260 lines
38 KiB
XML
260 lines
38 KiB
XML
<document id="6CF788C676467412A31E59DB1FBEC715" ID-CLB-Dataset="305481" ID-DOI="10.5852/cr-palevol2023v22a1" ID-GBIF-Dataset="09614d02-74d1-413d-9cd8-ae29a8cb089b" ID-ISSN="1777-571X" ID-Zenodo-Dep="14204179" ID-ZooBank="urn:lsid:zoobank.org:pub:896BCEBD-8547-4822-9F61-58E7BECC9469" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1732277592039" checkinUser="felipe" docAuthor="Brandoni, Diego, Barasoain, Daniel & González Ruiz, Laureano R." docDate="2023" docId="039A87DE1340D932FCDD0858FE4D0E39" docLanguage="en" docName="CRPalevol.22.1.1-16.pdf" docOrigin="Comptes Rendus Palevol 22 (1)" docSource="http://dx.doi.org/10.5852/cr-palevol2023v22a1" docStyle="DocumentStyle:DACF86F01658A8850E13A63D98C287FD.1:CRPalevol.2020-.journal_article" docStyleId="DACF86F01658A8850E13A63D98C287FD" docStyleName="CRPalevol.2020-.journal_article" docStyleVersion="1" docTitle="Paleuphractus argentinus" docType="treatment" docVersion="2" lastPageNumber="10" masterDocId="FFA3FFA61349D939FF9B0D6BFF800B57" masterDocTitle="Late Miocene Dasypodidae Gray, 1821 (Xenarthra, Cingulata) from the Toro Negro Formation (Central Andes, Argentina): diversity and chronological and biogeographical implications" masterLastPageNumber="16" masterPageNumber="1" pageNumber="8" updateTime="1732728938826" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4F836590E4F1865B1C5E7B4B787C8A55">Late Miocene Dasypodidae Gray, 1821 (Xenarthra, Cingulata) from the Toro Negro Formation (Central Andes, Argentina): diversity and chronological and biogeographical implications</mods:title>
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<mods:namePart id="C5626D8904B9DEAC3F14C48C830DB83A">Brandoni, Diego</mods:namePart>
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<mods:affiliation id="C26DEFE0BD23FBC19FDB73000B39E77C">Laboratorio de Paleontología de Vertebrados, Centro de Investigación Científica y de Transferencia Tecnológica a la Producción - Consejo Nacional de Investigaciones Científicas y Técnicas (CICYTTP, CONICET-Prov. ER-UADER), España 149, E 3105 BWA Diamante (Argentina) dbrandoni @ cicyttp. org. ar (corresponding author)</mods:affiliation>
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<mods:nameIdentifier id="0F934B1FE3C70227DDDA9B3CF09239CB" type="email">dbrandoni@cicyttp.org.ar</mods:nameIdentifier>
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<mods:namePart id="26706A65367A25517C7F2A9CDFE2E00C">Barasoain, Daniel</mods:namePart>
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<mods:affiliation id="D6BD77412E34FB35F0BC7FF463D28F3D">Laboratorio de Evolución de Vertebrados y Ambientes Cenozoicos, Centro de Ecología Aplicada del Litoral - Universidad Nacional de Nordeste - Consejo Nacional de Investigaciones Científicas y Técnicas (CECOAL-UNNE-CONICET) y Cátedra de Paleontología, Facultad de Ciencias Exactas, Naturales y Agrimensura, Universidad Nacional del Nordeste, RP 5 3400 Corrientes (Argentina) danielbarasoain @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="BB5B510EFF55D8DB001C6AD736AC84C5" type="email">danielbarasoain@gmail.com</mods:nameIdentifier>
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<mods:namePart id="64EC2C2740B13E5BA8C2FC6B81517356">González Ruiz, Laureano R.</mods:namePart>
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<mods:affiliation id="11219C01D53C3655D58197741D5F23E2">Laboratorio de Investigaciones en Evolución y Biodiversidad (LIEB-FCNyCS sede Esquel, UNPSJB) y Centro de Investigaciones Esquel de Montaña y Estepa Patagónica (CIEMEP), Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Universidad Nacional de La Patagonia San Juan Bosco (UNPSJB), Roca 780, 9200, Esquel, Chubut (Argentina) lrgonzalezruiz @ comahue-conicet. gob. ar</mods:affiliation>
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<mods:nameIdentifier id="B9B3B0E68D92D59C30965266DDC4D7E9" type="email">lrgonzalezruiz@comahue-conicet.gob.ar</mods:nameIdentifier>
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<mods:title id="DF258FAB8CE217B2825AEA7266C741D3">Comptes Rendus Palevol</mods:title>
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<mods:part id="7B82E1824B9193C4602046B64FEB020A">
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<mods:date id="F4E171F40FB92E2A681483166915B549">2023</mods:date>
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<mods:number id="FE78AAD8D30F8DDAB8C345CA8D58F643">2023-01-16</mods:number>
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<mods:url id="48CF22B8A9CA4B788CFFD7898714D3C2">http://dx.doi.org/10.5852/cr-palevol2023v22a1</mods:url>
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<mods:identifier id="5EF1342CB3D106CABA978E45234D726B" type="DOI">10.5852/cr-palevol2023v22a1</mods:identifier>
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<treatment id="039A87DE1340D932FCDD0858FE4D0E39" ID-DOI="http://doi.org/10.5281/zenodo.14204193" ID-GBIF-Taxon="247568580" ID-Zenodo-Dep="14204193" LSID="urn:lsid:plazi:treatment:039A87DE1340D932FCDD0858FE4D0E39" httpUri="http://treatment.plazi.org/id/039A87DE1340D932FCDD0858FE4D0E39" lastPageId="11" lastPageNumber="10" pageId="9" pageNumber="8" scope_class="Mammalia" scope_family="Dasypodidae" scope_order="Xenarthra" scope_subFamily="Euphractinae" scope_subOrder="Cingulata" scope_tribe="Euphractini">
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<subSubSection id="C32965431340D930FCDD0858FA150E19" box="[838,1429,1331,1358]" pageId="9" pageNumber="8" type="nomenclature">
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<paragraph id="8B8C36C81340D930FCDD0858FA150E19" blockId="9.[838,1429,1331,1358]" box="[838,1429,1331,1358]" pageId="9" pageNumber="8">
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<heading id="D0C481A41340D930FCDD0858FA150E19" box="[838,1429,1331,1358]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="8" reason="2">
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<taxonomicName id="4C334D4B1340D930FCDD0858FA150E19" authority="(Moreno & Mercerat, 1891)" baseAuthorityName="Moreno & Mercerat" baseAuthorityYear="1891" box="[838,1429,1331,1358]" class="Mammalia" family="Dasypodidae" genus="Paleuphractus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xenarthra" pageId="9" pageNumber="8" phylum="Chordata" rank="species" species="argentinus">
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<emphasis id="B947EADA1340D930FCDD0858FBD90E19" bold="true" box="[838,1113,1331,1358]" italics="true" pageId="9" pageNumber="8">Paleuphractus argentinus</emphasis>
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(
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<bibRefCitation id="EFA24B391340D930FBFC085FFA0E0E19" author="MORENO F. P. & MERCERAT A." box="[1127,1422,1332,1358]" pageId="9" pageNumber="8" pagination="222 - 236" refId="ref14616" refString="MORENO F. P. & MERCERAT A. 1891. - Exploracion arqueologica de la Provincia de Catamarca: paleontologia. Revista del Museo de La Plata 1: 222 - 236." type="journal article" year="1891">Moreno & Mercerat, 1891</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32965431340D930FB83083FFB440E39" box="[1048,1220,1364,1390]" pageId="9" pageNumber="8" type="description">
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<paragraph id="8B8C36C81340D930FB83083FFB440E39" blockId="9.[1048,1220,1364,1390]" box="[1048,1220,1364,1390]" pageId="9" pageNumber="8">
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(
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<figureCitation id="13082A4D1340D930FBBA083FFBDF0E39" box="[1057,1119,1364,1390]" captionStart="FIG" captionStartId="10.[132,143,885,902]" captionTargetBox="[133,1448,223,843]" captionTargetId="figure-641@10.[698,1267,213,754]" captionTargetPageId="10" captionText="FIG. 4. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-N, Paleuphractus argentinus (Moreno & Mercerat, 1891) (CRILAR-Pv 122). Scale bars: 1 cm." pageId="9" pageNumber="8">Fig. 4</figureCitation>
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;
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<tableCitation id="C6B103731340D930FBF0083FFB3B0E39" box="[1131,1211,1364,1390]" captionStart="TABLE" captionStartId="8.[526,537,219,236]" captionTargetPageId="8" captionText="TABLE 1. — Measurements (in mm) of the described osteoderms." httpUri="http://table.plazi.org/id/DF4C66401341D931FD950DB0FBA80BBB" pageId="9" pageNumber="8" tableUuid="DF4C66401341D931FD950DB0FBA80BBB">Table 1</tableCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32965431340D930FCB608E6FABA0EF2" box="[813,1338,1421,1445]" pageId="9" pageNumber="8" type="reference_group">
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<paragraph id="8B8C36C81340D930FCB608E6FABA0EF2" blockId="9.[813,1338,1421,1445]" box="[813,1338,1421,1445]" pageId="9" pageNumber="8">
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<treatmentCitationGroup id="AB2311E61340D930FCB608E6FABA0EF2" box="[813,1338,1421,1445]" pageId="9" pageNumber="8">
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<taxonomicName id="4C334D4B1340D930FCB608E6FC600EF3" authorityName="Moreno & Mercerat" authorityYear="1891" box="[813,992,1421,1444]" class="Mammalia" family="Dasypodidae" genus="Dasypus" kingdom="Animalia" order="Xenarthra" pageId="9" pageNumber="8" phylum="Chordata" rank="species" species="argentinus">
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<emphasis id="B947EADA1340D930FCB608E6FC600EF3" box="[813,992,1421,1444]" italics="true" pageId="9" pageNumber="8">Dasypus argentinus</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0A9210D91340D930FC6108E6FAB60EF2" author="MORENO F. P. & MERCERAT A." box="[1018,1334,1421,1445]" page="227" pageId="9" pageNumber="8" year="1891">
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<bibRefCitation id="EFA24B391340D930FC6108E6FAB60EF2" author="MORENO F. P. & MERCERAT A." box="[1018,1334,1421,1445]" pageId="9" pageNumber="8" pagination="222 - 236" refId="ref14616" refString="MORENO F. P. & MERCERAT A. 1891. - Exploracion arqueologica de la Provincia de Catamarca: paleontologia. Revista del Museo de La Plata 1: 222 - 236." type="journal article" year="1891">Moreno & Mercerat, 1891: 227</bibRefCitation>
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</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32965431340D930FCB608D6FB380D73" pageId="9" pageNumber="8" type="materials_examined">
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<paragraph id="8B8C36C81340D930FCB608D6FB380D73" blockId="9.[811,1458,1468,1572]" pageId="9" pageNumber="8">
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||
<smallCapsWord id="8D6AA0141340D930FCB608D6FC100E84" baselines="1486,1487" box="[813,912,1469,1492]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Material" pageId="9" pageNumber="8">MATERIAL</smallCapsWord>
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<smallCapsWord id="8D6AA0141340D930FC0308A9FC770E85" baselines="1486" box="[920,1015,1474,1490]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="referred" pageId="9" pageNumber="8">REFERRED</smallCapsWord>
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.
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<emphasis id="B947EADA1340D930FB9E08D6FB9E0E83" box="[1029,1054,1469,1492]" italics="true" pageId="9" pageNumber="8">—</emphasis>
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<materialsCitation id="3B5B3C951340D930FBB308D7FB380D73" ID-GBIF-Occurrence="4985208301" pageId="9" pageNumber="8" specimenCode="CRILAR-Pv 122" specimenCount="1">
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<specimenCode id="DB959EB31340D930FBB308D7FB520E83" box="[1064,1234,1468,1492]" pageId="9" pageNumber="8">CRILAR-Pv 122</specimenCode>
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, several associated elements, including 15 fixed osteoderms, nine mobile osteoderms, one nuchal osteoderm and seven osteoderms of the carapace borders, plus several broken postcranial elements.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32965431340D930FCB60B50FA320DF4" pageId="9" pageNumber="8" type="distribution">
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||
<paragraph id="8B8C36C81340D930FCB60B50FA320DF4" blockId="9.[806,1458,1595,1699]" pageId="9" pageNumber="8">
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<smallCapsWord id="8D6AA0141340D930FC200B2AFC660D05" baselines="1614" box="[955,998,1601,1618]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="and" pageId="9" pageNumber="8">AND</smallCapsWord>
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<smallCapsWord id="8D6AA0141340D930FC740B2AFB0A0D05" baselines="1614" box="[1007,1162,1601,1618]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="stratigraphic" pageId="9" pageNumber="8">STRATIGRAPHIC</smallCapsWord>
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<smallCapsWord id="8D6AA0141340D930FB080B2AFA980D05" baselines="1614" box="[1171,1304,1601,1618]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="provenance" pageId="9" pageNumber="8">PROVENANCE</smallCapsWord>
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.
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<emphasis id="B947EADA1340D930FABC0B57FAC10D04" box="[1319,1345,1596,1619]" italics="true" pageId="9" pageNumber="8">—</emphasis>
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Quebrada de Las Torrecillas (
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||
<geoCoordinate id="EE07500F1340D930FC7C0B3DFBDA0D39" box="[999,1114,1622,1646]" degrees="28" direction="south" minutes="36" orientation="latitude" pageId="9" pageNumber="8" precision="15" seconds="37" value="-28.61028">28°36’37”S</geoCoordinate>
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,
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<geoCoordinate id="EE07500F1340D930FBFD0B3DFB620D39" box="[1126,1250,1622,1646]" degrees="68" direction="west" minutes="14" orientation="longitude" pageId="9" pageNumber="8" precision="15" seconds="14" value="-68.23722">68°14’14”W</geoCoordinate>
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),
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<collectingRegion id="49F7F82A1340D930FB620B3DFA2B0D39" box="[1273,1451,1622,1646]" country="Argentina" name="La Rioja" pageId="9" pageNumber="8">La Rioja Province</collectingRegion>
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,
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<collectingCountry id="F32476581340D930FCBD0B1BFC070DDF" box="[806,903,1648,1672]" name="Argentina" pageId="9" pageNumber="8">Argentina</collectingCountry>
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||
(
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<figureCitation id="13082A4D1340D930FC060B1BFC560DDF" box="[925,982,1648,1672]" captionStart="FIG" captionStartId="4.[134,143,1284,1301]" captionTargetBox="[132,1455,215,1241]" captionTargetId="figure-487@4.[698,1267,213,774]" captionTargetPageId="4" captionText="FIG. 1. — A, Map showing the study area (black circle), La Rioja Province, Argentina; B, panoramic view of Toro Negro Formation at road 76 between Vinchina and Jagüel; C, panoramic view of Toro Negro Formation at Quebrada de Las Torrecillas; D, detail of the lower member of the Toro Negro Formation at Quebrada de Las Torrecillas. Credits: map modified from Google Earth." pageId="9" pageNumber="8">Fig. 1</figureCitation>
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). Upper levels of the lower member of the Toro Negro Formation (Messinian-Zanclean; Late Miocene-Early Pliocene).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32965431340D932FCB10BB8FD150867" lastPageId="11" lastPageNumber="10" pageId="9" pageNumber="8" type="description">
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<paragraph id="8B8C36C81340D930FCB10BB8FC3D0DBC" blockId="9.[810,1457,1747,2028]" box="[810,957,1747,1773]" pageId="9" pageNumber="8">D ESCRIPTION</paragraph>
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<paragraph id="8B8C36C81340D933FCB60B99FD9C0F78" blockId="9.[810,1457,1747,2028]" lastBlockId="10.[131,775,981,1071]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
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Nuchal osteoderm. A single osteoderm (
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<figureCitation id="13082A4D1340D930FB560B99FA9B0C5A" box="[1229,1307,1778,1805]" captionStart="FIG" captionStartId="10.[132,143,885,902]" captionTargetBox="[133,1448,223,843]" captionTargetId="figure-641@10.[698,1267,213,754]" captionTargetPageId="10" captionText="FIG. 4. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-N, Paleuphractus argentinus (Moreno & Mercerat, 1891) (CRILAR-Pv 122). Scale bars: 1 cm." pageId="9" pageNumber="8">Fig. 4A</figureCitation>
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) of the nuchal bands was recovered. As in mobile osteoderms (see below), it is divided into an articular and an ornamented portion. However, it is smaller and has a very reduced articular portion in relation to osteoderms of the mobile bands. It has a rectangular outline. The ornamentation includes flat and poorly developed central and lateral figures, which are only partially delimited by shallow sulci. At the posterior margin there is a single row of 6 piliferous foramina but, contrary to the osteoderms of the mobile bands, there are 3 to 4 much larger foramina at the lateral margins.
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</paragraph>
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<caption id="DF4C66401343D933FF1F0E1EFDB708CA" pageId="10" pageNumber="9" startId="10.[132,143,885,902]" targetBox="[133,1448,223,843]" targetPageId="10" targetType="figure">
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<paragraph id="8B8C36C81343D933FF1F0E1EFDB708CA" blockId="10.[132,1456,885,925]" pageId="10" pageNumber="9">
|
||
<smallCapsWord id="8D6AA0141343D933FF1F0E1EFF1D08D2" baselines="898,898" box="[132,157,885,902]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="10" pageNumber="9">FIG</smallCapsWord>
|
||
. 4. —
|
||
<taxonomicName id="4C334D4B1343D933FF4E0E1EFEC308D1" authorityName="Gray" authorityYear="1821" box="[213,323,885,902]" class="Mammalia" family="Dasypodidae" kingdom="Animalia" order="Cingulata" pageId="10" pageNumber="9" phylum="Chordata" rank="family">Dasypodidae</taxonomicName>
|
||
from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina:
|
||
<emphasis id="B947EADA1343D933FBCF0E1EFBF608D1" bold="true" box="[1108,1142,885,902]" pageId="10" pageNumber="9">A -N</emphasis>
|
||
,
|
||
<taxonomicName id="4C334D4B1343D933FB1B0E1EFE8808CA" authority="(Moreno & Mercerat, 1891)" baseAuthorityName="Moreno & Mercerat" baseAuthorityYear="1891" class="Mammalia" family="Dasypodidae" genus="Paleuphractus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="10" pageNumber="9" phylum="Chordata" rank="species" species="argentinus">
|
||
<emphasis id="B947EADA1343D933FB1B0E1EFAD408D1" box="[1152,1364,885,902]" italics="true" pageId="10" pageNumber="9">Paleuphractus argentinus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA24B391343D933FAC60E1EFE8308CA" author="MORENO F. P. & MERCERAT A." pageId="10" pageNumber="9" pagination="222 - 236" refId="ref14616" refString="MORENO F. P. & MERCERAT A. 1891. - Exploracion arqueologica de la Provincia de Catamarca: paleontologia. Revista del Museo de La Plata 1: 222 - 236." type="journal article" year="1891">Moreno & Mercerat, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(CRILAR-Pv 122). Scale bars: 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C36C81343D933FF1F093EFEB00F38" blockId="10.[132,304,1109,1135]" box="[132,304,1109,1135]" pageId="10" pageNumber="9">
|
||
<heading id="D0C481A41343D933FF1F093EFEB00F38" box="[132,304,1109,1135]" fontSize="11" level="2" pageId="10" pageNumber="9" reason="8">
|
||
<emphasis id="B947EADA1343D933FF1F093EFEB00F38" box="[132,304,1109,1135]" italics="true" pageId="10" pageNumber="9">Fixed osteoderms</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B8C36C81343D933FF1F091FFE390E79" blockId="10.[131,778,1140,2028]" pageId="10" pageNumber="9">
|
||
These osteoderms (
|
||
<figureCitation id="13082A4D1343D933FED3091FFE020FD8" box="[328,386,1140,1167]" captionStart="FIG" captionStartId="10.[132,143,885,902]" captionTargetBox="[133,1448,223,843]" captionTargetId="figure-641@10.[698,1267,213,754]" captionTargetPageId="10" captionText="FIG. 4. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-N, Paleuphractus argentinus (Moreno & Mercerat, 1891) (CRILAR-Pv 122). Scale bars: 1 cm." pageId="10" pageNumber="9">Fig. 4</figureCitation>
|
||
B-G) can be either rectangular or pentagonal in outline, and are slightly longer than wide (
|
||
<tableCitation id="C6B103731343D933FD3109FFFD770FF8" box="[682,759,1172,1199]" captionStart="TABLE" captionStartId="8.[526,537,219,236]" captionTargetPageId="8" captionText="TABLE 1. — Measurements (in mm) of the described osteoderms." httpUri="http://table.plazi.org/id/DF4C66401341D931FD950DB0FBA80BBB" pageId="10" pageNumber="9" tableUuid="DF4C66401341D931FD950DB0FBA80BBB">Table 1</tableCitation>
|
||
). The entire dorsal surface is covered by very small foramina, very abundant in some osteoderms while practically absent in others. All osteoderms have a constant thickness from the anterior to posterior margin.
|
||
</paragraph>
|
||
<paragraph id="8B8C36C81343D933FF00085FFEE70CDB" blockId="10.[131,778,1140,2028]" pageId="10" pageNumber="9">The ornamentation pattern includes a wide and sub-elliptical central figure surrounded laterally and anteriorly by a variable number of peripheral figures. It extends posteriorly to reach the posterior margin, and deviates slightly toward the external lateral margin of the osteoderm. Towards the posterior margin it widens progressively, occupying more than one third of the osteoderm width. It is delimited by a shallow main sulcus. Anteriorly, this sulcus becomes shallower and disappears, so that the central figure merges with the dorsal surface. In lateral view, the central figure progressively increases its elevation until reaching the posterior margin, where it is much more elevated than the peripheral figures. Peripheral figures vary from 5 to 8, being seven in most cases. These figures are delimited by such shallow sulci and their limits are often diffuse or mostly indistinguishable. Some osteoderms develop large dorsal foramina at all the boundaries between main and minor sulci, while in others they are only present at some boundary or absent.</paragraph>
|
||
<paragraph id="8B8C36C81343D933FF000AF9FB550F18" blockId="10.[131,778,1140,2028]" lastBlockId="10.[811,1457,981,1326]" pageId="10" pageNumber="9">
|
||
Two of the fixed osteoderms have a particular morphology, including a deep cavity placed at the anterior limit of the central figure (
|
||
<figureCitation id="13082A4D1343D933FECF0ABAFE480CBB" box="[340,456,2001,2028]" captionStart="FIG" captionStartId="10.[132,143,885,902]" captionTargetBox="[133,1448,223,843]" captionTargetId="figure-641@10.[698,1267,213,754]" captionTargetPageId="10" captionText="FIG. 4. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-N, Paleuphractus argentinus (Moreno & Mercerat, 1891) (CRILAR-Pv 122). Scale bars: 1 cm." pageId="10" pageNumber="9">Fig. 4F, G</figureCitation>
|
||
). Such cavities are related to the presence of odoriferous glands, which are present in the pelvic shield of many euphractine armadillos (
|
||
<bibRefCitation id="EFA24B391343D933FAB80E9EFCE50F78" author="SCILLATO-YANE G. J." pageId="10" pageNumber="9" refId="ref15627" refString="SCILLATO-YANE G. J. 1982. - Los Dasypodidae (Mammalia: Edentata) del Plioceno y Pleistoceno de Argentina. PhD Thesis, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, 127 p." type="book" year="1982">Scillato-Yané 1982</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA24B391343D933FCE9097EFBF50F67" author="REDFORD K. H. & WETZEL R. M." box="[882,1141,1045,1072]" pageId="10" pageNumber="9" pagination="1 - 4" refId="ref15216" refString="REDFORD K. H. & WETZEL R. M. 1985. - Euphractus sexcinctus. Mammalian Species 252: 1 - 4" type="journal article" year="1985">Redford & Wetzel 1985</bibRefCitation>
|
||
). This structure has not been previously described for
|
||
<taxonomicName id="4C334D4B1343D933FBAF095EFB4E0F18" baseAuthorityName="Moreno & Mercerat" baseAuthorityYear="1891" box="[1076,1230,1077,1103]" class="Mammalia" family="Dasypodidae" genus="Paleuphractus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="10" pageNumber="9" phylum="Chordata" rank="species" species="argentinus">
|
||
<emphasis id="B947EADA1343D933FBAF095EFB4E0F18" box="[1076,1230,1077,1103]" italics="true" pageId="10" pageNumber="9">Pal. argentinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8C36C81343D933FCD8093EFBCE0E79" blockId="10.[811,1457,981,1326]" pageId="10" pageNumber="9">At the posterior margin, there are several small piliferous foramina, arranged in an irregular row. These foramina are variable in both number (5-10) and size, as some are twice the size of others, and are arranged in a zig-zag row. Although the lateral margins do not bear foramina, the row of piliferous foramina may extend slightly towards the more distal area of the external lateral margin.</paragraph>
|
||
<paragraph id="8B8C36C81343D933FCB6083FFC680E39" blockId="10.[813,1000,1364,1390]" box="[813,1000,1364,1390]" pageId="10" pageNumber="9">
|
||
<heading id="D0C481A41343D933FCB6083FFC680E39" box="[813,1000,1364,1390]" fontSize="11" level="2" pageId="10" pageNumber="9" reason="8">
|
||
<emphasis id="B947EADA1343D933FCB6083FFC680E39" box="[813,1000,1364,1390]" italics="true" pageId="10" pageNumber="9">Mobile osteoderms</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B8C36C81343D933FCB6081FFB910D7A" blockId="10.[811,1457,1396,2028]" pageId="10" pageNumber="9">
|
||
These osteoderms (
|
||
<figureCitation id="13082A4D1343D933FC65081FFBBE0ED9" box="[1022,1086,1396,1422]" captionStart="FIG" captionStartId="10.[132,143,885,902]" captionTargetBox="[133,1448,223,843]" captionTargetId="figure-641@10.[698,1267,213,754]" captionTargetPageId="10" captionText="FIG. 4. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-N, Paleuphractus argentinus (Moreno & Mercerat, 1891) (CRILAR-Pv 122). Scale bars: 1 cm." pageId="10" pageNumber="9">Fig. 4</figureCitation>
|
||
H-K) are rectangular, longer than wide (
|
||
<tableCitation id="C6B103731343D933FCEF08F8FC470EF9" box="[884,967,1427,1454]" captionStart="TABLE" captionStartId="8.[526,537,219,236]" captionTargetPageId="8" captionText="TABLE 1. — Measurements (in mm) of the described osteoderms." httpUri="http://table.plazi.org/id/DF4C66401341D931FD950DB0FBA80BBB" pageId="10" pageNumber="9" tableUuid="DF4C66401341D931FD950DB0FBA80BBB">Table 1</tableCitation>
|
||
). The articular portion represents less than one third of the total length. The transitional area is flat and unornamented. Some osteoderms develop a transverse ridge at the end of the transitional area, delimiting it from the ornamented portion.
|
||
</paragraph>
|
||
<paragraph id="8B8C36C81343D933FCD80B58FB500CBC" blockId="10.[811,1457,1396,2028]" pageId="10" pageNumber="9">The ornamentation pattern is composed by a central figure surrounded by 2 elongated and undivided lateral figures. The central figure is rounded to slightly keeled in section and sub-elliptical, and extends to the posterior margin. In some osteoderms it is straight, while in others it deviates slightly towards the external lateral margin. In contrast to the condition in fixed osteoderms, it gets much narrower towards the posterior margin. The central figure is delimited by a main sulcus, along which there are several dorsal foramina.Towards the anterior area of the ornamented portion, this sulcus becomes progressively shallower, so that the central figure merges with the dorsal surface. Lateral figures are parallel to the central figure. In some cases, some of these peripheral figures are subdivided by shallow sulci.</paragraph>
|
||
<paragraph id="8B8C36C81342D932FF000DB3FC850AC6" blockId="11.[130,777,216,816]" pageId="11" pageNumber="10">At the posterior margin there is a single row of 4 to 7 piliferous foramina, much smaller than those of fixed osteoderms. Also, in contrast to those of fixed osteoderms, these foramina are arranged into a straight line. In those osteoderms in which the central figure is slightly deviated, there are 2 to 3 piliferous foramina in the more ventral area of the external lateral margin.</paragraph>
|
||
<paragraph id="8B8C36C81342D932FF000CFCFD0B0926" blockId="11.[130,777,216,816]" pageId="11" pageNumber="10">
|
||
Osteoderms of the marginal rows. These osteoderms (
|
||
<figureCitation id="13082A4D1342D932FF150CDCFF4F0A86" box="[142,207,439,465]" captionStart="FIG" captionStartId="10.[132,143,885,902]" captionTargetBox="[133,1448,223,843]" captionTargetId="figure-641@10.[698,1267,213,754]" captionTargetPageId="10" captionText="FIG. 4. — Dasypodidae from the Toro Negro Formation at Quebrada de Las Torrecillas, La Rioja Province, Argentina: A-N, Paleuphractus argentinus (Moreno & Mercerat, 1891) (CRILAR-Pv 122). Scale bars: 1 cm." pageId="11" pageNumber="10">Fig. 4</figureCitation>
|
||
L-N) belong to the most lateral rows of the dorsal carapace. They are more elongated than regular osteoderms, and develop an irregular wedge shape. This is a representative morphology of both fossil and extant euphractine armadillos, which usually develop serrated lateral carapace borders (see
|
||
<bibRefCitation id="EFA24B391342D932FF2E0F3DFE040926" author="SCILLATO-YANE G. J." box="[181,388,598,625]" pageId="11" pageNumber="10" refId="ref15627" refString="SCILLATO-YANE G. J. 1982. - Los Dasypodidae (Mammalia: Edentata) del Plioceno y Pleistoceno de Argentina. PhD Thesis, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, 127 p." type="book" year="1982">Scillato-Yané 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA24B391342D932FE080F3DFDFB0926" author="KRMPOTIC C. M. & CIANCIO M. R. & BARBEITO C. & MARIO R. C. & CARLINI A. A." box="[403,635,598,625]" pageId="11" pageNumber="10" pagination="339 - 351" refId="ref14104" refString="KRMPOTIC C. M., CIANCIO M. R., BARBEITO C., MARIO R. C. & CARLINI A. A. 2009. - Osteoderm morphology in recent and fossil euphractine xenarthrans. Acta Zoologica 90 (4): 339 - 351. https: // doi. org / 10.1111 / j. 1463 - 6395.2008.00359. x" type="journal article" year="2009">
|
||
Krmpotic
|
||
<emphasis id="B947EADA1342D932FD9D0F3CFDBB0927" box="[518,571,598,624]" italics="true" pageId="11" pageNumber="10">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8C36C81342D932FF000F1DFD150867" blockId="11.[130,777,216,816]" pageId="11" pageNumber="10">Ornamentation is nearly or entirely absent, although some osteoderms have poorly developed central and peripheral figures. Some osteoderms of the lateral area develop more similar morphologies to those of other carapace areas, having a single row of piliferous foramina at the posterior margin which is absent in those belonging to the carapace border.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32965431342D932FF1F0E3DFE4D0E39" pageId="11" pageNumber="10" type="discussion">
|
||
<paragraph id="8B8C36C81342D932FF1F0E3DFF680839" blockId="11.[131,777,854,1391]" box="[132,232,854,880]" pageId="11" pageNumber="10">
|
||
<smallCapsWord id="8D6AA0141342D932FF1F0E3DFF680839" baselines="873,873" box="[132,232,854,880]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="11" pageNumber="10">REMARKS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B8C36C81342D932FF1F0E1EFE4D0E39" blockId="11.[131,777,854,1391]" pageId="11" pageNumber="10">
|
||
<taxonomicName id="4C334D4B1342D932FF1F0E1EFE9608D8" authorityName="Kraglievich" authorityYear="1934" box="[132,278,885,911]" class="Mammalia" family="Dasypodidae" genus="Paleuphractus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="11" pageNumber="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="B947EADA1342D932FF1F0E1EFE9608D8" box="[132,278,885,911]" italics="true" pageId="11" pageNumber="10">Paleuphractus</emphasis>
|
||
</taxonomicName>
|
||
is monotypic, with
|
||
<taxonomicName id="4C334D4B1342D932FE630E1EFD1708D8" baseAuthorityName="Moreno & Mercerat" baseAuthorityYear="1891" box="[504,663,885,911]" class="Mammalia" family="Dasypodidae" genus="Paleuphractus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="11" pageNumber="10" phylum="Chordata" rank="species" species="argentinus">
|
||
<emphasis id="B947EADA1342D932FE630E1EFD1708D8" box="[504,663,885,911]" italics="true" pageId="11" pageNumber="10">Pal. argentinus</emphasis>
|
||
</taxonomicName>
|
||
being the only recognized species. The material here analyzed can be assigned to
|
||
<taxonomicName id="4C334D4B1342D932FE960EDEFE2F0898" baseAuthorityName="Moreno & Mercerat" baseAuthorityYear="1891" box="[269,431,949,975]" class="Mammalia" family="Dasypodidae" genus="Paleuphractus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="11" pageNumber="10" phylum="Chordata" rank="species" species="argentinus">
|
||
<emphasis id="B947EADA1342D932FE960EDEFE2F0898" box="[269,431,949,975]" italics="true" pageId="11" pageNumber="10">Pal. argentinus</emphasis>
|
||
</taxonomicName>
|
||
based on a low central figure, mobile osteoderms with undivided lateral figures, and small piliferous foramina located on the posterior and external lateral margins (see
|
||
<bibRefCitation id="EFA24B391342D932FEF5097EFDC80F78" author="SCILLATO-YANE G. J." box="[366,584,1045,1071]" pageId="11" pageNumber="10" refId="ref15627" refString="SCILLATO-YANE G. J. 1982. - Los Dasypodidae (Mammalia: Edentata) del Plioceno y Pleistoceno de Argentina. PhD Thesis, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, 127 p." type="book" year="1982">Scillato-Yané 1982</bibRefCitation>
|
||
). It differs from
|
||
<taxonomicName id="4C334D4B1342D932FF1F095EFD930F07" authority="Scillato-Yane, 1975" authorityName="Scillato-Yane" authorityYear="1975" box="[132,531,1077,1104]" class="Mammalia" family="Dasypodidae" genus="Paraeuphractus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="11" pageNumber="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="B947EADA1342D932FF1F095EFEA90F18" box="[132,297,1077,1103]" italics="true" pageId="11" pageNumber="10">Paraeuphractus</emphasis>
|
||
<bibRefCitation id="EFA24B391342D932FEAF095EFD930F07" author="SCILLATO-YANE G. J." box="[308,531,1077,1104]" pageId="11" pageNumber="10" pagination="449 - 461" refId="ref15478" refString="SCILLATO-YANE G. J. 1975. - Nuevo genero de Dasypodidae (Edentata, Xenarthra) del Plioceno de Catamarca (Argentina). Algunas consideraciones filogeneticas y zoogeograficas sobre los Euphractini, in I Congreso Argentino de Paleontologia y Bioestratigrafia, Tucuman 1974. Actas 2: 449 - 461." type="journal article" year="1975">
|
||
Scillato-Yané,
|
||
<quantity id="4CCB9B2D1342D932FE42095EFD930F07" box="[473,531,1077,1104]" metricMagnitude="1" metricUnit="m" metricValue="5.0165" pageId="11" pageNumber="10" unit="in" value="1975.0">1975</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in developing dorsal foramina at the intersections between the main sulcus and the minor sulci of fixed osteoderms (
|
||
<bibRefCitation id="EFA24B391342D932FD8D091FFE9D0FF8" author="RODRIGUEZ BRIZUELA R. & TAUBER A." pageId="11" pageNumber="10" pagination="257 - 272" refId="ref15381" refString="RODRIGUEZ BRIZUELA R. & TAUBER A. 2006. - Estratigrafia y mamiferos fosiles de la Formacion Toro Negro (Neogeno), Departamento Vinchina, noreste de la provincia de La Rioja. Ameghiniana 43: 257 - 272." type="journal article" year="2006">Rodríguez Brizuela & Tauber, 2006</bibRefCitation>
|
||
), and from
|
||
<taxonomicName id="4C334D4B1342D932FE3C09FFFD9C0FF9" authorityName="Reig" authorityYear="1958" box="[423,540,1172,1198]" class="Mammalia" family="Dasypodidae" genus="Chorobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="11" pageNumber="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="B947EADA1342D932FE3C09FFFD9C0FF9" box="[423,540,1172,1198]" italics="true" pageId="11" pageNumber="10">Chorobates</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C334D4B1342D932FDC309FFFC890FF9" authorityName="Scillato-Yane" authorityYear="1980" box="[600,777,1172,1198]" class="Mammalia" family="Dasypodidae" genus="Macrochorobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="11" pageNumber="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="B947EADA1342D932FDC309FFFC890FF9" box="[600,777,1172,1198]" italics="true" pageId="11" pageNumber="10">Macrochorobates</emphasis>
|
||
</taxonomicName>
|
||
in being smaller, having a less elevated and deviated central figure in both fixed and mobile osteoderms, and developing undivided lateral figures in mobile osteoderms (
|
||
<bibRefCitation id="EFA24B391342D932FDE0099FFF3D0E79" author="SCILLATO-YANE G. J." pageId="11" pageNumber="10" refId="ref15627" refString="SCILLATO-YANE G. J. 1982. - Los Dasypodidae (Mammalia: Edentata) del Plioceno y Pleistoceno de Argentina. PhD Thesis, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, 127 p." type="book" year="1982">Scillato-Yané 1982</bibRefCitation>
|
||
). It differs from
|
||
<taxonomicName id="4C334D4B1342D932FEF5087FFD820E79" authorityName="Ameghino" authorityYear="1886" box="[366,514,1300,1326]" class="Mammalia" family="Dasypodidae" genus="Proeuphractus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="11" pageNumber="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="B947EADA1342D932FEF5087FFD820E79" box="[366,514,1300,1326]" italics="true" pageId="11" pageNumber="10">Proeuphractus</emphasis>
|
||
</taxonomicName>
|
||
in having much smaller piliferous foramina at both the posterior and lateral margins (
|
||
<bibRefCitation id="EFA24B391342D932FF15083FFE3D0E38" author="PEREA D. & SCILLATO-YANE G. J." box="[142,445,1364,1391]" pageId="11" pageNumber="10" pagination="125 - 130" refId="ref14895" refString="PEREA D. & SCILLATO-YANE G. J. 1995. - Proephractus limpidus Ameghino, 1886 (Xenarthra, Dasypodidae, Euphractini): Osteologia comparada del craneo y elementos de la coraza asociados (Neogeno del Uruguay). Boletin de la Real Sociedad Espanola de Historia Natural, Seccion geologica 90: 125 - 130." type="journal article" year="1995">Perea & Scillato-Yané 1995</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |