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<document id="C60BC553B88120AFF6C67DB232574299" ID-CLB-Dataset="8114" ID-DOI="10.37520/aemnp.2020.24" ID-GBIF-Dataset="9cf87cc9-79ad-4a31-83ca-b635d495a0b9" ID-ISSN="1804-6487" ID-Zenodo-Dep="4489252" ID-ZooBank="AC387BAF-E7A8-40B2-9486-E5642074587D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612221266920" checkinUser="felipe" docAuthor="Lackner, Tomáš" docDate="2020" docId="039A87F1FFC3FFB6FF32B26AFB72FBF6" docLanguage="en" docName="ActaEntMusNatPrag.60.2.397-409.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Gnathoncus Jacquelin du Val 1857" docType="treatment" docVersion="8" lastPageNumber="398" masterDocId="FFA3FF89FFC2FFB7FFEBB448FFB2FFA3" masterDocTitle="A review of Gnathoncus of Southeast Asia (Coleoptera: Histeridae: Saprininae)" masterLastPageNumber="409" masterPageNumber="397" pageNumber="398" updateTime="1698926172273" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="1C611E74E6F9144B04BCA3A71541BB3C">A review of Gnathoncus of Southeast Asia (Coleoptera: Histeridae: Saprininae)</mods:title>
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<treatment id="039A87F1FFC3FFB6FF32B26AFB72FBF6" ID-DOI="http://doi.org/10.5281/zenodo.4549633" ID-GBIF-Taxon="177425804" ID-Zenodo-Dep="4549633" LSID="urn:lsid:plazi:treatment:039A87F1FFC3FFB6FF32B26AFB72FBF6" httpUri="http://treatment.plazi.org/id/039A87F1FFC3FFB6FF32B26AFB72FBF6" lastPageNumber="398" pageId="1" pageNumber="398">
<subSubSection id="C329656CFFC3FFB6FF32B26AFD04F99A" box="[217,694,1569,1593]" pageId="1" pageNumber="398" type="nomenclature">
<paragraph id="8B8C36E7FFC3FFB6FF32B26AFD04F99A" blockId="1.[217,694,1569,1593]" box="[217,694,1569,1593]" pageId="1" pageNumber="398">
<heading id="D0C4818BFFC3FFB6FF32B26AFD04F99A" box="[217,694,1569,1593]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="398" reason="3">
<emphasis id="B947EAF5FFC3FFB6FF32B26AFD04F99A" bold="true" box="[217,694,1569,1593]" pageId="1" pageNumber="398">
Genus
<taxonomicName id="4C334D64FFC3FFB6FEC3B269FD04F99A" ID-CoL="4PL8" authority="Jacquelin du Val, 1857" authorityName="Jacquelin du Val" authorityYear="1857" box="[296,694,1569,1593]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="B947EAF5FFC3FFB6FEC3B269FE1DF99A" bold="true" box="[296,431,1569,1593]" italics="true" pageId="1" pageNumber="398">Gnathoncus</emphasis>
Jacquelin du Val, 1857
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC3FFB6FF7CB20FFE57F909" pageId="1" pageNumber="398" type="reference_group">
<paragraph id="8B8C36E7FFC3FFB6FF7CB20FFE57F909" blockId="1.[151,765,1607,1706]" pageId="1" pageNumber="398">
<taxonomicName id="4C334D64FFC3FFB6FF7CB20FFF4CF9F9" ID-CoL="4PL8" authorityName="Jacquelin du Val" authorityYear="1857" box="[151,254,1607,1626]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="B947EAF5FFC3FFB6FF7CB20FFF4CF9F9" box="[151,254,1607,1626]" italics="true" pageId="1" pageNumber="398">Gnathoncus</emphasis>
</taxonomicName>
: MΑƵUR (2011): 175 (catalogue); VιΕΝΝΑ +| RΑττο (2013): 30 (description of a new species from
<collectingCountry id="F3247677FFC3FFB6FE15B22AFD96F9D6" box="[510,548,1634,1653]" name="Iran" pageId="1" pageNumber="398">Iran</collectingCountry>
); LΑΚΝΕR et al. (2015): 113 (catalogue); LΑΚΝΕR +| LΕඌർΗΕΝ (2017): 28 (diagnosis and key to the Australopacific
<taxonomicName id="4C334D64FFC3FFB6FE9AB2DFFE6AF909" ID-CoL="4PL8" authorityName="Jacquelin du Val" authorityYear="1857" box="[369,472,1687,1706]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="B947EAF5FFC3FFB6FE9AB2DFFE6AF909" box="[369,472,1687,1706]" italics="true" pageId="1" pageNumber="398">Gnathoncus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC3FFB6FF7CB28BFE25F8F9" pageId="1" pageNumber="398" type="description">
<paragraph id="8B8C36E7FFC3FFB6FF7CB28BFE25F8F9" blockId="1.[151,760,1731,2074]" pageId="1" pageNumber="398">
<emphasis id="B947EAF5FFC3FFB6FF7CB28BFF66F979" bold="true" box="[151,212,1731,1754]" pageId="1" pageNumber="398">Note.</emphasis>
Complete list of synonymies and literature references of this taxon are given in LΑΚΝΕR (2010: 105) and the reader is referred to them there. For the sake of completeness we only list the references published after LΑΚΝΕR (2010) above.
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC3FFB6FF7CB32BFB05FD36" pageId="1" pageNumber="398" type="diagnosis">
<paragraph id="8B8C36E7FFC3FFB6FF7CB32BFB05FD36" blockId="1.[151,760,1731,2074]" lastBlockId="1.[828,1441,190,1109]" pageId="1" pageNumber="398">
<emphasis id="B947EAF5FFC3FFB6FF7CB32BFEBCF8D9" bold="true" box="[151,270,1891,1914]" pageId="1" pageNumber="398">Diagnosis.</emphasis>
Members of the genus
<taxonomicName id="4C334D64FFC3FFB6FDCBB32BFD11F8D9" authorityName="Jacquelin du Val" authorityYear="1857" box="[544,675,1891,1914]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="B947EAF5FFC3FFB6FDCBB32BFD11F8D9" box="[544,675,1891,1914]" italics="true" pageId="1" pageNumber="398">Gnathoncus</emphasis>
</taxonomicName>
can be easily distinguished from all other SE Asian
<taxonomicName id="4C334D64FFC3FFB6FD6DB3CBFD4AF839" authorityName="Blanchard" authorityYear="1845" box="[646,760,1923,1946]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
by the following external and genitalic characters: body rather small (PEL=
<quantity id="4CCB9B02FFC3FFB6FEB4B38BFE4EF879" box="[351,508,1987,2010]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="3.2" metricValueMin="1.7" pageId="1" pageNumber="398" unit="mm" value="2.45" valueMax="3.2" valueMin="1.7">1.703.20 mm</quantity>
), uniformly dark-brown to black, without metallic hue; frontoclypeal suture (frontal stria) absent; dorsum punctate; pronotal hypomeron asetose, pronotum only with marginal pronotal stria (shortened in case of
<taxonomicName id="4C334D64FFC3FFB6FC4EB496FBDCFF56" authorityName="Bickhardt" authorityYear="1920" box="[933,1134,222,245]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="semimarginatus">
<emphasis id="B947EAF5FFC3FFB6FC4EB496FBDCFF56" box="[933,1134,222,245]" italics="true" pageId="1" pageNumber="398">G. semimarginatus</emphasis>
</taxonomicName>
); elytra with complete striae IIV; between base of elytral stria IV and sutural elytral stria (that can be interrupted or present only as a short basal fragment) present a characteristic short, hooked appendix; lateral prosternal striae very short, reaching approximately mid-length of carinal prosternal striae; outer lateral costa of prosternal process reaching prosternal keel. Ninth tergite of male terminalia longitudinally divided; VIII sternite and tergite not fused laterally. For a key to the Thai
<taxonomicName id="4C334D64FFC3FFB6FAC1B596FA2EFE56" authorityName="Blanchard" authorityYear="1845" box="[1322,1436,478,501]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
see MΑƵUR +| ÔΗΑRΑ (2003); for a key to the Indonesian
<taxonomicName id="4C334D64FFC3FFB6FCD7B656FC1CFD96" authorityName="Blanchard" authorityYear="1845" box="[828,942,542,565]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
see ÔΗΑRΑ +| HΑRιτιΝι (2008). As yet, there is no key to the SE Asian genera of
<taxonomicName id="4C334D64FFC3FFB6FB4DB676FAA9FDF6" authorityName="Blanchard" authorityYear="1845" box="[1190,1307,574,597]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
, mostly due to the existence of undescribed taxa from
<collectingCountry id="F3247677FFC3FFB6FAE6B616FADEFDD6" box="[1293,1388,606,629]" name="Thailand" pageId="1" pageNumber="398">Thailand</collectingCountry>
and
<collectingCountry id="F3247677FFC3FFB6FCD7B636FC24FD36" box="[828,918,638,661]" name="Vietnam" pageId="1" pageNumber="398">Vietnam</collectingCountry>
(T. Lackner, unpublished).
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC3FFB6FCD7B6D6FB74FCD6" pageId="1" pageNumber="398" type="biology_ecology">
<paragraph id="8B8C36E7FFC3FFB6FCD7B6D6FB74FCD6" blockId="1.[828,1441,190,1109]" pageId="1" pageNumber="398">
<emphasis id="B947EAF5FFC3FFB6FCD7B6D6FC2AFD16" bold="true" box="[828,920,670,693]" pageId="1" pageNumber="398">Biology.</emphasis>
<taxonomicName id="4C334D64FFC3FFB6FC49B6D6FB00FD16" authorityName="Lewis" authorityYear="1907" box="[930,1202,670,693]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="brevisternus">
<emphasis id="B947EAF5FFC3FFB6FC49B6D6FB00FD16" box="[930,1202,670,693]" italics="true" pageId="1" pageNumber="398">Gnathoncus brevisternus</emphasis>
</taxonomicName>
was collected inside caves;
<taxonomicName id="4C334D64FFC3FFB6FC6CB6F6FBAFFD76" baseAuthorityName="Kugelann" baseAuthorityYear="1792" box="[903,1053,702,725]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B947EAF5FFC3FFB6FC6CB6F6FBAFFD76" box="[903,1053,702,725]" italics="true" pageId="1" pageNumber="398">G. rotundatus</emphasis>
</taxonomicName>
is a synanthrope often collected in anthropogenic settings;
<taxonomicName id="4C334D64FFC3FFB6FBD5B696FB6CFD56" baseAuthorityName="Marseul" baseAuthorityYear="1862" box="[1086,1246,734,757]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="nannetensis">
<emphasis id="B947EAF5FFC3FFB6FBD5B696FB6CFD56" box="[1086,1246,734,757]" italics="true" pageId="1" pageNumber="398">G. nannetensis</emphasis>
</taxonomicName>
was collected on carrion and in bird nests;
<taxonomicName id="4C334D64FFC3FFB6FBA4B6B6FB52FCB6" authority="Lackner, 2020" authorityName="Lackner" authorityYear="2020" box="[1103,1248,766,789]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="sechuanus" status="sp. nov.">
<emphasis id="B947EAF5FFC3FFB6FBA4B6B6FB52FCB6" box="[1103,1248,766,789]" italics="true" pageId="1" pageNumber="398">G. sechuanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A274578EFFC3FFB6FB0CB6B6FA89FCB6" box="[1255,1339,766,789]" pageId="1" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
was collected under a mushroom on a tree trunk in a mixed forest. The biology of two SE Asian endemics (
<taxonomicName id="4C334D64FFC3FFB6FB11B776FA2EFCF6" authorityName="Kryzhanovskij" authorityYear="1972" box="[1274,1436,830,853]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B947EAF5FFC3FFB6FB11B776FA2EFCF6" box="[1274,1436,830,853]" italics="true" pageId="1" pageNumber="398">G. vietnamicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C334D64FFC3FFB6FC82B716FB80FCD6" authorityName="Bickhardt" authorityYear="1920" box="[873,1074,862,885]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="semimarginatus">
<emphasis id="B947EAF5FFC3FFB6FC82B716FB80FCD6" box="[873,1074,862,885]" italics="true" pageId="1" pageNumber="398">G. semimarginatus</emphasis>
</taxonomicName>
) is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC3FFB6FCD7B736FB72FBF6" pageId="1" pageNumber="398" type="distribution">
<paragraph id="8B8C36E7FFC3FFB6FCD7B736FB72FBF6" blockId="1.[828,1441,190,1109]" pageId="1" pageNumber="398">
<emphasis id="B947EAF5FFC3FFB6FCD7B736FC7CFC36" bold="true" box="[828,974,894,917]" pageId="1" pageNumber="398">Distribution.</emphasis>
In SE Asia,
<taxonomicName id="4C334D64FFC3FFB6FBB1B736FB69FC36" authorityName="Jacquelin du Val" authorityYear="1857" box="[1114,1243,894,917]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="B947EAF5FFC3FFB6FBB1B736FB69FC36" box="[1114,1243,894,917]" italics="true" pageId="1" pageNumber="398">Gnathoncus</emphasis>
</taxonomicName>
is present mainly in the north of the region: southern
<collectingCountry id="F3247677FFC3FFB6FB2CB7D6FAB5FC16" box="[1223,1287,926,949]" name="China" pageId="1" pageNumber="398">China</collectingCountry>
(
<collectingRegion id="49F7F805FFC3FFB6FAF3B7D6FA2BFC16" box="[1304,1433,926,949]" country="China" name="Guangdong" pageId="1" pageNumber="398">Guangdong</collectingRegion>
,
<collectingRegion id="49F7F805FFC3FFB6FCD7B7F6FC31FC76" box="[828,899,958,981]" country="China" name="Gansu" pageId="1" pageNumber="398">Gansu</collectingRegion>
,
<collectingRegion id="49F7F805FFC3FFB6FC7BB7F6FC58FC76" box="[912,1002,958,981]" country="China" name="Sichuan" pageId="1" pageNumber="398">Sichuan</collectingRegion>
),
<collectingCountry id="F3247677FFC3FFB6FC15B7F6FBFDFC76" box="[1022,1103,958,981]" name="Taiwan" pageId="1" pageNumber="398">Taiwan</collectingCountry>
,
<collectingCountry id="F3247677FFC3FFB6FBB0B7F6FB2FFC76" box="[1115,1181,958,981]" name="Nepal" pageId="1" pageNumber="398">Nepal</collectingCountry>
, northern
<collectingCountry id="F3247677FFC3FFB6FAE5B7F6FADFFC76" box="[1294,1389,958,981]" name="Thailand" pageId="1" pageNumber="398">Thailand</collectingCountry>
and
<collectingCountry id="F3247677FFC3FFB6FCD7B796FC28FC56" box="[828,922,990,1013]" name="Vietnam" pageId="1" pageNumber="398">Vietnam</collectingCountry>
. One unidentified species is known from
<collectingRegion id="49F7F805FFC3FFB6FA86B796FA2FFC56" box="[1389,1437,990,1013]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="398">Java</collectingRegion>
(
<collectingCountry id="F3247677FFC3FFB6FCA8B7B6FC03FBB6" box="[835,945,1022,1045]" name="Indonesia" pageId="1" pageNumber="398">Indonesia</collectingCountry>
). The exact distribution of
<taxonomicName id="4C334D64FFC3FFB6FB39B7B6FA2EFBB6" authorityName="Bickhardt" authorityYear="1920" box="[1234,1436,1022,1045]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="semimarginatus">
<emphasis id="B947EAF5FFC3FFB6FB39B7B6FA2EFBB6" box="[1234,1436,1022,1045]" italics="true" pageId="1" pageNumber="398">G. semimarginatus</emphasis>
</taxonomicName>
is unknown (its
<typeStatus id="54888845FFC3FFB6FC0CB056FBF6FB96" box="[999,1092,1054,1077]" pageId="1" pageNumber="398" type="holotype">holotype</typeStatus>
was either collected in northern
<collectingCountry id="F3247677FFC3FFB6FCD7B076FCC1FBF6" box="[828,883,1086,1109]" name="India" pageId="1" pageNumber="398">India</collectingCountry>
or southern
<collectingCountry id="F3247677FFC3FFB6FC13B076FB88FBF6" box="[1016,1082,1086,1109]" name="China" pageId="1" pageNumber="398">China</collectingCountry>
, see below).
</paragraph>
</subSubSection>
</treatment>
</document>