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<document id="335A4AED0BF155CC63B065140A48C5D1" ID-CLB-Dataset="53415" ID-DOI="10.11646/zootaxa.5296.2.1" ID-GBIF-Dataset="06dd5db0-bae0-40fe-8c7f-14b6d87ceabc" ID-ISSN="1175-5326" ID-Zenodo-Dep="7978135" ID-ZooBank="87CF3B54-A6EE-4650-8C4B-0D7378CB32C3" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1685085826471" checkinUser="plazi" docAuthor="Theischinger, Günther, Mitchell, Andrew, Richards, Stephen J. &amp; Polhemus, Dan A." docDate="2023" docId="039A8912ED68FF95B199EBDADE11F99D" docLanguage="en" docName="zootaxa.5296.2.1.pdf" docOrigin="Zootaxa 5296 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nososticta tagula Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="129" masterDocId="FFA3F16AED73FF89B10EEB4DDD38FFF3" masterDocTitle="Systematics of the Nososticta salomonis complex (Odonata: Zygoptera: Platycnemididae)" masterLastPageNumber="146" masterPageNumber="101" pageNumber="128" updateTime="1685370491969" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D9A1888EEE1E38C5DCDC9869C9FD581F">Systematics of the Nososticta salomonis complex (Odonata: Zygoptera: Platycnemididae)</mods:title>
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<mods:namePart id="8BEF15FAE900BD8EB9D271FA0BEB20E1">Mitchell, Andrew</mods:namePart>
<mods:affiliation id="F65F0D14C3617B30F259F33DC2F123D6">Australian Museum, Entomology, 1 William Street, Sydney, N. S. W. 2010, Australia</mods:affiliation>
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<mods:namePart id="90762E9A243F57514091B117670BE224">Richards, Stephen J.</mods:namePart>
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<mods:affiliation id="E0E64EF3E65692CCA5562B09AF8C97BD">Herpetology Department, South Australian Museum, North Terrace, Adelaide, S. A. 5000, Australia steve. richards @ samuseum. sa. gov. au; https: // orcid. org / 0000 - 0002 - 0251 - 3884</mods:affiliation>
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<mods:namePart id="5160A18C453A1104D9E74547D5070E7E">Polhemus, Dan A.</mods:namePart>
<mods:affiliation id="2527DE0C0B876C6D184B949DFFDD1F73">Dept. of Natural Sciences, Bishop Museum, 1525 Bernice St., Honolulu, HI, 96817 USA</mods:affiliation>
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<treatment id="039A8912ED68FF95B199EBDADE11F99D" ID-DOI="http://doi.org/10.5281/zenodo.7982156" ID-Zenodo-Dep="7982156" LSID="urn:lsid:plazi:treatment:039A8912ED68FF95B199EBDADE11F99D" httpUri="http://treatment.plazi.org/id/039A8912ED68FF95B199EBDADE11F99D" lastPageId="28" lastPageNumber="129" pageId="27" pageNumber="128">
<subSubSection id="C3296B8FED68FF92B199EBDADCF6FF42" box="[151,462,151,178]" pageId="27" pageNumber="128" type="nomenclature">
<paragraph id="8B8C3804ED68FF92B199EBDADCF6FF42" blockId="27.[151,531,151,214]" box="[151,462,151,178]" pageId="27" pageNumber="128">
<heading id="D0C48F68ED68FF92B199EBDADCF6FF42" bold="true" box="[151,462,151,178]" fontSize="11" level="1" pageId="27" pageNumber="128" reason="1">
<emphasis id="B947E416ED68FF92B199EBDADCF6FF42" bold="true" box="[151,462,151,178]" pageId="27" pageNumber="128">
<taxonomicName id="4C334387ED68FF92B199EBDADC51FF41" authority="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus, 2023" authorityName="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus" authorityYear="2023" box="[151,361,151,178]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="tagula" status="sp. nov.">
<emphasis id="B947E416ED68FF92B199EBDADC51FF41" bold="true" box="[151,361,151,178]" italics="true" pageId="27" pageNumber="128">Nososticta tagula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A274596DED68FF92B07EEBDADCF6FF42" box="[368,462,151,177]" pageId="27" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3296B8FED68FF92B199EBF6DF2BFF26" box="[151,531,187,214]" pageId="27" pageNumber="128" type="description">
<paragraph id="8B8C3804ED68FF92B199EBF6DF2BFF26" blockId="27.[151,531,151,214]" box="[151,531,187,214]" pageId="27" pageNumber="128">
<emphasis id="B947E416ED68FF92B199EBF6DD98FF26" bold="true" box="[151,160,187,213]" pageId="27" pageNumber="128">(</emphasis>
<figureCitation id="13082481ED68FF92B1AFEBF6DC19FF25" box="[161,289,187,214]" captionStart="FIGURES 5054" captionStartId="27.[151,264,1406,1430]" captionTargetBox="[151,1435,634,1383]" captionTargetId="figure-163@27.[151,1435,633,1383]" captionTargetPageId="27" captionText="FIGURES 5054. Nososticta tagula sp. nov. (5052, 54) male: (50) synthorax, schematically; (51) abdominal segment 2, dorsal; (52) superior anal appendage; (53) female, synthorax, schematically; (54) specimen as photographed in ethanol." figureDoi="http://doi.org/10.5281/zenodo.7978171" httpUri="https://zenodo.org/record/7978171/files/figure.png" pageId="27" pageNumber="128">Figs 5054</figureCitation>
,
<figureCitation id="13082481ED68FF92B022EBF6DC4CFF25" box="[300,372,187,214]" captionStart="FIGURES 5565" captionStartId="29.[151,264,1909,1933]" captionTargetBox="[169,1415,194,1879]" captionTargetId="figure-22@29.[169,1419,181,1884]" captionTargetPageId="29" captionText="FIGURES 5565. Nososticta species. (55, 56) male abdominal segment 2, dorsal: (55) N. africana (Schmidt); (56) N. tagula sp. nov.; (5765) synthorax, schematically: (57) N. africana (Schmidt), female; (58) N. tagula sp. nov., female; (59, 60) N. hedigeri sp. nov.: (59) male; (60) female; (61, 62) N. stueberi sp. nov.: (61) male; (62) female; (63) N. chrismulleri Theischinger &amp; Richards, male; (64, 65) N. boonei sp. nov.: (64) female; (65) male." figureDoi="http://doi.org/10.5281/zenodo.7978175" httpUri="https://zenodo.org/record/7978175/files/figure.png" pageId="27" pageNumber="128">56, 58</figureCitation>
,
<figureCitation id="13082481ED68FF92B08EEBF1DCF1FF26" box="[384,457,187,214]" captionStart="MAP 3" captionStartId="34.[151,207,1000,1024]" captionTargetBox="[153,1433,183,974]" captionTargetId="figure-42@34.[151,1435,181,976]" captionText="MAP 3.—Distribution of Nososticta tagula sp. nov. in the Louisiade Archipelago." figureDoi="http://doi.org/10.5281/zenodo.7982146" httpUri="https://zenodo.org/record/7982146/files/figure.png" pageId="27" pageNumber="128">Map 3</figureCitation>
, Pl. 6)
</paragraph>
</subSubSection>
<subSubSection id="C3296B8FED68FF92B199EA4EDCEFFE5E" pageId="27" pageNumber="128" type="materials_examined">
<paragraph id="8B8C3804ED68FF92B199EA4EDE51FE96" blockId="27.[151,1436,259,609]" pageId="27" pageNumber="128">
<materialsCitation id="3B5B3259ED68FF92B199EA4EDE51FE96" collectingDate="2002-08-29" collectionCode="T" collectorName="Tagula Is." country="Papua New Guinea" elevation="15" latitude="-11.364445" location="Kalitau Creek" longLatPrecision="21" longitude="153.24055" pageId="27" pageNumber="128" specimenCount="1" stateProvince="Milne Bay" typeStatus="holotype">
<emphasis id="B947E416ED68FF92B199EA4EDC33FEEE" bold="true" box="[151,267,259,285]" pageId="27" pageNumber="128">
<typeStatus id="548886A6ED68FF92B199EA4EDC3EFEEE" box="[151,262,259,285]" pageId="27" pageNumber="128">Holotype</typeStatus>
.
</emphasis>
♁, BPBMENT 0000081204,
<collectingCountry id="F3247894ED68FF92B36EEA49DE07FEEE" box="[608,831,259,286]" name="Papua New Guinea" pageId="27" pageNumber="128">Papua New Guinea</collectingCountry>
,
<collectingRegion id="49F7F6E6ED68FF92B242EA4ED932FEED" box="[844,1034,259,286]" country="Papua New Guinea" name="Milne Bay" pageId="27" pageNumber="128">Milne Bay Prov.</collectingRegion>
, Louisiade Archipelago,
<collectorName id="26C65DD2ED68FF92B428EA4ED8AFFEED" box="[1318,1431,259,286]" pageId="27" pageNumber="128">Tagula Is.</collectorName>
,
<location id="8EEC6EDFED68FF92B199EA6ADC0DFEB2" LSID="urn:lsid:plazi:treatment:039A8912ED68FF95B199EBDADE11F99D:8EEC6EDFED68FF92B199EA6ADC0DFEB2" box="[151,309,295,321]" country="Papua New Guinea" latitude="-11.364445" longLatPrecision="21" longitude="153.24055" name="Kalitau Creek" pageId="27" pageNumber="128" stateProvince="Milne Bay">Kalitau Creek</location>
, above estuary,
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<quantity id="4CCB95E1ED68FF92B0EFEA6ADF0DFEB1" box="[481,565,295,322]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="27" pageNumber="128" unit="m" value="15.0" valueMax="30.0" valueMin="0.0">030 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE075EC3ED68FF92B360EA6ADFC2FEB1" box="[622,762,295,322]" degrees="11" direction="south" minutes="21" orientation="latitude" pageId="27" pageNumber="128" precision="15" seconds="52" value="-11.364445">11°2152”S</geoCoordinate>
,
<geoCoordinate id="EE075EC3ED68FF92B20AEA6ADEA4FEB1" box="[772,924,295,322]" degrees="153" direction="east" minutes="14" orientation="longitude" pageId="27" pageNumber="128" precision="15" seconds="26" value="153.24055">153°1426”E</geoCoordinate>
;
<date id="FF8D1EC4ED68FF92B2ABEA6AD90CFEB2" box="[933,1076,295,321]" pageId="27" pageNumber="128" value="2002-08-29">
<collectingDate id="EFC9E72CED68FF92B2ABEA6AD90CFEB2" box="[933,1076,295,321]" pageId="27" pageNumber="128" value="2002-08-29">29-viii-2002</collectingDate>
</date>
, CL 7190, D.A. &amp; J.
<collectionCode id="ED22A0C1ED68FF92B41CEA65D819FEB1" box="[1298,1313,296,322]" name="Tavera, Department of Geology and Geophysics" pageId="27" pageNumber="128">T</collectionCode>
. Polhemus (CL 7190); deposited in the Bishop Museum, Honolulu, Hawaii.
</materialsCitation>
</paragraph>
<paragraph id="8B8C3804ED68FF92B1C9EA22DCEFFE5E" blockId="27.[151,1436,259,609]" pageId="27" pageNumber="128">
<materialsCitation id="3B5B3259ED68FF92B1C9EA22DCEBFE5E" collectingDate="2002-08-29" collectionCode="T" collectorName="Tagula Is." country="Papua New Guinea" elevation="15" latitude="-11.364445" location="Honolulu" longLatPrecision="21" longitude="153.24055" municipality="Bishop Museum" pageId="27" pageNumber="128" specimenCount="1" specimenCount-female="1" stateProvince="Hawaii" typeStatus="paratype">
<emphasis id="B947E416ED68FF92B1C9EA22DC7EFE7A" bold="true" box="[199,326,367,393]" pageId="27" pageNumber="128">
<typeStatus id="548886A6ED68FF92B1C9EA22DC79FE7A" box="[199,321,367,393]" pageId="27" pageNumber="128">Paratypes</typeStatus>
.
</emphasis>
6 ♁♁,
<specimenCount id="9D35F38DED68FF92B0A7EA22DCE0FE79" box="[425,472,367,394]" pageId="27" pageNumber="128" type="female">1 ♀</specimenCount>
, BPBMENT 000008120581211, same data as holotype; deposited in the
<collectingMunicipality id="6BE8A27EED68FF92B443EA22DDC4FE5D" pageId="27" pageNumber="128">Bishop Museum</collectingMunicipality>
,
<location id="8EEC6EDFED68FF92B009EADEDC4CFE5E" LSID="urn:lsid:plazi:treatment:039A8912ED68FF95B199EBDADE11F99D:8EEC6EDFED68FF92B009EADEDC4CFE5E" box="[263,372,403,429]" municipality="Bishop Museum" name="Honolulu" pageId="27" pageNumber="128" stateProvince="Hawaii">Honolulu</location>
,
<collectingRegion id="49F7F6E6ED68FF92B071EADEDCEBFE5E" box="[383,467,403,429]" country="United States of America" name="Hawaii" pageId="27" pageNumber="128">Hawaii</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3296B8FED68FF92B1C9EAFADCCFFE05" pageId="27" pageNumber="128" type="etymology">
<paragraph id="8B8C3804ED68FF92B1C9EAFADCCFFE05" blockId="27.[151,1436,259,609]" pageId="27" pageNumber="128">
<emphasis id="B947E416ED68FF92B1C9EAFADC76FE22" bold="true" box="[199,334,439,465]" pageId="27" pageNumber="128">Etymology.</emphasis>
Tagula after the
<typeStatus id="548886A6ED68FF92B302EAF5DF04FE21" box="[524,572,440,466]" pageId="27" pageNumber="128">type</typeStatus>
locality Tagula Island in the Louisiade Archipelago; it is treated as a noun in apposition to the generic name.
</paragraph>
</subSubSection>
<subSubSection id="C3296B8FED68FF95B1C9EAB2D966FABD" lastPageId="28" lastPageNumber="129" pageId="27" pageNumber="128" type="description">
<paragraph id="8B8C3804ED68FF92B1C9EAB2DDF5FD92" blockId="27.[151,1436,259,609]" pageId="27" pageNumber="128">
<emphasis id="B947E416ED68FF92B1C9EAB2DCE3FDEA" bold="true" box="[199,475,511,537]" pageId="27" pageNumber="128">
Description of
<typeStatus id="548886A6ED68FF92B07AEAB2DCE3FDEA" box="[372,475,511,537]" pageId="27" pageNumber="128">holotype</typeStatus>
</emphasis>
(all available specimens (type series) apparently turned pale by the way of preservation (ethanol), meaning that in particular the larger now pale greyish blue areas would have been largely bright blue in life).
</paragraph>
<caption id="DF4C688CED68FF92B199EE33D869FA48" ID-DOI="http://doi.org/10.5281/zenodo.7978171" ID-Zenodo-Dep="7978171" httpUri="https://zenodo.org/record/7978171/files/figure.png" pageId="27" pageNumber="128" startId="27.[151,264,1406,1430]" targetBox="[151,1435,634,1383]" targetPageId="27" targetType="figure">
<paragraph id="8B8C3804ED68FF92B199EE33D869FA48" blockId="27.[151,1437,1406,1467]" pageId="27" pageNumber="128">
<emphasis id="B947E416ED68FF92B199EE33DC6FFA65" bold="true" box="[151,343,1406,1430]" pageId="27" pageNumber="128">FIGURES 5054.</emphasis>
<taxonomicName id="4C334387ED68FF92B06EEE32DF2EFA64" authority="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus, 2023" authorityName="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus" authorityYear="2023" box="[352,534,1407,1431]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="tagula" status="sp. nov.">
<emphasis id="B947E416ED68FF92B06EEE32DF2EFA64" box="[352,534,1407,1431]" italics="true" pageId="27" pageNumber="128">Nososticta tagula</emphasis>
</taxonomicName>
<emphasis id="B947E416ED68FF92B311EE32DF4AFA64" bold="true" box="[543,626,1407,1431]" pageId="27" pageNumber="128">
<taxonomicNameLabel id="A274596DED68FF92B311EE32DF4AFA64" box="[543,626,1407,1431]" pageId="27" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B947E416ED68FF92B372EE32DFCDFA64" box="[636,757,1407,1431]" italics="true" pageId="27" pageNumber="128">(5052, 54)</emphasis>
male:
<emphasis id="B947E416ED68FF92B24FEE32DE53FA64" box="[833,875,1407,1431]" italics="true" pageId="27" pageNumber="128">(50)</emphasis>
synthorax, schematically;
<emphasis id="B947E416ED68FF92B58DEE32D996FA64" box="[1155,1198,1407,1431]" italics="true" pageId="27" pageNumber="128">(51)</emphasis>
abdominal segment 2, dorsal;
<emphasis id="B947E416ED68FF92B1ECEEEEDC35FA48" box="[226,269,1443,1467]" italics="true" pageId="27" pageNumber="128">(52)</emphasis>
superior anal appendage;
<emphasis id="B947E416ED68FF92B319EEEEDF79FA48" box="[535,577,1443,1467]" italics="true" pageId="27" pageNumber="128">(53)</emphasis>
female, synthorax, schematically;
<emphasis id="B947E416ED68FF92B2ACEEEEDEF4FA48" box="[930,972,1443,1467]" italics="true" pageId="27" pageNumber="128">(54)</emphasis>
specimen as photographed in ethanol.
</paragraph>
</caption>
<paragraph id="8B8C3804ED68FF92B1C9ED40DDDEF960" blockId="27.[151,1437,1549,2043]" pageId="27" pageNumber="128">
<emphasis id="B947E416ED68FF92B1C9ED40DC33F9D4" box="[199,267,1549,1575]" italics="true" pageId="27" pageNumber="128">Head.</emphasis>
Largely black; base of labium pale brownish yellow, anteclypeus greyish yellow marbled with grey, pleurostoma and transverse frontal bar from eye to eye very pale greyish blue, an ill-defined mark between antennal base and lateral ocellus pale brown as are apex of largely blackish brown scape and pedicel, antennal flagellum dark brown.
</paragraph>
<paragraph id="8B8C3804ED68FF95B1C9EDD0D801FF26" blockId="27.[151,1437,1549,2043]" lastBlockId="28.[151,1437,151,429]" lastPageId="28" lastPageNumber="129" pageId="27" pageNumber="128">
<emphasis id="B947E416ED68FF92B1C9EDD0DC20F944" box="[199,280,1693,1719]" italics="true" pageId="27" pageNumber="128">Thorax</emphasis>
. Pronotum black, anterior lobe with two pale greyish yellow to bluish lateral spots on each side, the larger inner more yellowish spot detached and smaller outer more bluish spot only marginally separated from large mark of same colour on median lobe; propleura largely pale greyish blue, black along sutures and at rear of epimeron. Synthorax with pleura largely black except following pale greyish blue areas: ante-humeral patch, very slightly longer than 1/2 length of mesanepisternum, approximately twice as wide basally as dorsally, tip evenly rounded; patch covering about dorsal 4/5 length of mesepimeron, partly connected with stripe across whole length and anterior 3/4 to 4/5 of metepisternum; a subtriangular narrow-tailed patch across posterior half of metepimeron and posterolateral tip of meso- and metakatepisternum. Postcoxae blackish brown to black and pale greyish blue; poststernum pale brownish blue with ill-defined small, narrow patches of greyish brown. Legs: Coxae largely pale greyish blue, meso-and metacoxa anterobasally brown to black, trochanters black and pale greyish blue, legs otherwise black, with only very base of femora and part of basal 1/2 to 2/3 of metafemur pale greyish blue. Wings: membrane almost clear, venation black; pterostigma greyish brown, slightly more than twice as long as wide; postnodals 16/1314.
</paragraph>
<paragraph id="8B8C3804ED6FFF95B1C9EB92DF4DFE7A" blockId="28.[151,1437,151,429]" pageId="28" pageNumber="129">
<emphasis id="B947E416ED6FFF95B1C9EB92DC01FF0A" box="[199,313,223,249]" italics="true" pageId="28" pageNumber="129">Abdomen.</emphasis>
S1 dorsally largely black, laterally pale greyish blue; S2 largely black, with distinct pale greyish blue apical dorsal mark, and pale greyish blue adjacent to genitalia; S37 dorsally black with blue basal mid-dorsal mark, the one on S7 very short, all ventrally margined greyish brown to dull yellow; S8 largely blue, narrowly lined black basally, more widely and irregularly lined black apically; S9 and S10 black. Superior anal appendages largely blue, inferiors brownish grey to brownish black.
</paragraph>
<paragraph id="8B8C3804ED6FFF95B1C9EADED95DFE5E" blockId="28.[151,1437,151,429]" box="[199,1125,403,429]" pageId="28" pageNumber="129">
<emphasis id="B947E416ED6FFF95B1C9EADEDC6BFE5F" box="[199,339,403,428]" italics="true" pageId="28" pageNumber="129">Dimensions.</emphasis>
Hind wing
<quantity id="4CCB95E1ED6FFF95B0D9EADEDF06FE5E" box="[471,574,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.88" pageId="28" pageNumber="129" unit="mm" value="18.8">18.8 mm</quantity>
; abdomen (including anal appendages)
<quantity id="4CCB95E1ED6FFF95B2F4EADED959FE5E" box="[1018,1121,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" pageId="28" pageNumber="129" unit="mm" value="31.3">31.3 mm</quantity>
.
</paragraph>
<paragraph id="8B8C3804ED6FFF95B199EAB3DC9AFDEB" blockId="28.[151,418,510,536]" box="[151,418,510,536]" pageId="28" pageNumber="129">
<heading id="D0C48F68ED6FFF95B199EAB3DC9AFDEB" bold="true" box="[151,418,510,536]" fontSize="11" level="1" pageId="28" pageNumber="129" reason="1">
<emphasis id="B947E416ED6FFF95B199EAB3DC9AFDEB" bold="true" box="[151,418,510,536]" pageId="28" pageNumber="129">Description of female</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C3804ED6FFF95B199E90DD920FD8D" blockId="28.[151,1437,576,1647]" pageId="28" pageNumber="129">
<emphasis id="B947E416ED6FFF95B199E90DDDE3FDA9" box="[151,219,576,602]" italics="true" pageId="28" pageNumber="129">Head.</emphasis>
Much as in male, but dark structures and areas darker, pale areas largely pale greyish yellow instead of pale greyish blue; no pale mark between antennal base and lateral ocellus detectable.
</paragraph>
<paragraph id="8B8C3804ED6FFF95B1C9E9C5D819FBF9" blockId="28.[151,1437,576,1647]" pageId="28" pageNumber="129">
<emphasis id="B947E416ED6FFF95B1C9E9C5DC18FD51" box="[199,288,648,674]" italics="true" pageId="28" pageNumber="129">Thorax.</emphasis>
Pronotum much as in male, but anterior lobe with ill-defined smaller outer mark only, vaguely connected to large mark of same colour on median lobe, posterior lobe with vertical dorsal and horizontal ventral lobe; propleura much as in male but pale areas largely pale greyish yellow instead of pale greyish blue. Synthorax with pleura largely black; pale greyish yellow areas restricted to: ante-humeral patch, very slightly longer than 1/2 length of mesanepisternum, very narrowly almond-shaped to almost parallel sided; mesepimeral patch reduced to tiny remnant completely fused with stripe across whole length and anterior 3/4 to 4/5 of metepisternum; subtriangular, narrow-tailed patch across posterior half of metepimeron; posterolateral tip of meso- and metakatepisternum. Postcoxae blackish brown to black and pale greyish yellow; poststernum pale greyish yellow with narrow elongate patches of dark greyish brown. Legs: Coxae largely pale greyish yellow, anterobasally brown to black, trochanters black and pale greyish yellow, legs otherwise black, with only very base of femora and part of basal 3/4 of metafemur pale greyish yellow. Wings: membrane rather clear, venation black; pterostigma black; postnodals 16/1314.
</paragraph>
<paragraph id="8B8C3804ED6FFF95B1C9EF59DFE1FB69" blockId="28.[151,1437,576,1647]" pageId="28" pageNumber="129">
<emphasis id="B947E416ED6FFF95B1C9EF59DC01FBDD" box="[199,313,1044,1070]" italics="true" pageId="28" pageNumber="129">Abdomen.</emphasis>
S1 and S2 dorsally black, laterally largely pale greyish yellow; S37 dorsally black with greyish yellow dorsal mark very close to base, very tiny on S7, ventrally margined greyish brown to dull yellow; S8 black, ventrally margined greyish brown to dull yellow; S9 and S10 black. Anal appendages and inferior anal lamina pale yellow; supra-anal plate and valves brown to black.
</paragraph>
<paragraph id="8B8C3804ED6FFF95B1C9EFE8DE1BFB4C" blockId="28.[151,1437,576,1647]" box="[199,803,1188,1215]" pageId="28" pageNumber="129">
<emphasis id="B947E416ED6FFF95B1C9EFE8DC6BFB4D" box="[199,339,1189,1214]" italics="true" pageId="28" pageNumber="129">Dimensions.</emphasis>
Hind wing
<quantity id="4CCB95E1ED6FFF95B0D9EFE9DF06FB4C" box="[471,574,1188,1215]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="28" pageNumber="129" unit="mm" value="19.5">19.5 mm</quantity>
; abdomen
<quantity id="4CCB95E1ED6FFF95B3B9EFE9DE26FB4C" box="[695,798,1188,1215]" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="28" pageNumber="129" unit="mm" value="29.3">29.3 mm</quantity>
.
</paragraph>
<paragraph id="8B8C3804ED6FFF95B1C9EF85D9FBFAF4" blockId="28.[151,1437,576,1647]" pageId="28" pageNumber="129">
<emphasis id="B947E416ED6FFF95B1C9EF85DF7FFB11" bold="true" box="[199,583,1224,1250]" pageId="28" pageNumber="129">
Variation in the male
<typeStatus id="548886A6ED6FFF95B0C4EF85DF7AFB11" box="[458,578,1224,1250]" pageId="28" pageNumber="129">paratypes</typeStatus>
.
</emphasis>
Small pale spots on anterior lobe of pronotum present or absent. Postnodals 1516/1214. Hind wing
<quantity id="4CCB95E1ED6FFF95B0B9EFA1DF65FAF4" box="[439,605,1260,1287]" metricMagnitude="-2" metricUnit="m" metricValue="1.875" metricValueMax="1.94" metricValueMin="1.81" pageId="28" pageNumber="129" unit="mm" value="18.75" valueMax="19.4" valueMin="18.1">18.119.4 mm</quantity>
; abdomen (including anal appendages)
<quantity id="4CCB95E1ED6FFF95B517EFA1D987FAF4" box="[1049,1215,1260,1287]" metricMagnitude="-2" metricUnit="m" metricValue="3.09" metricValueMax="3.15" metricValueMin="3.03" pageId="28" pageNumber="129" unit="mm" value="30.9" valueMax="31.5" valueMin="30.3">30.331.5 mm</quantity>
.
</paragraph>
<paragraph id="8B8C3804ED6FFF95B1C9EE5DD966FABD" blockId="28.[151,1437,576,1647]" pageId="28" pageNumber="129">
<emphasis id="B947E416ED6FFF95B1C9EE5DDC05FAD9" bold="true" box="[199,317,1296,1322]" pageId="28" pageNumber="129">Affinities.</emphasis>
Based on the present study (
<figureCitation id="13082481ED6FFF95B370EE5DDFFAFAD9" box="[638,706,1296,1322]" captionStart="FIGURE 6" captionStartId="8.[151,250,1705,1729]" captionTargetBox="[272,1316,187,1675]" captionTargetId="figure-42@8.[266,1321,181,1681]" captionTargetPageId="8" captionText="FIGURE 6. Bayesian tree from concatenated (all genes) data set. Numbers above branches are BI posterior probabilities and ML bootstrap values shown only if ≥ 0.9 and 70%, respectively." figureDoi="http://doi.org/10.5281/zenodo.7978141" httpUri="https://zenodo.org/record/7978141/files/figure.png" pageId="28" pageNumber="129">Fig. 6</figureCitation>
),
<taxonomicName id="4C334387ED6FFF95B3D9EE5CDE07FAD9" authority="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus, 2023" authorityName="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus" authorityYear="2023" box="[727,831,1296,1322]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="tagula" status="sp. nov.">
<emphasis id="B947E416ED6FFF95B3D9EE5CDE07FAD9" box="[727,831,1296,1322]" italics="true" pageId="28" pageNumber="129">N. tagula</emphasis>
</taxonomicName>
<emphasis id="B947E416ED6FFF95B24BEE5DDEA7FAD9" bold="true" box="[837,927,1296,1322]" pageId="28" pageNumber="129">
<taxonomicNameLabel id="A274596DED6FFF95B24BEE5DDEA7FAD9" box="[837,927,1296,1322]" pageId="28" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is closely allied with
<taxonomicName id="4C334387ED6FFF95B59CEE5CD835FAD9" authority="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus, 2023" authorityName="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus" authorityYear="2023" box="[1170,1293,1296,1322]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="stueberi" status="sp. nov.">
<emphasis id="B947E416ED6FFF95B59CEE5CD835FAD9" box="[1170,1293,1296,1322]" italics="true" pageId="28" pageNumber="129">N. stueberi</emphasis>
</taxonomicName>
<emphasis id="B947E416ED6FFF95B41AEE5DD855FAD9" bold="true" box="[1300,1389,1296,1322]" pageId="28" pageNumber="129">
<taxonomicNameLabel id="A274596DED6FFF95B41AEE5DD855FAD9" box="[1300,1389,1296,1322]" pageId="28" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C334387ED6FFF95B199EE78DC3CFABD" authority="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus, 2023" authorityName="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus" authorityYear="2023" box="[151,260,1332,1358]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="boonei" status="sp. nov.">
<emphasis id="B947E416ED6FFF95B199EE78DC3CFABD" box="[151,260,1332,1358]" italics="true" pageId="28" pageNumber="129">N. boonei</emphasis>
</taxonomicName>
<emphasis id="B947E416ED6FFF95B005EE79DC5DFABD" bold="true" box="[267,357,1332,1358]" pageId="28" pageNumber="129">
<taxonomicNameLabel id="A274596DED6FFF95B005EE79DC5DFABD" box="[267,357,1332,1358]" pageId="28" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, together these species comprise the sister group of
<emphasis id="B947E416ED6FFF95B2A3EE78D966FABD" box="[941,1118,1332,1358]" italics="true" pageId="28" pageNumber="129">
<taxonomicName id="4C334387ED6FFF95B2A3EE78D962FABD" box="[941,1114,1332,1358]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="chrismulleri">N. chrismulleri</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3296B8FED6FFF95B1C9EE15DE61F9F1" pageId="28" pageNumber="129" type="diagnosis">
<paragraph id="8B8C3804ED6FFF95B1C9EE15DE61F9F1" blockId="28.[151,1437,576,1647]" pageId="28" pageNumber="129">
<emphasis id="B947E416ED6FFF95B1C9EE15DCF5FA81" bold="true" box="[199,461,1368,1394]" pageId="28" pageNumber="129">Differential diagnosis.</emphasis>
Among members of the
<taxonomicName id="4C334387ED6FFF95B3ECEE14DE4CFA81" box="[738,884,1368,1394]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="salomonis">
<emphasis id="B947E416ED6FFF95B3ECEE14DE4CFA81" box="[738,884,1368,1394]" italics="true" pageId="28" pageNumber="129">N. salomonis</emphasis>
</taxonomicName>
complex, only
<taxonomicName id="4C334387ED6FFF95B52EEE14D998FA81" baseAuthorityName="Schmidt" baseAuthorityYear="1944" box="[1056,1184,1368,1394]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B947E416ED6FFF95B52EEE14D998FA81" box="[1056,1184,1368,1394]" italics="true" pageId="28" pageNumber="129">N. africana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C334387ED6FFF95B5DBEE14D805FA81" authority="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus, 2023" authorityName="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus" authorityYear="2023" box="[1237,1341,1368,1394]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="tagula" status="sp. nov.">
<emphasis id="B947E416ED6FFF95B5DBEE14D805FA81" box="[1237,1341,1368,1394]" italics="true" pageId="28" pageNumber="129">N. tagula</emphasis>
</taxonomicName>
<emphasis id="B947E416ED6FFF95B44DEE15D8A4FA81" bold="true" box="[1347,1436,1368,1394]" pageId="28" pageNumber="129">
<taxonomicNameLabel id="A274596DED6FFF95B44DEE15D8A4FA81" box="[1347,1436,1368,1394]" pageId="28" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have abdominal segment 2 dorsally marked with blue.
<taxonomicName id="4C334387ED6FFF95B3F0EE30DEE2FA65" baseAuthorityName="Schmidt" baseAuthorityYear="1944" box="[766,986,1404,1430]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B947E416ED6FFF95B3F0EE30DEE2FA65" box="[766,986,1404,1430]" italics="true" pageId="28" pageNumber="129">Nososticta africana</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C334387ED6FFF95B405EE30D84CFA65" authority="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus, 2023" authorityName="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus" authorityYear="2023" box="[1291,1396,1404,1430]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="tagula" status="sp. nov.">
<emphasis id="B947E416ED6FFF95B405EE30D84CFA65" box="[1291,1396,1404,1430]" italics="true" pageId="28" pageNumber="129">N. tagula</emphasis>
</taxonomicName>
<emphasis id="B947E416ED6FFF95B475EE31DDF0FA49" bold="true" pageId="28" pageNumber="129">
<taxonomicNameLabel id="A274596DED6FFF95B475EE31DDF0FA49" pageId="28" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having one or two blue antero-dorsal marks and one postero-dorsal mark on segment 2 (vs. only a blue postero-dorsal mark).
<taxonomicName id="4C334387ED6FFF95B09DEE88DF49FA2D" baseAuthorityName="Schmidt" baseAuthorityYear="1944" box="[403,625,1476,1502]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B947E416ED6FFF95B09DEE88DF49FA2D" box="[403,625,1476,1502]" italics="true" pageId="28" pageNumber="129">Nososticta africana</emphasis>
</taxonomicName>
is known almost exclusively from the
<collectingCountry id="F3247894ED6FFF95B521EE89D817FA2D" box="[1071,1327,1476,1502]" name="Papua New Guinea" pageId="28" pageNumber="129">Bismarck Archipelago</collectingCountry>
, while
<taxonomicName id="4C334387ED6FFF95B48DEE88DDE7F9F1" authority="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus, 2023" authorityName="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus" authorityYear="2023" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="tagula" status="sp. nov.">
<emphasis id="B947E416ED6FFF95B48DEE88DDE7F9F1" italics="true" pageId="28" pageNumber="129">N. tagula</emphasis>
</taxonomicName>
<emphasis id="B947E416ED6FFF95B1E8EEA5DC07F9F1" bold="true" box="[230,319,1512,1538]" pageId="28" pageNumber="129">
<taxonomicNameLabel id="A274596DED6FFF95B1E8EEA5DC07F9F1" box="[230,319,1512,1538]" pageId="28" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known only from the Louisiade Archipelago.
</paragraph>
</subSubSection>
<subSubSection id="C3296B8FED6FFF95B1C9ED41DE11F99D" pageId="28" pageNumber="129" type="distribution">
<paragraph id="8B8C3804ED6FFF95B1C9ED41DE11F99D" blockId="28.[151,1437,576,1647]" pageId="28" pageNumber="129">
<emphasis id="B947E416ED6FFF95B1C9ED41DF3AF9D5" bold="true" box="[199,514,1548,1574]" pageId="28" pageNumber="129">Distribution and habitats.</emphasis>
<taxonomicName id="4C334387ED6FFF95B302ED40DFEDF9D5" authorityName="Theischinger &amp; Mitchell &amp; Richards &amp; Polhemus" authorityYear="2023" box="[524,725,1548,1574]" class="Insecta" family="Protoneuridae" genus="Nososticta" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="tagula">
<emphasis id="B947E416ED6FFF95B302ED40DFEDF9D5" box="[524,725,1548,1574]" italics="true" pageId="28" pageNumber="129">Nososticta tagula</emphasis>
</taxonomicName>
is currently known only from Tagula Island in the Louisiade Archipelago (
<figureCitation id="13082481ED6FFF95B03DED7DDCBAF9B9" box="[307,386,1584,1610]" captionStart="MAP 3" captionStartId="34.[151,207,1000,1024]" captionTargetBox="[153,1433,183,974]" captionTargetId="figure-42@34.[151,1435,181,976]" captionText="MAP 3.—Distribution of Nososticta tagula sp. nov. in the Louisiade Archipelago." figureDoi="http://doi.org/10.5281/zenodo.7982146" httpUri="https://zenodo.org/record/7982146/files/figure.png" pageId="28" pageNumber="129">Map 3</figureCitation>
). It has not been detected on other
<collectingRegion id="49F7F6E6ED6FFF95B22FED7DDEA3F9B8" box="[801,923,1584,1611]" country="Papua New Guinea" name="Milne Bay" pageId="28" pageNumber="129">Milne Bay</collectingRegion>
Islands despite extensive surveys across the region and may be range-restricted Tagula Island endemic.
</paragraph>
</subSubSection>
</treatment>
</document>