287 lines
40 KiB
XML
287 lines
40 KiB
XML
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<mods:title id="DB56F425607218B2C010CF172D04CE6C">Late Miocene Dasypodidae Gray, 1821 (Xenarthra, Cingulata) from the Toro Negro Formation (Central Andes, Argentina): diversity and chronological and biogeographical implications</mods:title>
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<mods:namePart id="EBC5176DA7ECDA7D09189444999BB965">Brandoni, Diego</mods:namePart>
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<mods:affiliation id="EA344BA32EA8AA487122916502B2741D">Laboratorio de Paleontología de Vertebrados, Centro de Investigación Científica y de Transferencia Tecnológica a la Producción - Consejo Nacional de Investigaciones Científicas y Técnicas (CICYTTP, CONICET-Prov. ER-UADER), España 149, E 3105 BWA Diamante (Argentina) dbrandoni @ cicyttp. org. ar (corresponding author)</mods:affiliation>
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<mods:nameIdentifier id="EB0F3BCE780212DB279D5F31DD25EC97" type="email">dbrandoni@cicyttp.org.ar</mods:nameIdentifier>
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<mods:namePart id="E11E88875FA69EA824305B638D2C2268">Barasoain, Daniel</mods:namePart>
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<mods:affiliation id="EDDFDE01AFFDD88BD63E8418C45EE244">Laboratorio de Evolución de Vertebrados y Ambientes Cenozoicos, Centro de Ecología Aplicada del Litoral - Universidad Nacional de Nordeste - Consejo Nacional de Investigaciones Científicas y Técnicas (CECOAL-UNNE-CONICET) y Cátedra de Paleontología, Facultad de Ciencias Exactas, Naturales y Agrimensura, Universidad Nacional del Nordeste, RP 5 3400 Corrientes (Argentina) danielbarasoain @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="2A15136060DB887F8C34B0C4A934FDF9" type="email">danielbarasoain@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B009FE2FA69D1D6CC9F5461CE8FFA1F7">González Ruiz, Laureano R.</mods:namePart>
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<mods:affiliation id="A9412AAE732B6C32D56FEBB377B065B3">Laboratorio de Investigaciones en Evolución y Biodiversidad (LIEB-FCNyCS sede Esquel, UNPSJB) y Centro de Investigaciones Esquel de Montaña y Estepa Patagónica (CIEMEP), Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Universidad Nacional de La Patagonia San Juan Bosco (UNPSJB), Roca 780, 9200, Esquel, Chubut (Argentina) lrgonzalezruiz @ comahue-conicet. gob. ar</mods:affiliation>
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<mods:nameIdentifier id="E3D22E5EEBB6E1CC9D790B80ED53A1DA" type="email">lrgonzalezruiz@comahue-conicet.gob.ar</mods:nameIdentifier>
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<mods:title id="23EA22B39CB2ACC64BEF8846447FEAC3">Comptes Rendus Palevol</mods:title>
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<mods:part id="307D233F2BA49D02D7E3818A855090E1">
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<mods:date id="783C6802055022707EB67F9885A9642E">2023</mods:date>
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<mods:number id="7EA1BB83C161A764E9BADDECBCACBD13">2023-01-16</mods:number>
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<treatment id="039A87DE134CD93EFC0008B8FC870FD8" ID-GBIF-Taxon="247568569" LSID="urn:lsid:plazi:treatment:039A87DE134CD93EFC0008B8FC870FD8" httpUri="http://treatment.plazi.org/id/039A87DE134CD93EFC0008B8FC870FD8" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4" scope_class="Mammalia" scope_family="Dasypodidae" scope_order="Xenarthra" scope_subFamily="Tolypeutinae" scope_subOrder="Cingulata" scope_tribe="Priodontini">
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<subSubSection id="C3296543134CD93CFC0008B8FAC10EBA" box="[923,1345,1491,1518]" pageId="5" pageNumber="4" type="nomenclature">
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<paragraph id="8B8C36C8134CD93CFC0008B8FAC10EBA" blockId="5.[923,1345,1491,1518]" box="[923,1345,1491,1518]" pageId="5" pageNumber="4">
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<heading id="D0C481A4134CD93CFC0008B8FAC10EBA" box="[923,1345,1491,1518]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="4" reason="2">
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<taxonomicName id="4C334D4B134CD93CFC0008B8FAC10EBA" authority="Esteban & Nasif, 1996" authorityName="Esteban & Nasif" authorityYear="1996" box="[923,1345,1491,1518]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xenarthra" pageId="5" pageNumber="4" phylum="Chordata" rank="species" species="gandhii">
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<emphasis id="B947EADA134CD93CFC0008B8FBC00EB9" bold="true" box="[923,1088,1491,1518]" italics="true" pageId="5" pageNumber="4">Vetelia gandhii</emphasis>
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<bibRefCitation id="EFA24B39134CD93CFBDC08B8FAC10EBA" author="ESTEBAN G. I. & NASIF N. L." box="[1095,1345,1491,1517]" pageId="5" pageNumber="4" pagination="327 - 334" refId="ref12563" refString="ESTEBAN G. I. & NASIF N. L. 1996. - Nuevos Dasypodidae (Mammalia, Xenarthra) del Mioceno tardio del Valle del Cajon, Catamarca, Argentina. Ameghiniana 33 (3): 327 - 334." type="journal article" year="1996">Esteban & Nasif, 1996</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3296543134CD93CFB980898FB590D59" box="[1027,1241,1523,1550]" pageId="5" pageNumber="4" type="description">
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<paragraph id="8B8C36C8134CD93CFB980898FB590D59" blockId="5.[1027,1241,1523,1550]" box="[1027,1241,1523,1550]" pageId="5" pageNumber="4">
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(
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<figureCitation id="13082A4D134CD93CFB970898FBCA0D5A" box="[1036,1098,1523,1549]" captionStart="FIG" captionStartId="6.[132,143,1507,1524]" captionTargetBox="[132,1456,185,1474]" captionTargetId="figure-276@6.[698,1267,743,1285]" captionTargetPageId="6" captionText="FIG. 3. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-F, Vetelia gandhii Esteban & Nasif, 1996 (CRILAR-Pv 116; CRILAR-Pv 117; CRILAR-Pv 118); G-L, Chasicotatus peiranoi Esteban & Nasif, 1996 (CRILAR-Pv 119); M, N, Macrochorobates scalabrinii (Moreno & Mercerat, 1891) (CRILAR-Pv 120); O-R, Prozaedyus sp. (CRILAR-Pv 121). Scale bars: 1 cm." pageId="5" pageNumber="4">Fig. 3</figureCitation>
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A-F;
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<tableCitation id="C6B10373134CD93CFB1B0898FB500D5A" box="[1152,1232,1523,1549]" captionStart="TABLE" captionStartId="8.[526,537,219,236]" captionTargetPageId="8" captionText="TABLE 1. — Measurements (in mm) of the described osteoderms." httpUri="http://table.plazi.org/id/DF4C66401341D931FD950DB0FBA80BBB" pageId="5" pageNumber="4" tableUuid="DF4C66401341D931FD950DB0FBA80BBB">Table 1</tableCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3296543134CD93CFCB60B47FB520D13" box="[813,1234,1580,1604]" pageId="5" pageNumber="4" type="reference_group">
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<paragraph id="8B8C36C8134CD93CFCB60B47FB520D13" blockId="5.[813,1234,1580,1604]" box="[813,1234,1580,1604]" pageId="5" pageNumber="4">
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<treatmentCitationGroup id="AB2311E6134CD93CFCB60B47FB520D13" box="[813,1234,1580,1604]" pageId="5" pageNumber="4">
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<treatmentCitation id="0A9210D9134CD93CFCB60B47FB4E0D13" author="ESTEBAN G. I. & NASIF N. L." box="[813,1230,1580,1604]" page="329" pageId="5" pageNumber="4" year="1996">
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||
<taxonomicName id="4C334D4B134CD93CFCB60B47FB4E0D13" authority="Esteban & Nasif, 1996: 329" authorityName="Esteban & Nasif" authorityPageNumber="329" authorityYear="1996" box="[813,1230,1580,1604]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xenarthra" pageId="5" pageNumber="4" phylum="Chordata" rank="species" species="puncta">
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<emphasis id="B947EADA134CD93CFCB60B47FC320D13" box="[813,946,1580,1604]" italics="true" pageId="5" pageNumber="4">Vetelia puncta</emphasis>
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<bibRefCitation id="EFA24B39134CD93CFC2C0B47FB4E0D13" author="ESTEBAN G. I. & NASIF N. L." box="[951,1230,1580,1604]" pageId="5" pageNumber="4" pagination="327 - 334" refId="ref12563" refString="ESTEBAN G. I. & NASIF N. L. 1996. - Nuevos Dasypodidae (Mammalia, Xenarthra) del Mioceno tardio del Valle del Cajon, Catamarca, Argentina. Ameghiniana 33 (3): 327 - 334." type="journal article" year="1996">Esteban & Nasif, 1996: 329</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3296543134CD93CFCB60B37FB1F0D89" pageId="5" pageNumber="4" type="materials_examined">
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||
<paragraph id="8B8C36C8134CD93CFCB60B37FB1F0D89" blockId="5.[812,1456,1628,1758]" pageId="5" pageNumber="4">
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<smallCapsWord id="8D6AA014134CD93CFC0F0B09FC730D25" baselines="1646" box="[916,1011,1634,1650]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="referred" pageId="5" pageNumber="4">REFERRED</smallCapsWord>
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.
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<emphasis id="B947EADA134CD93CFC650B36FB970D23" box="[1022,1047,1629,1652]" italics="true" pageId="5" pageNumber="4">—</emphasis>
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<materialsCitation id="3B5B3C95134CD93CFB810B37FCE30DD9" ID-GBIF-Occurrence="4985208304" pageId="5" pageNumber="4" specimenCode="CRILAR-Pv 116" specimenCount="1">
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<specimenCode id="DB959EB3134CD93CFB810B37FB400D23" box="[1050,1216,1628,1652]" pageId="5" pageNumber="4">CRILAR-Pv 116</specimenCode>
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, one isolated fixed osteoderm
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</materialsCitation>
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;
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<materialsCitation id="3B5B3C95134CD93CFCF00B1DFA2C0DFF" ID-GBIF-Occurrence="4985208303" pageId="5" pageNumber="4" specimenCode="CRILAR-Pv 117" specimenCount="1">
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<specimenCode id="DB959EB3134CD93CFCF00B1DFB900DD9" box="[875,1040,1654,1678]" pageId="5" pageNumber="4">CRILAR-Pv 117</specimenCode>
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, one fixed osteoderm associated with a fragment of an osteoderm from the right lateral border of the carapace
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</materialsCitation>
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; and
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<materialsCitation id="3B5B3C95134CD93CFCCC0BC0FB1B0D89" ID-GBIF-Occurrence="4985208305" pageId="5" pageNumber="4" specimenCode="CRILAR-Pv 118" specimenCount="1" specimenCount-juvenile="1">
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<specimenCode id="DB959EB3134CD93CFCCC0BC0FB810D94" box="[855,1025,1707,1731]" pageId="5" pageNumber="4">CRILAR-Pv 118</specimenCode>
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, an association of seven broken osteoderms interpreted as belonging to a juvenile
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C3296543134CD93CFCB60B9EFA1C0C35" pageId="5" pageNumber="4" type="distribution">
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<paragraph id="8B8C36C8134CD93CFCB60B9EFA1C0C35" blockId="5.[813,1456,1781,1890]" pageId="5" pageNumber="4">
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.
|
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<emphasis id="B947EADA134CD93CFA940B9DFAA80C5A" box="[1295,1320,1782,1805]" italics="true" pageId="5" pageNumber="4">—</emphasis>
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Quebrada de Las Torrecillas (
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<geoCoordinate id="EE07500F134CD93CFC510A78FBA30C7A" box="[970,1059,1811,1837]" degrees="28" direction="south" minutes="36" orientation="latitude" pageId="5" pageNumber="4" precision="925" value="-28.6">28°36’S</geoCoordinate>
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,
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<geoCoordinate id="EE07500F134CD93CFBAB0A78FB130C7A" box="[1072,1171,1811,1837]" degrees="68" direction="west" minutes="14" orientation="longitude" pageId="5" pageNumber="4" precision="925" value="-68.23333">68°14’W</geoCoordinate>
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),
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<collectingRegion id="49F7F82A134CD93CFB310A7EFADF0C7A" box="[1194,1375,1813,1837]" country="Argentina" name="La Rioja" pageId="5" pageNumber="4">La Rioja Province</collectingRegion>
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||
,
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<collectingCountry id="F3247658134CD93CFAF00A7EFCD30C10" name="Argentina" pageId="5" pageNumber="4">Argentina</collectingCountry>
|
||
(
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||
<figureCitation id="13082A4D134CD93CFCFA0A44FC190C10" box="[865,921,1839,1863]" captionStart="FIG" captionStartId="4.[134,143,1284,1301]" captionTargetBox="[132,1455,215,1241]" captionTargetId="figure-487@4.[698,1267,213,774]" captionTargetPageId="4" captionText="FIG. 1. — A, Map showing the study area (black circle), La Rioja Province, Argentina; B, panoramic view of Toro Negro Formation at road 76 between Vinchina and Jagüel; C, panoramic view of Toro Negro Formation at Quebrada de Las Torrecillas; D, detail of the lower member of the Toro Negro Formation at Quebrada de Las Torrecillas. Credits: map modified from Google Earth." pageId="5" pageNumber="4">Fig. 1</figureCitation>
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). Upper levels of the lower member of the Toro Negro Formation (Messinian-Zanclean; Late Miocene-Early Pliocene).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3296543134CD93EFCB60AF9FD7609A7" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4" type="description">
|
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<paragraph id="8B8C36C8134CD93CFCB60AF9FC3D0CFD" blockId="5.[812,1455,1938,2028]" box="[813,957,1938,1964]" pageId="5" pageNumber="4">
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</paragraph>
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<paragraph id="8B8C36C8134CD93FFCB60AD9FD2F0C3B" blockId="5.[812,1455,1938,2028]" lastBlockId="6.[130,777,1619,2028]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
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Fixed and lateral osteoderms. The size of fixed osteoderms is similar to that of the extant
|
||
<taxonomicName id="4C334D4B134CD93CFBD50ABAFA2A0CBB" authority="(Kerr, 1792)" baseAuthorityName="Kerr" baseAuthorityYear="1792" box="[1102,1450,2001,2028]" class="Mammalia" family="Dasypodidae" genus="Priodontes" kingdom="Animalia" order="Cingulata" pageId="5" pageNumber="4" phylum="Chordata" rank="species" species="maximus">
|
||
<emphasis id="B947EADA134CD93CFBD50ABAFA9D0CBC" box="[1102,1309,2001,2027]" italics="true" pageId="5" pageNumber="4">Priodontes maximus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA24B39134CD93CFAB20AB9FA220CBB" author="KERR R." box="[1321,1442,2002,2028]" pageId="5" pageNumber="4" refId="ref13887" refString="KERR R. 1792. - The Animal Kingdom or Zoological System, of the Celebrated Sir Charles Linnaeus. Class I. Mammalia A. Edinburgh Press, Edinburgh, 400 p." type="book" year="1792">Kerr, 1792</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The fixed osteoderm CRILAR-Pv 116 (
|
||
<figureCitation id="13082A4D134FD93FFDDD0B38FD1C0D3A" box="[582,668,1619,1645]" captionStart="FIG" captionStartId="6.[132,143,1507,1524]" captionTargetBox="[132,1456,185,1474]" captionTargetId="figure-276@6.[698,1267,743,1285]" captionTargetPageId="6" captionText="FIG. 3. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-F, Vetelia gandhii Esteban & Nasif, 1996 (CRILAR-Pv 116; CRILAR-Pv 117; CRILAR-Pv 118); G-L, Chasicotatus peiranoi Esteban & Nasif, 1996 (CRILAR-Pv 119); M, N, Macrochorobates scalabrinii (Moreno & Mercerat, 1891) (CRILAR-Pv 120); O-R, Prozaedyus sp. (CRILAR-Pv 121). Scale bars: 1 cm." pageId="6" pageNumber="5">Fig. 3A</figureCitation>
|
||
) is thick, wide, and has a pentagonal outline, and probably belongs to the more dorsal region of the carapace. In turn, the fixed osteoderm CRILAR-Pv 117 (
|
||
<figureCitation id="13082A4D134FD93FFE220BD9FD860D9A" box="[441,518,1714,1741]" captionStart="FIG" captionStartId="6.[132,143,1507,1524]" captionTargetBox="[132,1456,185,1474]" captionTargetId="figure-276@6.[698,1267,743,1285]" captionTargetPageId="6" captionText="FIG. 3. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-F, Vetelia gandhii Esteban & Nasif, 1996 (CRILAR-Pv 116; CRILAR-Pv 117; CRILAR-Pv 118); G-L, Chasicotatus peiranoi Esteban & Nasif, 1996 (CRILAR-Pv 119); M, N, Macrochorobates scalabrinii (Moreno & Mercerat, 1891) (CRILAR-Pv 120); O-R, Prozaedyus sp. (CRILAR-Pv 121). Scale bars: 1 cm." pageId="6" pageNumber="5">Fig. 3B</figureCitation>
|
||
) is thinner and narrower than CRILAR-Pv 116 and has a sub-rectangular in outline, so it probably occupied a more lateral position within the carapace. The whole dorsal surface of the osteoderms is full of small foramina. In lateral view, osteoderms have a constant thickness from the anterior to the posterior margin.
|
||
</paragraph>
|
||
<caption id="DF4C6640134FD93FFF1F0888FC5C0D76" pageId="6" pageNumber="5" startId="6.[132,143,1507,1524]" targetBox="[132,1456,185,1474]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B8C36C8134FD93FFF1F0888FC5C0D76" blockId="6.[132,1456,1507,1569]" pageId="6" pageNumber="5">
|
||
<smallCapsWord id="8D6AA014134FD93FFF1F0888FF1D0EA4" baselines="1520,1520" box="[132,157,1507,1524]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="6" pageNumber="5">FIG</smallCapsWord>
|
||
. 3. —
|
||
<taxonomicName id="4C334D4B134FD93FFF4F0888FEC20EA3" authorityName="Gray" authorityYear="1821" box="[212,322,1507,1524]" class="Mammalia" family="Dasypodidae" kingdom="Animalia" order="Cingulata" pageId="6" pageNumber="5" phylum="Chordata" rank="family">Dasypodidae</taxonomicName>
|
||
from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina:
|
||
<emphasis id="B947EADA134FD93FFBD40888FBEE0EA3" bold="true" box="[1103,1134,1507,1524]" pageId="6" pageNumber="5">A -F</emphasis>
|
||
,
|
||
<taxonomicName id="4C334D4B134FD93FFBE20888FA300EA3" authority="Esteban & Nasif, 1996" authorityName="Esteban & Nasif" authorityYear="1996" box="[1145,1456,1507,1524]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="6" pageNumber="5" phylum="Chordata" rank="species" species="gandhii">
|
||
<emphasis id="B947EADA134FD93FFBE20888FB720EA3" box="[1145,1266,1507,1524]" italics="true" pageId="6" pageNumber="5">Vetelia gandhii</emphasis>
|
||
<bibRefCitation id="EFA24B39134FD93FFB6C0888FA300EA3" author="ESTEBAN G. I. & NASIF N. L." box="[1271,1456,1507,1524]" pageId="6" pageNumber="5" pagination="327 - 334" refId="ref12563" refString="ESTEBAN G. I. & NASIF N. L. 1996. - Nuevos Dasypodidae (Mammalia, Xenarthra) del Mioceno tardio del Valle del Cajon, Catamarca, Argentina. Ameghiniana 33 (3): 327 - 334." type="journal article" year="1996">Esteban & Nasif, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
(CRILAR-Pv 116; CRILAR-Pv 117; CRILAR-Pv 118);
|
||
<emphasis id="B947EADA134FD93FFDD90892FDE30D5D" bold="true" box="[578,611,1529,1546]" pageId="6" pageNumber="5">G -L</emphasis>
|
||
,
|
||
<taxonomicName id="4C334D4B134FD93FFDF40892FC6B0D5D" authority="Esteban & Nasif, 1996" authorityName="Esteban & Nasif" authorityYear="1996" box="[623,1003,1529,1546]" class="Mammalia" family="Dasypodidae" genus="Chasicotatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="6" pageNumber="5" phylum="Chordata" rank="species" species="peiranoi">
|
||
<emphasis id="B947EADA134FD93FFDF40892FCA50D5D" box="[623,805,1529,1546]" italics="true" pageId="6" pageNumber="5">Chasicotatus peiranoi</emphasis>
|
||
<bibRefCitation id="EFA24B39134FD93FFCB60892FC6B0D5D" author="ESTEBAN G. I. & NASIF N. L." box="[813,1003,1529,1546]" pageId="6" pageNumber="5" pagination="327 - 334" refId="ref12563" refString="ESTEBAN G. I. & NASIF N. L. 1996. - Nuevos Dasypodidae (Mammalia, Xenarthra) del Mioceno tardio del Valle del Cajon, Catamarca, Argentina. Ameghiniana 33 (3): 327 - 334." type="journal article" year="1996">Esteban & Nasif, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
(CRILAR-Pv 119);
|
||
<emphasis id="B947EADA134FD93FFB100892FB1C0D5D" bold="true" box="[1163,1180,1529,1546]" pageId="6" pageNumber="5">M</emphasis>
|
||
,
|
||
<emphasis id="B947EADA134FD93FFB330892FB360D5D" bold="true" box="[1192,1206,1529,1546]" pageId="6" pageNumber="5">N</emphasis>
|
||
,
|
||
<taxonomicName id="4C334D4B134FD93FFB590892FEE50D76" authority="(Moreno & Mercerat, 1891)" baseAuthorityName="Moreno & Mercerat" baseAuthorityYear="1891" class="Mammalia" family="Dasypodidae" genus="Macrochorobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="6" pageNumber="5" phylum="Chordata" rank="species" species="scalabrinii">
|
||
<emphasis id="B947EADA134FD93FFB590892FA2F0D5D" box="[1218,1455,1529,1546]" italics="true" pageId="6" pageNumber="5">Macrochorobates scalabrinii</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA24B39134FD93FFF110B7BFEE00D76" author="MORENO F. P. & MERCERAT A." box="[138,352,1552,1569]" pageId="6" pageNumber="5" pagination="222 - 236" refId="ref14616" refString="MORENO F. P. & MERCERAT A. 1891. - Exploracion arqueologica de la Provincia de Catamarca: paleontologia. Revista del Museo de La Plata 1: 222 - 236." type="journal article" year="1891">Moreno & Mercerat, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(CRILAR-Pv 120);
|
||
<emphasis id="B947EADA134FD93FFD9B0B7BFDA20D76" bold="true" box="[512,546,1552,1569]" pageId="6" pageNumber="5">O -R</emphasis>
|
||
,
|
||
<taxonomicName id="4C334D4B134FD93FFDB60B7BFD2E0D76" box="[557,686,1552,1569]" class="Mammalia" family="Dasypodidae" genus="Prozaedyus" kingdom="Animalia" order="Cingulata" pageId="6" pageNumber="5" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B947EADA134FD93FFDB60B7BFD0D0D76" box="[557,653,1552,1569]" italics="true" pageId="6" pageNumber="5">Prozaedyus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
(CRILAR-Pv 121). Scale bars: 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C36C8134FD93FFF000A19FB670C3B" blockId="6.[130,777,1619,2028]" lastBlockId="6.[811,1457,1619,2028]" pageId="6" pageNumber="5">The ornamentation pattern of the dorsal surface is composed by a wide and slightly convex central figure, and a variable number (4-5) of minor peripheral figures restricted to the anterior half of the osteoderm. The central figure is sub-elliptical, and is delimited by a main sulcus containing several small dorsal foramina, which are more numerous in the proximal area. Towards the posterior margin of the osteoderm, this sulcus becomes progressively shallower until disappearing, so that the central figure becomes unified with the remaining dorsal surface of the osteoderm. The peripheral figures, anteriorly placed, are sub-pentagonal, and delimited by minor sulci. Contrary to the main sulcus, these sulci have no dorsal foramina and are much deeper.</paragraph>
|
||
<paragraph id="8B8C36C8134FD93EFCD80A19FEF90A26" blockId="6.[811,1457,1619,2028]" lastBlockId="7.[131,777,216,752]" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5">
|
||
In the posterior margin, there is a variable number of large piliferous foramina, posteriorly oriented and arranged into a single row. Osteoderm CRILAR-Pv 116 has just one foramen, while osteoderm CRILAR-Pv 117 has a row of three piliferous foramina. There are no foramina at the lateral margins of the fixed osteoderms. By contrast, there are two piliferous foramina in the external margin of the preserved fragment of the osteoderm from the right lateral border of the carapace (
|
||
<figureCitation id="13082A4D134ED93EFE830C3CFEEC0A26" box="[280,364,343,369]" captionStart="FIG" captionStartId="6.[132,143,1507,1524]" captionTargetBox="[132,1456,185,1474]" captionTargetId="figure-276@6.[698,1267,743,1285]" captionTargetPageId="6" captionText="FIG. 3. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-F, Vetelia gandhii Esteban & Nasif, 1996 (CRILAR-Pv 116; CRILAR-Pv 117; CRILAR-Pv 118); G-L, Chasicotatus peiranoi Esteban & Nasif, 1996 (CRILAR-Pv 119); M, N, Macrochorobates scalabrinii (Moreno & Mercerat, 1891) (CRILAR-Pv 120); O-R, Prozaedyus sp. (CRILAR-Pv 121). Scale bars: 1 cm." pageId="7" pageNumber="6">Fig. 3C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8C36C8134ED93EFF000C1CFD7609A7" blockId="7.[131,777,216,752]" pageId="7" pageNumber="6">
|
||
Juvenile osteoderms. These osteoderms (CRILAR-Pv 118;
|
||
<figureCitation id="13082A4D134ED93EFF180CFCFF410AE6" box="[131,193,407,433]" captionStart="FIG" captionStartId="6.[132,143,1507,1524]" captionTargetBox="[132,1456,185,1474]" captionTargetId="figure-276@6.[698,1267,743,1285]" captionTargetPageId="6" captionText="FIG. 3. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-F, Vetelia gandhii Esteban & Nasif, 1996 (CRILAR-Pv 116; CRILAR-Pv 117; CRILAR-Pv 118); G-L, Chasicotatus peiranoi Esteban & Nasif, 1996 (CRILAR-Pv 119); M, N, Macrochorobates scalabrinii (Moreno & Mercerat, 1891) (CRILAR-Pv 120); O-R, Prozaedyus sp. (CRILAR-Pv 121). Scale bars: 1 cm." pageId="7" pageNumber="6">Fig. 3</figureCitation>
|
||
D-F) are of similar shape to those of fixed osteoderms of adult individuals, but can be recognized as belonging to juvenile stage due to several features, such as a much smaller size (
|
||
<tableCitation id="C6B10373134ED93EFF250C9CFE910946" box="[190,273,503,529]" captionStart="TABLE" captionStartId="8.[526,537,219,236]" captionTargetPageId="8" captionText="TABLE 1. — Measurements (in mm) of the described osteoderms." httpUri="http://table.plazi.org/id/DF4C66401341D931FD950DB0FBA80BBB" pageId="7" pageNumber="6" tableUuid="DF4C66401341D931FD950DB0FBA80BBB">Table 1</tableCitation>
|
||
); undeveloped central and peripheral figures and very punctuate dorsal and ventral surfaces. The same morphological differences have been reported for juveniles of other cingulate groups such as glyptodonts (see
|
||
<bibRefCitation id="EFA24B39134ED93EFD2A0F3CFECF09C7" author="LUNA C. A. & KRAPOVICKAS J. M." pageId="7" pageNumber="6" pagination="135 - 141" refId="ref14253" refString="LUNA C. A. & KRAPOVICKAS J. M. 2011. - Primer registro de un ejemplar juvenil de Glyptodon sp. (Cingulata, Glyptodontidae) del Cuaternario de la provincia de Cordoba, Argentina. Mastozoologia Neotropical 18: 135 - 141." type="journal article" year="2011">Luna & Krapovickas 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA24B39134ED93EFEC40F1DFDA909C7" author="ZURITA A. E. & OLIVA C. & DONDAS A. & SOIBELZON E. & ISLA F. I." box="[351,553,630,656]" pageId="7" pageNumber="6" pagination="69 - 75" refId="ref16467" refString="ZURITA A. E., OLIVA C., DONDAS A., SOIBELZON E. & ISLA F. I. 2011. - El registro mas completo de un Hoplophorini (Xenarthra: Glyptodontidae) para los Pisos / Edades Chapadmalalense- Marplatense (Plioceno tardio-Pleistoceno temprano). Revista del Museo Argentino de Ciencias Naturales " Bernardino Rivadavia " 13: 69 - 75." type="journal article" year="2011">
|
||
Zurita
|
||
<emphasis id="B947EADA134ED93EFE2B0F1CFE6709C7" box="[432,487,630,656]" italics="true" pageId="7" pageNumber="6">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA24B39134ED93EFDA10F1CFD7609C7" author="LUNA C. A. & CERDA I. A. & ZURITA A. E. & GONZALEZ R. & PRIETO M. C. & MOTHE D. & AVILLA L. S." box="[570,758,630,657]" pageId="7" pageNumber="6" pagination="59 - 170" refId="ref14300" refString="LUNA C. A., CERDA I. A., ZURITA A. E., GONZALEZ R., PRIETO M. C., MOTHE D. & AVILLA L. S. 2018. - Distinguishing Quaternary glyptodontine cingulates in South America: how informative are juvenile specimens?. Acta Palaeontologica Polonica 63 (1): 59 - 170. https: // doi. org / 10.4202 / app. 00409.2017" type="journal article" year="2018">
|
||
Luna
|
||
<emphasis id="B947EADA134ED93EFDE70F1CFD3409C7" box="[636,692,630,656]" italics="true" pageId="7" pageNumber="6">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). Additionally, these osteoderms have a series of conspicuous dorsal foramina which correspond with the location of the main sulcus in osteoderms belonging to adult individuals.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3296543134ED93EFF1F0E7DFC870FD8" pageId="7" pageNumber="6" type="discussion">
|
||
<paragraph id="8B8C36C8134ED93EFF1F0E7DFF680879" blockId="7.[130,776,790,1167]" box="[132,232,790,816]" pageId="7" pageNumber="6">
|
||
<smallCapsWord id="8D6AA014134ED93EFF1F0E7DFF680879" baselines="809,809" box="[132,232,790,816]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="7" pageNumber="6">REMARKS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B8C36C8134ED93EFF1F0E5DFC870FD8" blockId="7.[130,776,790,1167]" pageId="7" pageNumber="6">
|
||
<taxonomicName id="4C334D4B134ED93EFF1F0E5DFF470807" authorityName="Ameghino" authorityYear="1891" box="[132,199,822,848]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="7" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="B947EADA134ED93EFF1F0E5DFF470807" box="[132,199,822,848]" italics="true" pageId="7" pageNumber="6">Vetelia</emphasis>
|
||
</taxonomicName>
|
||
includes
|
||
<taxonomicName id="4C334D4B134ED93EFEB30E5DFE050818" authorityName="Esteban & Nasif" authorityYear="1996" box="[296,389,822,847]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="7" pageNumber="6" phylum="Chordata" rank="species" species="puncta">
|
||
<emphasis id="B947EADA134ED93EFEB30E5DFE050818" box="[296,389,822,847]" italics="true" pageId="7" pageNumber="6">V. puncta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D4B134ED93EFE0A0E5DFD560807" authority="Scillato-Yane, 1977" authorityName="Scillato-Yane" authorityYear="1977" box="[401,726,822,848]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="7" pageNumber="6" phylum="Chordata" rank="species" species="perforata">
|
||
<emphasis id="B947EADA134ED93EFE0A0E5DFD850807" box="[401,517,822,848]" italics="true" pageId="7" pageNumber="6">V. perforata</emphasis>
|
||
<bibRefCitation id="EFA24B39134ED93EFD920E5DFD560807" author="SCILLATO-YANE G. J." box="[521,726,822,848]" pageId="7" pageNumber="6" pagination="133 - 144" refId="ref15531" refString="SCILLATO-YANE G. J. 1977. - Notas sobre los Dasypodidae (Mammalia, Edentata) del Plioceno del territorio argentino I. Los restos de edad Chasiquense (Plioceno inferior) del sur de la Provincia de Buenos Aires. Ameghiniana 14: 133 - 144." type="journal article" year="1977">Scillato-Yané, 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C334D4B134ED93EFF190E3DFF6C0838" authorityName="Esteban & Nasif" authorityYear="1996" box="[130,236,853,879]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="7" pageNumber="6" phylum="Chordata" rank="species" species="ghandii">
|
||
<emphasis id="B947EADA134ED93EFF190E3DFF6C0838" box="[130,236,853,879]" italics="true" pageId="7" pageNumber="6">V. ghandii</emphasis>
|
||
</taxonomicName>
|
||
. The analyzed specimens are assigned to
|
||
<taxonomicName id="4C334D4B134ED93EFD000E3DFC850838" authorityName="Esteban & Nasif" authorityYear="1996" box="[667,773,853,879]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="7" pageNumber="6" phylum="Chordata" rank="species" species="ghandii">
|
||
<emphasis id="B947EADA134ED93EFD000E3DFC850838" box="[667,773,853,879]" italics="true" pageId="7" pageNumber="6">V. ghandii</emphasis>
|
||
</taxonomicName>
|
||
for having large and scarce piliferous foramina (1-3) at the posterior margin. In
|
||
<taxonomicName id="4C334D4B134ED93EFEFD0EFDFE5E08F8" authorityName="Scillato-Yane" authorityYear="1977" box="[358,478,917,943]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="7" pageNumber="6" phylum="Chordata" rank="species" species="perforata">
|
||
<emphasis id="B947EADA134ED93EFEFD0EFDFE5E08F8" box="[358,478,917,943]" italics="true" pageId="7" pageNumber="6">V. perforata</emphasis>
|
||
</taxonomicName>
|
||
foramina are much smaller and numerous (6-14) (see
|
||
<bibRefCitation id="EFA24B39134ED93EFE0B0EDEFDD20898" author="SCILLATO-YANE G. J." box="[400,594,949,975]" pageId="7" pageNumber="6" pagination="133 - 144" refId="ref15531" refString="SCILLATO-YANE G. J. 1977. - Notas sobre los Dasypodidae (Mammalia, Edentata) del Plioceno del territorio argentino I. Los restos de edad Chasiquense (Plioceno inferior) del sur de la Provincia de Buenos Aires. Ameghiniana 14: 133 - 144." type="journal article" year="1977">Scillato-Yané 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA24B39134ED93EFDC40EDEFD150898" author="SCILLATO-YANE G. J." box="[607,661,949,975]" pageId="7" pageNumber="6" refId="ref15627" refString="SCILLATO-YANE G. J. 1982. - Los Dasypodidae (Mammalia: Edentata) del Plioceno y Pleistoceno de Argentina. PhD Thesis, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, 127 p." type="book" year="1982">1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA24B39134ED93EFD390EDEFF7A08B8" author="BARASOAIN D. & GONZALEZ RUIZ L. R. & TOMASSINI R. L. & ZURITA A. E. & CONTRERAS V. H. & MONTALVO C. I." pageId="7" pageNumber="6" pagination="2791 - 2797" refId="ref10187" refString="BARASOAIN D., GONZALEZ RUIZ L. R., TOMASSINI R. L., ZURITA A. E., CONTRERAS V. H. & MONTALVO C. I. 2021. - First phylogenetic analysis of the Miocene armadillo Vetelia reveals novel affinities with Tolypeutinae. Acta Paleontologica Polonica 66: 2791 - 2797" type="journal article" year="2021">
|
||
Barasoain
|
||
<emphasis id="B947EADA134ED93EFF1F0EBDFF3A08B8" box="[132,186,981,1007]" italics="true" pageId="7" pageNumber="6">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
). In turn,
|
||
<taxonomicName id="4C334D4B134ED93EFEEE0EBDFE5A08B8" authorityName="Esteban & Nasif" authorityYear="1996" box="[373,474,982,1007]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="7" pageNumber="6" phylum="Chordata" rank="species" species="puncta">
|
||
<emphasis id="B947EADA134ED93EFEEE0EBDFE5A08B8" box="[373,474,982,1007]" italics="true" pageId="7" pageNumber="6">V. puncta</emphasis>
|
||
</taxonomicName>
|
||
osteoderms develop several small piliferous foramina at the posterior margin (
|
||
<bibRefCitation id="EFA24B39134ED93EFD0D0E9EFF3C0F78" author="AMEGHINO F." pageId="7" pageNumber="6" pagination="240 - 288" refId="ref9779" refString="AMEGHINO F. 1891. - Mamiferos y aves fosiles argentinas. Revista Argentina de Historia Natural 1: 240 - 288." type="journal article" year="1891">Ameghino 1891</bibRefCitation>
|
||
), several dorsal foramina in both main and minor sulci, and smaller piliferous foramina are present at the anterior and lateral margins of the osteoderm, which are absent in
|
||
<taxonomicName id="4C334D4B134ED93EFF1D091EFF700FD9" authorityName="Esteban & Nasif" authorityYear="1996" box="[134,240,1140,1166]" class="Mammalia" family="Dasypodidae" genus="Vetelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="7" pageNumber="6" phylum="Chordata" rank="species" species="ghandii">
|
||
<emphasis id="B947EADA134ED93EFF1D091EFF700FD9" box="[134,240,1140,1166]" italics="true" pageId="7" pageNumber="6">V. ghandii</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFA24B39134ED93EFEBA091FFD8E0FD9" author="ESTEBAN G. I. & NASIF N. L." box="[289,526,1140,1167]" pageId="7" pageNumber="6" pagination="327 - 334" refId="ref12563" refString="ESTEBAN G. I. & NASIF N. L. 1996. - Nuevos Dasypodidae (Mammalia, Xenarthra) del Mioceno tardio del Valle del Cajon, Catamarca, Argentina. Ameghiniana 33 (3): 327 - 334." type="journal article" year="1996">Esteban & Nasif 1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA24B39134ED93EFD81091FFD770FD8" author="BARASOAIN D. & GONZALEZ RUIZ L. R. & TOMASSINI R. L. & ZURITA A. E. & CONTRERAS V. H. & MONTALVO C. I." box="[538,759,1140,1167]" pageId="7" pageNumber="6" pagination="2791 - 2797" refId="ref10187" refString="BARASOAIN D., GONZALEZ RUIZ L. R., TOMASSINI R. L., ZURITA A. E., CONTRERAS V. H. & MONTALVO C. I. 2021. - First phylogenetic analysis of the Miocene armadillo Vetelia reveals novel affinities with Tolypeutinae. Acta Paleontologica Polonica 66: 2791 - 2797" type="journal article" year="2021">
|
||
Barasoain
|
||
<emphasis id="B947EADA134ED93EFD13091EFD3A0FD9" box="[648,698,1140,1166]" italics="true" pageId="7" pageNumber="6">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |