259 lines
33 KiB
XML
259 lines
33 KiB
XML
<document id="0E75296BF695DD693DBE35D1DCB0C388" ID-CLB-Dataset="20378" ID-DOI="10.11646/zootaxa.5104.4.4" ID-GBIF-Dataset="c3a77909-4443-494a-a3f2-cf57eed196c2" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332181" ID-ZooBank="44C79B8A-12BC-4066-9864-E42DCBA0EF6B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646595398227" checkinUser="plazi" docAuthor="Poorani, J." docDate="2022" docId="039FCF4AFFB0FF8D2AB26A69FAE1FD18" docLanguage="en" docName="zootaxa.5104.4.4.pdf" docOrigin="Zootaxa 5104 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Horniolus wiolettae Poorani 2022, sp. n." docType="treatment" docVersion="7" lastPageNumber="553" masterDocId="FFA6B732FFB5FF852A256D22FFF6FFC4" masterDocTitle="A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species" masterLastPageNumber="556" masterPageNumber="545" pageNumber="550" updateTime="1699316983446" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="28242494ABA3F69FA0F039D000200822">
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<mods:title id="46524A362949051BFC5357FF847B2ADC">A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species</mods:title>
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<mods:name id="CBED7107D9E7036B0A29FA9EF8EF70C3" type="personal">
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<mods:roleTerm id="056E3C607DB6F447A0784BC4359E80D6">Author</mods:roleTerm>
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<mods:namePart id="5AC6AD361B685CE4235D54ED74FE5E27">Poorani, J.</mods:namePart>
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<mods:typeOfResource id="47364F7A8B3F2BE3C789551803F238F4">text</mods:typeOfResource>
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<mods:title id="464192AA4BF0E15BA03719742966DE79">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FAC46C1F83A5BE4CD7A12E6D80331F2F">
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<mods:date id="CC5F45E9BEF230FBDF9CB3038766C725">2022</mods:date>
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<mods:number id="73A6644C50572AC2F256C0A86BD5CE8F">2022-03-01</mods:number>
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<mods:number id="209850907FF29149050E75F382143A7A">5104</mods:number>
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<mods:number id="0B92D3933301E23640ED8930E0103444">4</mods:number>
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<mods:start id="E853BC40E85AF00AB10F38EA8C40321B">545</mods:start>
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<mods:identifier id="CDEB8820E65F169BFA9F1B5CC1EB17AD" type="DOI">10.11646/zootaxa.5104.4.4</mods:identifier>
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<mods:identifier id="F2885B0DD6ADC718D46AEC1D293ECE4C" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="039FCF4AFFB0FF8D2AB26A69FAE1FD18" ID-DOI="http://doi.org/10.5281/zenodo.6333334" ID-GBIF-Taxon="193878401" ID-Zenodo-Dep="6333334" LSID="urn:lsid:plazi:treatment:039FCF4AFFB0FF8D2AB26A69FAE1FD18" httpUri="http://treatment.plazi.org/id/039FCF4AFFB0FF8D2AB26A69FAE1FD18" lastPageId="8" lastPageNumber="553" pageId="5" pageNumber="550">
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<subSubSection id="C32C2DD7FFB0FF802AB26A69FE39F8A2" box="[151,463,1867,1894]" pageId="5" pageNumber="550" type="nomenclature">
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<paragraph id="8B897E5CFFB0FF802AB26A69FE39F8A2" blockId="5.[151,463,1867,1930]" box="[151,463,1867,1894]" pageId="5" pageNumber="550">
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<heading id="D0C1C930FFB0FF802AB26A69FE39F8A2" bold="true" box="[151,463,1867,1894]" fontSize="11" level="1" pageId="5" pageNumber="550" reason="1">
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<emphasis id="B942A24EFFB0FF802AB26A69FE39F8A2" bold="true" box="[151,463,1867,1894]" pageId="5" pageNumber="550">
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<taxonomicName id="4C3605DFFFB0FF802AB26A69FE70F8A2" authority="Poorani, 2022" authorityName="Poorani" authorityYear="2022" box="[151,390,1867,1894]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="wiolettae" status="sp. nov.">
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<emphasis id="B942A24EFFB0FF802AB26A69FE70F8A2" bold="true" box="[151,390,1867,1894]" italics="true" pageId="5" pageNumber="550">Horniolus wiolettae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2711F35FFB0FF802BA86A6EFE39F8A2" box="[397,463,1868,1894]" pageId="5" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C2DD7FFB0FF802AB26A4DFF1DF84E" box="[151,235,1903,1930]" pageId="5" pageNumber="550" type="description">
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<paragraph id="8B897E5CFFB0FF802AB26A4DFF1DF84E" blockId="5.[151,463,1867,1930]" box="[151,235,1903,1930]" pageId="5" pageNumber="550">
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(
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<figureCitation id="130D62D9FFB0FF802A856A4DFF15F84E" box="[160,227,1903,1930]" captionStart="FIGURE 4" captionStartId="7.[151,250,1682,1706]" captionTargetBox="[164,1412,181,1647]" captionTargetId="figure-15@7.[164,1423,181,1659]" captionTargetPageId="7" captionText="FIGURE 4. Horniolus wiolettae sp. n.: a. holotype male, dorsal view; b. abdomen; c–f. male genitalia: c. tegmen, ventral view; d. tegmen, lateral view; e. penis; f. penis apex." figureDoi="http://doi.org/10.5281/zenodo.6332189" httpUri="https://zenodo.org/record/6332189/files/figure.png" pageId="5" pageNumber="550">Fig. 4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C2DD7FFB0FF832AB26A95FCBEFE4D" lastPageId="6" lastPageNumber="551" pageId="5" pageNumber="550" type="diagnosis">
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<paragraph id="8B897E5CFFB0FF832AB26A95FCBEFE4D" blockId="5.[151,1436,1975,2038]" lastBlockId="6.[151,1437,151,393]" lastPageId="6" lastPageNumber="551" pageId="5" pageNumber="550">
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<emphasis id="B942A24EFFB0FF802AB26A95FEE4F815" bold="true" box="[151,274,1975,2001]" pageId="5" pageNumber="550">Diagnosis.</emphasis>
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<taxonomicName id="4C3605DFFFB0FF802B3C6A95FE0EF815" authority="Poorani, 2022" authorityName="Poorani" authorityYear="2022" box="[281,504,1975,2001]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="wiolettae" status="sp. nov.">
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<emphasis id="B942A24EFFB0FF802B3C6A95FE0EF815" box="[281,504,1975,2001]" italics="true" pageId="5" pageNumber="550">Horniolus wiolettae</emphasis>
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</taxonomicName>
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<emphasis id="B942A24EFFB0FF802BDA6A9AFDC9F816" bold="true" box="[511,575,1976,2002]" pageId="5" pageNumber="550">
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<taxonomicNameLabel id="A2711F35FFB0FF802BDA6A9AFDC9F816" box="[511,575,1976,2002]" pageId="5" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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resembles
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<taxonomicName id="4C3605DFFFB0FF8028986A9AFC5BF815" authority="Miyatake" authorityName="Miyatake" box="[701,941,1975,2001]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="nigripes">
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<emphasis id="B942A24EFFB0FF8028986A9AFCCAF815" box="[701,828,1976,2001]" italics="true" pageId="5" pageNumber="550">H. nigripes</emphasis>
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Miyatake
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</taxonomicName>
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and the male genitalia are also superficially similar. However, the head and pronotum in
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<taxonomicName id="4C3605DFFFB0FF8028A86AFEFCFDF831" box="[653,779,2012,2037]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="nigripes">
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<emphasis id="B942A24EFFB0FF8028A86AFEFCFDF831" box="[653,779,2012,2037]" italics="true" pageId="5" pageNumber="550">H. nigripes</emphasis>
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</taxonomicName>
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are black and the male genitalia also differ as follows: the penis guide is as long as parameres and distinctly narrower in the apical one-third and the penis is more strongly curved in
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<taxonomicName id="4C3605DFFFB3FF832B206D99FE12FF11" authority="Poorani, 2022" authorityName="Poorani" authorityYear="2022" box="[261,484,187,213]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="wiolettae" status="sp. nov.">
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<emphasis id="B942A24EFFB3FF832B206D99FE12FF11" box="[261,484,187,213]" italics="true" pageId="6" pageNumber="551">Horniolus wiolettae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2711F35FFB3FF832BCC6D99FDD2FF11" box="[489,548,187,213]" pageId="6" pageNumber="551" rank="species">sp. n.</taxonomicNameLabel>
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while in
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<taxonomicName id="4C3605DFFFB3FF8328AF6D99FCFEFF10" box="[650,776,187,212]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="nigripes">
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<emphasis id="B942A24EFFB3FF8328AF6D99FCFEFF10" box="[650,776,187,212]" italics="true" pageId="6" pageNumber="551">H. nigripes</emphasis>
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</taxonomicName>
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, the penis guide is shorter than parameres, elongate lanceolate and not narrowed in the apical one-third, and distinctly narrower in lateral view. It also resembles
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<taxonomicName id="4C3605DFFFB3FF832F386DFDFA6AFF3C" authorityName="Poorani" authorityYear="2015" box="[1309,1436,223,248]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="sororius">
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<emphasis id="B942A24EFFB3FF832F386DFDFA6AFF3C" box="[1309,1436,223,248]" italics="true" pageId="6" pageNumber="551">H. sororius</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C3605DFFFB3FF832AED6C21FE93FED9" authorityName=": Canepari" authorityYear="2003" box="[200,357,259,285]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="hisamatsui">
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<emphasis id="B942A24EFFB3FF832AED6C21FE93FED9" box="[200,357,259,285]" italics="true" pageId="6" pageNumber="551">H. hisamatsui</emphasis>
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</taxonomicName>
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in having four-spotted elytra but the latter two species differ from it in having distinctly more transverse, narrower and smaller elytral maculae and different male genitalia.
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<taxonomicName id="4C3605DFFFB3FF832E3E6C05FB28FE85" authorityName="Weise, R. G. Booth" authorityYear="1901" box="[1051,1246,295,321]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="dispar">
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<emphasis id="B942A24EFFB3FF832E3E6C05FB28FE85" box="[1051,1246,295,321]" italics="true" pageId="6" pageNumber="551">Horniolus dispar</emphasis>
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</taxonomicName>
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differs from the newly described species in having a much darker brown head and pronotum and anterior pair of elytral maculae differently shaped and more deeply constricted in the middle.
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</paragraph>
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</subSubSection>
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<caption id="DF492ED4FFB3FF832AB26AABFC39F801" ID-DOI="http://doi.org/10.5281/zenodo.6332187" ID-Zenodo-Dep="6332187" httpUri="https://zenodo.org/record/6332187/files/figure.png" pageId="6" pageNumber="551" startId="6.[151,250,1929,1953]" targetBox="[167,1423,432,1847]" targetPageId="6">
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<paragraph id="8B897E5CFFB3FF832AB26AABFC39F801" blockId="6.[151,1436,1929,1990]" pageId="6" pageNumber="551">
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<emphasis id="B942A24EFFB3FF832AB26AABFEE4F865" bold="true" box="[151,274,1929,1953]" pageId="6" pageNumber="551">FIGURE 3.</emphasis>
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<taxonomicName id="4C3605DFFFB3FF832B326AABFDE8F865" authority="Poorani, 2022" authorityName="Poorani" authorityYear="2022" box="[279,542,1929,1953]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="darjeelingensis" status="sp. nov.">
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<emphasis id="B942A24EFFB3FF832B326AABFDE8F865" box="[279,542,1929,1953]" italics="true" pageId="6" pageNumber="551">Horniolus darjeelingensis</emphasis>
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</taxonomicName>
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<emphasis id="B942A24EFFB3FF8328066AABFDADF865" bold="true" box="[547,603,1929,1953]" pageId="6" pageNumber="551">
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<taxonomicNameLabel id="A2711F35FFB3FF8328066AABFDADF865" box="[547,603,1929,1953]" pageId="6" pageNumber="551" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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: a. holotype male, dorsal view; b. abdomen; c. abdominal postcoxal line; d–g. male genitalia: d. tegmen, ventral view; e. tegmen, lateral view; f. penis; g. penis apex.
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</paragraph>
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</caption>
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<caption id="DF492ED4FFB2FF822AB26BB0FD99F90A" ID-DOI="http://doi.org/10.5281/zenodo.6332189" ID-Zenodo-Dep="6332189" httpUri="https://zenodo.org/record/6332189/files/figure.png" pageId="7" pageNumber="552" startId="7.[151,250,1682,1706]" targetBox="[164,1412,181,1647]" targetPageId="7">
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||
<paragraph id="8B897E5CFFB2FF822AB26BB0FD99F90A" blockId="7.[151,1436,1682,1743]" pageId="7" pageNumber="552">
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||
<emphasis id="B942A24EFFB2FF822AB26BB0FEE4F96F" bold="true" box="[151,274,1682,1707]" pageId="7" pageNumber="552">FIGURE 4.</emphasis>
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<taxonomicName id="4C3605DFFFB2FF822B3D6BB1FE14F96F" authority="Poorani, 2022" authorityName="Poorani" authorityYear="2022" box="[280,482,1683,1707]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="wiolettae" status="sp. nov.">
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<emphasis id="B942A24EFFB2FF822B3D6BB1FE14F96F" box="[280,482,1683,1707]" italics="true" pageId="7" pageNumber="552">Horniolus wiolettae</emphasis>
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</taxonomicName>
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<emphasis id="B942A24EFFB2FF822BC26BB1FDE9F96F" bold="true" box="[487,543,1683,1707]" pageId="7" pageNumber="552">
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<taxonomicNameLabel id="A2711F35FFB2FF822BC26BB1FDE9F96F" box="[487,543,1683,1707]" pageId="7" pageNumber="552" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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: a. holotype male, dorsal view; b. abdomen; c–f. male genitalia: c. tegmen, ventral view; d. tegmen, lateral view; e. penis; f. penis apex.
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</paragraph>
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</caption>
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<subSubSection id="C32C2DD7FFB2FF8D2AE26BDFFE4EFE45" lastPageId="8" lastPageNumber="553" pageId="7" pageNumber="552" type="description">
|
||
<paragraph id="8B897E5CFFB2FF8D2AE26BDFFAB7FE9A" blockId="7.[151,1437,1788,2024]" lastBlockId="8.[151,1437,151,732]" lastPageId="8" lastPageNumber="553" pageId="7" pageNumber="552">
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<emphasis id="B942A24EFFB2FF822AE26BDFFE6CF8D3" bold="true" box="[199,410,1788,1815]" pageId="7" pageNumber="552">
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Description.
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<emphasis id="B942A24EFFB2FF822B456BDEFE6CF8D3" bold="true" box="[352,410,1788,1815]" italics="true" pageId="7" pageNumber="552">Male</emphasis>
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</emphasis>
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: Length:
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<quantity id="4CCED3B9FFB2FF82282F6BDFFD5AF8D3" box="[522,684,1789,1815]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.68" metricValueMin="1.42" pageId="7" pageNumber="552" unit="mm" value="1.5499999999999998" valueMax="1.68" valueMin="1.42">1.42–1.68 mm</quantity>
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; width:
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<quantity id="4CCED3B9FFB2FF8229296BDFFC47F8D3" box="[780,945,1789,1815]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.19" metricValueMin="1.01" pageId="7" pageNumber="552" unit="mm" value="1.1" valueMax="1.19" valueMin="1.01">1.01–1.19 mm</quantity>
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. TL/TW: 1.41; PL/PW: 0.49–0.53; EL/EW: 1.00–1.03. Form (
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<figureCitation id="130D62D9FFB2FF822B466A3DFE42F8FE" box="[355,436,1823,1850]" captionStart="FIGURE 4" captionStartId="7.[151,250,1682,1706]" captionTargetBox="[164,1412,181,1647]" captionTargetId="figure-15@7.[164,1423,181,1659]" captionTargetPageId="7" captionText="FIGURE 4. Horniolus wiolettae sp. n.: a. holotype male, dorsal view; b. abdomen; c–f. male genitalia: c. tegmen, ventral view; d. tegmen, lateral view; e. penis; f. penis apex." figureDoi="http://doi.org/10.5281/zenodo.6332189" httpUri="https://zenodo.org/record/6332189/files/figure.png" pageId="7" pageNumber="552">Fig. 4a</figureCitation>
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) elongate oval, dorsum moderately convex, densely pubescent, head and pronotum with yellowish hairs, lateral sides of pronotum with a mixure of dark brown and paler hairs, elytra with bright yellowish hairs on yellow maculae, rest of elytra with dark brown pubescence. Head and pronotum dark reddish brown; scutellar shield black; elytra black, each elytron with a pair of yellowish maculae, anterior one large, broad with a wavy margin, just below basal margin, not reaching suture, almost touching lateral margin, posterior spot smaller, elongate, oblique oval. Head about 3.5x as wide as an eye, interocular distance about 1.6x as wide as an eye; Pronotum densely punctate, punctures separated by 1–2 diameters. Elytra densely punctate, punctures shallowly impressed, separated by 1–3 diameters, interspaces smooth. Abdominal postcoxal lines (
|
||
<figureCitation id="130D62D9FFBDFF8D29026D9BFC87FF10" box="[807,881,185,212]" captionStart="FIGURE 4" captionStartId="7.[151,250,1682,1706]" captionTargetBox="[164,1412,181,1647]" captionTargetId="figure-15@7.[164,1423,181,1659]" captionTargetPageId="7" captionText="FIGURE 4. Horniolus wiolettae sp. n.: a. holotype male, dorsal view; b. abdomen; c–f. male genitalia: c. tegmen, ventral view; d. tegmen, lateral view; e. penis; f. penis apex." figureDoi="http://doi.org/10.5281/zenodo.6332189" httpUri="https://zenodo.org/record/6332189/files/figure.png" pageId="8" pageNumber="553">Fig.4b</figureCitation>
|
||
) complete, broadly and deeply semicircular. Male genitalia (
|
||
<figureCitation id="130D62D9FFBDFF8D2B216DFEFE82FF33" box="[260,372,220,247]" captionStart="FIGURE 4" captionStartId="7.[151,250,1682,1706]" captionTargetBox="[164,1412,181,1647]" captionTargetId="figure-15@7.[164,1423,181,1659]" captionTargetPageId="7" captionText="FIGURE 4. Horniolus wiolettae sp. n.: a. holotype male, dorsal view; b. abdomen; c–f. male genitalia: c. tegmen, ventral view; d. tegmen, lateral view; e. penis; f. penis apex." figureDoi="http://doi.org/10.5281/zenodo.6332189" httpUri="https://zenodo.org/record/6332189/files/figure.png" pageId="8" pageNumber="553">Figs. 4c–f</figureCitation>
|
||
) with penis guide in ventral view (
|
||
<figureCitation id="130D62D9FFBDFF8D28D56DFEFCC8FF32" box="[752,830,220,246]" captionStart="FIGURE 4" captionStartId="7.[151,250,1682,1706]" captionTargetBox="[164,1412,181,1647]" captionTargetId="figure-15@7.[164,1423,181,1659]" captionTargetPageId="7" captionText="FIGURE 4. Horniolus wiolettae sp. n.: a. holotype male, dorsal view; b. abdomen; c–f. male genitalia: c. tegmen, ventral view; d. tegmen, lateral view; e. penis; f. penis apex." figureDoi="http://doi.org/10.5281/zenodo.6332189" httpUri="https://zenodo.org/record/6332189/files/figure.png" pageId="8" pageNumber="553">Fig. 4c</figureCitation>
|
||
) basally broadest, its sides gradually narrowed towards middle, abruptly narrowed posteriorly into a triangular apex; penis guide in lateral view (
|
||
<figureCitation id="130D62D9FFBDFF8D2E426DDDFB40FEDE" box="[1127,1206,255,282]" captionStart="FIGURE 4" captionStartId="7.[151,250,1682,1706]" captionTargetBox="[164,1412,181,1647]" captionTargetId="figure-15@7.[164,1423,181,1659]" captionTargetPageId="7" captionText="FIGURE 4. Horniolus wiolettae sp. n.: a. holotype male, dorsal view; b. abdomen; c–f. male genitalia: c. tegmen, ventral view; d. tegmen, lateral view; e. penis; f. penis apex." figureDoi="http://doi.org/10.5281/zenodo.6332189" httpUri="https://zenodo.org/record/6332189/files/figure.png" pageId="8" pageNumber="553">Fig. 4d</figureCitation>
|
||
) basally curved, apically strongly narrowed to an acute tip; parameres (
|
||
<figureCitation id="130D62D9FFBDFF8D28EE6C03FCC3FEF8" box="[715,821,289,316]" captionStart="FIGURE 4" captionStartId="7.[151,250,1682,1706]" captionTargetBox="[164,1412,181,1647]" captionTargetId="figure-15@7.[164,1423,181,1659]" captionTargetPageId="7" captionText="FIGURE 4. Horniolus wiolettae sp. n.: a. holotype male, dorsal view; b. abdomen; c–f. male genitalia: c. tegmen, ventral view; d. tegmen, lateral view; e. penis; f. penis apex." figureDoi="http://doi.org/10.5281/zenodo.6332189" httpUri="https://zenodo.org/record/6332189/files/figure.png" pageId="8" pageNumber="553">Fig. 4c, d</figureCitation>
|
||
) with inner and apical margins densely setose, subequal in length to penis guide; penis (
|
||
<figureCitation id="130D62D9FFBDFF8D2BCB6C66FDCDFE9A" box="[494,571,324,350]" captionStart="FIGURE 4" captionStartId="7.[151,250,1682,1706]" captionTargetBox="[164,1412,181,1647]" captionTargetId="figure-15@7.[164,1423,181,1659]" captionTargetPageId="7" captionText="FIGURE 4. Horniolus wiolettae sp. n.: a. holotype male, dorsal view; b. abdomen; c–f. male genitalia: c. tegmen, ventral view; d. tegmen, lateral view; e. penis; f. penis apex." figureDoi="http://doi.org/10.5281/zenodo.6332189" httpUri="https://zenodo.org/record/6332189/files/figure.png" pageId="8" pageNumber="553">Fig. 4e</figureCitation>
|
||
) with a distinct capsule, strongly curved, apex as illustrated on
|
||
<figureCitation id="130D62D9FFBDFF8D2ED46C66FAC8FE9A" box="[1265,1342,324,350]" captionStart="FIGURE 4" captionStartId="7.[151,250,1682,1706]" captionTargetBox="[164,1412,181,1647]" captionTargetId="figure-15@7.[164,1423,181,1659]" captionTargetPageId="7" captionText="FIGURE 4. Horniolus wiolettae sp. n.: a. holotype male, dorsal view; b. abdomen; c–f. male genitalia: c. tegmen, ventral view; d. tegmen, lateral view; e. penis; f. penis apex." figureDoi="http://doi.org/10.5281/zenodo.6332189" httpUri="https://zenodo.org/record/6332189/files/figure.png" pageId="8" pageNumber="553">
|
||
Fig.
|
||
<specimenCount id="9D30B5D5FFBDFF8D2F066C66FAC8FE9A" box="[1315,1342,324,350]" pageId="8" pageNumber="553" type="generic">4f</specimenCount>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B897E5CFFBDFF8D2AE26C45FE4EFE45" blockId="8.[151,1437,151,732]" box="[199,440,359,385]" pageId="8" pageNumber="553">
|
||
<emphasis id="B942A24EFFBDFF8D2AE26C45FEDEFE45" bold="true" box="[199,296,359,385]" pageId="8" pageNumber="553">Female:</emphasis>
|
||
Not studied.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C2DD7FFBDFF8D2AE26CABFBEBFE02" pageId="8" pageNumber="553" type="materials_examined">
|
||
<paragraph id="8B897E5CFFBDFF8D2AE26CABFBEBFE02" blockId="8.[151,1437,151,732]" pageId="8" pageNumber="553">
|
||
<emphasis id="B942A24EFFBDFF8D2AE26CABFE47FE67" bold="true" box="[199,433,393,419]" pageId="8" pageNumber="553">Material examined.</emphasis>
|
||
<materialsCitation id="3B5E7401FFBDFF8D2B9C6CABFEA0FE02" ID-GBIF-Occurrence="3697318304" collectionCode="NBAIR" country="India" latitude="8.26" location="Kanyakumari" longLatPrecision="781" longitude="77.18" municipality="Pechiparai" pageId="8" pageNumber="553" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="548DC0FEFFBDFF8D2B9C6CABFDD7FE67" box="[441,545,393,419]" pageId="8" pageNumber="553" type="holotype">Holotype</typeStatus>
|
||
male; “
|
||
<collectingCountry id="F3213ECCFFBDFF8D285D6CABFD3BFE67" box="[632,717,393,419]" name="India" pageId="8" pageNumber="553">INDIA</collectingCountry>
|
||
, TN,
|
||
<location id="8EE92887FFBDFF8D29296CABFC51FE67" LSID="urn:lsid:plazi:treatment:039FCF4AFFB0FF8D2AB26A69FAE1FD18:8EE92887FFBDFF8D29296CABFC51FE67" box="[780,935,393,419]" country="India" latitude="8.26" longLatPrecision="781" longitude="77.18" municipality="Pechiparai" name="Kanyakumari" pageId="8" pageNumber="553">Kanyakumari</location>
|
||
,
|
||
<collectingMunicipality id="6BEDE426FFBDFF8D29966CABFBDAFE67" box="[947,1068,393,419]" pageId="8" pageNumber="553">Pechiparai</collectingMunicipality>
|
||
, 18.vi.15,
|
||
<geoCoordinate id="EE02189BFFBDFF8D2E866CABFB08FE53" box="[1187,1278,393,419]" degrees="08.26" direction="north" orientation="latitude" pageId="8" pageNumber="553" precision="555" value="8.26">
|
||
N08.26
|
||
<superScript id="7C43D314FFBDFF8D2ED36CABFB08FE53" attach="left" box="[1270,1278,393,407]" fontSize="6" pageId="8" pageNumber="553">o</superScript>
|
||
</geoCoordinate>
|
||
<geoCoordinate id="EE02189BFFBDFF8D2F236CABFA9AFE53" box="[1286,1388,393,419]" degrees="077.18" direction="east" orientation="longitude" pageId="8" pageNumber="553" precision="555" value="77.18">
|
||
E077.18
|
||
<superScript id="7C43D314FFBDFF8D2F416CABFA9AFE53" attach="left" box="[1380,1388,393,407]" fontSize="6" pageId="8" pageNumber="553">o</superScript>
|
||
</geoCoordinate>
|
||
SM Team” (
|
||
<collectionCode id="ED27E699FFBDFF8D2AD56C8EFEB8FE02" box="[240,334,428,454]" pageId="8" pageNumber="553">NBAIR</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5E7401FFBDFF8D2B446C8FFBEFFE02" ID-GBIF-Occurrence="3697318302" box="[353,1049,428,455]" collectionCode="NBAIR" country="India" latitude="8.26" location="Kanyakumari" longLatPrecision="781" longitude="77.18" municipality="Pechiparai" pageId="8" pageNumber="553" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="548DC0FEFFBDFF8D2B446C8FFE31FE03" box="[353,455,429,455]" pageId="8" pageNumber="553" type="paratype">Paratype</typeStatus>
|
||
: One male, with the same data as holotype (
|
||
<collectionCode id="ED27E699FFBDFF8D29966C8EFBE7FE02" box="[947,1041,428,454]" pageId="8" pageNumber="553">NBAIR</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C2DD7FFBDFF8D2AE26CECFDCDFE2D" box="[199,571,462,489]" pageId="8" pageNumber="553" type="distribution">
|
||
<paragraph id="8B897E5CFFBDFF8D2AE26CECFDCDFE2D" blockId="8.[151,1437,151,732]" box="[199,571,462,489]" pageId="8" pageNumber="553">
|
||
<emphasis id="B942A24EFFBDFF8D2AE26CECFE97FE2C" bold="true" box="[199,353,462,488]" pageId="8" pageNumber="553">Distribution.</emphasis>
|
||
<collectingCountry id="F3213ECCFFBDFF8D2B426CEDFE53FE2D" box="[359,421,463,489]" name="India" pageId="8" pageNumber="553">India</collectingCountry>
|
||
:
|
||
<collectingRegion id="49F2B0BEFFBDFF8D2B8A6CEDFDC1FE2D" box="[431,567,463,489]" country="India" name="Tamil Nadu" pageId="8" pageNumber="553">Tamil Nadu</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C2DD7FFBDFF8D2AE26CD3FAE1FD18" pageId="8" pageNumber="553" type="etymology">
|
||
<paragraph id="8B897E5CFFBDFF8D2AE26CD3FC76FDEA" blockId="8.[151,1437,151,732]" pageId="8" pageNumber="553">
|
||
<emphasis id="B942A24EFFBDFF8D2AE26CD3FEB8FDCF" bold="true" box="[199,334,497,523]" pageId="8" pageNumber="553">Etymology.</emphasis>
|
||
This species is named in honour of Dr. Wioletta Tomaszewska, Polish Academy of Sciences, Warsaw, for her significant contributions to
|
||
<taxonomicName id="4C3605DFFFBDFF8D28736F36FD06FDEA" authorityName="Latreille" authorityYear="1807" box="[598,752,532,558]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="family">Coccinellidae</taxonomicName>
|
||
systematics.
|
||
</paragraph>
|
||
<paragraph id="8B897E5CFFBDFF8D2AE26F15FAE1FD18" blockId="8.[151,1437,151,732]" pageId="8" pageNumber="553">
|
||
<emphasis id="B942A24EFFBDFF8D2AE26F15FEF0FD95" bold="true" box="[199,262,567,593]" pageId="8" pageNumber="553">Note.</emphasis>
|
||
A single specimen identified as
|
||
<taxonomicName id="4C3605DFFFBDFF8D28546F15FD15FD95" authorityName="Weise" authorityYear="1901" box="[625,739,567,593]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942A24EFFBDFF8D28546F15FD15FD95" box="[625,739,567,593]" italics="true" pageId="8" pageNumber="553">Horniolus</emphasis>
|
||
</taxonomicName>
|
||
nr.
|
||
<taxonomicName id="4C3605DFFFBDFF8D292B6F15FC9CFD94" box="[782,874,567,592]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="nigripes">
|
||
<emphasis id="B942A24EFFBDFF8D292B6F15FC9CFD94" box="[782,874,567,592]" italics="true" pageId="8" pageNumber="553">nigripes</emphasis>
|
||
</taxonomicName>
|
||
(BMNH,
|
||
<figureCitation id="130D62D9FFBDFF8D29F96F15FBD1FD96" box="[988,1063,567,594]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1965]" captionTargetBox="[219,1368,378,1915]" captionTargetId="figure-96@2.[219,1368,378,1915]" captionTargetPageId="2" captionText="FIGURE 1. Horniolus spp.: a–d. Horniolus dispar Weise: a, b. dorsal habitus; c, d. male genitalia: c. tegmen, ventral view; d. penis; e. Horniolus guimeti (Mulsant); f. Horniolus nigripes Miyatake (Non-type material from BMNH; Image credits for c, d. Keita Matsumoto, BMNH)." figureDoi="http://doi.org/10.5281/zenodo.6332183" httpUri="https://zenodo.org/record/6332183/files/figure.png" pageId="8" pageNumber="553">Fig. 1f</figureCitation>
|
||
) corresponds to this species. The specimen bears two identification labels, one as
|
||
<emphasis id="B942A24EFFBDFF8D28896F7BFCCAFDB7" box="[684,828,601,627]" italics="true" pageId="8" pageNumber="553">
|
||
“
|
||
<taxonomicName id="4C3605DFFFBDFF8D28936F7BFCDDFDB7" authorityName="Weise" authorityYear="1901" box="[694,811,601,627]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="genus">Horniolus</taxonomicName>
|
||
?
|
||
</emphasis>
|
||
<taxonomicName id="4C3605DFFFBDFF8D29186F7BFC24FDB7" authority="Weise" authorityName="Weise, R. G. Booth" authorityYear="1901" box="[829,978,601,627]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="dispar">
|
||
<emphasis id="B942A24EFFBDFF8D29186F7BFC70FDB7" box="[829,902,601,627]" italics="true" pageId="8" pageNumber="553">dispar</emphasis>
|
||
Weise
|
||
</taxonomicName>
|
||
, det. R. B. Madge 1985” and the other as “
|
||
<taxonomicName id="4C3605DFFFBDFF8D2A866F5EFE15FD52" authority="Miyatake" authorityName="Miyatake" box="[163,483,636,662]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="nigripes">
|
||
<emphasis id="B942A24EFFBDFF8D2A866F5EFE85FD52" box="[163,371,636,662]" italics="true" pageId="8" pageNumber="553">Horniolus nigripes</emphasis>
|
||
Miyatake
|
||
</taxonomicName>
|
||
, det. R.G. Booth 1985” [“
|
||
<collectingCountry id="F3213ECCFFBDFF8D28DE6F5EFCA4FD52" box="[763,850,636,662]" name="India" pageId="8" pageNumber="553">INDIA</collectingCountry>
|
||
: Chrys S.
|
||
<collectingRegion id="49F2B0BEFFBDFF8D29E46F5EFBFCFD52" box="[961,1034,636,662]" country="India" name="Kerala" pageId="8" pageNumber="553">Kerala</collectingRegion>
|
||
/ on Lagerstroemia sp., CIE
|
||
<accessionNumber id="9465E3BFFFBDFF8D2F666F5EFA6AFD53" box="[1347,1436,636,663]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A16725" pageId="8" pageNumber="553" type="EnaNcbi">A16725</accessionNumber>
|
||
/ Nymph on leaf, Lagerstroemia sp., Vazhani, 16.8.84, K.M. Das C.M., P.P., KFRI / genitalia in glass vial /
|
||
<taxonomicName id="4C3605DFFFBDFF8D2F0A6FBDFA6BFD7D" authorityName="Weise" authorityYear="1901" box="[1327,1437,671,697]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="genus">Horniolus</taxonomicName>
|
||
?
|
||
<taxonomicName id="4C3605DFFFBDFF8D2A846FE3FEC7FD1F" authority="Weise" authorityName="Weise, R. G. Booth" authorityYear="1901" box="[161,305,705,731]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="dispar">dispar Weise</taxonomicName>
|
||
, det. R. B. Madge 1985 /
|
||
<taxonomicName id="4C3605DFFFBDFF8D286C6FE3FD40FD1F" authorityName="Weise" authorityYear="1901" box="[585,694,705,731]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="genus">Horniolus</taxonomicName>
|
||
?
|
||
<taxonomicName id="4C3605DFFFBDFF8D28ED6FE3FC65FD1F" authority="Miyatake" authorityName="Miyatake" box="[712,915,705,731]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="nigripes">nigripes Miyatake</taxonomicName>
|
||
, det. R. G. Booth 1985” (BMNH)].
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |