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<document id="18D0BB9FCA76CD3F450CD8565D568C8C" ID-DOI="10.5852/ejt.2020.691" ID-GBIF-Dataset="918d86bb-f11d-414e-9c1e-21ec17737792" ID-ISSN="2118-9773" ID-Zenodo-Dep="3961840" ID-ZooBank="CC8487E0-169D-4B32-A2AB-F9B34DFB3F3B" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1595862647445" checkinUser="valdenar" docAuthor="Grego, Jozef" docDate="2020" docId="039F79458425FFFEBC00FC7965C72486" docLanguage="en" docName="ejt-691_grego.pdf" docOrigin="European Journal of Taxonomy 691" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Plagigeyeria inflata " docType="treatment" docVersion="11" lastPageNumber="12" masterDocId="FFA6013D842EFFF5BE26FFE6663C236D" masterDocTitle="Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego &amp; Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions" masterLastPageNumber="56" masterPageNumber="1" pageNumber="12" updateTime="1733502183861" updateUser="guilherme" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="18DB0D76EDDA731761B6B3C99DB02B06">Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego &amp; Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions</mods:title>
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<mods:namePart id="B454E837F4C0F6EC860857279B9F8238">Grego, Jozef</mods:namePart>
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<mods:affiliation id="8550A59270E8FF359F34017096B9C6E7">Horná Mičiná, SK- 97401 Banská Bystrica, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="FEA7801743CCEC5E4E1D80C16491CC5A" type="email">jozef.grego@gmail.com</mods:nameIdentifier>
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<mods:title id="5C940DD7A7CF1BDF55FFFE8D9368B2D1">European Journal of Taxonomy</mods:title>
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<mods:date id="475C6630657B0AFB2680F9B9A8D3BC3B">2020</mods:date>
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<treatment id="039F79458425FFFEBC00FC7965C72486" ID-DOI="http://doi.org/10.5281/zenodo.4336060" ID-GBIF-Taxon="166003120" ID-Zenodo-Dep="4336060" LSID="urn:lsid:plazi:treatment:039F79458425FFFEBC00FC7965C72486" httpUri="http://treatment.plazi.org/id/039F79458425FFFEBC00FC7965C72486" lastPageNumber="12" pageId="11" pageNumber="12" scope_class="Gastropoda" scope_family="Moitessieriidae" scope_phylum="Mollusca" scope_subFamily="Belgrandiellinae" scope_superFamily="Truncatelloidea">
<subSubSection id="C32C9BD88425FFFEBC00FC79623120D7" box="[550,1037,927,954]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B89C8538425FFFEBC00FC79623120D7" blockId="11.[550,1037,927,991]" box="[550,1037,927,954]" pageId="11" pageNumber="12">
<heading id="D0C17F3F8425FFFEBC00FC79623120D7" box="[550,1037,927,954]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<taxonomicName id="4C36B3D08425FFFEBC00FC79623120D7" ID-CoL="7SQKD" authority="(A. J. Wagner, 1928)" authorityName="" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1928" box="[550,1037,927,954]" class="Gastropoda" family="Hydrobiidae" genus="Plagigeyeria" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="inflata">
<emphasis id="B94214418425FFFEBC00FC79652F20D7" bold="true" box="[550,787,927,954]" italics="true" pageId="11" pageNumber="12">Plagigeyeria inflata</emphasis>
(A. J.
<bibRefCitation id="EFA7B5A28425FFFEBD47FC46623920D7" author="Wagner A. J." box="[865,1029,928,954]" pageId="11" pageNumber="12" pagination="263 - 264" refId="ref26540" refString="Wagner A. J. 1928. Studien der Molluskenfauna der Balkanhalbinsel mit besonderer Berucksichtigung Bulgariens and Thraziens, nebst monographisher Bearbeitung einzelner Gruppen. Prace Zoologiczne Polskiego Panstvowiego Muzeum Przyrodniczego, 1. XII. 1927, 6: 263 - 264." type="journal article" year="1928">Wagner, 1928</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32C9BD88425FFFEBCF4FC23655D20B3" box="[722,865,964,991]" pageId="11" pageNumber="12" type="description">
<paragraph id="8B89C8538425FFFEBCF4FC23655D20B3" blockId="11.[550,1037,927,991]" box="[722,865,964,991]" pageId="11" pageNumber="12">
<figureCitation id="130DD4D68425FFFEBCF4FC23650D20B2" box="[722,817,964,991]" captionStart="Fig" captionStartId="9.[189,230,1754,1780]" captionTargetBox="[189,1398,265,1918]" captionTargetId="figure@10.[189,1398,265,1918]" captionTargetPageId="10" captionText="Fig. 5 (next page). A. Plagigeyeria plagiostoma (A. J. Wagner, 1914), from Vrelo Bosne in Ilidža, (topotype JG F1116). B. Travunijana vruljakensis Grego &amp; Glöer, 2019, Gorica near Trebinje, spring Studenac (holotype HNHM-MOLL-104416). C. P. inflata (A. J. Wagner, 1928), from Vrelo Bosne in Ilidža (topotype JG F1117). D. P. ljutaensis sp. nov., Konjic district, spring of River Ljuta (holotype HNHM-MOLL-104180). E. P. konjicensis sp. nov., small spring at left bank of Ljuta River, 600 m downstream of the main spring (holotype HNHM-MOLL-104174); F–G. P. olsavskyi sp. nov., Studenci, spring Kajtazovina F. Holotype (HNHM-MOLL-104178). G. Paratype (JG F1192). H–I. P. olsavskyi sp. nov., Ljubuški district, Donji Proložac, spring Mali Prokop (JG F1209)." figureDoi="http://doi.org/10.5281/zenodo.3962586" httpUri="https://zenodo.org/record/3962586/files/figure.png" pageId="11" pageNumber="12">Figs 5C</figureCitation>
,
<figureCitation id="130DD4D68425FFFEBD1BFC22655D20B3" box="[829,865,964,990]" captionStart="Fig" captionStartId="14.[193,236,1895,1921]" captionTargetBox="[246,1342,268,1814]" captionTargetId="figure@14.[246,1342,268,1818]" captionTargetPageId="14" captionText="Fig. 6. A. Plagigeyeria ljutaensissp. nov., Konjic,Vrelo Ljuta, paratype (SBMNH 626401).B. P.konjicensis sp. nov., Konjic, left side spring of Ljuta River, paratype (SBMNH 626249). C. P. plagiostoma (A. J. Wagner, 1914), Sarajevo, Ilidža, Vrelo Bosne, (SBMNH 632718). D. P. inflata (A. J. Wagner, 1928), Sarajevo, Ilidža, Vrelo Bosne (SBMNH 33032). Scale bars = 1 mm (SEM SBMNH Vanessa Delnavaz)." figureDoi="http://doi.org/10.5281/zenodo.3961848" httpUri="https://zenodo.org/record/3961848/files/figure.png" pageId="11" pageNumber="12">6D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32C9BD88425FFFEBE9BFBEE633F27E6" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8B89C8538425FFFEBE9BFBEE6204274F" blockId="11.[189,1080,1032,1059]" box="[189,1080,1032,1059]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB26EF7D8425FFFEBE9BFBEE6204274F" box="[189,1080,1032,1059]" pageId="11" pageNumber="12">
<treatmentCitation id="0A97EE428425FFFEBE9BFBEE6510274E" author="Wagner A. J." box="[189,812,1032,1059]" page="285" pageId="11" pageNumber="12" year="1928">
<taxonomicName id="4C36B3D08425FFFEBE9BFBEE6510274E" ID-CoL="7TVKH" authority="A. J. Wagner, 1928: 285" authorityName="A. J. Wagner" authorityPageNumber="285" authorityYear="1928" box="[189,812,1032,1059]" class="Gastropoda" family="Hydrobiidae" genus="Geyeria" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="subSpecies" species="plagiostoma" subSpecies="inflata">
<emphasis id="B94214418425FFFEBE9BFBEE643A274F" box="[189,518,1032,1058]" italics="true" pageId="11" pageNumber="12">Geyeria plagiostoma inflata</emphasis>
A. J.
<bibRefCitation id="EFA7B5A28425FFFEBC6DFBEE6510274E" author="Wagner A. J." box="[587,812,1032,1059]" pageId="11" pageNumber="12" pagination="263 - 264" refId="ref26540" refString="Wagner A. J. 1928. Studien der Molluskenfauna der Balkanhalbinsel mit besonderer Berucksichtigung Bulgariens and Thraziens, nebst monographisher Bearbeitung einzelner Gruppen. Prace Zoologiczne Polskiego Panstvowiego Muzeum Przyrodniczego, 1. XII. 1927, 6: 263 - 264." type="journal article" year="1928">Wagner, 1928: 285</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 13. figs 68–69, 72.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B89C8538425FFFEBE9BFBAD65B22708" blockId="11.[189,1283,1099,1163]" box="[189,910,1099,1126]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB26EF7D8425FFFEBE9BFBAD65B22708" box="[189,910,1099,1126]" pageId="11" pageNumber="12">
<taxonomicName id="4C36B3D08425FFFEBE9BFBAD64642708" ID-CoL="6VMZL" authority="(A. J. Wagner, 1928)" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1928" box="[189,600,1099,1125]" class="Gastropoda" family="Hydrobiidae" form="inflata" genus="Plagigeyeria" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="form" species="plagiostoma">
<emphasis id="B94214418425FFFEBE9BFBAD67D12708" box="[189,493,1099,1125]" italics="true" pageId="11" pageNumber="12">Plagigeyeria plagiostoma</emphasis>
f.
<emphasis id="B94214418425FFFEBC2BFBAD64642708" box="[525,600,1099,1125]" italics="true" pageId="11" pageNumber="12">inflata</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A97EE428425FFFEBC53FBAD65B22708" box="[629,910,1099,1126]" pageId="11" pageNumber="12">Schütt 1972: 119, fig. 2.</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B89C8538425FFFEBE9BFB96633F27E6" blockId="11.[189,1283,1099,1163]" box="[189,1283,1136,1163]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB26EF7D8425FFFEBE9BFB96633F27E6" box="[189,1283,1136,1163]" pageId="11" pageNumber="12">
<taxonomicName id="4C36B3D08425FFFEBE9BFB9667D127E7" ID-CoL="6VMZL" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1914" box="[189,493,1136,1162]" class="Gastropoda" family="Hydrobiidae" genus="Plagigeyeria" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="plagiostoma">
<emphasis id="B94214418425FFFEBE9BFB9667D127E7" box="[189,493,1136,1162]" italics="true" pageId="11" pageNumber="12">Plagigeyeria plagiostoma</emphasis>
</taxonomicName>
–
<bibRefCitation id="EFA7B5A28425FFFEBC2CFB9664B227E7" author="Bank R. A." box="[522,654,1136,1162]" pageId="11" pageNumber="12" refId="ref24339" refString="Bank R. A. 2013. Fauna Europaea: Mollusca. Fauna Europaea version 2.6. 2. https: // fauna-eu. org / [accessed 11 Nov. 2019]" type="url" year="2013">Bank 2013</bibRefCitation>
: Fauna Europaea v. 2.6 —
<treatmentCitation id="0A97EE428425FFFEBDEDFB9662C327E6" author="Bank R. A. &amp; Neubert E." box="[971,1279,1136,1163]" page="25" pageId="11" pageNumber="12" year="2017">
<bibRefCitation id="EFA7B5A28425FFFEBDEDFB9662C327E6" author="Bank R. A. &amp; Neubert E." box="[971,1279,1136,1163]" pageId="11" pageNumber="12" refId="ref24372" refString="Bank R. A. &amp; Neubert E. 2017. Checklist of the land and freshwater Gastropoda of Europe. Available from http: // www. molluscabase. org / aphia. php? p = sourcedetails &amp; id = 279050 [last update: 16 Jul. 2017; accessed 11 Nov. 2019]." type="url" year="2017">Bank &amp; Neubert 2017: 25</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32C9BD88425FFFEBE9BFB5567352600" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8B89C8538425FFFEBE9BFB55670A27A0" blockId="11.[189,310,1203,1229]" box="[189,310,1203,1229]" pageId="11" pageNumber="12">
<heading id="D0C17F3F8425FFFEBE9BFB55670A27A0" bold="true" box="[189,310,1203,1229]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B94214418425FFFEBE9BFB55670A27A0" bold="true" box="[189,310,1203,1229]" pageId="11" pageNumber="12">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B89C8538425FFFEBE9BFB0267352600" blockId="11.[189,1398,1251,1389]" pageId="11" pageNumber="12">
The species can be separated from
<taxonomicName id="4C36B3D08425FFFEBC45FB0265A92793" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1914" box="[611,917,1252,1278]" class="Gastropoda" family="Hydrobiidae" genus="Plagigeyeria" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="plagiostoma">
<emphasis id="B94214418425FFFEBC45FB0265A92793" box="[611,917,1252,1278]" italics="true" pageId="11" pageNumber="12">Plagigeyeria plagiostoma</emphasis>
</taxonomicName>
by its larger shell (
<quantity id="4CCE65B68425FFFEBAA6FB0562E12793" box="[1152,1245,1251,1278]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="12" unit="mm" value="2.5">2.5 mm</quantity>
high against
<quantity id="4CCE65B68425FFFEBE9BFAEF6724264E" box="[189,280,1289,1315]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="12" unit="mm" value="1.9">1.9 mm</quantity>
) with a more robust and more conical shell shape with more open umbilicus and less blunt apex, by larger, more expanded aperture and by coarser ribs with different cancellate surface of the nepionic whorl.
</paragraph>
</subSubSection>
<subSubSection id="C32C9BD88425FFFEBE9BFA706325268D" box="[189,1305,1430,1505]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8B89C8538425FFFEBE9BFA70676A26DD" blockId="11.[189,342,1430,1456]" box="[189,342,1430,1456]" pageId="11" pageNumber="12">
<heading id="D0C17F3F8425FFFEBE9BFA70676A26DD" bold="true" box="[189,342,1430,1456]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B94214418425FFFEBE9BFA70676A26DD" bold="true" box="[189,342,1430,1456]" pageId="11" pageNumber="12">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B89C8538425FFFEBE9BFA206325268D" blockId="11.[189,1305,1478,1505]" box="[189,1305,1478,1505]" pageId="11" pageNumber="12">
Only known from the
<typeStatus id="548D76F18425FFFEBFE4FA2167C9268C" box="[450,501,1479,1505]" pageId="11" pageNumber="12">type</typeStatus>
locality in Vrelo Bosne (
<quantity id="4CCE65B68425FFFEBD3CFA20655A268C" box="[794,870,1478,1505]" metricMagnitude="2" metricUnit="m" metricValue="5.81" pageId="11" pageNumber="12" unit="m" value="581.0">581 m</quantity>
a.s.l.) near Ilidža, South of Sarajevo.
</paragraph>
</subSubSection>
<subSubSection id="C32C9BD88425FFFEBE9BF9EC65C72486" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8B89C8538425FFFEBE9BF9EC67132549" blockId="11.[189,303,1546,1572]" box="[189,303,1546,1572]" pageId="11" pageNumber="12">
<heading id="D0C17F3F8425FFFEBE9BF9EC67132549" bold="true" box="[189,303,1546,1572]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B94214418425FFFEBE9BF9EC67132549" bold="true" box="[189,303,1546,1572]" pageId="11" pageNumber="12">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B89C8538425FFFEBE9BF9DC65C72486" blockId="11.[189,1399,1594,2027]" pageId="11" pageNumber="12">
<taxonomicName id="4C36B3D08425FFFEBE9BF9DC64942539" authority="(A. J. Wagner, 1928)" authorityName="" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1928" box="[189,680,1594,1620]" class="Gastropoda" family="Hydrobiidae" genus="Plagigeyeria" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="inflata">
<emphasis id="B94214418425FFFEBE9BF9DC67942539" box="[189,424,1594,1620]" italics="true" pageId="11" pageNumber="12">Plagigeyeria inflata</emphasis>
(A. J.
<bibRefCitation id="EFA7B5A28425FFFEBFDCF9DC649C2539" author="Wagner A. J." box="[506,672,1594,1620]" pageId="11" pageNumber="12" pagination="263 - 264" refId="ref26540" refString="Wagner A. J. 1928. Studien der Molluskenfauna der Balkanhalbinsel mit besonderer Berucksichtigung Bulgariens and Thraziens, nebst monographisher Bearbeitung einzelner Gruppen. Prace Zoologiczne Polskiego Panstvowiego Muzeum Przyrodniczego, 1. XII. 1927, 6: 263 - 264." type="journal article" year="1928">Wagner, 1928</bibRefCitation>
)
</taxonomicName>
was described from the same spring where the empty shells of
<taxonomicName id="4C36B3D08425FFFEBEC6F98667AC2514" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1914" box="[224,400,1631,1657]" class="Gastropoda" family="Hydrobiidae" genus="Plagigeyeria" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="plagiostoma">
<emphasis id="B94214418425FFFEBEC6F98667AC2514" box="[224,400,1631,1657]" italics="true" pageId="11" pageNumber="12">P. plagiostoma</emphasis>
</taxonomicName>
had been present and, supposing their subspecific status and identical locality, the two species were synonymized by Schütt (1972) and then again in the Fauna Europaea (
<bibRefCitation id="EFA7B5A28425FFFEBAC5F962635A25F3" author="Bank R. A." box="[1251,1382,1668,1694]" pageId="11" pageNumber="12" refId="ref24339" refString="Bank R. A. 2013. Fauna Europaea: Mollusca. Fauna Europaea version 2.6. 2. https: // fauna-eu. org / [accessed 11 Nov. 2019]" type="url" year="2013">Bank 2013</bibRefCitation>
). Nevertheless, I have to note their conchological distinctness without intermediates, and that both species were found in the same locality only as empty shells washed together within the recent thanatocoenoses of the spring zone. We have no idea about the underground aquifer range and about its mutually separated cave branches. It is very common in the morphology of many spring caves, that the large isolated, hydrologically active cave passages converge their flow just upstream of the spring outlet. Thus we cannot exclude the presence of two separated species or populations existing within one cave system without being sympatric. The recent thanatocoenoses do not represent their true habitat. Both species are very well differentiated by their shell shape and size (
<bibRefCitation id="EFA7B5A28425FFFEBDEAF84A625624AB" author="Wagner A. J." box="[972,1130,1964,1990]" pageId="11" pageNumber="12" pagination="263 - 264" refId="ref26540" refString="Wagner A. J. 1928. Studien der Molluskenfauna der Balkanhalbinsel mit besonderer Berucksichtigung Bulgariens and Thraziens, nebst monographisher Bearbeitung einzelner Gruppen. Prace Zoologiczne Polskiego Panstvowiego Muzeum Przyrodniczego, 1. XII. 1927, 6: 263 - 264." type="journal article" year="1928">Wagner 1928</bibRefCitation>
; Schütt 1972) without any intermediate and I prefer to treat them as two independent species.
</paragraph>
</subSubSection>
</treatment>
</document>
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