,1667]" pageId="71" pageNumber="72">HYPOPROCT. Semicircular, no marginal tubercles.</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38F921FAA5F95F" blockId="71.[189,1398,1706,1768]" pageId="71" pageNumber="72">LEGS. Length 1.5 × body width. Relative lengths of podomeres: tarsus (>) femur> tibia (>) prefemur (>) postfemur. Scopulae poorly developed, only on tibia and tarsus, absent from posteriormost leg pairs.</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38F885FB7EF892" blockId="71.[189,1164,1807,1834]" box="[189,1164,1807,1834]" pageId="71" pageNumber="72">STERNUM 5. With two small median knobs, one between legs 4, one between legs 5.</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38F8DAFBB1F8D2" blockId="71.[189,1091,1872,1899]" box="[189,1091,1872,1899]" pageId="71" pageNumber="72">STERNUM 6. Not excavated, coxae more separated than on normal body rings.</paragraph>
|
||
<paragraph id="8BB7BA51FFBFFFC1FF38F818FA84F877" blockId="71.[189,1399,1938,1999]" pageId="71" pageNumber="72">
|
||
GONOPODS (
|
||
<figureCitation id="1333A6D4FFBFFFC1FEC2F818FE50F815" box="[327,418,1938,1965]" captionStart="Fig. 35" captionStartId="72.[189,232,1892,1918]" captionTargetBox="[190,1402,265,1863]" captionTargetId="figure@72.[180,1427,249,1874]" captionTargetPageId="72" captionText="Fig. 35. Eviulisoma biquintum sp. nov., holotype. A–D. Right gonopod. A. Mesal view. B. Ventral view. C. Lateral view. D. Acropodite, subapical view. E. Sternum 5. Abbreviations: cxl = coxal lobe; map = mesal acropodital process; prf = prefemoral part; slm = solenomere; sph = solenophore. Scale bars: A–C, E = 0.1 mm; D = 0.05 mm." httpUri="https://zenodo.org/record/1489668/files/figure.png" pageId="71" pageNumber="72">Fig. 35</figureCitation>
|
||
). Coxal lobe (
|
||
<emphasis id="B97C6643FFBFFFC1FDD6F818FD87F814" box="[595,629,1938,1964]" italics="true" pageId="71" pageNumber="72">cxl</emphasis>
|
||
) moderate, irregular, not evenly rounded. Prefemoral part (
|
||
<emphasis id="B97C6643FFBFFFC1FACFF818FA9EF814" box="[1354,1388,1938,1964]" italics="true" pageId="71" pageNumber="72">prf</emphasis>
|
||
) ca 0.4× as long as acropodite. Mesal acropodital process (
|
||
<emphasis id="B97C6643FFBFFFC1FC04F83CFC46F877" box="[897,948,1974,1999]" italics="true" pageId="71" pageNumber="72">map</emphasis>
|
||
) long, narrow, straight sides tapering
|
||
</paragraph>
|
||
<caption id="DF77EAD9FFB0FFCEFF38F8EEFD73F851" httpUri="https://zenodo.org/record/1489668/files/figure.png" pageId="72" pageNumber="73" startId="72.[189,232,1892,1918]" targetBox="[190,1402,265,1863]" targetPageId="72">
|
||
<paragraph id="8BB7BA51FFB0FFCEFF38F8EEFD73F851" blockId="72.[189,1399,1892,2025]" pageId="72" pageNumber="73">
|
||
<emphasis id="B97C6643FFB0FFCEFF38F8EEFEECF8C6" bold="true" box="[189,286,1892,1918]" pageId="72" pageNumber="73">Fig. 35.</emphasis>
|
||
<taxonomicName id="4C08C1D2FFB0FFCEFEACF8EFFDDDF8C7" authority="Enghoff, 2018" authorityName="Enghoff" authorityYear="2018" box="[297,559,1893,1919]" class="Diplopoda" family="Paradoxosomatidae" genus="Eviulisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="biquintum" status="sp. nov.">
|
||
<emphasis id="B97C6643FFB0FFCEFEACF8EFFDDDF8C7" box="[297,559,1893,1919]" italics="true" pageId="72" pageNumber="73">Eviulisoma biquintum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24FDB38FFB0FFCEFDBEF8EFFD6EF8C7" box="[571,668,1893,1919]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype.
|
||
<emphasis id="B97C6643FFB0FFCEFCA3F8EEFCADF8C6" bold="true" box="[806,863,1892,1918]" pageId="72" pageNumber="73">A–D</emphasis>
|
||
. Right gonopod. A. Mesal view. B. Ventral view. C. Lateral view. D. Acropodite, subapical view.
|
||
<emphasis id="B97C6643FFB0FFCEFCC0F802FCABF81A" bold="true" box="[837,857,1928,1954]" pageId="72" pageNumber="73">E</emphasis>
|
||
. Sternum 5. Abbreviations:
|
||
<emphasis id="B97C6643FFB0FFCEFB2EF802FB18F81A" box="[1195,1258,1928,1954]" italics="true" pageId="72" pageNumber="73">cxl =</emphasis>
|
||
coxal lobe;
|
||
<emphasis id="B97C6643FFB0FFCEFF38F826FF02F87D" box="[189,240,1964,1989]" italics="true" pageId="72" pageNumber="73">map</emphasis>
|
||
= mesal acropodital process;
|
||
<emphasis id="B97C6643FFB0FFCEFDD2F821FD8BF87D" box="[599,633,1963,1989]" italics="true" pageId="72" pageNumber="73">prf</emphasis>
|
||
= prefemoral part;
|
||
<emphasis id="B97C6643FFB0FFCEFCE3F821FC7DF87D" box="[870,911,1963,1989]" italics="true" pageId="72" pageNumber="73">slm</emphasis>
|
||
= solenomere;
|
||
<emphasis id="B97C6643FFB0FFCEFBCEF821FB86F87D" box="[1099,1140,1963,1989]" italics="true" pageId="72" pageNumber="73">sph</emphasis>
|
||
= solenophore. Scale bars: A–C, E = 0.1 mm; D = 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB7BA51FFB1FFCFFF38FE81FB74FEF1" blockId="73.[189,1398,267,329]" pageId="73" pageNumber="74">
|
||
towards tip, apically regularly rounded, dorsally concave, forming gutter for solenophore. Solenophore (
|
||
<emphasis id="B97C6643FFB1FFCFFF42FEA5FF02FEF1" box="[199,240,303,329]" italics="true" pageId="73" pageNumber="74">sph</emphasis>
|
||
) slender, simple, a little longer than
|
||
<emphasis id="B97C6643FFB1FFCFFD18FEA5FD22FEF0" box="[669,720,303,328]" italics="true" pageId="73" pageNumber="74">map</emphasis>
|
||
, apically with two slender processes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312E9DAFFB1FFCFFF38FEFAFCA3FE64" pageId="73" pageNumber="74" type="distribution">
|
||
<paragraph id="8BB7BA51FFB1FFCFFF38FEFAFE1CFE32" blockId="73.[189,494,368,394]" box="[189,494,368,394]" pageId="73" pageNumber="74">
|
||
<heading id="D0FF0D3DFFB1FFCFFF38FEFAFE1CFE32" bold="true" box="[189,494,368,394]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
|
||
<emphasis id="B97C6643FFB1FFCFFF38FEFAFE1CFE32" bold="true" box="[189,494,368,394]" pageId="73" pageNumber="74">Distribution and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB7BA51FFB1FFCFFF38FE15FCA3FE64" blockId="73.[189,1398,414,476]" pageId="73" pageNumber="74">
|
||
Known from two parts of the Udzungwa Mts: Udzungwa Mountains National Park (incl. Mwanihana FR) and Nyambanike Mts in
|
||
<collectingRegion id="49CC74B3FFB1FFCFFD96FE48FDBFFE64" box="[531,589,450,476]" country="Tanzania" name="Mjini Magharibi" pageId="73" pageNumber="74">West</collectingRegion>
|
||
Kilombero Scarp FR.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312E9DAFFB1FFCFFF38FD89FD12FCD8" pageId="73" pageNumber="74" type="discussion">
|
||
<paragraph id="8BB7BA51FFB1FFCFFF38FD89FEDDFDA5" blockId="73.[189,303,515,541]" box="[189,303,515,541]" pageId="73" pageNumber="74">
|
||
<heading id="D0FF0D3DFFB1FFCFFF38FD89FEDDFDA5" bold="true" box="[189,303,515,541]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
|
||
<emphasis id="B97C6643FFB1FFCFFF38FD89FEDDFDA5" bold="true" box="[189,303,515,541]" pageId="73" pageNumber="74">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB7BA51FFB1FFCFFF38FDB8FA85FD0D" blockId="73.[189,1399,562,694]" pageId="73" pageNumber="74">
|
||
The newly collected specimen from the Udzungwa National Park is strikingly coloured – black body and white legs (
|
||
<figureCitation id="1333A6D4FFB1FFCFFEC2FDDFFE50FDD7" box="[327,418,597,623]" captionStart="Fig. 3" captionStartId="11.[189,232,1860,1886]" captionTargetBox="[190,1399,670,1825]" captionTargetId="figure@11.[170,1424,661,1834]" captionTargetPageId="11" captionText="Fig. 3. Eviulisoma spp., examples of colour patterns. A. E. akkariae sp. nov., paratype, showing a banded pattern (pale prozonites, dark metazonites) similar to that shown in Fig. 1. B. E. coxale sp. nov., tentatively referred female from type locality, showing dark lateral spots on a pale background. C. E. biquintum sp. nov., paratype from Mito Mitatu, showing a black body contrasting with white legs. Not to scale. Photographs by F. Vad and A. Illum." httpUri="https://zenodo.org/record/1489604/files/figure.png" pageId="73" pageNumber="74">Fig. 3C</figureCitation>
|
||
) – whereas the one from Nyambenike seems to be less remarkable in this respect. Fading with age of the specimen may explain the difference, but more specimens from each of these sites would be highly desirable in order to ascertain whether we are really dealing with only one species.
|
||
</paragraph>
|
||
<paragraph id="8BB7BA51FFB1FFCFFF38FD57FD12FCD8" blockId="73.[189,1399,733,865]" pageId="73" pageNumber="74">
|
||
The gonopods resemble those of
|
||
<taxonomicName id="4C08C1D2FFB1FFCFFDEFFD54FB39FD4F" authority="VandenSpiegel & Golovatch, 2014" authorityName="VandenSpiegel & Golovatch" authorityYear="2014" box="[618,1227,733,759]" class="Diplopoda" family="Paradoxosomatidae" genus="Eviulisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="taitaorum">
|
||
<emphasis id="B97C6643FFB1FFCFFDEFFD54FCF5FD4F" box="[618,775,734,759]" italics="true" pageId="73" pageNumber="74">E. taitaorum</emphasis>
|
||
<bibRefCitation id="EF99C7A0FFB1FFCFFC93FD57FB39FD4F" box="[790,1227,733,759]" pageId="73" pageNumber="74" refId="ref34882" refString="VandenSpiegel D. & Golovatch S. I. 2014. The millipede genus Eviulisoma Silvestri, 1910 in Kenya, with descriptions of new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 459: 11 - 34. https: // doi. org / 10.3897 / zookeys. 459.8621" type="journal article">VandenSpiegel & Golovatch, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C08C1D2FFB1FFCFFA9DFD54FD96FCA2" authority="VandenSpiegel & Golovatch, 2014" authorityName="VandenSpiegel & Golovatch" authorityYear="2014" class="Diplopoda" family="Paradoxosomatidae" genus="Eviulisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="taita">
|
||
<emphasis id="B97C6643FFB1FFCFFA9DFD54FA85FD4F" box="[1304,1399,734,759]" italics="true" pageId="73" pageNumber="74">E. taita</emphasis>
|
||
<bibRefCitation id="EF99C7A0FFB1FFCFFF38FC8AFD96FCA2" box="[189,612,768,794]" pageId="73" pageNumber="74" refId="ref34882" refString="VandenSpiegel D. & Golovatch S. I. 2014. The millipede genus Eviulisoma Silvestri, 1910 in Kenya, with descriptions of new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 459: 11 - 34. https: // doi. org / 10.3897 / zookeys. 459.8621" type="journal article">VandenSpiegel & Golovatch, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
. These two species, however, have an excavated sternum 6, and
|
||
<taxonomicName id="4C08C1D2FFB1FFCFFF38FCAEFEA4FC85" authorityName="VandenSpiegel & Golovatch" authorityYear="2014" box="[189,342,804,829]" class="Diplopoda" family="Paradoxosomatidae" genus="Eviulisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="taitaorum">
|
||
<emphasis id="B97C6643FFB1FFCFFF38FCAEFEA4FC85" box="[189,342,804,829]" italics="true" pageId="73" pageNumber="74">E. taitaorum</emphasis>
|
||
</taxonomicName>
|
||
lacks a lobe on sternum 5. No other species has two processes behind each other on sternum 5, according to available descriptions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |