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<document id="7692188987CDC1155C1AC5AAB048BBAC" ID-DOI="10.11646/zootaxa.3209.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5248266" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629763376848" checkinUser="felipe" docAuthor="Zahniser, J. N. &amp; Nielson, M. W." docDate="2012" docId="03A1071DFFFFFFEBFF16F8D66AD0FAF7" docLanguage="en" docName="zt03209p052.pdf" docOrigin="Zootaxa 3209 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3209.1.2" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ikelibeloha Zahniser &amp; Nielson 2012, gen.n." docType="treatment" docVersion="4" lastPageNumber="35" masterDocId="FF987F65FFFAFFECFF81F9616A1FFF9D" masterDocTitle="An extraordinary new genus and three new species of Acostemmini (Hemiptera: Cicadellidae: Deltocephalinae) from Madagascar with comments on the morphology and classification of the tribe" masterLastPageNumber="52" masterPageNumber="28" pageNumber="33" updateTime="1699108751254" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3470DB11084BC66D1AAB2B4D5A98AC80">An extraordinary new genus and three new species of Acostemmini (Hemiptera: Cicadellidae: Deltocephalinae) from Madagascar with comments on the morphology and classification of the tribe</mods:title>
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<mods:namePart id="823BD2135B911E2641537C11FBDD236D">Zahniser, J. N.</mods:namePart>
<mods:affiliation id="75C225872EB5B81F9F1EA669724B9B4F">Illinois Natural History Survey Prairie Research Institute University of Illinois at Urbana / Champaign 1816 S. Oak St. Champaign, IL 61820 USA. E-mail: zahniser @ illinois. edu</mods:affiliation>
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<mods:namePart id="9D2217A8CF784528F805935C07775186">Nielson, M. W.</mods:namePart>
<mods:affiliation id="6188583EC4AF4257AFC272864935FF3E">Affiliate Faculty, Monte L. Bean Museum Brigham Young University Provo, UT 84602 USA. E-mail: mwnielz @ gmail. com Corresponding author. E-mail: zahniser @ illinois. edu</mods:affiliation>
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<taxonomicName id="4C08CD88FFFFFFE9FF16F8D66B3FFE4C" ID-CoL="8RGN6" authority="Zahniser &amp; Nielson, 2012" authorityName="Zahniser &amp; Nielson" authorityYear="2012" box="[151,288,439,465]" class="Insecta" family="Cicadellidae" genus="Ikelibeloha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97C6A19FFFFFFE9FF16F8D66B3FFE4C" bold="true" box="[151,288,439,465]" italics="true" pageId="5" pageNumber="33">Ikelibeloha</emphasis>
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<paragraph id="8BB7B60BFFFFFFE9FF17F8BB6B00FE6F" blockId="5.[150,369,439,498]" box="[150,287,474,498]" pageId="5" pageNumber="33">
(
<figureCitation id="1333AA8EFFFFFFE9FF1EF8BB6B08FE6F" box="[159,279,474,498]" captionStart-0="FIGURES 19" captionStart-1="FIGURES 1013" captionStartId-0="6.[151,263,1856,1879]" captionStartId-1="7.[151,263,869,892]" captionTargetBox-0="[156,1433,214,1796]" captionTargetBox-1="[209,1378,201,842]" captionTargetId-0="figure-61@6.[151,1436,211,1802]" captionTargetId-1="figure-373@7.[201,1386,193,848]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." captionText-1="FIGURES 1013. Ikelibeloha cristata gen.n., sp.n. 10. pygofer and 1st valvulae; 11. detail of 1st valvula dorsal sculpturing; 12. second valvulae; 13. detail of second valvulae teeth." figureDoi-0="http://doi.org/10.5281/zenodo.5248268" figureDoi-1="http://doi.org/10.5281/zenodo.5248270" httpUri-0="https://zenodo.org/record/5248268/files/figure.png" httpUri-1="https://zenodo.org/record/5248270/files/figure.png" pageId="5" pageNumber="33">Figs. 113</figureCitation>
)
</paragraph>
<paragraph id="8BB7B60BFFFFFFE9FF16FB42683AFDA5" blockId="5.[151,549,546,568]" box="[151,549,546,568]" pageId="5" pageNumber="33">
<typeStatus id="54B308A9FFFFFFE9FF16FB426AD4FDA5" box="[151,203,547,568]" pageId="5" pageNumber="33">Type</typeStatus>
species:
<taxonomicName id="4C08CD88FFFFFFE9FEA7FB436BF4FDA5" authority="Zahniser &amp; Nielson, 2012" authorityName="Zahniser &amp; Nielson" authorityYear="2012" box="[294,491,546,568]" class="Insecta" family="Cicadellidae" genus="Ikelibeloha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="cristata" status="sp. nov.">
<emphasis id="B97C6A19FFFFFFE9FEA7FB436BF4FDA5" box="[294,491,546,568]" italics="true" pageId="5" pageNumber="33">Ikelibeloha cristata</emphasis>
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<paragraph id="8BB7B60BFFFFFFE9FF16FB046F40FBCC" blockId="5.[151,1437,613,2005]" pageId="5" pageNumber="33">
<emphasis id="B97C6A19FFFFFFE9FF16FB046B37FDE3" bold="true" box="[151,296,613,638]" inLineHeading="true" pageId="5" pageNumber="33" reason="1">Description.</emphasis>
Head. Head (Figs. 1,2,7) distinctly narrower than pronotum; anteriorly-posteriorly compressed. Crown extremely short; median length less than length next to eyes; delimited anteriorly by a single distinct transverse carina turning ventrad laterally and not reaching eyes; yellowish with darker spots medially on each side and next to eyes; surface irregular; texture finely shagreen in darker areas, glabrous or coarsely striate in lighter pigmented areas. Ocelli (
<figureCitation id="1333AA8EFFFFFFE9FE0FFB946BCDFC90" box="[398,466,757,781]" captionStart="FIGURES 19" captionStartId="6.[151,263,1856,1879]" captionTargetBox="[156,1433,214,1796]" captionTargetId="figure-61@6.[151,1436,211,1802]" captionTargetPageId="6" captionText="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." figureDoi="http://doi.org/10.5281/zenodo.5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="5" pageNumber="33">Fig. 7</figureCitation>
) distant from eyes, closer to each other than to eyes; bases stalked, arising from deep circular pits. Face (
<figureCitation id="1333AA8EFFFFFFE9FEF2FA786BA7FCAC" box="[371,440,793,817]" captionStart="FIGURES 19" captionStartId="6.[151,263,1856,1879]" captionTargetBox="[156,1433,214,1796]" captionTargetId="figure-61@6.[151,1436,211,1802]" captionTargetPageId="6" captionText="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." figureDoi="http://doi.org/10.5281/zenodo.5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="5" pageNumber="33">Fig. 7</figureCitation>
) below anterior carina finely shagreen with shallow oblique furrows; texture laterad of ocelli glabrous to rugose with coarse oblique furrows. Frontoclypeus (
<figureCitation id="1333AA8EFFFFFFE9FC5AFA5C6E3CFCC8" box="[987,1059,829,853]" captionStart="FIGURES 19" captionStartId="6.[151,263,1856,1879]" captionTargetBox="[156,1433,214,1796]" captionTargetId="figure-61@6.[151,1436,211,1802]" captionTargetPageId="6" captionText="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." figureDoi="http://doi.org/10.5281/zenodo.5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="5" pageNumber="33">Fig. 7</figureCitation>
) long, narrow; with median pit between ocelli; with short, curved lateral carinae next to antennae meeting lateral frontal suture and antennal ledge; texture shagreen or glabrous to rugose. Lateral frontal suture distinctly sulcate below antenna; weakly carinate above antenna; sharply angled at meeting point with antennal ledge; extending or nearly extending to ocellus. Antennal ledges sharply carinate. Antennal bases stalked; antennae long, about 2x width of head. Clypellar suture obscure. Clypellus narrow at base, abruptly expanded apically; apex slightly notched. Lora somewhat tumid, elevated above genae; lateral margins deeply sulcate. Genae broad; margin obtusely angulate, not emarginate below eyes; with deep oblique and swirling furrows; with fine erect seta somewhat distant from laterofrontal suture.
</paragraph>
<paragraph id="8BB7B60BFFFFFFE9FF46FD3C68DCFB20" blockId="5.[151,1437,613,2005]" pageId="5" pageNumber="33">Thorax. Pronotum (Figs. 1,2) very large, produced into angulate crest dorsally; texture punctate; with median longitudinal keel. Mesonotum (Figs. 1,2) large, bulbous; deep reddish-brown. Mesoscutellum (Figs. 1,2) produced, with median longitudinal carina on posterior half.</paragraph>
<paragraph id="8BB7B60BFFFFFFE9FF46FDA86F51FA08" blockId="5.[151,1437,613,2005]" pageId="5" pageNumber="33">
<materialsCitation id="3B60BC56FFFFFFE9FF46FDA86F56FA08" collectionCode="R" country="Oman" location="Hindwing" municipality="Plate" pageId="5" pageNumber="33" specimenCode="R1, R2, R4" specimenCount="1">
Wings. Forewing (
<figureCitation id="1333AA8EFFFFFFE9FE1DFDA86BFDFB7C" box="[412,482,1225,1249]" captionStart="FIGURES 19" captionStartId="6.[151,263,1856,1879]" captionTargetBox="[156,1433,214,1796]" captionTargetId="figure-61@6.[151,1436,211,1802]" captionTargetPageId="6" captionText="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." figureDoi="http://doi.org/10.5281/zenodo.5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="5" pageNumber="33">Fig. 2</figureCitation>
) squarish apically; appendix extending around apex; bases of C, claval suture, A1, and A2 thick, with punctate indentations; r-m crossvein obliquely meeting
<collectionCode id="ED192ECEFFFFFFE9FC7CFD8F6E10FA98" box="[1021,1039,1262,1285]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="33">R</collectionCode>
before separation of
<collectionCode id="ED192ECEFFFFFFE9FA8BFD8F6F02FA98" box="[1290,1309,1262,1285]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="33">R</collectionCode>
veins;
<specimenCode id="DBAE1E70FFFFFFE9FAF5FD8C6F88FA98" box="[1396,1431,1261,1285]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="33">R1</specimenCode>
,
<specimenCode id="DBAE1E70FFFFFFE9FF16FC706AA4FAB4" box="[151,187,1297,1321]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="33">R2</specimenCode>
+3, and
<specimenCode id="DBAE1E70FFFFFFE9FE97FC706B27FAB4" box="[278,312,1297,1321]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="33">R4</specimenCode>
+5 diverging from nearly same point; base of outer anteapical cell sharply angled; with or without extra costal vein; inner anteapical cell closed (m-cu crossvein present), somewhat expanded apically; with A1-A2 crossvein.
<location id="8ED7E0D0FFFFFFE9FE90FC386B9EFAEC" LSID="urn:lsid:plazi:treatment:03A1071DFFFFFFEBFF16F8D66AD0FAF7:8ED7E0D0FFFFFFE9FE90FC386B9EFAEC" box="[273,385,1369,1393]" country="Oman" municipality="Plate" name="Hindwing" pageId="5" pageNumber="33">Hindwing</location>
venation of normal deltocephaline
<typeStatus id="54B308A9FFFFFFE9FC99FC3B6957FAEF" box="[792,840,1370,1394]" pageId="5" pageNumber="33">type</typeStatus>
(e.g.
<collectingMunicipality id="6BD32C71FFFFFFE9FC0DFC3869DBFAEC" box="[908,964,1369,1393]" pageId="5" pageNumber="33">Plate</collectingMunicipality>
39,
<figureCitation id="1333AA8EFFFFFFE9FC78FC386E56FAEC" box="[1017,1097,1369,1393]" captionStart="FIGURES 19" captionStartId="6.[151,263,1856,1879]" captionTargetBox="[156,1433,214,1796]" captionTargetId="figure-61@6.[151,1436,211,1802]" captionTargetPageId="6" captionText="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." figureDoi="http://doi.org/10.5281/zenodo.5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="5" pageNumber="33">Fig. 9a</figureCitation>
of
<taxonomicName id="4C08CD88FFFFFFE9FBF3FC386F2FFAEC" authorityName="Stal" authorityYear="1864" box="[1138,1328,1369,1393]" class="Insecta" family="Cicadellidae" genus="Fieberiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="florii">
<emphasis id="B97C6A19FFFFFFE9FBF3FC386F2FFAEC" box="[1138,1328,1369,1393]" italics="true" pageId="5" pageNumber="33">Fieberiella florii</emphasis>
</taxonomicName>
in
<bibRefCitation id="EF99CBFAFFFFFFE9FAD6FC386AFAFA08" author="Oman, P. W." pageId="5" pageNumber="33" pagination="1 - 253" refId="ref14712" refString="Oman, P. W. (1949) The Nearctic leafhoppers. A generic classification and check list. Memoirs of the Entomological Society of Washington, 3, 1 - 253." type="journal article" year="1949">
<collectingCountry id="F31FF69BFFFFFFE9FAD6FC386F84FAEC" box="[1367,1435,1369,1393]" name="Oman" pageId="5" pageNumber="33">Oman</collectingCountry>
[1949]
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); with 4 apical cells; with perpendicular r4+5 - m1+2 crossvein; with oblique m3+4 - cuA crossvein
</materialsCitation>
.
</paragraph>
<paragraph id="8BB7B60BFFFFFFE9FF46FCC36804F944" blockId="5.[151,1437,613,2005]" pageId="5" pageNumber="33">Legs. Profemur row AV with only scattered fine hairlike setae; intercalary row with one row of long fine setae; row AM with AM1 present near midheight of femur and with or without additional fine setae proximally. Protibia somewhat flattened dorsally; without large macrosetae. Mesotrochanter with scattered fine setae. Mesofemur row AV without distinct setae. Metafemur apical macrosetae 2+1+1+1 (females) or 2+1+1 (male); sometimes with few smaller scattered setae. Metatibia somewhat expanded proximally; dorsal surface flat, without setae between rows; row PD setae not alternating much in length, entire row situated on distinctly raised ledge; row AD with macrosetae intercalated with smaller setae; row AV setae extending from apex to base ~3/4 length of tibia; apical pecten setae relatively small, tapered, of even length. Metatarsomere plantar surface with setae indistinct; apex with 45 platellae, outermost seta platellate.</paragraph>
<paragraph id="8BB7B60BFFFFFFE9FF46FF846851F810" blockId="5.[151,1437,613,2005]" pageId="5" pageNumber="33">
Female. Pygofer subquadrate; without macrosetae (
<figureCitation id="1333AA8EFFFFFFE9FC88FF846942F960" box="[777,861,1765,1789]" captionStart="FIGURES 1013" captionStartId="7.[151,263,869,892]" captionTargetBox="[209,1378,201,842]" captionTargetId="figure-373@7.[201,1386,193,848]" captionTargetPageId="7" captionText="FIGURES 1013. Ikelibeloha cristata gen.n., sp.n. 10. pygofer and 1st valvulae; 11. detail of 1st valvula dorsal sculpturing; 12. second valvulae; 13. detail of second valvulae teeth." figureDoi="http://doi.org/10.5281/zenodo.5248270" httpUri="https://zenodo.org/record/5248270/files/figure.png" pageId="5" pageNumber="33">Fig. 10</figureCitation>
). First valvifer rhomboidal in shape (
<figureCitation id="1333AA8EFFFFFFE9FA81FF846F4BF960" box="[1280,1364,1765,1789]" captionStart="FIGURES 1013" captionStartId="7.[151,263,869,892]" captionTargetBox="[209,1378,201,842]" captionTargetId="figure-373@7.[201,1386,193,848]" captionTargetPageId="7" captionText="FIGURES 1013. Ikelibeloha cristata gen.n., sp.n. 10. pygofer and 1st valvulae; 11. detail of 1st valvula dorsal sculpturing; 12. second valvulae; 13. detail of second valvulae teeth." figureDoi="http://doi.org/10.5281/zenodo.5248270" httpUri="https://zenodo.org/record/5248270/files/figure.png" pageId="5" pageNumber="33">Fig. 10</figureCitation>
). First valvula not strongly convex (
<figureCitation id="1333AA8EFFFFFFE9FE5EFE68682CF8BC" box="[479,563,1801,1825]" captionStart="FIGURES 1013" captionStartId="7.[151,263,869,892]" captionTargetBox="[209,1378,201,842]" captionTargetId="figure-373@7.[201,1386,193,848]" captionTargetPageId="7" captionText="FIGURES 1013. Ikelibeloha cristata gen.n., sp.n. 10. pygofer and 1st valvulae; 11. detail of 1st valvula dorsal sculpturing; 12. second valvulae; 13. detail of second valvulae teeth." figureDoi="http://doi.org/10.5281/zenodo.5248270" httpUri="https://zenodo.org/record/5248270/files/figure.png" pageId="5" pageNumber="33">Fig. 10</figureCitation>
); dorsal sculpturing pattern strigate, reaching dorsal margin (
<figureCitation id="1333AA8EFFFFFFE9FB63FE686F2BF8BC" box="[1250,1332,1801,1825]" captionStart="FIGURES 1013" captionStartId="7.[151,263,869,892]" captionTargetBox="[209,1378,201,842]" captionTargetId="figure-373@7.[201,1386,193,848]" captionTargetPageId="7" captionText="FIGURES 1013. Ikelibeloha cristata gen.n., sp.n. 10. pygofer and 1st valvulae; 11. detail of 1st valvula dorsal sculpturing; 12. second valvulae; 13. detail of second valvulae teeth." figureDoi="http://doi.org/10.5281/zenodo.5248270" httpUri="https://zenodo.org/record/5248270/files/figure.png" pageId="5" pageNumber="33">Fig. 11</figureCitation>
). Second valvula broadening subbasally, broad throughout length (
<figureCitation id="1333AA8EFFFFFFE9FCABFE4C699FF8D8" box="[810,896,1837,1861]" captionStart="FIGURES 1013" captionStartId="7.[151,263,869,892]" captionTargetBox="[209,1378,201,842]" captionTargetId="figure-373@7.[201,1386,193,848]" captionTargetPageId="7" captionText="FIGURES 1013. Ikelibeloha cristata gen.n., sp.n. 10. pygofer and 1st valvulae; 11. detail of 1st valvula dorsal sculpturing; 12. second valvulae; 13. detail of second valvulae teeth." figureDoi="http://doi.org/10.5281/zenodo.5248270" httpUri="https://zenodo.org/record/5248270/files/figure.png" pageId="5" pageNumber="33">Fig. 12</figureCitation>
); dorsal teeth obtusely triangular, not serrated, present on over 2/3 of length; in profile, teeth from left and right valvulae alternating (
<figureCitation id="1333AA8EFFFFFFE9FBE3FE306EA9F8F4" box="[1122,1206,1873,1897]" captionStart="FIGURES 1013" captionStartId="7.[151,263,869,892]" captionTargetBox="[209,1378,201,842]" captionTargetId="figure-373@7.[201,1386,193,848]" captionTargetPageId="7" captionText="FIGURES 1013. Ikelibeloha cristata gen.n., sp.n. 10. pygofer and 1st valvulae; 11. detail of 1st valvula dorsal sculpturing; 12. second valvulae; 13. detail of second valvulae teeth." figureDoi="http://doi.org/10.5281/zenodo.5248270" httpUri="https://zenodo.org/record/5248270/files/figure.png" pageId="5" pageNumber="33">Fig. 13</figureCitation>
). Third valvula with several small setae ventrally near apex.
</paragraph>
<paragraph id="8BB7B60BFFFFFFEBFF46FEF8690FFB2B" blockId="5.[151,1437,613,2005]" lastBlockId="7.[151,1436,966,1386]" lastPageId="7" lastPageNumber="35" pageId="5" pageNumber="33">
Male. Pygofer posterior margin incised dorsally to about midlength; apex with group of short stout setae; without basolateral membranous cleft (Figs. 3,8). Valve large; lateral margin long, articulating with pygofer; produced, pointed apically; surface with distinctive microtrichia (
<figureCitation id="1333AA8EFFFDFFEBFD7AFAA76920FC43" box="[763,831,966,990]" captionStart="FIGURES 19" captionStartId="6.[151,263,1856,1879]" captionTargetBox="[156,1433,214,1796]" captionTargetId="figure-61@6.[151,1436,211,1802]" captionTargetPageId="6" captionText="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." figureDoi="http://doi.org/10.5281/zenodo.5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="7" pageNumber="35">Fig. 9</figureCitation>
). Subgenital plates more or less triangular; ~2x longer than wide; without macrosetae; surface with distinctive microtrichia (
<figureCitation id="1333AA8EFFFDFFEBFC28FA8B69F0FB9F" box="[937,1007,1002,1026]" captionStart="FIGURES 19" captionStartId="6.[151,263,1856,1879]" captionTargetBox="[156,1433,214,1796]" captionTargetId="figure-61@6.[151,1436,211,1802]" captionTargetPageId="6" captionText="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." figureDoi="http://doi.org/10.5281/zenodo.5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="7" pageNumber="35">Fig. 9</figureCitation>
). Connective with long stem; anterior arms widely divergent; with ventrally directed anteromedial lobe (
<figureCitation id="1333AA8EFFFDFFEBFC04FD6F69D4FBBB" box="[901,971,1038,1062]" captionStart="FIGURES 19" captionStartId="6.[151,263,1856,1879]" captionTargetBox="[156,1433,214,1796]" captionTargetId="figure-61@6.[151,1436,211,1802]" captionTargetPageId="6" captionText="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." figureDoi="http://doi.org/10.5281/zenodo.5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="7" pageNumber="35">Fig. 4</figureCitation>
). Style linear, anteromedial lobe not pronounced; apophysis with well developed preapical lobe separated from distal hook by weakly sclerotized area; distal hook strongly curved, cheliform, forming narrow angle with preapical lobe (
<figureCitation id="1333AA8EFFFDFFEBFB91FD376E4AFBF3" box="[1040,1109,1110,1134]" captionStart="FIGURES 19" captionStartId="6.[151,263,1856,1879]" captionTargetBox="[156,1433,214,1796]" captionTargetId="figure-61@6.[151,1436,211,1802]" captionTargetPageId="6" captionText="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." figureDoi="http://doi.org/10.5281/zenodo.5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="7" pageNumber="35">Fig. 4</figureCitation>
). Aedeagus short, stout; with one shaft and gonopore (Figs. 5,6). Segment X short; with darkly sclerotized band dorsally and laterally 45x wider than long; remaining parts lightly sclerotized (Figs. 3,8).
</paragraph>
</subSubSection>
<caption id="DF77E683FFFCFFEAFF16FE216892F809" ID-DOI="http://doi.org/10.5281/zenodo.5248268" ID-Zenodo-Dep="5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="6" pageNumber="34" startId="6.[151,263,1856,1879]" targetBox="[156,1433,214,1796]" targetPageId="6">
<paragraph id="8BB7B60BFFFCFFEAFF16FE216892F809" blockId="6.[151,1436,1856,1940]" pageId="6" pageNumber="34">
<emphasis id="B97C6A19FFFCFFEAFF16FE216B24F8CA" bold="true" box="[151,315,1856,1879]" pageId="6" pageNumber="34">FIGURES 19.</emphasis>
<taxonomicName id="4C08CD88FFFCFFEAFEC3FE206817F8CA" authority="Zahniser &amp; Nielson, 2012" authorityName="Zahniser &amp; Nielson" authorityYear="2012" box="[322,520,1857,1879]" class="Insecta" family="Cicadellidae" genus="Ikelibeloha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="34" phylum="Arthropoda" rank="species" species="cristata" status="gen.n., sp.n.">
<emphasis id="B97C6A19FFFCFFEAFEC3FE206817F8CA" box="[322,520,1857,1879]" italics="true" pageId="6" pageNumber="34">Ikelibeloha cristata</emphasis>
</taxonomicName>
<emphasis id="B97C6A19FFFCFFEAFD91FE20688EF8CB" bold="true" box="[528,657,1857,1878]" pageId="6" pageNumber="34">
<taxonomicNameLabel id="A24FD762FFFCFFEAFD91FE20688EF8CB" box="[528,657,1857,1878]" pageId="6" pageNumber="34" rank="species">gen.n., sp.n.</taxonomicNameLabel>
</emphasis>
12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally.
</paragraph>
</caption>
<caption id="DF77E683FFFDFFEBFF16FA0468D1FC04" ID-DOI="http://doi.org/10.5281/zenodo.5248270" ID-Zenodo-Dep="5248270" httpUri="https://zenodo.org/record/5248270/files/figure.png" pageId="7" pageNumber="35" startId="7.[151,263,869,892]" targetBox="[209,1378,201,842]" targetPageId="7">
<paragraph id="8BB7B60BFFFDFFEBFF16FA0468D1FC04" blockId="7.[151,1436,869,921]" pageId="7" pageNumber="35">
<emphasis id="B97C6A19FFFDFFEBFF16FA046B4AFCE1" bold="true" box="[151,341,869,892]" pageId="7" pageNumber="35">FIGURES 1013.</emphasis>
<taxonomicName id="4C08CD88FFFDFFEBFEDDFA04683DFCE6" authority="Zahniser &amp; Nielson, 2012" authorityName="Zahniser &amp; Nielson" authorityYear="2012" box="[348,546,869,891]" class="Insecta" family="Cicadellidae" genus="Ikelibeloha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="35" phylum="Arthropoda" rank="species" species="cristata" status="gen.n., sp.n.">
<emphasis id="B97C6A19FFFDFFEBFEDDFA04683DFCE6" box="[348,546,869,891]" italics="true" pageId="7" pageNumber="35">Ikelibeloha cristata</emphasis>
</taxonomicName>
<emphasis id="B97C6A19FFFDFFEBFDABFA0768B4FCE6" bold="true" box="[554,683,870,891]" pageId="7" pageNumber="35">
<taxonomicNameLabel id="A24FD762FFFDFFEBFDABFA0768B4FCE6" box="[554,683,870,891]" pageId="7" pageNumber="35" rank="species">gen.n., sp.n.</taxonomicNameLabel>
</emphasis>
10. pygofer and 1st valvulae; 11. detail of 1st valvula dorsal sculpturing; 12. second valvulae; 13. detail of second valvulae teeth.
</paragraph>
</caption>
<subSubSection id="C312E580FFFDFFEBFF46FDA3691BFB47" box="[199,772,1218,1243]" pageId="7" pageNumber="35" type="distribution">
<paragraph id="8BB7B60BFFFDFFEBFF46FDA3691BFB47" blockId="7.[151,1436,966,1386]" box="[199,772,1218,1243]" pageId="7" pageNumber="35">
<emphasis id="B97C6A19FFFDFFEBFF46FDA36B7FFB46" bold="true" box="[199,352,1218,1243]" inLineHeading="true" pageId="7" pageNumber="35" reason="1">Distribution.</emphasis>
<collectingCountry id="F31FF69BFFFDFFEBFEE6FDA36BEFFB47" box="[359,496,1218,1242]" name="Madagascar" pageId="7" pageNumber="35">Madagascar</collectingCountry>
, Province dAntsiranana
</paragraph>
</subSubSection>
<subSubSection id="C312E580FFFDFFEBFF46FD8668C8FABF" pageId="7" pageNumber="35" type="discussion">
<paragraph id="8BB7B60BFFFDFFEBFF46FD8668C8FABF" blockId="7.[151,1436,966,1386]" pageId="7" pageNumber="35">
<emphasis id="B97C6A19FFFDFFEBFF46FD866B24FB62" bold="true" box="[199,315,1255,1279]" inLineHeading="true" pageId="7" pageNumber="35" reason="1">Remarks.</emphasis>
The specimen illustrated in
<figureCitation id="1333AA8EFFFDFFEBFDF7FD8768A9FB63" box="[630,694,1254,1278]" captionStart="FIGURES 19" captionStartId="6.[151,263,1856,1879]" captionTargetBox="[156,1433,214,1796]" captionTargetId="figure-61@6.[151,1436,211,1802]" captionTargetPageId="6" captionText="FIGURES 19. Ikelibeloha cristata gen.n., sp.n. 12: female paratype; 39: male holotype. 1. dorsal habitus; 2. lateral habitus; 3. pygofer, laterally; 4. connective and style, ventrally; 5. aedeagus, caudally; 6. aedeagus, laterally; 7. face; 8. pygofer, dorsally; 9. valve and subgenital plates, ventrally." figureDoi="http://doi.org/10.5281/zenodo.5248268" httpUri="https://zenodo.org/record/5248268/files/figure.png" pageId="7" pageNumber="35">Fig. 7</figureCitation>
has the frontoclypeus texture glabrous to rugose, while the other
<specimenCount id="9D0E7D82FFFDFFEBFA0FFD876B13FABF" pageId="7" pageNumber="35" type="generic">3 specimens</specimenCount>
have the frontoclypeus mostly shagreen.
</paragraph>
</subSubSection>
<subSubSection id="C312E580FFFDFFEBFF46FC4E6AD0FAF7" pageId="7" pageNumber="35" type="etymology">
<paragraph id="8BB7B60BFFFDFFEBFF46FC4E6AD0FAF7" blockId="7.[151,1436,966,1386]" pageId="7" pageNumber="35">
<emphasis id="B97C6A19FFFDFFEBFF46FC4E6B51FADA" bold="true" box="[199,334,1327,1351]" inLineHeading="true" pageId="7" pageNumber="35" reason="1">Etymology.</emphasis>
The genus is named for the Malagasy word meaning “little big headed one”. The gender is feminine.
</paragraph>
</subSubSection>
</treatment>
</document>