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<document id="34E6B699863885975BDF412B3BBCE687" ID-CLB-Dataset="20081" ID-DOI="10.11646/zootaxa.5120.1.5" ID-GBIF-Dataset="af0cd02f-e97d-4480-873d-c0e7f7bdb299" ID-ISSN="1175-5326" ID-Zenodo-Dep="6388909" ID-ZooBank="6BD3DEE0-A7CF-4B83-A397-E8B54B943387" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648452540938" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Ferreira, Ana B. M. &amp; Génier, François" docDate="2022" docId="039A2A76FFDEFFA809C9E5BDFE2FFA94" docLanguage="en" docName="zootaxa.5120.1.5.pdf" docOrigin="Zootaxa 5120 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltepilissus Pereira 1949" docType="treatment" docVersion="7" lastPageNumber="86" masterDocId="FFA3520EFFDCFFAB095EE13BFF89FFED" masterDocTitle="Taxonomic revision of the genus Deltepilissus Pereira, 1949 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)" masterLastPageNumber="96" masterPageNumber="83" pageNumber="85" updateTime="1699318576615" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AFC0D310BF421393652B85A3F26974FC">Taxonomic revision of the genus Deltepilissus Pereira, 1949 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:roleTerm id="DB97A12F8EDFA11570E281ADD4A7306D">Author</mods:roleTerm>
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<mods:namePart id="06116955C1B21D9292D28EA429ED8EC6">Silva, Fernando A. B.</mods:namePart>
<mods:affiliation id="20DE72589FD827D70A0D6CBCBB253EFB">Universidade Federal Rural de Pernambuco, Unidade Acadêmica de Serra Talhada. Avenida Gregório Ferraz Nogueira, s / n. Serra Talhada, PE, 56909 - 535, Brazil. &amp; Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01, Guamá. Belém, PA, 66075 - 110, Brazil.</mods:affiliation>
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<mods:namePart id="AF315075964A2B9EAE4D959CDDAEB0F8">Ferreira, Ana B. M.</mods:namePart>
<mods:affiliation id="E15AE8C0DC98A7B32C283FBD9C0B90E9">Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01, Guamá. Belém, PA, 66075 - 110, Brazil.</mods:affiliation>
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<mods:namePart id="D0B29247B5ADF5C8B9998585A23EA56F">Génier, François</mods:namePart>
<mods:nameIdentifier id="BF619E54CC3854EF90691A83B564EF24" type="ORCID">0000-0002-5399-8412</mods:nameIdentifier>
<mods:affiliation id="0EEF1FE631BBB3D359D12DF335F3416F">Beaty Centre for Species Discovery, Canadian Museum of Nature, 1740 Chemin Pink, Gatineau, Quebec, J 9 J 3 N 7, Canada. fgenier @ nature. ca; https: // orcid. org / 0000 - 0002 - 5399 - 8412</mods:affiliation>
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<mods:date id="52E17BC9B988EFD44C8D33BC540E99CF">2022</mods:date>
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<taxonomicName id="4C33E0E3FFDEFFA909C9E5BDFE57FB4D" ID-CoL="42SD" authority="Pereira, 1949" authorityName="Pereira" authorityYear="1949" box="[151,478,1158,1185]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
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<emphasis id="B9474772FFDEFFA909C9E5BDFEB9FB4C" bold="true" box="[151,304,1158,1185]" italics="true" pageId="2" pageNumber="85">Deltepilissus</emphasis>
<bibRefCitation id="EFA2E691FFDEFFA90866E5BDFE57FB4D" author="Pereira, F." box="[312,478,1158,1184]" pageId="2" pageNumber="85" pagination="231 - 246" refId="ref9277" refString="Pereira, F. (1949) Escarabeideos americanos II (Coleopt. - Scarabaeidae). Arquivos do Museu Paranaense, 7, 231 - 246." type="journal article" year="1949">Pereira, 1949</bibRefCitation>
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<treatmentCitationGroup id="AB23BC4EFFDEFFA909C9E5F0FEB0FA76" pageId="2" pageNumber="85">
<treatmentCitation id="0A92BD71FFDEFFA909C9E5F0FE44FB0E" author="Pereira, F." box="[151,461,1227,1251]" page="231" pageId="2" pageNumber="85" year="1949">
<taxonomicName id="4C33E0E3FFDEFFA909C9E5F0FE44FB0E" ID-CoL="42SD" authority="Pereira 1949: 231" authorityName="Pereira" authorityPageNumber="231" authorityYear="1949" box="[151,461,1227,1251]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B9474772FFDEFFA909C9E5F0FE93FB0E" box="[151,282,1227,1251]" italics="true" pageId="2" pageNumber="85">Deltepilissus</emphasis>
<bibRefCitation id="EFA2E691FFDEFFA90840E5F0FE44FB0E" author="Pereira, F." box="[286,461,1227,1251]" pageId="2" pageNumber="85" pagination="231 - 246" refId="ref9277" refString="Pereira, F. (1949) Escarabeideos americanos II (Coleopt. - Scarabaeidae). Arquivos do Museu Paranaense, 7, 231 - 246." type="journal article" year="1949">Pereira 1949: 231</bibRefCitation>
</taxonomicName>
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(original description);
<treatmentCitation id="0A92BD71FFDEFFA90BEFE5F0FC44FB0E" author="Pereira, F. &amp; Martinez, A." box="[689,973,1227,1251]" page="96" pageId="2" pageNumber="85" year="1956">
<bibRefCitation id="EFA2E691FFDEFFA90BEFE5F0FC44FB0E" author="Pereira, F. &amp; Martinez, A." box="[689,973,1227,1251]" pageId="2" pageNumber="85" pagination="91 - 192" refId="ref9328" refString="Pereira, F. &amp; Martinez, A. (1956) Os generos de Canthonini americanos. Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez 1956: 96</bibRefCitation>
</treatmentCitation>
, 125, 184 (identification key, catalog);
<treatmentCitation id="0A92BD71FFDEFFA90C12E5F0FEB9FAEF" author="Halffter, G." page="231" pageId="2" pageNumber="85" year="1961">
<bibRefCitation id="EFA2E691FFDEFFA90C12E5F0FEB9FAEF" author="Halffter, G." pageId="2" pageNumber="85" pagination="225 - 320" refId="ref8907" refString="Halffter, G. (1961) Monografia de las especies norteamericanas del genero Canthon Hoffsg. (Coleopt., Scarab.). Ciencia, 20, 225 - 320." type="journal article" year="1961">Halffter 1961: 231</bibRefCitation>
</treatmentCitation>
(identification key);
<treatmentCitation id="0A92BD71FFDEFFA90B4EE5D1FCCCFAEF" author="Vulcano, M. A. &amp; Pereira, F." box="[528,837,1258,1282]" page="660" pageId="2" pageNumber="85" year="1964">
<bibRefCitation id="EFA2E691FFDEFFA90B4EE5D1FCCCFAEF" author="Vulcano, M. A. &amp; Pereira, F." box="[528,837,1258,1282]" pageId="2" pageNumber="85" pagination="570 - 685" refId="ref10315" refString="Vulcano, M. A. &amp; Pereira, F. (1964) Catalogue of the Canthonini (Col. Scarab.) inhabiting the Western Hemisphere. Entomologische Arbeiten aus dem Museum G. Frey, 15, 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 660</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="0A92BD71FFDEFFA90AE7E5D1FA8FFAEF" author="Halffter, G. &amp; Matthews, E. G." box="[953,1286,1258,1282]" page="261" pageId="2" pageNumber="85" year="1966">
<bibRefCitation id="EFA2E691FFDEFFA90AE7E5D1FA8FFAEF" author="Halffter, G. &amp; Matthews, E. G." box="[953,1286,1258,1282]" pageId="2" pageNumber="85" pagination="1 - 312" refId="ref9020" refString="Halffter, G. &amp; Matthews, E. G. (1966). The natural history of dung beetles of the subfamily Scarabaeinae. Folia Entomologica Mexicana, 12 - 14, 1 - 312." type="journal article" year="1966">Halffter &amp; Matthews 1966: 261</bibRefCitation>
</treatmentCitation>
(distribution);
<treatmentCitation id="0A92BD71FFDEFFA90999E433FE73FACC" author="Halffter, G. &amp; Martinez, A." box="[199,506,1288,1313]" page="37" pageId="2" pageNumber="85" year="1977">
<bibRefCitation id="EFA2E691FFDEFFA90999E433FE73FACC" author="Halffter, G. &amp; Martinez, A." box="[199,506,1288,1313]" pageId="2" pageNumber="85" pagination="29 - 107" refId="ref8982" refString="Halffter, G. &amp; Martinez, A. (1977) Revision monografica de los Canthonina Americanos, IV parte. Clave para generos y subgeneros. Folia Entomologica Mexicana, 38, 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 37</bibRefCitation>
</treatmentCitation>
, 52, 66 (identification key, taxonomic remarks, checklist); Halffter &amp; Edmonds 1982: 139 (distribution);
<treatmentCitation id="0A92BD71FFDEFFA90803E41CFDD5FAD2" author="Vaz-de-Mello, F." box="[349,604,1319,1343]" page="186" pageId="2" pageNumber="85" year="2000">
<bibRefCitation id="EFA2E691FFDEFFA90803E41CFDD5FAD2" author="Vaz-de-Mello, F." box="[349,604,1319,1343]" pageId="2" pageNumber="85" pagination="183 - 195" refId="ref9891" refString="Vaz-de-Mello, F. (2000) Estado atual de conhecimento dos Scarabaeinae s. str. (Coleoptera: Scarabeoidea) do Brasil. In: Martin- Piera, F., Morrone, J. &amp; Melic, A. (Eds.), Hacia un Proyecto CYTED para el inventario Y Estimacion de la Diversidad Entomologica en Iberoamerica: PrIBES- 2000. Monografias Tercer Milenio &amp; Sociedad Entomologica Aragonesa, Zaragoza, Spain, pp. 183 - 195." type="book chapter" year="2000">Vaz-de-Mello 2000: 186</bibRefCitation>
</treatmentCitation>
, 192 (checklist);
<treatmentCitation id="0A92BD71FFDEFFA90A4BE41CFC4FFAD2" author="Krajcik, M." box="[789,966,1319,1343]" page="47" pageId="2" pageNumber="85" year="2006">
<bibRefCitation id="EFA2E691FFDEFFA90A4BE41CFC4FFAD2" author="Krajcik, M." box="[789,966,1319,1343]" pageId="2" pageNumber="85" pagination="1 - 189" refId="ref9158" refString="Krajcik, M. (2006) Checklist of Scarabeoidea of the World. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma. X, Supplement 3, 1 - 189." type="journal article" year="2006">Krajcik 2006: 47</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="0A92BD71FFDEFFA90D14E41CFAE7FAD2" author="Vaz-de-Mello, F. &amp; Edmonds, W. &amp; Ocampo, F. &amp; Schoolmeesters, P." box="[1098,1390,1319,1343]" page="5" pageId="2" pageNumber="85" year="2011">
<bibRefCitation id="EFA2E691FFDEFFA90D14E41CFAE7FAD2" author="Vaz-de-Mello, F. &amp; Edmonds, W. &amp; Ocampo, F. &amp; Schoolmeesters, P." box="[1098,1390,1319,1343]" pageId="2" pageNumber="85" pagination="1 - 73" refId="ref10018" refString="Vaz-de-Mello, F., Edmonds, W., Ocampo, F. &amp; Schoolmeesters, P. (2011) A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa, 2854 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 2854.1.1" type="journal article" year="2011">
Vaz-de-Mello
<emphasis id="B9474772FFDEFFA90D81E413FA86FAD2" box="[1247,1295,1319,1343]" italics="true" pageId="2" pageNumber="85">et al</emphasis>
. 2011: 5
</bibRefCitation>
</treatmentCitation>
, 10, 18, 25, 32, 40, 55 (identification key);
<treatmentCitation id="0A92BD71FFDEFFA90B07E47DFC8EFAB3" author="Krajcik, M." box="[601,775,1350,1374]" page="88" pageId="2" pageNumber="85" year="2012">
<bibRefCitation id="EFA2E691FFDEFFA90B07E47DFC8EFAB3" author="Krajcik, M." box="[601,775,1350,1374]" pageId="2" pageNumber="85" pagination="1 - 278" refId="ref9194" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. X. Supplement 5, 1 - 278." type="journal article" year="2012">Krajcik 2012: 88</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="0A92BD71FFDEFFA90AD6E47DFB33FAB3" author="Tarasov, S. &amp; Dimitrov, D." box="[904,1210,1350,1374]" page="15" pageId="2" pageNumber="85" year="2016">
<bibRefCitation id="EFA2E691FFDEFFA90AD6E47DFB33FAB3" author="Tarasov, S. &amp; Dimitrov, D." box="[904,1210,1350,1374]" pageId="2" pageNumber="85" pagination="1 - 19" refId="ref9717" refString="Tarasov, S. &amp; Dimitrov, D. (2016) Multigene phylogenetic analysis redefines dung beetles relationships and classification (Coleoptera: Scarabaeidae: Scarabaeinae). BMC Evolionary Biology, 16 (257), 1 - 19. https: // doi. org / 10.1186 / s 12862 - 016 - 0822 - x" type="journal article" year="2016">Tarasov &amp; Dimitrov 2016: 15</bibRefCitation>
</treatmentCitation>
(new delimitation for
<taxonomicName id="4C33E0E3FFDEFFA90999E45FFECCFA91" ID-CoL="KS3" authorityName="Lacordaire" authorityYear="1856" box="[199,325,1380,1404]" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="86" rank="tribe" tribe="Deltochilini">Deltochilini</taxonomicName>
);
<treatmentCitation id="0A92BD71FFDEFFA90808E45FFD27FA90" author="Cupello, M. &amp; Vaz-de-Mello, F." box="[342,686,1380,1405]" page="18" pageId="2" pageNumber="85" year="2018">
<bibRefCitation id="EFA2E691FFDEFFA90808E45FFD27FA90" author="Cupello, M. &amp; Vaz-de-Mello, F." box="[342,686,1380,1405]" pageId="2" pageNumber="85" pagination="1 - 205" refId="ref8610" refString="Cupello, M. &amp; Vaz-de-Mello, F. (2018) A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter &amp; Martinez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of ' Canthon sensu lato'. European Journal of Taxonomy, 467, 1 - 205. https: // doi. org / 10.5852 / ejt. 2018.467" type="journal article" year="2018">Cupello &amp; Vaz-de-Mello 2018: 18</bibRefCitation>
</treatmentCitation>
(taxonomic remarks);
<bibRefCitation id="EFA2E691FFDEFFA90ACBE45FFBE5FA90" author="Schoolmeesters, P." box="[917,1132,1380,1405]" pageId="2" pageNumber="85" pagination="2405 - 8858" refId="ref9492" refString="Schoolmeesters, P. (2020) Scarabs: world Scarabaeidae database (version 2019 - 11 - 02). In: Roskov Y. et al., (Eds.), Species 2000 &amp; ITIS Catalogue of Life, 2020 - 02 - 24. ISSN 2405 - 8858. Species 2000: Naturalis, Leiden, the Netherlands. Available from: http: // www. catalogueoflife. org / col (accessed 12 January 2022)" type="book chapter" year="2020">Schoolmeesters 2020</bibRefCitation>
(catalog);
<bibRefCitation id="EFA2E691FFDEFFA90D86E45FFA14FA90" author="Vaz-de-Mello, F." box="[1240,1437,1380,1405]" pageId="2" pageNumber="85" refId="ref9975" refString="Vaz-de-Mello, F. (2021) Scarabaeinae. In: Catalogo Taxonomico da Fauna do Brasil. Available from: http: // fauna. jbrj. gov. br / fauna / faunadobrasil / 127498 (accessed 5 March 2021)" type="url" year="2021">Vaz-de-Mello 2021</bibRefCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="C329C8EBFFDEFFA909C9E4F2FDBDF9EA" pageId="2" pageNumber="85" type="type_taxon">
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<emphasis id="B9474772FFDEFFA909C9E4F2FEBBFA0E" bold="true" box="[151,306,1481,1507]" pageId="2" pageNumber="85">
<typeStatus id="548825C2FFDEFFA909C9E4F2FF5BFA0E" box="[151,210,1481,1507]" pageId="2" pageNumber="85">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C33E0E3FFDEFFA90867E4F2FD54FA0E" ID-CoL="34K64" authority="Pereira, 1949" authorityName="Pereira" authorityYear="1949" box="[313,733,1481,1507]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="B9474772FFDEFFA90867E4F2FDCBFA0E" box="[313,578,1481,1507]" italics="true" pageId="2" pageNumber="85">Deltepilissus travassosi</emphasis>
<bibRefCitation id="EFA2E691FFDEFFA90B16E4F2FD54FA0E" author="Pereira, F." box="[584,733,1481,1507]" pageId="2" pageNumber="85" pagination="231 - 246" refId="ref9277" refString="Pereira, F. (1949) Escarabeideos americanos II (Coleopt. - Scarabaeidae). Arquivos do Museu Paranaense, 7, 231 - 246." type="journal article" year="1949">Pereira, 1949</bibRefCitation>
</taxonomicName>
by original designation (currently a junior subjective synonym of
<taxonomicName id="4C33E0E3FFDEFFA909E8E4D6FDAEF9EA" ID-CoL="QPCX" authority="Harold, 1880" authorityName="Harold" authorityYear="1880" box="[182,551,1517,1543]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDEFFA909E8E4D6FE01F9EA" box="[182,392,1517,1543]" italics="true" pageId="2" pageNumber="85">Canthon infernalis</emphasis>
<bibRefCitation id="EFA2E691FFDEFFA908D1E4D6FDAEF9EA" author="Harold, E." box="[399,551,1517,1543]" pageId="2" pageNumber="85" pagination="148 - 171" refId="ref9058" refString="Harold, E. (1880) Einige Neue Coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 148 - 171." type="journal article" year="1880">Harold, 1880</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFDEFFA90999E72AFD0BF97A" pageId="2" pageNumber="85" type="diagnosis">
<paragraph id="8B8C9B60FFDEFFA90999E72AFD0BF97A" blockId="2.[151,1437,1481,2012]" pageId="2" pageNumber="85">
<emphasis id="B9474772FFDEFFA90999E72AFECBF9C6" bold="true" box="[199,322,1553,1579]" pageId="2" pageNumber="85">Diagnosis.</emphasis>
Among the genera included in the tribe
<taxonomicName id="4C33E0E3FFDEFFA90BA0E72AFC0DF9C6" authorityName="Lacordaire" authorityYear="1856" box="[766,900,1553,1579]" pageId="2" pageNumber="85" rank="tribe" tribe="Deltochilini">Deltochilini</taxonomicName>
(sensu
<bibRefCitation id="EFA2E691FFDEFFA90A89E729FB70F9C1" author="Tarasov, S. &amp; Dimitrov, D." box="[983,1273,1553,1580]" pageId="2" pageNumber="85" pagination="1 - 19" refId="ref9717" refString="Tarasov, S. &amp; Dimitrov, D. (2016) Multigene phylogenetic analysis redefines dung beetles relationships and classification (Coleoptera: Scarabaeidae: Scarabaeinae). BMC Evolionary Biology, 16 (257), 1 - 19. https: // doi. org / 10.1186 / s 12862 - 016 - 0822 - x" type="journal article" year="2016">Tarasov &amp; Dimitrov 2016</bibRefCitation>
),
<taxonomicName id="4C33E0E3FFDEFFA90C52E72AFA14F9C6" authorityName="Pereira" authorityYear="1949" box="[1292,1437,1553,1579]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B9474772FFDEFFA90C52E72AFA14F9C6" box="[1292,1437,1553,1579]" italics="true" pageId="2" pageNumber="85">Deltepilissus</emphasis>
</taxonomicName>
can be distinguished by reduced tarsal claws (
<figureCitation id="130887E5FFDEFFA90BCAE70EFC81F9A2" box="[660,776,1589,1615]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="2" pageNumber="85">Figs 1G–I</figureCitation>
,
<figureCitation id="130887E5FFDEFFA90A4AE70EFCC7F9A2" box="[788,846,1589,1615]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="2" pageNumber="85">2I–L</figureCitation>
); protibiae long, weakly widened apically, with three small and spaced, lateral teeth (
<figureCitation id="130887E5FFDEFFA908A9E762FDFDF99E" box="[503,628,1625,1651]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="2" pageNumber="85">Figs 1A–B</figureCitation>
,
<figureCitation id="130887E5FFDEFFA90BDEE762FD40F99E" box="[640,713,1625,1651]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="2" pageNumber="85">2A–D</figureCitation>
); metatibial spur short, widened apically, with spiniform process at least on internal edge (
<figureCitation id="130887E5FFDEFFA908EEE746FDADF97A" box="[432,548,1661,1687]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="2" pageNumber="85">Figs 1H–I</figureCitation>
,
<figureCitation id="130887E5FFDEFFA90B6EE746FDFFF97A" box="[560,630,1661,1687]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="2" pageNumber="85">2K–L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFDEFFA80999E79AFBAAFC00" lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85" type="description">
<paragraph id="8B8C9B60FFDEFFA80999E79AFBAAFC00" blockId="2.[151,1437,1481,2012]" lastBlockId="3.[151,1437,151,1401]" lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85">
<emphasis id="B9474772FFDEFFA90999E79AFEE4F956" bold="true" box="[199,365,1697,1723]" pageId="2" pageNumber="85">Redescription</emphasis>
.
<emphasis id="B9474772FFDEFFA908DEE79AFC89F956" bold="true" box="[384,768,1697,1723]" pageId="2" pageNumber="85">Colour and tegument sculpture</emphasis>
(
<figureCitation id="130887E5FFDEFFA90A4DE79AFC1CF956" box="[787,917,1697,1723]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="2" pageNumber="85">Figs 1A–B</figureCitation>
,
<figureCitation id="130887E5FFDEFFA90AFBE79AFC67F956" box="[933,1006,1697,1723]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="2" pageNumber="85">2A–D</figureCitation>
). Head, pronotum, elytra and ventral surface with black, copper or red coloration. Body densely punctate.
<emphasis id="B9474772FFDEFFA90A9DE7FEFB90F932" bold="true" box="[963,1049,1733,1759]" pageId="2" pageNumber="85">Length</emphasis>
.
<quantity id="4CCB3685FFDEFFA90D74E7FEFB15F90D" box="[1066,1180,1733,1760]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="2" pageNumber="85" unit="mm" value="12.0" valueMax="15.0" valueMin="9.0">9–15 mm</quantity>
.
<emphasis id="B9474772FFDEFFA90DF5E7FEFB62F932" bold="true" box="[1195,1259,1733,1759]" pageId="2" pageNumber="85">Head</emphasis>
(
<figureCitation id="130887E5FFDEFFA90DA3E7FEFAD5F932" box="[1277,1372,1733,1759]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="2" pageNumber="85">Figs 1C</figureCitation>
,
<figureCitation id="130887E5FFDEFFA90C33E7FEFA19F932" box="[1389,1424,1733,1759]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="2" pageNumber="85">2E</figureCitation>
). Surface with strong, deep and dense punctures. Clypeus with two widely spaced, small teeth. Eye comma shaped in dorsal view. Dorsal interocular space at least 10 times eye width. Lateral margin of head regularly curved outward.
<emphasis id="B9474772FFDEFFA909C9E60AFF79F8A6" bold="true" box="[151,240,1841,1867]" pageId="2" pageNumber="85">Thorax</emphasis>
(
<figureCitation id="130887E5FFDEFFA9085FE60AFEE9F8A6" box="[257,352,1841,1867]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="2" pageNumber="85">Figs 1A</figureCitation>
,
<figureCitation id="130887E5FFDEFFA90833E60AFE4DF8A6" box="[365,452,1841,1867]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="2" pageNumber="85">2A, 2C</figureCitation>
). Pronotum convex; anterior angles acute, directed forward. Lateral edge of pronotum regularly curved outward; posterior angle obtuse. Pronotum with a longitudinal sulcus on midline posteriorly. Hypomera not excavated (
<figureCitation id="130887E5FFDEFFA9089AE642FDABF87E" box="[452,546,1913,1939]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="2" pageNumber="85">Figs 1D</figureCitation>
,
<figureCitation id="130887E5FFDEFFA90B6EE642FDDBF87E" box="[560,594,1913,1939]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="2" pageNumber="85">2F</figureCitation>
). Mesometasternal suture slightly arched. Metaventrite punctures variable in size; smaller and denser medially (
<figureCitation id="130887E5FFDEFFA90B61E6A6FD12F85A" box="[575,667,1949,1975]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="2" pageNumber="85">Figs 1B</figureCitation>
,
<figureCitation id="130887E5FFDEFFA90BF7E6A6FD76F85A" box="[681,767,1949,1975]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="2" pageNumber="85">2B, 2D</figureCitation>
).
<emphasis id="B9474772FFDEFFA90A4DE6A6FCD6F85A" bold="true" box="[787,863,1949,1975]" pageId="2" pageNumber="85">Elytra</emphasis>
(
<figureCitation id="130887E5FFDEFFA90A2EE6A6FC47F85A" box="[880,974,1949,1975]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="2" pageNumber="85">Figs 1A</figureCitation>
,
<figureCitation id="130887E5FFDEFFA90A85E6A6FBB8F85A" box="[987,1073,1949,1975]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="2" pageNumber="85">2A, 2C</figureCitation>
). Lateral margin slightly curved outward. Elytral striae visibly impressed. Pseudepipleural carina complete, visible on entire length of epipleuron. Elytral lateral edge formed by the pseudepipleural carina. Interstriae flattened, without carinae or tubercles basally and apically.
<emphasis id="B9474772FFDFFFA8086DE181FE21FF39" bold="true" box="[307,424,186,212]" pageId="3" pageNumber="86">Abdomen</emphasis>
(
<figureCitation id="130887E5FFDFFFA808E4E180FD91FF38" box="[442,536,187,213]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1E</figureCitation>
,
<figureCitation id="130887E5FFDFFFA80B76E180FD3CFF38" box="[552,693,187,213]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2B, 2D, 2G</figureCitation>
). Punctures denser on lateral surface of ventrites. Sixth ventrite longer than 2nd, 3rd, 4th and 5th. Pygidium rounded apically, densely punctate, separated from propygidium by carina (
<figureCitation id="130887E5FFDFFFA809B3E038FECCFEF0" box="[237,325,259,285]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1F</figureCitation>
,
<figureCitation id="130887E5FFDFFFA8080DE038FEF0FEF0" box="[339,377,259,285]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2H</figureCitation>
).
<emphasis id="B9474772FFDFFFA808D1E038FE4EFEF0" bold="true" box="[399,455,259,285]" pageId="3" pageNumber="86">Legs</emphasis>
. Femora densely punctate (
<figureCitation id="130887E5FFDFFFA80A5BE038FCEBFEF0" box="[773,866,259,285]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1B</figureCitation>
,
<figureCitation id="130887E5FFDFFFA80A2FE038FC41FEF0" box="[881,968,259,285]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2B, 2D</figureCitation>
); anterior and posterior edges marginate. Protibiae long, thin and curved. Apical one-quarter of protibiae abruptly expanded along inner edge (
<figureCitation id="130887E5FFDFFFA80C46E01CFA1EFEAC" box="[1304,1431,295,321]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1A–B</figureCitation>
,
<figureCitation id="130887E5FFDFFFA809C9E070FF69FE88" box="[151,224,331,357]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2A–D</figureCitation>
). Apical one-third of protibiae with three distinct lateral teeth; medial lateral tooth closer to apical tooth than to basal tooth (
<figureCitation id="130887E5FFDFFFA80818E054FE4FFE64" box="[326,454,367,393]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1A–B</figureCitation>
,
<figureCitation id="130887E5FFDFFFA8088BE054FD97FE64" box="[469,542,367,393]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2A–D</figureCitation>
). Meso- and metatibiae smoothly arched toward body, not abruptly expanded along inner edge (
<figureCitation id="130887E5FFDFFFA8083DE0A8FE6BFE40" box="[355,482,403,429]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1A–B</figureCitation>
,
<figureCitation id="130887E5FFDFFFA808B1E0A8FDB1FE40" box="[495,568,403,429]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2A–D</figureCitation>
). Longitudinal carinae of meso- and metatibiae with row of setae. Metatibial spur short, sinuous and spatulate; bifurcate apically (
<figureCitation id="130887E5FFDFFFA80BB8E08CFCB1FE3C" box="[742,824,439,465]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1I</figureCitation>
,
<figureCitation id="130887E5FFDFFFA80A1AE08CFCEFFE3C" box="[836,870,439,465]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2L</figureCitation>
), almost rounded (
<figureCitation id="130887E5FFDFFFA80D66E08CFB19FE3C" box="[1080,1168,439,465]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Fig. 1H</figureCitation>
), or with a conspicuous denticle (spiniform process) on inner edge (
<figureCitation id="130887E5FFDFFFA80BD2E0E0FD60FE18" box="[652,745,475,501]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">Fig. 2K</figureCitation>
). First meso- and metatarsomeres triangular; tarsomeres 2–4 trapezoidal, the last subrectangular (
<figureCitation id="130887E5FFDFFFA80B72E0C4FD28FDF4" box="[556,673,511,537]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1H–I</figureCitation>
,
<figureCitation id="130887E5FFDFFFA80BF3E0C4FD7BFDF4" box="[685,754,511,537]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2K–L</figureCitation>
). Tarsal claws reduced (
<figureCitation id="130887E5FFDFFFA80AA0E0C4FBFAFDF4" box="[1022,1139,511,537]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1G–I</figureCitation>
,
<figureCitation id="130887E5FFDFFFA80D21E0C4FB33FDF4" box="[1151,1210,511,537]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2I–L</figureCitation>
).
<emphasis id="B9474772FFDFFFA80D93E0C5FE9CFDD0" bold="true" pageId="3" pageNumber="86">Secondary sexual characters</emphasis>
. Females can be distinguished from males, in general, by the last abdominal ventrite evenly wide (
<figureCitation id="130887E5FFDFFFA80C31E318FF34FD8C" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">Fig. 2D</figureCitation>
); in males, the last abdominal ventrite is subtly narrower medially (
<figureCitation id="130887E5FFDFFFA80A96E37CFBAAFD8C" box="[968,1059,583,609]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1E</figureCitation>
,
<figureCitation id="130887E5FFDFFFA80D6CE37CFBD1FD8C" box="[1074,1112,583,609]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2G</figureCitation>
). Females have the protibial spur bifurcate apically (
<figureCitation id="130887E5FFDFFFA808FAE350FE7EFD68" box="[420,503,619,645]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">Fig. 2J</figureCitation>
); in males, protibial spur is wide and rounded, lacking denticle apically (
<figureCitation id="130887E5FFDFFFA80C6AE350FA04FD68" box="[1332,1421,619,645]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Fig. 1G</figureCitation>
), or with a denticle on external edge (
<figureCitation id="130887E5FFDFFFA80B75E3B4FDF2FD44" box="[555,635,655,681]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">Fig. 2I</figureCitation>
). Females have also sinuous and bifurcate apex of metatibial spur, with denticle (spiniform process) on internal and external edges (
<figureCitation id="130887E5FFDFFFA80A72E388FCF5FD20" box="[812,892,691,717]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1I</figureCitation>
,
<figureCitation id="130887E5FFDFFFA80AD9E388FC23FD20" box="[903,938,691,717]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2L</figureCitation>
); in males, metatibial spur only have denticle on inner edge (
<figureCitation id="130887E5FFDFFFA80861E3ECFE10FD1C" box="[319,409,727,753]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">Fig. 2K</figureCitation>
), or it is rounded apically, with inner denticle almost inconspicuous (
<figureCitation id="130887E5FFDFFFA80DC5E3ECFB7DFD1C" box="[1179,1268,727,753]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Fig. 1H</figureCitation>
). Furthermore, females have the apical third of metatibiae almost straight (
<figureCitation id="130887E5FFDFFFA80A0AE3C0FC23FCF8" box="[852,938,763,789]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1I</figureCitation>
,
<figureCitation id="130887E5FFDFFFA80AE4E3C0FC55FCF8" box="[954,988,763,789]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2L</figureCitation>
), whereas in males the apical third of metatibiae is distinctly curved inward (
<figureCitation id="130887E5FFDFFFA80B19E224FD2BFCD4" box="[583,674,799,825]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="3" pageNumber="86">Figs 1H</figureCitation>
,
<figureCitation id="130887E5FFDFFFA80BF0E224FD5DFCD4" box="[686,724,799,825]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. A–B, Body, male, dorsal and ventral views, respectively; C–D, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; I–J, detail of protibial spur, male and female, respectively; K–L, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="3" pageNumber="86">2K</figureCitation>
).
<emphasis id="B9474772FFDFFFA80BB9E224FCCCFCD4" bold="true" box="[743,837,799,825]" pageId="3" pageNumber="86">Tegmen</emphasis>
. Parameres symmetrical, subtriangular in lateral view (
<figureCitation id="130887E5FFDFFFA809C1E278FEAEFCB0" box="[159,295,835,861]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="3" pageNumber="86">Figs 3C, 3F</figureCitation>
), pointed and curved inward apically (
<figureCitation id="130887E5FFDFFFA80B88E278FCDAFCB0" box="[726,851,835,861]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="3" pageNumber="86">Figs 3A–F</figureCitation>
).
<emphasis id="B9474772FFDFFFA80A38E278FC74FCB0" bold="true" box="[870,1021,835,861]" pageId="3" pageNumber="86">Endophallus</emphasis>
. Lamella copulatrix absent. Superior right peripheral endophallite circular, with curved handle-shaped extension and ring with thin border (
<figureCitation id="130887E5FFDFFFA80C51E25CFA19FC6C" box="[1295,1424,871,897]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="3" pageNumber="86">Figs 3G–H</figureCitation>
). Frontolateral peripheral endophallite with irregular shape (
<figureCitation id="130887E5FFDFFFA80A76E2B0FC1CFC48" box="[808,917,907,933]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="3" pageNumber="86">Figs 3I–J</figureCitation>
). Complex of axial and subaxial endophallites elongate, with irregular shape (
<figureCitation id="130887E5FFDFFFA808AFE294FDFEFC24" box="[497,631,943,969]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="3" pageNumber="86">Figs 3M–N</figureCitation>
). There are also two short additional endophallites (
<figureCitation id="130887E5FFDFFFA80DE8E294FAB9FC24" box="[1206,1328,943,969]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="3" pageNumber="86">Figs 3K–L</figureCitation>
) between the frontolateral peripheral endophallite and the axial and subaxial endophallites.
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFDFFFA80999E2CCFE2FFA94" pageId="3" pageNumber="86" type="discussion">
<paragraph id="8B8C9B60FFDFFFA80999E2CCFE2FFA94" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="86">
<emphasis id="B9474772FFDFFFA80999E2CCFEBDFBFC" bold="true" box="[199,308,1015,1041]" pageId="3" pageNumber="86">Remarks</emphasis>
. Although none of the specimens with collecting data provide indications on the natural history of the genus, we suspect that species of
<taxonomicName id="4C33E0E3FFDFFFA80B16E520FD50FBD8" authorityName="Pereira" authorityYear="1949" box="[584,729,1051,1077]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B9474772FFDFFFA80B16E520FD50FBD8" box="[584,729,1051,1077]" italics="true" pageId="3" pageNumber="86">Deltepilissus</emphasis>
</taxonomicName>
are inquilines. We base this assumption on the fact that none specimen was collected in dung traps or any other
<typeStatus id="548825C2FFDFFFA80B96E504FD71FBB4" box="[712,760,1087,1113]" pageId="3" pageNumber="86">type</typeStatus>
of baited pitfall traps. The reduced tarsal claws is a common adaptation for inquiline, however no other obvious adaptations are present such as concealed mouthparts, highly modified legs or trichome. The rather dense, small swallow punctures with a minute seta present in
<taxonomicName id="4C33E0E3FFDFFFA80C52E5BCFA14FB4C" authorityName="Pereira" authorityYear="1949" box="[1292,1437,1159,1185]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B9474772FFDFFFA80C52E5BCFA14FB4C" box="[1292,1437,1159,1185]" italics="true" pageId="3" pageNumber="86">Deltepilissus</emphasis>
</taxonomicName>
are eerly similar to those found in
<taxonomicName id="4C33E0E3FFDFFFA80B42E590FBD6FB28" authority="(Genier, 2010)" baseAuthorityName="Genier" baseAuthorityYear="2010" box="[540,1119,1195,1221]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="punctatissimus" subGenus="Lobidion">
<emphasis id="B9474772FFDFFFA80B42E590FD0AFB28" box="[540,643,1195,1221]" italics="true" pageId="3" pageNumber="86">Ateuchus</emphasis>
(
<emphasis id="B9474772FFDFFFA80BCAE590FD70FB28" box="[660,761,1195,1221]" italics="true" pageId="3" pageNumber="86">Lobidion</emphasis>
)
<emphasis id="B9474772FFDFFFA80A54E590FC39FB29" box="[778,944,1195,1220]" italics="true" pageId="3" pageNumber="86">punctatissimus</emphasis>
(
<bibRefCitation id="EFA2E691FFDFFFA80AE1E590FBDEFB28" author="Genier, F." box="[959,1111,1195,1221]" pageId="3" pageNumber="86" pagination="35 - 48" refId="ref8767" refString="Genier, F. (2010) A review of the Neotropical dung beetle genera Deltorhinum Harold, 1869, and Lobidion gen. nov. (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 2693 (1), 35 - 48. https: // doi. org / 10.11646 / zootaxa. 2693.1.3" type="journal article" year="2010">Génier, 2010</bibRefCitation>
)
</taxonomicName>
, a documented ant inquiline species (
<bibRefCitation id="EFA2E691FFDFFFA809A6E5F4FE18FB04" author="Genier, F." box="[248,401,1231,1257]" pageId="3" pageNumber="86" pagination="35 - 48" refId="ref8767" refString="Genier, F. (2010) A review of the Neotropical dung beetle genera Deltorhinum Harold, 1869, and Lobidion gen. nov. (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 2693 (1), 35 - 48. https: // doi. org / 10.11646 / zootaxa. 2693.1.3" type="journal article" year="2010">Génier, 2010</bibRefCitation>
). Another possibility, considering its fully functional wings and rather elongate legs, would be a predatory behavior on social insects. Meliponine bees are common in the Atlantic rainforest and nests in trees. Flying higher in the canopy would explain the fact that species of
<taxonomicName id="4C33E0E3FFDFFFA80AD4E42CFB92FADC" authorityName="Pereira" authorityYear="1949" box="[906,1051,1303,1329]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B9474772FFDFFFA80AD4E42CFB92FADC" box="[906,1051,1303,1329]" italics="true" pageId="3" pageNumber="86">Deltepilissus</emphasis>
</taxonomicName>
have also never been collected in flight interception traps set on the ground. This is purely speculative at the moment, but it would be interesting to verify these hypotheses.
</paragraph>
</subSubSection>
</treatment>
</document>
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