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<document id="1D58BEFAD5E6F81C69883E829E575C7C" ID-DOI="10.11646/zootaxa.3852.4.2" ID-GBIF-Dataset="e2b6c068-3fba-4911-b0f0-5211b9480417" ID-ISSN="1175-5326" ID-Zenodo-Dep="228687" ID-ZooBank="FABC5498-D812-450C-B8CB-149487DDB803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460884980021" checkinUser="plazi" docAuthor="Han, Qingxi &amp; Li, Xinzheng" docDate="2014" docId="0399D6297F63562EFCEDFDF0FE71F83A" docLanguage="en" docName="zt03852p444.pdf" docOrigin="Zootaxa 3852 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glyphocrangon denticulata Han &amp; Li, 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="443" masterDocId="FFA0AE517F60562BFC7AFFD4FFA6FFD3" masterDocTitle="Two new species of Glyphocrangon (Decapoda: Caridea: Glyphocrangonidae) from the East China Sea and the Philippines" masterLastPageNumber="444" masterPageNumber="438" pageNumber="441" updateTime="1698358869074" updateUser="plazi">
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<mods:title id="E91B9925572A7B32503428EB6B2D631F">Two new species of Glyphocrangon (Decapoda: Caridea: Glyphocrangonidae) from the East China Sea and the Philippines</mods:title>
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<mods:name id="81789B565D38C75F4A9B6036C33C58E9" type="personal">
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<mods:namePart id="B877F23985FF5BFAD8E5650C3B52ED73">Han, Qingxi</mods:namePart>
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<mods:namePart id="B84F5961D373C2E8B206AD1EDDFFC30A">Li, Xinzheng</mods:namePart>
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<mods:title id="D23C16A0DA380D06DA623B46020C9C89">Zootaxa</mods:title>
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<mods:date id="0B25CDA674DC0CE111E781163AC45429">2014</mods:date>
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<mods:number id="183237E6C9558F8BF3F15A3176F5B5B6">3852</mods:number>
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<treatment id="0399D6297F63562EFCEDFDF0FE71F83A" ID-DOI="http://doi.org/10.5281/zenodo.6139387" ID-GBIF-Taxon="119586246" ID-Zenodo-Dep="6139387" LSID="urn:lsid:plazi:treatment:0399D6297F63562EFCEDFDF0FE71F83A" httpUri="http://treatment.plazi.org/id/0399D6297F63562EFCEDFDF0FE71F83A" lastPageId="5" lastPageNumber="443" pageId="3" pageNumber="441">
<subSubSection id="C32A34B47F635628FCEDFDF0FCE6FD10" pageId="3" pageNumber="441" type="nomenclature">
<paragraph id="8B8F673F7F635628FCEDFDF0FDE1FDED" blockId="3.[151,583,548,607]" box="[151,583,548,574]" pageId="3" pageNumber="441">
<heading id="D0C7D0537F635628FCEDFDF0FDE1FDED" bold="true" box="[151,583,548,574]" fontSize="11" level="1" pageId="3" pageNumber="441" reason="1">
<emphasis id="B944BB2D7F635628FCEDFDF0FDE1FDED" bold="true" box="[151,583,548,574]" pageId="3" pageNumber="441">
<taxonomicName id="4C301CBC7F635628FCEDFDF0FE44FDED" ID-CoL="9CR6H" box="[151,482,548,574]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="441" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
<emphasis id="B944BB2D7F635628FCEDFDF0FE44FDED" bold="true" box="[151,482,548,574]" italics="true" pageId="3" pageNumber="441">Glyphocrangon denticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27706567F635628FD93FDF0FDE1FDED" box="[489,583,548,574]" pageId="3" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F673F7F635628FCEDFD92FEB7FD8C" blockId="3.[151,583,548,607]" box="[151,273,582,607]" pageId="3" pageNumber="441">
(
<figureCitation id="130B7BBA7F635628FCE5FD92FEAFFD8C" box="[159,265,582,607]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="3.[151,250,1641,1663]" captionStartId-1="4.[151,250,1900,1922]" captionStartId-2="5.[151,250,895,917]" captionTargetBox-0="[158,1423,871,1615]" captionTargetBox-1="[165,1426,671,1869]" captionTargetBox-2="[281,1305,193,873]" captionTargetId-0="figure@3.[151,1436,865,1620]" captionTargetId-1="figure@4.[150,1436,661,1879]" captionTargetId-2="figure@5.[281,1305,193,873]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 3. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines, body. A, lateral view; B, dorsal view. Scale bar = 1 mm." captionText-1="FIGURE 4. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines. A, left scaphocerite, setae omitted; B, left first pereopod, lateral view; C, left second pereopod, lateral view; D, palm of second pereopod, lateral view; E, left third pereopod, lateral view; F, left fourth pereopod, lateral view; G, dactylus of fourth pereopod, lateral view; H, left fifth pereopod, lateral view; I, left second pleopod, dorsal view. Scale bars = 1 mm (A C, E I); 0.2 mm (D)." captionText-2="FIGURE 5. Glyphocrangon denticulata sp. nov., paratype, ovigerous female (cl 21.1 mm), the Philippines, colour photo. Photographed by Dr. Tin-Yam Chan." httpUri-0="https://zenodo.org/record/228690/files/figure.png" httpUri-1="https://zenodo.org/record/228691/files/figure.png" httpUri-2="https://zenodo.org/record/228692/files/figure.png" pageId="3" pageNumber="441">Figs. 35</figureCitation>
)
</paragraph>
<paragraph id="8B8F673F7F635628FCEDFD5BFD03FD77" blockId="3.[151,832,654,707]" box="[151,677,654,676]" pageId="3" pageNumber="441">
<treatmentCitationGroup id="AB2040117F635628FCEDFD5BFD03FD77" box="[151,677,654,676]" pageId="3" pageNumber="441">
<taxonomicName id="4C301CBC7F635628FCEDFD5BFE21FD77" ID-CoL="9CR7W" box="[151,391,655,676]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="441" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B944BB2D7F635628FCEDFD5BFE21FD77" box="[151,391,655,676]" italics="true" pageId="3" pageNumber="441">Glyphocrangon pugnax</emphasis>
</taxonomicName>
<treatmentCitation id="0A91412E7F635628FDDBFD5AFDE3FD77" author="Chace" box="[417,581,654,676]" page="19" pageId="3" pageNumber="441" year="1984">
<bibRefCitation id="EFA11ACE7F635628FDDBFD5AFD84FD77" author="Chace" box="[417,546,654,676]" pageId="3" pageNumber="441" refString="Chace, F. A. Jr. (1984) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 2: families Glyphocrangonidae and Crangonidae. Smithsonian Contributions to Zoology, 397, 1 - 63. http: // dx. doi. org / 10.5479 / si. 00810282.397" type="journal article" year="1984">Chace, 1984</bibRefCitation>
: 19
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8F673F7F635628FCEDFD79FCE6FD10" blockId="3.[151,832,654,707]" box="[151,832,685,707]" pageId="3" pageNumber="441">
<treatmentCitationGroup id="AB2040117F635628FCEDFD79FCE6FD10" box="[151,832,685,707]" pageId="3" pageNumber="441">
<taxonomicName id="4C301CBC7F635628FCEDFD79FE21FD10" ID-CoL="9CR7W" box="[151,391,685,707]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="441" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B944BB2D7F635628FCEDFD79FE21FD10" box="[151,391,685,707]" italics="true" pageId="3" pageNumber="441">Glyphocrangon pugnax</emphasis>
</taxonomicName>
<treatmentCitation id="0A91412E7F635628FDDAFD79FD05FD10" author="Komai" box="[416,675,685,707]" page="52" pageId="3" pageNumber="441" year="2008">
<bibRefCitation id="EFA11ACE7F635628FDDAFD79FDDBFD10" author="Komai" box="[416,637,685,707]" pageId="3" pageNumber="441" refString="Komai, T. &amp; Chan, T. - Y. (2008) Further records of deep-sea shrimps of the genus Glyphocrangon A. Milne-Edwards, 1881 (Crustacea: Decapoda: Caridea) from the Philippines, with descriptions of three new species. Raffles Bulletin of Zoology, Supplement 19, 39 - 62." type="journal article" year="2008">Komai &amp; Chan, 2008</bibRefCitation>
: 52
</treatmentCitation>
, fig. 13D1, D2.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32A34B47F635628FCEDFD3BFB39FCFE" pageId="3" pageNumber="441" type="materials_examined">
<paragraph id="8B8F673F7F635628FCEDFD3BFB39FCFE" blockId="3.[151,1436,751,813]" pageId="3" pageNumber="441">
<emphasis id="B944BB2D7F635628FCEDFD3BFEDEFCDB" bold="true" box="[151,376,751,776]" pageId="3" pageNumber="441">Material examined</emphasis>
. The
<collectingCountry id="F32727AF7F635628FDC2FD3BFD9CFCDB" box="[440,570,751,776]" name="Philippines" pageId="3" pageNumber="441">Philippines</collectingCountry>
.
<typeStatus id="548BD99D7F635628FE3EFD3BFD09FCDB" box="[580,687,751,776]" pageId="3" pageNumber="441" type="holotype">Holotype</typeStatus>
, male (cl
<quantity id="4CC8CADA7F635628FF61FD24FC24FCDB" box="[795,898,752,776]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="441" unit="mm" value="13.1">13.1 mm</quantity>
),
<typeStatus id="548BD99D7F635628FFEFFD24FBA5FCDB" box="[917,1027,752,776]" pageId="3" pageNumber="441" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9D36ACB67F635628F874FD24FBFEFCDB" box="[1038,1112,751,776]" pageId="3" pageNumber="441" type="male">1 male</specimenCount>
(cl
<quantity id="4CC8CADA7F635628F8F9FD24FB4CFCDB" box="[1155,1258,752,776]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="3" pageNumber="441" unit="mm" value="11.4">11.4 mm</quantity>
), 1 ovig. female (cl
<quantity id="4CC8CADA7F635628FCC1FCC0FE85FCFF" box="[187,291,788,812]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="3" pageNumber="441" unit="mm" value="21.1">21.1 mm</quantity>
), stn. CP2390,
<geoCoordinate id="EE0401F87F635628FDB7FCC0FD93FCFE" box="[461,565,788,813]" direction="north" orientation="latitude" pageId="3" pageNumber="441" precision="92" value="9.456667">9°27.4N</geoCoordinate>
,
<geoCoordinate id="EE0401F87F635628FE3AFCC0FD66FCFE" box="[576,704,788,813]" direction="east" orientation="longitude" pageId="3" pageNumber="441" precision="92" value="123.71833">123°43.1E</geoCoordinate>
,
<quantity id="4CC8CADA7F635628FEB1FCC0FCEAFCFF" box="[715,844,788,813]" metricMagnitude="2" metricUnit="m" metricValue="6.2" metricValueMax="6.27" metricValueMin="6.13" pageId="3" pageNumber="441" unit="m" value="620.0" valueMax="627.0" valueMin="613.0">613627 m</quantity>
, rock and sand,
<date id="FF8E41FF7F635628F87BFCC0FB3DFCFE" box="[1025,1179,788,813]" pageId="3" pageNumber="441" value="2005-05-30">30 May 2005</date>
.
</paragraph>
</subSubSection>
<caption id="DF4F37B77F635628FCEDF9BDFE2BF94E" httpUri="https://zenodo.org/record/228690/files/figure.png" pageId="3" pageNumber="441" targetBox="[158,1423,871,1615]" targetPageId="3">
<paragraph id="8B8F673F7F635628FCEDF9BDFE2BF94E" blockId="3.[151,1436,1641,1693]" pageId="3" pageNumber="441">
<emphasis id="B944BB2D7F635628FCEDF9BDFEB4F9AD" bold="true" box="[151,274,1641,1663]" pageId="3" pageNumber="441">FIGURE 3.</emphasis>
<taxonomicName id="4C301CBC7F635628FD62F9BDFD89F9AD" box="[280,559,1641,1662]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="441" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
<emphasis id="B944BB2D7F635628FD62F9BDFD89F9AD" box="[280,559,1641,1662]" italics="true" pageId="3" pageNumber="441">Glyphocrangon denticulata</emphasis>
</taxonomicName>
<emphasis id="B944BB2D7F635628FE4FF9BDFD23F9AD" bold="true" box="[565,645,1641,1662]" pageId="3" pageNumber="441">
<taxonomicNameLabel id="A27706567F635628FE4FF9BDFD23F9AD" box="[565,645,1641,1662]" pageId="3" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 13.1 mm), the Philippines, body. A, lateral view; B, dorsal view. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="C32A34B47F63562EFCBDF910FA26FA56" lastPageId="5" lastPageNumber="443" pageId="3" pageNumber="441" type="description">
<paragraph id="8B8F673F7F635628FCBDF910FB71F90E" blockId="3.[151,1437,1732,2010]" box="[199,1239,1732,1758]" pageId="3" pageNumber="441">
<emphasis id="B944BB2D7F635628FCBDF910FEF7F90E" bold="true" box="[199,337,1732,1757]" pageId="3" pageNumber="441">Description</emphasis>
. Body slender. Integument of carapace and abdomen naked, without short setae.
</paragraph>
<paragraph id="8B8F673F7F63562FFCBDF93EFF42FF2B" blockId="3.[151,1437,1732,2010]" lastBlockId="4.[151,1437,151,608]" lastPageId="4" lastPageNumber="442" pageId="3" pageNumber="441">
Rostrum (
<figureCitation id="130B7BBA7F635628FD42F93DFEDCF8D2" box="[312,378,1769,1794]" captionStart="FIGURE 3" captionStartId="3.[151,250,1641,1663]" captionTargetBox="[158,1423,871,1615]" captionTargetId="figure@3.[151,1436,865,1620]" captionTargetPageId="3" captionText="FIGURE 3. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines, body. A, lateral view; B, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/228690/files/figure.png" pageId="3" pageNumber="441">Fig. 3</figureCitation>
A, B) almost as long as postorbital carapace length, dorsal surface with long middorsal ridge and two lateral ridges; lateral margin with 2 pairs of teeth, anterior pair anterior to cornea. Carapace (
<figureCitation id="130B7BBA7F635628F963F8D9FAFAF8F6" box="[1305,1372,1805,1830]" captionStart="FIGURE 3" captionStartId="3.[151,250,1641,1663]" captionTargetBox="[158,1423,871,1615]" captionTargetId="figure@3.[151,1436,865,1620]" captionTargetPageId="3" captionText="FIGURE 3. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines, body. A, lateral view; B, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/228690/files/figure.png" pageId="3" pageNumber="441">Fig. 3</figureCitation>
A, B) with first anterior carina composed of 56 acute tubercles, first posterior carina composed of 45 low and blunt tubercles. Second anterior carina composed of 3 acute teeth, anteriormost tooth largest and most acute. Second posterior carina usually composed of some continuous and low tubercles. Third anterior carina continuous with antennal spine, entire, posterior part represented by 23 small and inconspicuous tubercles. Third posterior carina entire, terminating anteriorly in blunt angle, and paralleled with dorsal surface of carapace. Fourth anterior carina high and entire, terminating anteriorly in acute spine. Branchial region bearing some inconspicuous tubercles, arranged at almost 1 row. Two antennal spines smooth, without small spine and almost as long as branchiostegal spines.
</paragraph>
<paragraph id="8B8F673F7F64562FFCBDFED0FA9CFDB3" blockId="4.[151,1437,151,608]" pageId="4" pageNumber="442">
All abdominal somites (
<figureCitation id="130B7BBA7F64562FFDA9FED0FDB5FECF" box="[467,531,260,285]" captionStart="FIGURE 3" captionStartId="3.[151,250,1641,1663]" captionTargetBox="[158,1423,871,1615]" captionTargetId="figure@3.[151,1436,865,1620]" captionTargetPageId="3" captionText="FIGURE 3. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines, body. A, lateral view; B, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/228690/files/figure.png" pageId="4" pageNumber="442">Fig. 3</figureCitation>
A, B) bearing middorsal carinae. Middorsal carinae of second to sixth abdominal somites distinctly notched. First abdominal somite with groove at posterior quarter and distinct middorsal carina, terminating anterior with large and acute spine, same as lateral dorsal carina. Intercarinal space between middorsl carina and submedian carina bearing some small and low tubercles. Middorsal carina of second abdominal somite notched by transverse groove, pleuron armed with 2 ventral spines. Pleuron of third abdominal somite armed with 2 ventral spines, posterior spine larger and more acute. Middorsal ridge of fourth abdominal somite notched at anterior 1/3 by transverse groove, pleuron armed with 2 ventral spines, anterior spine more acute. Pleuron of fifth abdominal somite armed with 2 ventral spines, posterior spine about 2 times longer than anterior one. Middorsal ridge of sixth abdominal somite notched at anterior 1/3 by transverse groove, lateral carina notched at posterior 1/5 to 1/4. Telson about 0.7 times as long as carapace length, bearing blunt protuberance at about anterior 1/5.
</paragraph>
<caption id="DF4F37B77F64562FFCEDF8B8FB0BF80D" httpUri="https://zenodo.org/record/228691/files/figure.png" pageId="4" pageNumber="442" targetBox="[165,1426,671,1869]" targetPageId="4">
<paragraph id="8B8F673F7F64562FFCEDF8B8FB0BF80D" blockId="4.[151,1436,1900,2014]" pageId="4" pageNumber="442">
<emphasis id="B944BB2D7F64562FFCEDF8B8FEB0F852" bold="true" box="[151,278,1900,1922]" pageId="4" pageNumber="442">FIGURE 4.</emphasis>
<taxonomicName id="4C301CBC7F64562FFD65F8B8FD9CF852" box="[287,570,1900,1921]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="442" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
<emphasis id="B944BB2D7F64562FFD65F8B8FD9CF852" box="[287,570,1900,1921]" italics="true" pageId="4" pageNumber="442">Glyphocrangon denticulata</emphasis>
</taxonomicName>
<emphasis id="B944BB2D7F64562FFE39F8B8FD31F852" bold="true" box="[579,663,1900,1921]" pageId="4" pageNumber="442">
<taxonomicNameLabel id="A27706567F64562FFE39F8B8FD31F852" box="[579,663,1900,1921]" pageId="4" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 13.1 mm), the Philippines. A, left scaphocerite, setae omitted; B, left first pereopod, lateral view; C, left second pereopod, lateral view; D, palm of second pereopod, lateral view; E, left third pereopod, lateral view; F, left fourth pereopod, lateral view; G, dactylus of fourth pereopod, lateral view; H, left fifth pereopod, lateral view; I, left second pleopod, dorsal view. Scale bars = 1 mm (AC, EI); 0.2 mm (D).
</paragraph>
</caption>
<caption id="DF4F37B77F65562EFCEDFCABFDAAFC60" httpUri="https://zenodo.org/record/228692/files/figure.png" pageId="5" pageNumber="443" targetBox="[281,1305,193,873]" targetPageId="5">
<paragraph id="8B8F673F7F65562EFCEDFCABFDAAFC60" blockId="5.[151,1437,895,947]" pageId="5" pageNumber="443">
<emphasis id="B944BB2D7F65562EFCEDFCABFEB0FC47" bold="true" box="[151,278,895,917]" pageId="5" pageNumber="443">FIGURE 5.</emphasis>
<taxonomicName id="4C301CBC7F65562EFD5AFCABFD9CFC47" box="[288,570,895,916]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="443" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
<emphasis id="B944BB2D7F65562EFD5AFCABFD9CFC47" box="[288,570,895,916]" italics="true" pageId="5" pageNumber="443">Glyphocrangon denticulata</emphasis>
</taxonomicName>
<emphasis id="B944BB2D7F65562EFE3EFCABFD3EFC47" bold="true" box="[580,664,895,916]" pageId="5" pageNumber="443">
<taxonomicNameLabel id="A27706567F65562EFE3EFCABFD3EFC47" box="[580,664,895,916]" pageId="5" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, ovigerous female (cl 21.1 mm), the Philippines, colour photo. Photographed by Dr. Tin-Yam Chan.
</paragraph>
</caption>
<paragraph id="8B8F673F7F65562EFCBDFC35FE81FBCE" blockId="5.[151,1437,992,2025]" pageId="5" pageNumber="443">
Eyes (
<figureCitation id="130B7BBA7F65562EFD75FC34FEF4FC2B" box="[271,338,992,1017]" captionStart="FIGURE 3" captionStartId="3.[151,250,1641,1663]" captionTargetBox="[158,1423,871,1615]" captionTargetId="figure@3.[151,1436,865,1620]" captionTargetPageId="3" captionText="FIGURE 3. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines, body. A, lateral view; B, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/228690/files/figure.png" pageId="5" pageNumber="443">Fig. 3</figureCitation>
A, B) moderately large, maximum diameter about 0.25 times of carapace length, dark brown in preservative.
</paragraph>
<paragraph id="8B8F673F7F65562EFCBDFBFCFEA4FBB6" blockId="5.[151,1437,992,2025]" pageId="5" pageNumber="443">
Antennal scaphocerite (
<figureCitation id="130B7BBA7F65562EFDAFFBFCFD82FB93" box="[469,548,1064,1089]" captionStart="FIGURE 3" captionStartId="3.[151,250,1641,1663]" captionTargetBox="[158,1423,871,1615]" captionTargetId="figure@3.[151,1436,865,1620]" captionTargetPageId="3" captionText="FIGURE 3. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines, body. A, lateral view; B, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/228690/files/figure.png" pageId="5" pageNumber="443">Figs. 3</figureCitation>
A, B, 4A) about 0.45 times of carapace length, bearing lateral tooth at about basal 1/3.
</paragraph>
<paragraph id="8B8F673F7F65562EFCBDFBA4FED3FAEE" blockId="5.[151,1437,992,2025]" pageId="5" pageNumber="443">
First pereopod (
<figureCitation id="130B7BBA7F65562EFD01FBA4FE19FB5B" box="[379,447,1136,1161]" captionStart="FIGURE 4" captionStartId="4.[151,250,1900,1922]" captionTargetBox="[165,1426,671,1869]" captionTargetId="figure@4.[150,1436,661,1879]" captionTargetPageId="4" captionText="FIGURE 4. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines. A, left scaphocerite, setae omitted; B, left first pereopod, lateral view; C, left second pereopod, lateral view; D, palm of second pereopod, lateral view; E, left third pereopod, lateral view; F, left fourth pereopod, lateral view; G, dactylus of fourth pereopod, lateral view; H, left fifth pereopod, lateral view; I, left second pleopod, dorsal view. Scale bars = 1 mm (A C, E I); 0.2 mm (D)." httpUri="https://zenodo.org/record/228691/files/figure.png" pageId="5" pageNumber="443">Fig. 4</figureCitation>
B) with numerous tufts of short setae on lateral surface of palm. Second pereopod (
<figureCitation id="130B7BBA7F65562EF914FBA4FF00FB7E" captionStart="FIGURE 4" captionStartId="4.[151,250,1900,1922]" captionTargetBox="[165,1426,671,1869]" captionTargetId="figure@4.[150,1436,661,1879]" captionTargetPageId="4" captionText="FIGURE 4. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines. A, left scaphocerite, setae omitted; B, left first pereopod, lateral view; C, left second pereopod, lateral view; D, palm of second pereopod, lateral view; E, left third pereopod, lateral view; F, left fourth pereopod, lateral view; G, dactylus of fourth pereopod, lateral view; H, left fifth pereopod, lateral view; I, left second pleopod, dorsal view. Scale bars = 1 mm (A C, E I); 0.2 mm (D)." httpUri="https://zenodo.org/record/228691/files/figure.png" pageId="5" pageNumber="443">Fig. 4</figureCitation>
C) unequal with right longer than left; left carpus composed of 19 articles and right composed of 30 articles. The last 3 pair of pereopods much stronger than the second pereopod. Third pereopod (
<figureCitation id="130B7BBA7F65562EF8EFFB6CFB79FB03" box="[1173,1247,1208,1233]" captionStart="FIGURE 4" captionStartId="4.[151,250,1900,1922]" captionTargetBox="[165,1426,671,1869]" captionTargetId="figure@4.[150,1436,661,1879]" captionTargetPageId="4" captionText="FIGURE 4. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines. A, left scaphocerite, setae omitted; B, left first pereopod, lateral view; C, left second pereopod, lateral view; D, palm of second pereopod, lateral view; E, left third pereopod, lateral view; F, left fourth pereopod, lateral view; G, dactylus of fourth pereopod, lateral view; H, left fifth pereopod, lateral view; I, left second pleopod, dorsal view. Scale bars = 1 mm (A C, E I); 0.2 mm (D)." httpUri="https://zenodo.org/record/228691/files/figure.png" pageId="5" pageNumber="443">Fig. 4</figureCitation>
E) only slightly overreaching distal margin of scaphocerite, propodus without distal tuft of setae. Fourth and fifth pereopods (
<figureCitation id="130B7BBA7F65562EF915FB08FF03FACB" captionStart="FIGURE 4" captionStartId="4.[151,250,1900,1922]" captionTargetBox="[165,1426,671,1869]" captionTargetId="figure@4.[150,1436,661,1879]" captionTargetPageId="4" captionText="FIGURE 4. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines. A, left scaphocerite, setae omitted; B, left first pereopod, lateral view; C, left second pereopod, lateral view; D, palm of second pereopod, lateral view; E, left third pereopod, lateral view; F, left fourth pereopod, lateral view; G, dactylus of fourth pereopod, lateral view; H, left fifth pereopod, lateral view; I, left second pleopod, dorsal view. Scale bars = 1 mm (A C, E I); 0.2 mm (D)." httpUri="https://zenodo.org/record/228691/files/figure.png" pageId="5" pageNumber="443">Fig. 4</figureCitation>
F, G, H) with dactylus simple and acuminate. Second pleopod (
<figureCitation id="130B7BBA7F65562EFF17FAD4FC17FACB" box="[877,945,1280,1305]" captionStart="FIGURE 4" captionStartId="4.[151,250,1900,1922]" captionTargetBox="[165,1426,671,1869]" captionTargetId="figure@4.[150,1436,661,1879]" captionTargetPageId="4" captionText="FIGURE 4. Glyphocrangon denticulata sp. nov., holotype, male (cl 13.1 mm), the Philippines. A, left scaphocerite, setae omitted; B, left first pereopod, lateral view; C, left second pereopod, lateral view; D, palm of second pereopod, lateral view; E, left third pereopod, lateral view; F, left fourth pereopod, lateral view; G, dactylus of fourth pereopod, lateral view; H, left fifth pereopod, lateral view; I, left second pleopod, dorsal view. Scale bars = 1 mm (A C, E I); 0.2 mm (D)." httpUri="https://zenodo.org/record/228691/files/figure.png" pageId="5" pageNumber="443">
Fig.
<date id="FF8E41FF7F65562EFFD8FAD4FC17FACB" box="[930,945,1280,1304]" pageId="5" pageNumber="443">4</date>
</figureCitation>
I) with endopod almost as long as appendix masculine in males.
</paragraph>
<paragraph id="8B8F673F7F65562EFCBDFA9CFA26FA56" blockId="5.[151,1437,992,2025]" pageId="5" pageNumber="443">
<emphasis id="B944BB2D7F65562EFCBDFA9CFE3CFAB2" bold="true" box="[199,410,1352,1377]" pageId="5" pageNumber="443">Coloration in life.</emphasis>
Body orange dorsally, whitish translucent laterally. Rostrum with anterior 1/3 cardinal red. Carina and tubercles on carapace and abdomen palm yellow. Most pereopod whitish. Eggs pale blue. See
<figureCitation id="130B7BBA7F65562EF94DFAB8FADDFA56" box="[1335,1403,1388,1413]" captionStart="FIGURE 5" captionStartId="5.[151,250,895,917]" captionTargetBox="[281,1305,193,873]" captionTargetId="figure@5.[281,1305,193,873]" captionTargetPageId="5" captionText="FIGURE 5. Glyphocrangon denticulata sp. nov., paratype, ovigerous female (cl 21.1 mm), the Philippines, colour photo. Photographed by Dr. Tin-Yam Chan." httpUri="https://zenodo.org/record/228692/files/figure.png" pageId="5" pageNumber="443">Fig. 5</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A34B47F65562EFCBDFA44FAE8FA7B" box="[199,1358,1424,1449]" pageId="5" pageNumber="443" type="distribution">
<paragraph id="8B8F673F7F65562EFCBDFA44FAE8FA7B" blockId="5.[151,1437,992,2025]" box="[199,1358,1424,1449]" pageId="5" pageNumber="443">
<emphasis id="B944BB2D7F65562EFCBDFA44FEC6FA7A" bold="true" box="[199,352,1424,1449]" pageId="5" pageNumber="443">Distribution.</emphasis>
So far only known from the
<typeStatus id="548BD99D7F65562EFEDEFA45FD72FA7A" box="[676,724,1425,1449]" pageId="5" pageNumber="443">type</typeStatus>
locality in the
<collectingCountry id="F32727AF7F65562EFF07FA44FC58FA7A" box="[893,1022,1424,1449]" name="Philippines" pageId="5" pageNumber="443">Philippines</collectingCountry>
, at depths of 613627 meters.
</paragraph>
</subSubSection>
<subSubSection id="C32A34B47F65562EFCBDFA60FE1FFA22" pageId="5" pageNumber="443" type="etymology">
<paragraph id="8B8F673F7F65562EFCBDFA60FE1FFA22" blockId="5.[151,1437,992,2025]" pageId="5" pageNumber="443">
<emphasis id="B944BB2D7F65562EFCBDFA60FEE8FA1E" bold="true" box="[199,334,1460,1485]" pageId="5" pageNumber="443">Etymology.</emphasis>
Derived from the Latin
<emphasis id="B944BB2D7F65562EFE27FA60FD40FA1F" box="[605,742,1460,1484]" italics="true" pageId="5" pageNumber="443">denticulatus</emphasis>
, means bearing small tooth, in referring to the lateral tooth on the antennal scaphocerite.
</paragraph>
</subSubSection>
<subSubSection id="C32A34B47F65562EFCBDFA28FE71F83A" pageId="5" pageNumber="443" type="discussion">
<paragraph id="8B8F673F7F65562EFCBDFA28FE71F83A" blockId="5.[151,1437,992,2025]" pageId="5" pageNumber="443">
<emphasis id="B944BB2D7F65562EFCBDFA28FE9DF9C6" bold="true" box="[199,315,1532,1557]" pageId="5" pageNumber="443">Remarks.</emphasis>
The new species closely resembles
<taxonomicName id="4C301CBC7F65562EFEA2FA28FB29F9C6" authority="De Man, 1918" authorityName="De Man" authorityYear="1918" box="[728,1167,1532,1557]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="443" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B944BB2D7F65562EFEA2FA28FC44F9C7" box="[728,994,1532,1556]" italics="true" pageId="5" pageNumber="443">Glyphocrangon pugnax</emphasis>
<bibRefCitation id="EFA11ACE7F65562EFF91FA29FB29F9C6" author="De" box="[1003,1167,1533,1557]" pageId="5" pageNumber="443" refString="De Man, J. G. (1918) Diagnoses of new species of macrurous decapod Crustacea from the Siboga Expedition. Tijdschrift der Nederlandsche Dierkundige Vereeniging, Series 2, 16, 293 - 306." type="journal article" year="1918">
<collectingCountry id="F32727AF7F65562EFF91FA29FBAAF9C7" box="[1003,1036,1533,1556]" name="South Georgia and the South Sandwich Islands" pageId="5" pageNumber="443">De</collectingCountry>
<collectingCountry id="F32727AF7F65562EF86EFA29FBECF9C7" box="[1044,1098,1533,1556]" name="Isle of Man" pageId="5" pageNumber="443">Man</collectingCountry>
, 1918
</bibRefCitation>
</taxonomicName>
with the entire anterior third carina on the carapace. It differs from
<taxonomicName id="4C301CBC7F65562EFEFBF9F5FD55F9EB" box="[641,755,1569,1592]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="443" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B944BB2D7F65562EFEFBF9F5FD55F9EB" box="[641,755,1569,1592]" italics="true" pageId="5" pageNumber="443">G. pugnax</emphasis>
</taxonomicName>
by: 1) the antennal scaphocerite bears a distinct lateral tooth at about basal 1/3, whereas in
<taxonomicName id="4C301CBC7F65562EFD8FF991FDCEF98F" box="[501,616,1605,1628]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="443" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B944BB2D7F65562EFD8FF991FDCEF98F" box="[501,616,1605,1628]" italics="true" pageId="5" pageNumber="443">G. pugnax</emphasis>
</taxonomicName>
, the antennal scaphocerite has no trace of lateral tooth (
<bibRefCitation id="EFA11ACE7F65562EF897F991FA31F98E" author="De" box="[1261,1431,1605,1629]" pageId="5" pageNumber="443" refString="De Man, J. G. (1918) Diagnoses of new species of macrurous decapod Crustacea from the Siboga Expedition. Tijdschrift der Nederlandsche Dierkundige Vereeniging, Series 2, 16, 293 - 306." type="journal article" year="1918">
<collectingCountry id="F32727AF7F65562EF897F991FAA9F98E" box="[1261,1295,1605,1629]" name="South Georgia and the South Sandwich Islands" pageId="5" pageNumber="443">De</collectingCountry>
<collectingCountry id="F32727AF7F65562EF962F991FAE8F98F" box="[1304,1358,1605,1628]" name="Isle of Man" pageId="5" pageNumber="443">Man</collectingCountry>
, 1918
</bibRefCitation>
;
<bibRefCitation id="EFA11ACE7F65562EFCEDF9BCFF75F953" author="Man" box="[151,211,1640,1664]" pageId="5" pageNumber="443" refString="De Man, J. G. (1920) The Decapoda of the Siboga Expedition. Part IV. Families Pasiphaeidae, Stylodactylidae, Hoplophoridae, Nematocarcinidae, Thalassocaridae, Pandalidae, Psalidopodidae, Gnathophyllidae, Processidae, Glyphocrangonidae and Crangonidae. Siboga Expeditie, 39 a 3, 1 - 318." type="journal article" year="1920">1920</bibRefCitation>
;
<bibRefCitation id="EFA11ACE7F65562EFCA7F9BCFED2F953" author="Komai" box="[221,372,1640,1665]" pageId="5" pageNumber="443" refString="Komai, T. (2004) A review of the Indo-West Pacific species of the genus Glyphocrangon A. Milne Edwards, 1881 (excluding the G. caeca species group) (Crustacea: Decapoda: Caridea: Glyphocrangonidae). Memoires du Museum national d'Histoire naturelle, 191, 375 - 610." type="journal article" year="2004">Komai, 2004</bibRefCitation>
); 2) the rostrum is about as long as (male) or 0.81 times as long as (female) the carapace length, whereas in
<taxonomicName id="4C301CBC7F65562EFD67F959FE37F977" box="[285,401,1677,1700]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="443" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B944BB2D7F65562EFD67F959FE37F977" box="[285,401,1677,1700]" italics="true" pageId="5" pageNumber="443">G. pugnax</emphasis>
</taxonomicName>
, the rostrum is about 0.680.77 times as long as the carapace length; 3) the carinae and intercarinae are less unconspicuous on the carapace, the posterior dorsolateral region bears only 311 small and low tubercles arranged in 1 row, and the upper part of the branchial region bears 710 tubercles arrangd in 1 row, whereas in
<taxonomicName id="4C301CBC7F65562EFD6FF92DFE20F8C3" box="[277,390,1785,1808]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="443" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B944BB2D7F65562EFD6FF92DFE20F8C3" box="[277,390,1785,1808]" italics="true" pageId="5" pageNumber="443">G. pugnax</emphasis>
</taxonomicName>
, the posterior dorsolateral region bears 2030 small pointed tubercles arranged in 3 rows, and the branchial region with the upper part bears 3040 tubercles arranged in 3 irregular rows; 4) the posterior spine on the pleuron of the fifth abdominal somite is about 2 times longer than the anterior one, whereas in
<taxonomicName id="4C301CBC7F65562EF886F895FACAF88B" box="[1276,1388,1857,1880]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="443" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B944BB2D7F65562EF886F895FACAF88B" box="[1276,1388,1857,1880]" italics="true" pageId="5" pageNumber="443">G. pugnax</emphasis>
</taxonomicName>
, the 2 spines are of equal in size; 5) the carapace is orange dorsally, whitish translucent laterally, whereas in
<taxonomicName id="4C301CBC7F65562EF95EF8B1FA32F8AF" box="[1316,1428,1893,1916]" class="Malacostraca" family="Glyphocrangonidae" genus="Glyphocrangon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="443" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B944BB2D7F65562EF95EF8B1FA32F8AF" box="[1316,1428,1893,1916]" italics="true" pageId="5" pageNumber="443">G. pugnax</emphasis>
</taxonomicName>
, it is salmon pink throughout (Jones &amp; Morgan, 1994;
<bibRefCitation id="EFA11ACE7F65562EFF71F85CFC02F873" author="Komai" box="[779,932,1928,1953]" pageId="5" pageNumber="443" refString="Komai, T. (2004) A review of the Indo-West Pacific species of the genus Glyphocrangon A. Milne Edwards, 1881 (excluding the G. caeca species group) (Crustacea: Decapoda: Caridea: Glyphocrangonidae). Memoires du Museum national d'Histoire naturelle, 191, 375 - 610." type="journal article" year="2004">Komai, 2004</bibRefCitation>
),
<bibRefCitation id="EFA11ACE7F65562EFFC0F85CFB19F873" author="Komai" box="[954,1215,1928,1953]" pageId="5" pageNumber="443" refString="Komai, T. &amp; Chan, T. - Y. (2008) Further records of deep-sea shrimps of the genus Glyphocrangon A. Milne-Edwards, 1881 (Crustacea: Decapoda: Caridea) from the Philippines, with descriptions of three new species. Raffles Bulletin of Zoology, Supplement 19, 39 - 62." type="journal article" year="2008">Komai &amp; Chan (2008)</bibRefCitation>
explained that this difference in colouration may be due to the perservation condition of the photographed specimen or the printing condition of the photograph.
</paragraph>
</subSubSection>
</treatment>
</document>