668 lines
151 KiB
XML
668 lines
151 KiB
XML
<document id="13EE39F79628E450693A44E29455A6C7" ID-DOI="10.5281/zenodo.205966" ID-GBIF-Dataset="4f63be5b-6550-40d8-8e17-7cf7b455ec03" ID-ISSN="1175-5326" ID-Zenodo-Dep="205966" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226899271" checkinUser="plazi" docAuthor="López-Arbarello, Adriana & Alvarado-Ortega, Jesús" docDate="2011" docId="0399C467FFE3021EFF42FF1C6FA0FA93" docLanguage="en" docName="zt02749p024.pdf" docOrigin="Zootaxa 2749" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tlayuamichin itztli" docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FFA0BC1FFFE7020FFFD5FF8B6A2FFD66" masterDocTitle="New semionotiform (Neopterygii) from the Tlayúa Quarry (Early Cretaceous, Albian), Mexico" masterLastPageNumber="24" masterPageNumber="1" pageNumber="5" updateTime="1698238591706" updateUser="plazi">
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<mods:titleInfo id="E5A097437F27ADFC4D6BAA3814BC4B40">
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<mods:title id="3C5C72DA8C1165796762C25C6E1C2390">New semionotiform (Neopterygii) from the Tlayúa Quarry (Early Cretaceous, Albian), Mexico</mods:title>
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</mods:titleInfo>
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<mods:name id="38FB311BC536A93A8F1ABD942991D6B9" type="personal">
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<mods:roleTerm id="1ACACF16173257480FBEF0BD0EC83005">Author</mods:roleTerm>
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<mods:namePart id="5D9599AE1F88E23DA1F1FF9C6366F011">López-Arbarello, Adriana</mods:namePart>
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<mods:roleTerm id="241B0307701799137C72F2CD607449EB">Author</mods:roleTerm>
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<mods:namePart id="12AB8FBB23317770C9DA6F02B57E3567">Alvarado-Ortega, Jesús</mods:namePart>
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<mods:typeOfResource id="49016A9274BEF9DB1B0F931ACBEDD1DF">text</mods:typeOfResource>
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<mods:title id="63D6456CB55E333470DBE550A497BD9C">Zootaxa</mods:title>
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<mods:part id="05575615E593915AC62168C96B62FE63">
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<mods:date id="9FEC42E389B6652F6B031F9DAA1DDAE6">2011</mods:date>
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<mods:number id="B4041D84DAAAFF1FEE58582476FFECF0">2749</mods:number>
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<mods:classification id="027637AF6A21EA20E094FFABBDE487D8">journal article</mods:classification>
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<mods:identifier id="0A6E415516ACF9E47EDF4C2DD8A5002E" type="DOI">10.5281/zenodo.205966</mods:identifier>
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<treatment id="0399C467FFE3021EFF42FF1C6FA0FA93" ID-DOI="http://doi.org/10.5281/zenodo.6184215" ID-GBIF-Taxon="119386649" ID-Zenodo-Dep="6184215" LSID="urn:lsid:plazi:treatment:0399C467FFE3021EFF42FF1C6FA0FA93" httpUri="http://treatment.plazi.org/id/0399C467FFE3021EFF42FF1C6FA0FA93" lastPageId="17" lastPageNumber="18" pageId="4" pageNumber="5">
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<subSubSection id="C32A26FAFFE3020BFF42FF1C6B37FDB4" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="8B8F7571FFE3020BFF42FF1C6816FDD7" blockId="4.[151,569,151,210]" box="[151,569,151,177]" pageId="4" pageNumber="5">
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<heading id="D0C7C21DFFE3020BFF42FF1C6816FDD7" bold="true" box="[151,569,151,177]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
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<emphasis id="B944A963FFE3020BFF42FF1C6816FDD7" bold="true" box="[151,569,151,177]" pageId="4" pageNumber="5">
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<taxonomicName id="4C300EF2FFE3020BFF42FF1C6B54FDD7" box="[151,379,151,177]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="itztli">
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<emphasis id="B944A963FFE3020BFF42FF1C6B54FDD7" bold="true" box="[151,379,151,177]" italics="true" pageId="4" pageNumber="5">Tlayuamichin itztli</emphasis>
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</taxonomicName>
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gen. et sp. nov.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8F7571FFE3020BFF42FF316B37FDB4" blockId="4.[151,569,151,210]" box="[151,280,186,210]" pageId="4" pageNumber="5">
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(
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<figureCitation id="130B69F4FFE3020BFF4AFF316B3EFDB4" box="[159,273,186,210]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-10="FIGURE 12" captionStart-11="FIGURE 13" captionStart-12="FIGURE 14" captionStart-13="FIGURE 15" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStart-8="FIGURE 10" captionStart-9="FIGURE 11" captionStartId-0="4.[151,249,1014,1037]" captionStartId-1="6.[151,249,1801,1824]" captionStartId-10="14.[153,251,1156,1179]" captionStartId-11="16.[151,249,1619,1642]" captionStartId-12="17.[151,249,1236,1259]" captionStartId-13="18.[151,249,807,830]" captionStartId-2="7.[151,249,938,961]" captionStartId-3="7.[151,249,1636,1659]" captionStartId-4="8.[151,249,1202,1225]" captionStartId-5="9.[151,249,1517,1540]" captionStartId-6="10.[151,249,1361,1384]" captionStartId-7="11.[151,249,1832,1855]" captionStartId-8="12.[151,249,1074,1097]" captionStartId-9="12.[151,249,1938,1961]" captionTargetBox-0="[151,1436,427,993]" captionTargetBox-1="[151,1436,193,1780]" captionTargetBox-10="[155,1436,177,1140]" captionTargetBox-11="[151,1436,1154,1598]" captionTargetBox-12="[375,1211,505,1215]" captionTargetBox-13="[152,1435,181,786]" captionTargetBox-2="[151,1436,193,916]" captionTargetBox-3="[151,1436,1010,1614]" captionTargetBox-4="[278,1307,179,1187]" captionTargetBox-5="[188,1398,625,1463]" captionTargetBox-6="[151,1436,193,1339]" captionTargetBox-7="[151,1436,841,1811]" captionTargetBox-8="[151,1436,193,1053]" captionTargetBox-9="[151,1431,1181,1915]" captionTargetId-0="figure@4.[151,1436,427,993]" captionTargetId-1="figure@6.[151,1436,193,1780]" captionTargetId-10="figure@14.[155,1440,177,1141]" captionTargetId-11="figure@16.[151,1436,1154,1598]" captionTargetId-12="figure@17.[375,1211,505,1215]" captionTargetId-13="figure@18.[152,1437,181,786]" captionTargetId-2="figure@7.[151,1436,193,917]" captionTargetId-3="figure@7.[151,1436,1010,1615]" captionTargetId-4="figure@8.[278,1308,179,1188]" captionTargetId-5="figure@9.[151,1436,589,1496]" captionTargetId-6="figure@10.[151,1436,193,1340]" captionTargetId-7="figure@11.[151,1436,841,1811]" captionTargetId-8="figure@12.[151,1436,193,1053]" captionTargetId-9="figure@12.[151,1436,1177,1917]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-10="14" captionTargetPageId-11="16" captionTargetPageId-12="17" captionTargetPageId-13="18" captionTargetPageId-2="7" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="12" captionTargetPageId-9="12" captionText-0="FIGURE 2. Tlayuamichin itztli gen. et sp. nov. Holotype (IGM 6716) preserved in left lateral view. Standard length of the specimen = 403 mm. Scale bar equals 70 mm." captionText-1="FIGURE 3. Tlayuamichin itztli gen. et sp. nov. A, Specimen IGM 6717 preserved in right lateroventral view. B, specimen IGM 6718 preserved in right lateral view. Scale bars equal 70 mm." captionText-10="FIGURE 12. Tlayuamichin itztli gen. et sp. nov., detailed photograph of the anterior part of the skull preserved in ventral view in IGM 6717. Scale bar equals = 10 mm." captionText-11="FIGURE 13. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of: A, the dorsal fin in IGM 6718; B, the anal fin in IGM 6718. Scale bars equal = 20 mm." captionText-12="FIGURE 14. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the caudal fin in IGM 6718. Scale bar equals = 20 mm." captionText-13="FIGURE 15. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of: A, the dorsal ridge scales in IGM 6719; B, several partially articulated scales in IGM 6717 showing the well-developed articular, anterodorsal and anteroventral processes. Scale bars equal = 10 mm." captionText-2="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." captionText-3="FIGURE 5. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6720 preserved in right lateral view. Scale bar equals = 50 mm." captionText-4="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-5="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-6="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." captionText-7="FIGURE 9. Tlayuamichin itztli gen. et sp. nov., details of the skull in IGM 6719. A, posterior part of the skull showing multiple opercular ossifications and the peculiar shape of the suboperculum in this specimen. B, detailed photograph of the dermopterotic bone; the arrow points to the peculiar anteroventral process of this bone in the new taxon. C, same photograph as in B, but in this case the arrows point to the pores of the preopercular sensory canal in the preoperculum (below), suborbital (middle) and dermopterotic (above). Scale bars equal = 10 mm." captionText-8="FIGURE 10. Tlayuamichin itztli gen. et sp. nov., ethmoidal region of the skull in IGM 6716 (holotype) showing the paraethmoids as indicated in Figure 7 (labeled as “ pet ”). Scale bar equals = 10 mm." captionText-9="FIGURE 11. Tlayuamichin itztli gen. et sp. nov., jaws in IGM 6719. A, photograph; B, line drawing. Note that the retroarticular bone is a little displaced posteriorly. Scale bar equals = 10 mm." httpUri-0="https://zenodo.org/record/205968/files/figure.png" httpUri-1="https://zenodo.org/record/205969/files/figure.png" httpUri-10="https://zenodo.org/record/205979/files/figure.png" httpUri-11="https://zenodo.org/record/205980/files/figure.png" httpUri-12="https://zenodo.org/record/205981/files/figure.png" httpUri-13="https://zenodo.org/record/205982/files/figure.png" httpUri-2="https://zenodo.org/record/205970/files/figure.png" httpUri-3="https://zenodo.org/record/205971/files/figure.png" httpUri-4="https://zenodo.org/record/205972/files/figure.png" httpUri-5="https://zenodo.org/record/205973/files/figure.png" httpUri-6="https://zenodo.org/record/205975/files/figure.png" httpUri-7="https://zenodo.org/record/205976/files/figure.png" httpUri-8="https://zenodo.org/record/205977/files/figure.png" httpUri-9="https://zenodo.org/record/205978/files/figure.png" pageId="4" pageNumber="5">Figs 2–15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C32A26FAFFE3020BFF42FF76686FFC74" box="[151,576,253,275]" pageId="4" pageNumber="5" type="description">
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<paragraph id="8B8F7571FFE3020BFF42FF76686FFC74" blockId="4.[151,576,253,275]" box="[151,576,253,275]" pageId="4" pageNumber="5">
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<taxonomicName id="4C300EF2FFE3020BFF42FF766AD6FC75" box="[151,249,253,275]" class="Actinopterygii" family="Semionotidae" genus="Lepidotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
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<emphasis id="B944A963FFE3020BFF42FF766AD6FC75" box="[151,249,253,275]" italics="true" pageId="4" pageNumber="5">Lepidotes</emphasis>
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</taxonomicName>
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sp. Applegate, 1996: 531, fig. 1
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A26FAFFE3020BFF42FEB06908FB46" pageId="4" pageNumber="5" type="materials_examined">
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<paragraph id="8B8F7571FFE3020BFF42FEB06952FC10" blockId="4.[151,1436,314,374]" pageId="4" pageNumber="5">
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<emphasis id="B944A963FFE3020BFF42FEB06B24FC35" bold="true" box="[151,267,315,339]" pageId="4" pageNumber="5">
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<typeStatus id="548BCBD3FFE3020BFF42FEB06B29FC35" box="[151,262,315,339]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED21EDB4FFE3020BFEC1FEB16B65FC34" box="[276,330,314,338]" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="4" pageNumber="5">IGM</collectionCode>
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6716; fairly complete specimen of
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<quantity id="4CC8D894FFE3020BFD31FEB1696CFC34" box="[740,835,314,338]" metricMagnitude="-1" metricUnit="m" metricValue="4.029999999999999" pageId="4" pageNumber="5" unit="mm" value="403.0">403 mm</quantity>
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SL (
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<figureCitation id="130B69F4FFE3020BFCABFEB169EAFC34" box="[894,965,314,338]" captionStart="FIGURE 2" captionStartId="4.[151,249,1014,1037]" captionTargetBox="[151,1436,427,993]" captionTargetId="figure@4.[151,1436,427,993]" captionTargetPageId="4" captionText="FIGURE 2. Tlayuamichin itztli gen. et sp. nov. Holotype (IGM 6716) preserved in left lateral view. Standard length of the specimen = 403 mm. Scale bar equals 70 mm." httpUri="https://zenodo.org/record/205968/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
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). Although missing parts of the unpaired fins, the specimen is almost completely and excellently preserved.
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</paragraph>
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<caption id="DF4F25F9FFE3020BFF42FC7D6844F94D" httpUri="https://zenodo.org/record/205968/files/figure.png" pageId="4" pageNumber="5" targetBox="[151,1436,427,993]" targetPageId="4">
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<paragraph id="8B8F7571FFE3020BFF42FC7D6844F94D" blockId="4.[151,1436,1014,1067]" pageId="4" pageNumber="5">
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<emphasis id="B944A963FFE3020BFF42FC7D6B3BF96A" bold="true" box="[151,276,1014,1037]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
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<taxonomicName id="4C300EF2FFE3020BFEC9FC7C6BF2F96B" box="[284,477,1015,1037]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="itztli">
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<emphasis id="B944A963FFE3020BFEC9FC7C6BF2F96B" box="[284,477,1015,1037]" italics="true" pageId="4" pageNumber="5">Tlayuamichin itztli</emphasis>
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</taxonomicName>
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<emphasis id="B944A963FFE3020BFE33FC7C68A6F96A" bold="true" box="[486,649,1015,1036]" pageId="4" pageNumber="5">gen. et sp. nov.</emphasis>
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Holotype (IGM 6716) preserved in left lateral view. Standard length of the specimen = 403 mm. Scale bar equals 70 mm.
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</paragraph>
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</caption>
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<paragraph id="8B8F7571FFE3020BFF12FBD268F3F82E" blockId="4.[151,1436,1112,1820]" pageId="4" pageNumber="5">
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<emphasis id="B944A963FFE3020BFF12FBD26B69F917" bold="true" box="[199,326,1113,1137]" pageId="4" pageNumber="5">
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<typeStatus id="548BCBD3FFE3020BFF12FBD26B6EF917" box="[199,321,1113,1137]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED21EDB4FFE3020BFE99FBD36BADF916" box="[332,386,1112,1136]" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="4" pageNumber="5">IGM</collectionCode>
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6717–5 (
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<figureCitation id="130B69F4FFE3020BFE39FBD3687DF916" box="[492,594,1112,1136]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="6.[151,249,1801,1824]" captionStartId-1="7.[151,249,938,961]" captionStartId-2="7.[151,249,1636,1659]" captionTargetBox-0="[151,1436,193,1780]" captionTargetBox-1="[151,1436,193,916]" captionTargetBox-2="[151,1436,1010,1614]" captionTargetId-0="figure@6.[151,1436,193,1780]" captionTargetId-1="figure@7.[151,1436,193,917]" captionTargetId-2="figure@7.[151,1436,1010,1615]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="7" captionText-0="FIGURE 3. Tlayuamichin itztli gen. et sp. nov. A, Specimen IGM 6717 preserved in right lateroventral view. B, specimen IGM 6718 preserved in right lateral view. Scale bars equal 70 mm." captionText-1="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." captionText-2="FIGURE 5. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6720 preserved in right lateral view. Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205969/files/figure.png" httpUri-1="https://zenodo.org/record/205970/files/figure.png" httpUri-2="https://zenodo.org/record/205971/files/figure.png" pageId="4" pageNumber="5">Figs 3–5</figureCitation>
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).
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||
<collectionCode id="ED21EDB4FFE3020BFDB3FBD368B3F916" box="[614,668,1112,1136]" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="4" pageNumber="5">IGM</collectionCode>
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6717 is an almost complete specimen, but preserved in lateroventral view and, although the bones are not largely displaced, the skull is disarticulated (
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<figureCitation id="130B69F4FFE3020BFBEEFBF76E51F9F2" box="[1083,1150,1148,1172]" captionStart="FIGURE 3" captionStartId="6.[151,249,1801,1824]" captionTargetBox="[151,1436,193,1780]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tlayuamichin itztli gen. et sp. nov. A, Specimen IGM 6717 preserved in right lateroventral view. B, specimen IGM 6718 preserved in right lateral view. Scale bars equal 70 mm." httpUri="https://zenodo.org/record/205969/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
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A).
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<collectionCode id="ED21EDB4FFE3020BFB7EFBF76ECEF9F2" box="[1195,1249,1148,1172]" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="4" pageNumber="5">IGM</collectionCode>
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6718 is missing the head and pectoral girdle, but the dorsal, anal, and caudal fins are very well preserved (
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<figureCitation id="130B69F4FFE3020BFB57FB2B6EEBF9DE" box="[1154,1220,1184,1208]" captionStart="FIGURE 3" captionStartId="6.[151,249,1801,1824]" captionTargetBox="[151,1436,193,1780]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tlayuamichin itztli gen. et sp. nov. A, Specimen IGM 6717 preserved in right lateroventral view. B, specimen IGM 6718 preserved in right lateral view. Scale bars equal 70 mm." httpUri="https://zenodo.org/record/205969/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
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B).
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<collectionCode id="ED21EDB4FFE3020BFB3BFB2B6F0BF9DE" box="[1262,1316,1184,1208]" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="4" pageNumber="5">IGM</collectionCode>
|
||
6719 contains a very well preserved and articulated skull with the anterodorsal portion of the body without fins (
|
||
<figureCitation id="130B69F4FFE3020BFA92FB4F6FA0F9BA" box="[1351,1423,1220,1244]" captionStart="FIGURE 4" captionStartId="7.[151,249,938,961]" captionTargetBox="[151,1436,193,916]" captionTargetId="figure@7.[151,1436,193,917]" captionTargetPageId="7" captionText="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." httpUri="https://zenodo.org/record/205970/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
|
||
).
|
||
<collectionCode id="ED21EDB4FFE3020BFF42FB636AE2F866" box="[151,205,1256,1280]" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="4" pageNumber="5">IGM</collectionCode>
|
||
6720 is partially incomplete (the paired fins and part of the unpaired fins and squamation are missing) but, although some of the bones of the head are crushed, its general preservation is excellent displaying some important skull features not preserved in the
|
||
<typeStatus id="548BCBD3FFE3020BFDCDFABB6855F82E" box="[536,634,1328,1352]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="130B69F4FFE3020BFD5FFABB68E1F82E" box="[650,718,1328,1352]" captionStart="FIGURE 5" captionStartId="7.[151,249,1636,1659]" captionTargetBox="[151,1436,1010,1614]" captionTargetId="figure@7.[151,1436,1010,1615]" captionTargetPageId="7" captionText="FIGURE 5. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6720 preserved in right lateral view. Scale bar equals = 50 mm." httpUri="https://zenodo.org/record/205971/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE3020BFF12FADE6E7AF80A" blockId="4.[151,1436,1112,1820]" box="[199,1109,1364,1389]" pageId="4" pageNumber="5">
|
||
<emphasis id="B944A963FFE3020BFF12FADE6B4AF80B" bold="true" box="[199,357,1364,1389]" pageId="4" pageNumber="5">
|
||
<typeStatus id="548BCBD3FFE3020BFF12FADE6B2DF80B" box="[199,258,1365,1389]" pageId="4" pageNumber="5">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Tlayúa quarry, near Tepexi de Rodríguez, Puebla,
|
||
<collectingCountry id="F32735E1FFE3020BFC4BFADF69DCF80A" box="[926,1011,1364,1388]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
|
||
(
|
||
<figureCitation id="130B69F4FFE3020BFBD6FADF6E6BF80A" box="[1027,1092,1364,1388]" captionStart="FIGURE 1" captionStartId="1.[151,249,1532,1555]" captionTargetBox="[454,1131,394,1500]" captionTargetId="figure@1.[437,1149,375,1509]" captionTargetPageId="1" captionText="FIGURE 1. Location map of the Tlayúa Quarry near Tepexi de Rodríguez, Puebla State, central Mexico." httpUri="https://zenodo.org/record/205967/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE3020BFF12FAF26F69F8D2" blockId="4.[151,1436,1112,1820]" pageId="4" pageNumber="5">
|
||
<emphasis id="B944A963FFE3020BFF12FAF26B42F8F7" bold="true" box="[199,365,1400,1425]" pageId="4" pageNumber="5">
|
||
<typeStatus id="548BCBD3FFE3020BFF12FAF26B2DF8F7" box="[199,258,1401,1425]" pageId="4" pageNumber="5">Type</typeStatus>
|
||
horizon.
|
||
</emphasis>
|
||
Middle Member of the Tlayúa Formation (
|
||
<bibRefCitation id="EFA10880FFE3020BFCB7FAF36E6FF8F6" author="Pantoja-Alor" box="[866,1088,1400,1424]" pageId="4" pageNumber="24" refString="Pantoja-Alor, J. (1992) Geologia y paleoambientes de la Cantera Tlayua, Tepexi de Rodriguez, estado de Puebla. Revista - Instituto de Geologia, 9, 156 - 169." type="journal article" year="1992">Pantoja-Alor, 1992</bibRefCitation>
|
||
); Middle-Upper Albian (Early Cretaceous) (
|
||
<bibRefCitation id="EFA10880FFE3020BFEFEFA17686BF8D2" author="Seibertz" box="[299,580,1436,1460]" pageId="4" pageNumber="24" refString="Seibertz, E. & Buitron, B. E. (1987) Paleontologia y estratigrafia de los Neohibolites del Albiano de Tepexi de Rodri´guez, Estado de Puebla (Cretacico Medio, Mexico). Sociedad Mexicana de Paleontologia, 1, 285 - 299." type="journal article" year="1987">Seibertz & Buitrón 1987</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA10880FFE3020BFD85FA176909F8D2" author="Cantu-Chapa" box="[592,806,1436,1460]" pageId="4" pageNumber="22" refString="Cantu-Chapa, A. (1987) Las amonitas del Albiano Superior de Tepexi de Rodriguez, Puebla. Sociedad Mexicana de Paleontologia, Revista, 1, 159 - 160." type="journal article" year="1987">Cantú-Chapa 1987</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA10880FFE3020BFCE4FA176E10F8D2" author="Seibertz" box="[817,1087,1436,1460]" pageId="4" pageNumber="24" refString="Seibertz, E. & Spaeth, C. (2002) Cretaceous belemnites of Mexico III: The Albian Neo- and Mesohibolites of the '' Mexican Solnhofen' ' Tepexi de Rodriguez (state of Puebla) and their biostratinomy (Lower Cretaceous). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 225, 55 - 74." type="journal article" year="2002">Seibertz & Spaeth 2002</bibRefCitation>
|
||
; Benammi
|
||
<emphasis id="B944A963FFE3020BFB6BFA166ED8F8D2" box="[1214,1271,1436,1460]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2006).
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE3020BFF12FA4B6908FB46" blockId="4.[151,1436,1112,1820]" pageId="4" pageNumber="5">
|
||
<emphasis id="B944A963FFE3020BFF12FA4B6B99F8BF" bold="true" box="[199,438,1472,1497]" pageId="4" pageNumber="5">Derivation of name.</emphasis>
|
||
The name of the fish is also written in the Náhuatl language. The species epithet “
|
||
<taxonomicName id="4C300EF2FFE3020BFA8FFA4B6FBEF8BE" box="[1370,1425,1472,1496]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="itztli">itztli</taxonomicName>
|
||
” means “obsidian” and it refers to the scale color in this fish. The complete generic and species name of this semionotid means “the fish of Tlayúa with obsidian color scales”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A26FAFFE3020BFF12F9A7681EFAEE" pageId="4" pageNumber="5" type="diagnosis">
|
||
<paragraph id="8B8F7571FFE3020BFF12F9A76EBCFA7A" blockId="4.[151,1436,1112,1820]" pageId="4" pageNumber="5">
|
||
<emphasis id="B944A963FFE3020BFF12F9A76B6EFB23" bold="true" box="[199,321,1580,1605]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
|
||
Frontals very long, about 4,5 times longer than wide, and narrowing anteriorly up to 37% of their maximal width; antorbital process absent; dermopterotic with anteroventral process articulating with the sphenotic*; operculum does not contact the dermopterotic; a series of paraethmoids anterior to the supraorbital series*; supraorbital series extends beyond the anterior rim of the orbit*; large pentagonal supraorbital partially closing the anterior rim of the orbit*; seven anterior infraorbitals*; infraorbitals at the ventral border of the orbit expanded dorsally, acuminating anterodorsally*; dorsalmost suborbital separating the preoperculum from the dermopterotic; premaxillary teeth smaller than other jaw or palatal teeth*; modified pectoral scale present.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE3020BFF12F8C768B6FA02" blockId="4.[199,665,1868,1928]" box="[199,665,1868,1892]" pageId="4" pageNumber="5">Pterygial formula: D23 T34</paragraph>
|
||
<paragraph id="8B8F7571FFE3020BFE72F8FA681EFAEE" blockId="4.[199,665,1868,1928]" box="[423,561,1904,1928]" pageId="4" pageNumber="5">P7 A18 C28</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A26FAFFE3021EFF12F8336FA0FA93" lastPageId="17" lastPageNumber="18" pageId="4" pageNumber="5" type="description">
|
||
<paragraph id="8B8F7571FFE3020AFF12F8336F6BFCC9" blockId="4.[151,1436,1976,2036]" lastBlockId="5.[151,1437,151,1943]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
<emphasis id="B944A963FFE3020BFF12F8336B76FAB7" bold="true" box="[199,345,1976,2001]" pageId="4" pageNumber="5">Description.</emphasis>
|
||
<taxonomicName id="4C300EF2FFE3020BFE8AF833681CFAB6" box="[351,563,1976,2000]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFE3020BFE8AF833681CFAB6" box="[351,563,1976,2000]" italics="true" pageId="4" pageNumber="5">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
is a fusiform fish with gently curved dorsal and ventral borders. Although the SL in IGM CT 1 and IGM CT 2, the most complete specimens available, is 42.2 and 27.6 cm respectively;
|
||
<taxonomicName id="4C300EF2FFE3020AFA90F8576B33FDD6" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="5" lastPageNumber="6" order="Amiiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFE3020AFA90F8576B33FDD6" italics="true" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
could be larger as shown by IGM CT 2, in which the SL could be around 44.6 cm. The maximal depth of the body (MBD), measured midway between the insertion of the pectoral and pelvic fins, is 0.39–0.4 of the SL. The head is triangular and slightly deeper than long. The maximal depth of the head (MDH) enters about 3.6 times in the SL and is almost 1.7–1.9 times the depth of the caudal peduncle. The length of the head (HL) is 0.3 of the SL and 5–5.5 times the longitudinal diameter of the orbit. The paired fins are sub-abdominal, the pelvic fin being located just behind the middle of the body, at 0.52 of the SL. Both unpaired fins are short and located in the posterior half of the body. The dorsal fin inserts at the 24rd raw of scales, at 0.6 of the SL, and the anal fin at the 17th raw of scales, at 0.68–0.74 of the SL. Body proportions are based on the specimens IGM 6716, 6718 and 6720.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE2020AFF12FE316B1FF951" blockId="5.[151,1437,151,1943]" pageId="5" pageNumber="6">
|
||
<emphasis id="B944A963FFE2020AFF12FE316B67FCB5" bold="true" box="[199,328,442,467]" pageId="5" pageNumber="6">Skull roof.</emphasis>
|
||
The bones of the skull roof are densely ornamented with tubercles. The ornamentation is more dense in the posterior part of the skull, on the extrascapulars, parietals, dermopterotics and posterior part of the frontals. Several tubercles in this area preserve a very thin layer of ganoin. There are two (left side of IGM 6716;
|
||
<figureCitation id="130B69F4FFE2020AFF42FDAC6AD4FF59" box="[151,251,551,575]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="5" pageNumber="6">Figs 6–7</figureCitation>
|
||
) or three (right side of IGM 6719 and 6720;
|
||
<figureCitation id="130B69F4FFE2020AFCD6FDAC6963FF59" box="[771,844,551,575]" captionStart="FIGURE 4" captionStartId="7.[151,249,938,961]" captionTargetBox="[151,1436,193,916]" captionTargetId="figure@7.[151,1436,193,917]" captionTargetPageId="7" captionText="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." httpUri="https://zenodo.org/record/205970/files/figure.png" pageId="5" pageNumber="6">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFE2020AFC8CFDAC69ABFF59" box="[857,900,551,575]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="10.[151,249,1361,1384]" captionStartId-1="11.[151,249,1832,1855]" captionTargetBox-0="[151,1436,193,1339]" captionTargetBox-1="[151,1436,841,1811]" captionTargetId-0="figure@10.[151,1436,193,1340]" captionTargetId-1="figure@11.[151,1436,841,1811]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." captionText-1="FIGURE 9. Tlayuamichin itztli gen. et sp. nov., details of the skull in IGM 6719. A, posterior part of the skull showing multiple opercular ossifications and the peculiar shape of the suboperculum in this specimen. B, detailed photograph of the dermopterotic bone; the arrow points to the peculiar anteroventral process of this bone in the new taxon. C, same photograph as in B, but in this case the arrows point to the pores of the preopercular sensory canal in the preoperculum (below), suborbital (middle) and dermopterotic (above). Scale bars equal = 10 mm." httpUri-0="https://zenodo.org/record/205975/files/figure.png" httpUri-1="https://zenodo.org/record/205976/files/figure.png" pageId="5" pageNumber="6">8–9</figureCitation>
|
||
) extrascapular bones on each side of the skull. When two extrascapulars are present, the medial extrascapulars are larger than the lateral ones, and are mediolaterally elongate, narrower medially than laterally. When three extrascapulars are present, the two most medial elements occupy an area equivalent to the single medial extrascapular described before. The lateral extrascapulars are trapezoidal, a little narrower dorsally than ventrally. The parietals are rectangular, twice as long as wide, and about a third of the length of the frontals. They form a sinuous inter-parietal suture. In the
|
||
<typeStatus id="548BCBD3FFE2020AFBB8FD516EFDFF94" box="[1133,1234,730,754]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
, the inter-parietal suture has an indentation to the left, but in IGM 6719 there is no indentation in this suture (
|
||
<figureCitation id="130B69F4FFE2020AFB79FD746EDCFE71" box="[1196,1267,767,791]" captionStart="FIGURE 4" captionStartId="7.[151,249,938,961]" captionTargetBox="[151,1436,193,916]" captionTargetId="figure@7.[151,1436,193,917]" captionTargetPageId="7" captionText="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." httpUri="https://zenodo.org/record/205970/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
|
||
). The parietals articulate with the frontals anteriorly, laterally with the dermopterotic, and posteriorly they meet the median extrascapulars only. In IGM 6719 the lateralmost extrascapular apparently articulates with the parietal, but this is due to the post-mortem ventral displacement of the parietal and dermopterotic (
|
||
<figureCitation id="130B69F4FFE2020AFC18FCE16E3DFEE4" box="[973,1042,874,898]" captionStart="FIGURE 9" captionStartId="11.[151,249,1832,1855]" captionTargetBox="[151,1436,841,1811]" captionTargetId="figure@11.[151,1436,841,1811]" captionTargetPageId="11" captionText="FIGURE 9. Tlayuamichin itztli gen. et sp. nov., details of the skull in IGM 6719. A, posterior part of the skull showing multiple opercular ossifications and the peculiar shape of the suboperculum in this specimen. B, detailed photograph of the dermopterotic bone; the arrow points to the peculiar anteroventral process of this bone in the new taxon. C, same photograph as in B, but in this case the arrows point to the pores of the preopercular sensory canal in the preoperculum (below), suborbital (middle) and dermopterotic (above). Scale bars equal = 10 mm." httpUri="https://zenodo.org/record/205976/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
|
||
). The frontals are very long, about 4,5 times longer than wide and articulate with each other through a sinuous suture. The frontals are widest at the posterodorsal corner of the orbit and the posterior border is rounded. Anteriorly, the frontal narrows gradually up to 37% of it maximal width and there is no antorbital process. The nasal bones are well preserved in the
|
||
<typeStatus id="548BCBD3FFE2020AFAEFFC5C6FB3FE89" box="[1338,1436,983,1007]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
(IGM 6716;
|
||
<figureCitation id="130B69F4FFE2020AFEF0FC716BA6F974" box="[293,393,1018,1042]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="5" pageNumber="6">Figs 6–7</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFE2020AFE40FC716B9DF974" box="[405,434,1018,1042]" captionStart="FIGURE 10" captionStartId="12.[151,249,1074,1097]" captionTargetBox="[151,1436,193,1053]" captionTargetId="figure@12.[151,1436,193,1053]" captionTargetPageId="12" captionText="FIGURE 10. Tlayuamichin itztli gen. et sp. nov., ethmoidal region of the skull in IGM 6716 (holotype) showing the paraethmoids as indicated in Figure 7 (labeled as “ pet ”). Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205977/files/figure.png" pageId="5" pageNumber="6">10</figureCitation>
|
||
) and in IGM 6719. They are delicate, longitudinally elongate bones with laterally curved anterior ends.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE2020AFF12FBC96E07F869" blockId="5.[151,1437,151,1943]" pageId="5" pageNumber="6">
|
||
The exposed portion of the dermopterotic is rectangular, about 3 times longer than wide. This bone articulates with the frontal anterodorsally, with the lateral extrascapular posteriorly, with the parietal dorsally, and with a large suborbital anteroventrally. In
|
||
<taxonomicName id="4C300EF2FFE2020AFE31FB006896F9C5" box="[484,697,1163,1187]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFE2020AFE31FB006896F9C5" box="[484,697,1163,1187]" italics="true" pageId="5" pageNumber="6">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
the dermopterotic does not contact the anterodorsal corner of the operculum, as it is the case in many other semionotiforms. The anterior portion of the ventral margin of the dermopterotic is distinctly concave and the bone is expanded in an anteroventral process that articulates medially with the sphenotic and is superficially overlapped by the dermosphenotic (
|
||
<figureCitation id="130B69F4FFE2020AFC75FB7C69CDF869" box="[928,994,1271,1295]" captionStart="FIGURE 9" captionStartId="11.[151,249,1832,1855]" captionTargetBox="[151,1436,841,1811]" captionTargetId="figure@11.[151,1436,841,1811]" captionTargetPageId="11" captionText="FIGURE 9. Tlayuamichin itztli gen. et sp. nov., details of the skull in IGM 6719. A, posterior part of the skull showing multiple opercular ossifications and the peculiar shape of the suboperculum in this specimen. B, detailed photograph of the dermopterotic bone; the arrow points to the peculiar anteroventral process of this bone in the new taxon. C, same photograph as in B, but in this case the arrows point to the pores of the preopercular sensory canal in the preoperculum (below), suborbital (middle) and dermopterotic (above). Scale bars equal = 10 mm." httpUri="https://zenodo.org/record/205976/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
|
||
A–B).
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE2020AFF12FA916870F8A4" blockId="5.[151,1437,151,1943]" pageId="5" pageNumber="6">
|
||
<emphasis id="B944A963FFE2020AFF12FA916804F855" bold="true" box="[199,555,1306,1331]" pageId="5" pageNumber="6">Braincase and parasphenoid.</emphasis>
|
||
Although no isolated braincase is preserved, some braincase structures are exposed in the
|
||
<typeStatus id="548BCBD3FFE2020AFEEAFAB46B8EF831" box="[319,417,1343,1367]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
mainly. The sphenotic forms a strong lateral process that is projected laterally and presents a dermal component, which is superficially exposed between the dermosphenotic, the last infraorbital bone, and the suborbital bones (
|
||
<figureCitation id="130B69F4FFE2020AFE88FA0C6BECF8F9" box="[349,451,1415,1439]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="5" pageNumber="6">Figs 6–7</figureCitation>
|
||
). This small tear-shape dermal component of the sphenotic is ornamented with tubercles covered with a very thin layer of ganoin.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE2020AFF12FA4469B1FAF1" blockId="5.[151,1437,151,1943]" pageId="5" pageNumber="6">
|
||
The ethmoidal and orbital regions of the braincase are partially preserved.
|
||
<taxonomicName id="4C300EF2FFE2020AFBC5FA446ECBF881" box="[1040,1252,1487,1511]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFE2020AFBC5FA446ECBF881" box="[1040,1252,1487,1511]" italics="true" pageId="5" pageNumber="6">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
peculiarly has a series of apparently endochondral bones aligned lateral to the very elongate ethmoidal region and above the parasphenoid. The series of paired bones extend from the first supraorbital to the tip of the snout dorsal to the antorbital, and these bones, six in total, show a serial correspondence with the anterior infraorbital bones (
|
||
<figureCitation id="130B69F4FFE2020AFB1FF9B16F1FFB34" box="[1226,1328,1594,1618]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="5" pageNumber="6">Figs 6–7</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFE2020AFAE9F9B16F74FB34" box="[1340,1371,1594,1618]" captionStart="FIGURE 10" captionStartId="12.[151,249,1074,1097]" captionTargetBox="[151,1436,193,1053]" captionTargetId="figure@12.[151,1436,193,1053]" captionTargetPageId="12" captionText="FIGURE 10. Tlayuamichin itztli gen. et sp. nov., ethmoidal region of the skull in IGM 6716 (holotype) showing the paraethmoids as indicated in Figure 7 (labeled as “ pet ”). Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205977/files/figure.png" pageId="5" pageNumber="6">10</figureCitation>
|
||
). The endochondral nature of these bones is indicated by the state of preservation, but it is not possible to certify this condition. Also, their actual origin is unknown and very difficult to interpret due to the lack of information. Such bones are so far unknown in other actinopterygians and, thus, comparisons are not possible. They might be ossifications originated from the ethmoidal cartilage, probably homologous with the lateral ethmoids. Since these bones are placed laterally along the elongate ethmoidal region, we propose to name them paraethmoids, indicating their topographic relationships. The paraethmoids are well preserved in the
|
||
<typeStatus id="548BCBD3FFE2020AFC5FF89969C3FA4C" box="[906,1004,1810,1834]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
(IGM 6716) only, where the paraethmoids 3 and 4 are very damaged due to the intrusion of two displaced anterior infraorbitals from the right side of the skull. In IGM 6720 (
|
||
<figureCitation id="130B69F4FFE2020AFE72F8D16BC6FA14" box="[423,489,1882,1906]" captionStart="FIGURE 8" captionStartId="10.[151,249,1361,1384]" captionTargetBox="[151,1436,193,1339]" captionTargetId="figure@10.[151,1436,193,1340]" captionTargetPageId="10" captionText="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri="https://zenodo.org/record/205975/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation>
|
||
) the most posterior of these elements is probably laterally covered by the most anterior dermal supraorbital (this supraorbital is missing in the
|
||
<typeStatus id="548BCBD3FFE2020AFCFEF8F469BDFAF1" box="[811,914,1919,1943]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF4F25F9FFE10209FF42F8826914FA58" httpUri="https://zenodo.org/record/205969/files/figure.png" pageId="6" pageNumber="7" targetBox="[151,1436,193,1780]" targetPageId="6">
|
||
<paragraph id="8B8F7571FFE10209FF42F8826914FA58" blockId="6.[151,1436,1801,1854]" pageId="6" pageNumber="7">
|
||
<emphasis id="B944A963FFE10209FF42F8826B3BFA79" bold="true" box="[151,276,1801,1824]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C300EF2FFE10209FEC8F8816BF1FA46" box="[285,478,1802,1824]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFE10209FEC8F8816BF1FA46" box="[285,478,1802,1824]" italics="true" pageId="6" pageNumber="7">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFE10209FE33F88168A5FA79" bold="true" box="[486,650,1802,1823]" pageId="6" pageNumber="7">gen. et sp. nov.</emphasis>
|
||
A, Specimen IGM 6717 preserved in right lateroventral view. B, specimen IGM 6718 preserved in right lateral view. Scale bars equal 70 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F25F9FFE00208FF42FC216FA0FEA6" box="[151,1423,938,961]" httpUri="https://zenodo.org/record/205970/files/figure.png" pageId="7" pageNumber="8" targetBox="[151,1436,193,916]" targetPageId="7">
|
||
<paragraph id="8B8F7571FFE00208FF42FC216FA0FEA6" blockId="7.[151,1423,938,961]" box="[151,1423,938,961]" pageId="7" pageNumber="8">
|
||
<emphasis id="B944A963FFE00208FF42FC216B3DFEA6" bold="true" box="[151,274,938,961]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C300EF2FFE00208FECDFC216BF8FEA6" box="[280,471,938,960]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFE00208FECDFC216BF8FEA6" box="[280,471,938,960]" italics="true" pageId="7" pageNumber="8">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFE00208FE08FC206855FEA6" bold="true" box="[477,634,939,960]" pageId="7" pageNumber="8">gen. et sp. nov.</emphasis>
|
||
Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F25F9FFE00208FF42F9EF6FB2FB1C" box="[151,1437,1636,1659]" httpUri="https://zenodo.org/record/205971/files/figure.png" pageId="7" pageNumber="8" targetBox="[151,1436,1010,1614]" targetPageId="7">
|
||
<paragraph id="8B8F7571FFE00208FF42F9EF6FB2FB1C" blockId="7.[151,1437,1636,1659]" box="[151,1437,1636,1659]" pageId="7" pageNumber="8">
|
||
<emphasis id="B944A963FFE00208FF42F9EF6B3DFB1C" bold="true" box="[151,274,1636,1659]" pageId="7" pageNumber="8">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C300EF2FFE00208FECDF9EF6BF9FB1C" box="[280,470,1636,1658]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFE00208FECDF9EF6BF9FB1C" box="[280,470,1636,1658]" italics="true" pageId="7" pageNumber="8">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFE00208FE09F9EE6857FB1C" bold="true" box="[476,632,1637,1658]" pageId="7" pageNumber="8">gen. et sp. nov.</emphasis>
|
||
Specimen IGM 6720 preserved in right lateral view. Scale bar equals = 50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F7571FFE00208FF12F92C6B85FAB9" blockId="7.[151,1436,1703,2016]" pageId="7" pageNumber="8">
|
||
Some partially preserved chondral elements within the orbit most probably represent remains of the orbitosphenoid. The orbital part of the parasphenoid is exposed in the
|
||
<typeStatus id="548BCBD3FFE00208FC8AF94069EEFB85" box="[863,961,1739,1763]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
and in IGM 6719 and 6720 (
|
||
<figureCitation id="130B69F4FFE00208FADCF9406F7CFB85" box="[1289,1363,1739,1763]" captionStart="FIGURE 4" captionStartId="7.[151,249,938,961]" captionTargetBox="[151,1436,193,916]" captionTargetId="figure@7.[151,1436,193,917]" captionTargetPageId="7" captionText="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." httpUri="https://zenodo.org/record/205970/files/figure.png" pageId="7" pageNumber="8">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFE00208FA8AF9406F40FB85" box="[1375,1391,1739,1763]" captionStart="FIGURE 8" captionStartId="10.[151,249,1361,1384]" captionTargetBox="[151,1436,193,1339]" captionTargetId="figure@10.[151,1436,193,1340]" captionTargetPageId="10" captionText="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri="https://zenodo.org/record/205975/files/figure.png" pageId="7" pageNumber="8">8</figureCitation>
|
||
). In IGM 6717 the parasphenoid is completely exposed, though poorly preserved in ventral view. The parasphenoid is edentulous and extends back throughout the area corresponding to the prootics and ends at the central portion of the basioccipital forming two posterior processes. There is a longitudinal deep median groove along the dorsal surface of the orbital section of the parasphenoid. Within the orbital region, the parasphenoid broadens posteriorly, reaching its maximal breadth at the posterior wall of the orbit, where there are two long and narrow basipterygoid processes. Dermal and possible chondral portions of the basipterygoid processes cannot be distinguished, but the processes are very large.
|
||
</paragraph>
|
||
<caption id="DF4F25F9FFEF0207FF42FB396AEAF98E" httpUri="https://zenodo.org/record/205972/files/figure.png" pageId="8" pageNumber="9" targetBox="[278,1307,179,1187]" targetPageId="8">
|
||
<paragraph id="8B8F7571FFEF0207FF42FB396AEAF98E" blockId="8.[151,1437,1202,1256]" pageId="8" pageNumber="9">
|
||
<emphasis id="B944A963FFEF0207FF42FB396B3EF9AE" bold="true" box="[151,273,1202,1225]" pageId="8" pageNumber="9">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C300EF2FFEF0207FEC2FB386BFAF9AF" box="[279,469,1203,1225]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFEF0207FEC2FB386BFAF9AF" box="[279,469,1203,1225]" italics="true" pageId="8" pageNumber="9">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFEF0207FE0EFB386859F9AE" bold="true" box="[475,630,1203,1224]" pageId="8" pageNumber="9">gen. et sp. nov.</emphasis>
|
||
Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F7571FFEF0207FF12FA826968FA58" blockId="8.[151,1437,1289,2034]" pageId="8" pageNumber="9">
|
||
<emphasis id="B944A963FFEF0207FF12FA826BFAF844" bold="true" box="[199,469,1289,1314]" pageId="8" pageNumber="9">Circumborbital series.</emphasis>
|
||
The circumorbital series, which is completely preserved in the
|
||
<typeStatus id="548BCBD3FFEF0207FB7FFA816F23F844" box="[1194,1292,1290,1314]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
(IGM 6716;
|
||
<figureCitation id="130B69F4FFEF0207FF42FAA56AD3F820" box="[151,252,1326,1350]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="8" pageNumber="9">Figs 6–7</figureCitation>
|
||
) and IGM 6720 (
|
||
<figureCitation id="130B69F4FFEF0207FE1DFAA56821F820" box="[456,526,1326,1350]" captionStart="FIGURE 8" captionStartId="10.[151,249,1361,1384]" captionTargetBox="[151,1436,193,1339]" captionTargetId="figure@10.[151,1436,193,1340]" captionTargetPageId="10" captionText="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri="https://zenodo.org/record/205975/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
||
), involves five supraorbitals, the dermosphenotic, six infraorbitals (io1––io
|
||
<quantity id="4CC8D894FFEF0207FABAFAA56FB3F820" box="[1391,1436,1326,1350]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="9" unit="in" value="6.0">6 in</quantity>
|
||
anterior to posterior direction), seven anterior infraorbitals (a.io1–a.io
|
||
<quantity id="4CC8D894FFEF0207FC64FAD969F0F80C" box="[945,991,1362,1386]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="8" pageNumber="9" unit="in" value="7.0">7 in</quantity>
|
||
anterior to posterior direction) and the antorbital. The circumborbital series is open (IGM 6716 and 6719;
|
||
<figureCitation id="130B69F4FFEF0207FC59FAFD69FBF8E8" box="[908,980,1398,1422]" captionStart="FIGURE 4" captionStartId="7.[151,249,938,961]" captionTargetBox="[151,1436,193,916]" captionTargetId="figure@7.[151,1436,193,917]" captionTargetPageId="7" captionText="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." httpUri="https://zenodo.org/record/205970/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFEF0207FC35FAFD6E22F8E8" box="[992,1037,1398,1422]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="8" pageNumber="9">6–7</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFEF0207FBCCFAFD6E18F8E8" box="[1049,1079,1398,1422]" captionStart="FIGURE 10" captionStartId="12.[151,249,1074,1097]" captionTargetBox="[151,1436,193,1053]" captionTargetId="figure@12.[151,1436,193,1053]" captionTargetPageId="12" captionText="FIGURE 10. Tlayuamichin itztli gen. et sp. nov., ethmoidal region of the skull in IGM 6716 (holotype) showing the paraethmoids as indicated in Figure 7 (labeled as “ pet ”). Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205977/files/figure.png" pageId="8" pageNumber="9">10</figureCitation>
|
||
), though it appears to be closed in IGM 6720 (
|
||
<figureCitation id="130B69F4FFEF0207FEEBFA116BADF8D4" box="[318,386,1434,1458]" captionStart="FIGURE 8" captionStartId="10.[151,249,1361,1384]" captionTargetBox="[151,1436,193,1339]" captionTargetId="figure@10.[151,1436,193,1340]" captionTargetPageId="10" captionText="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri="https://zenodo.org/record/205975/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
||
) due to the anterodorsal displacement of the bones in the anteroventral portion of the skull.
|
||
<taxonomicName id="4C300EF2FFEF0207FF42FA356B43F8B0" box="[151,364,1470,1494]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFEF0207FF42FA356B43F8B0" box="[151,364,1470,1494]" italics="true" pageId="8" pageNumber="9">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
is very peculiar in having the most anterior supraorbital element very large, placed exactly at the anterior rim of the orbit so that the bone, which has the shape of an inverted pentagon, has equally large orbital and anteorbital portions (it should be noticed that in IGM 6716 the bone is fractured right at the level of the anterior rim of the orbit, giving the appearance of two separate ossifications;
|
||
<figureCitation id="130B69F4FFEF0207FC7AF9A16E3AFB24" box="[943,1045,1578,1602]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="8" pageNumber="9">Figs 6–7</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFEF0207FBF7F9A16E6EFB24" box="[1058,1089,1578,1602]" captionStart="FIGURE 10" captionStartId="12.[151,249,1074,1097]" captionTargetBox="[151,1436,193,1053]" captionTargetId="figure@12.[151,1436,193,1053]" captionTargetPageId="12" captionText="FIGURE 10. Tlayuamichin itztli gen. et sp. nov., ethmoidal region of the skull in IGM 6716 (holotype) showing the paraethmoids as indicated in Figure 7 (labeled as “ pet ”). Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205977/files/figure.png" pageId="8" pageNumber="9">10</figureCitation>
|
||
). This pentagonal supraorbital forms a ventral wedge that faces a dorsal wedge formed by the last anterior infraorbital (a.io7) and the first infraorbital (io1) together, so that the two opposite wedges are partially closing the orbit anteriorly. In IGM 6720 there is another additional supraorbital anterior to this pentagonal bone (
|
||
<figureCitation id="130B69F4FFEF0207FCA4F91D6999FBC8" box="[881,950,1686,1710]" captionStart="FIGURE 8" captionStartId="10.[151,249,1361,1384]" captionTargetBox="[151,1436,193,1339]" captionTargetId="figure@10.[151,1436,193,1340]" captionTargetPageId="10" captionText="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri="https://zenodo.org/record/205975/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
||
). There are no traces of such an additional supraorbital in the other specimens, but it might have been present and not preserved. In any case,
|
||
<taxonomicName id="4C300EF2FFEF0207FAD7F9316AE6FB90" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFEF0207FAD7F9316AE6FB90" italics="true" pageId="8" pageNumber="9">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
is very peculiar in having a long series of supraorbital bones, which extends beyond the orbit anteriorly. The remaining supraorbitals are rectangular and relatively small bones. The supraorbital bones are ornamented with tubercles, which are covered with a very thin layer of ganoin.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFEF0207FF10F8C169F2FACC" blockId="8.[151,1437,1289,2034]" pageId="8" pageNumber="9">
|
||
<collectingCountry id="F32735E1FFEF0207FF10F8C16ACBFA04" box="[197,228,1866,1890]" name="American Samoa" pageId="8" pageNumber="9">As</collectingCountry>
|
||
usual in semionotiforms, the dermosphenotic is located at the posterodorsal corner of the orbit. This bone is rectangular, about twice as deep as long, and as long as the adjacent supraorbital (
|
||
<figureCitation id="130B69F4FFEF0207FBF7F8E56E44FAE0" box="[1058,1131,1902,1926]" captionStart="FIGURE 4" captionStartId="7.[151,249,938,961]" captionTargetBox="[151,1436,193,916]" captionTargetId="figure@7.[151,1436,193,917]" captionTargetPageId="7" captionText="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." httpUri="https://zenodo.org/record/205970/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFEF0207FBA3F8E56E8BFAE0" box="[1142,1188,1902,1926]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionStartId-2="10.[151,249,1361,1384]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetBox-2="[151,1436,193,1339]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetId-2="figure@10.[151,1436,193,1340]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-2="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" httpUri-2="https://zenodo.org/record/205975/files/figure.png" pageId="8" pageNumber="9">6–8</figureCitation>
|
||
). The dermosphenotic is also ornamented with tubercles covered with a very thin layer of ganoin.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFEF0206FF10F83D69B5FF71" blockId="8.[151,1437,1289,2034]" lastBlockId="9.[151,1437,151,535]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
The infraorbital bones are best preserved in the
|
||
<typeStatus id="548BCBD3FFEF0207FD33F83D6967FAA8" box="[742,840,1974,1998]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
(IGM 6716;
|
||
<figureCitation id="130B69F4FFEF0207FC34F83D6E68FAA8" box="[993,1095,1974,1998]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="8" pageNumber="9">Figs 6–7</figureCitation>
|
||
). Among the six infraorbitals, three form the ventral margin of the orbit (infraorbitals io1–io3), the fourth infraorbital (io4) is placed at the posteroventral corner of the orbit and the last two infraorbitals (io5–io6) form the posterior margin of the orbit. The infraorbitals three to six (io3–io6) articulate with suborbital bones posteriorly. The infraorbitals five and six (io5– io6) are relatively small, deeper than long, rectangular bones. The infraorbital four (io4) is about twice the size of the infraorbital five (io5) and smaller than the infraorbital three (io3); it has a concave orbital border and is convex posteriorly. The infraorbitals one to three (io1–io3) are irregularly triangular, acuminated anterodorsally. In the
|
||
<typeStatus id="548BCBD3FFEE0206FF42FEC76AD3FC02" box="[151,252,332,356]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
, the infraorbital three (io3) is slightly rotated clockwise and displaced dorsally because of the extrusion of the basipterygoid process of the parasphenoid. When moved back to its natural position, the infraorbital three (io3) alone forms most of the ventral margin of the orbit. The same happens in IGM 6720. The infraorbital two (io2) is the deepest bone of the whole series, followed by the infraorbital one (io1), which is only slightly shallower, and the infraorbital three (io3), which is about 20% shallower than the infraorbital two. Accordingly, the ventral margins of the infraorbitals 1–3 together describe a gently convex curve.
|
||
</paragraph>
|
||
<caption id="DF4F25F9FFEE0206FF42FA666FB9FB65" box="[151,1430,1517,1540]" httpUri="https://zenodo.org/record/205973/files/figure.png" pageId="9" pageNumber="10" targetBox="[188,1398,625,1463]" targetPageId="9">
|
||
<paragraph id="8B8F7571FFEE0206FF42FA666FB9FB65" blockId="9.[151,1430,1517,1540]" box="[151,1430,1517,1540]" pageId="9" pageNumber="10">
|
||
<emphasis id="B944A963FFEE0206FF42FA666B3DFB65" bold="true" box="[151,274,1517,1540]" pageId="9" pageNumber="10">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C300EF2FFEE0206FECDFA666BF8FB65" box="[280,471,1517,1539]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFEE0206FECDFA666BF8FB65" box="[280,471,1517,1539]" italics="true" pageId="9" pageNumber="10">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFEE0206FE08FA656855FB65" bold="true" box="[477,634,1518,1539]" pageId="9" pageNumber="10">gen. et sp. nov.</emphasis>
|
||
Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F7571FFEE0206FF10F9BB6F4DFAD6" blockId="9.[151,1436,1584,1969]" pageId="9" pageNumber="10">
|
||
The seven anterior infraorbitals are aligned ventral to the level of the parasphenoid (
|
||
<figureCitation id="130B69F4FFEE0206FBBAF9BB6EFBFB2E" box="[1135,1236,1584,1608]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="9" pageNumber="10">Figs 6–7</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFEE0206FB0AF9BB6ED1FB2E" box="[1247,1278,1584,1608]" captionStart="FIGURE 10" captionStartId="12.[151,249,1074,1097]" captionTargetBox="[151,1436,193,1053]" captionTargetId="figure@12.[151,1436,193,1053]" captionTargetPageId="12" captionText="FIGURE 10. Tlayuamichin itztli gen. et sp. nov., ethmoidal region of the skull in IGM 6716 (holotype) showing the paraethmoids as indicated in Figure 7 (labeled as “ pet ”). Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205977/files/figure.png" pageId="9" pageNumber="10">10</figureCitation>
|
||
). The anterior infraorbitals four to seven (a.io4–a.io7) are approximately rectangular, deeper than long, with their depths gradually decreasing anteriorly. The anterior infraorbital 7 is slightly acuminated dorsally, in the anterior infraorbital 6 the dorsal border is only slightly shorter than the ventral border, and the dorsal and ventral borders of the infraorbital 5 are approximately equal. The anterior infraorbital 4 is more irregular in shape, with the posterior border deeper than the anterior border and has, thus, anteroventrally inclined dorsal border. The anterior infraorbital 3 is approximately rectangular, about 1,5 times longer than deep, with the dorsal border a little longer than the ventral border. The anterior infraorbital 2 is smaller than its adjacent elements in the series. It is slightly narrower dorsally than ventrally and has a small laminar outgrowth ventral to the passage of the infraorbital sensory canal. Such an outgrowth is not present in the more posterior anterior infraorbitals. The first anterior infraorbital is somewhat boomerang-shaped and also has a well-ossified outgrowth ventral to the passage of the infraorbital sensory canal.
|
||
</paragraph>
|
||
<caption id="DF4F25F9FFED0205FF42FADA6F41F801" box="[151,1390,1361,1384]" httpUri="https://zenodo.org/record/205975/files/figure.png" pageId="10" pageNumber="11" targetBox="[151,1436,193,1339]" targetPageId="10">
|
||
<paragraph id="8B8F7571FFED0205FF42FADA6F41F801" blockId="10.[151,1390,1361,1384]" box="[151,1390,1361,1384]" pageId="10" pageNumber="11">
|
||
<emphasis id="B944A963FFED0205FF42FADA6B3DF801" bold="true" box="[151,274,1361,1384]" pageId="10" pageNumber="11">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C300EF2FFED0205FECDFADA6BF8F801" box="[280,471,1361,1383]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFED0205FECDFADA6BF8F801" box="[280,471,1361,1383]" italics="true" pageId="10" pageNumber="11">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFED0205FE08FAD96855F801" bold="true" box="[477,634,1362,1383]" pageId="10" pageNumber="11">gen. et sp. nov.</emphasis>
|
||
Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F7571FFED0205FF10FA1F6F10F8B6" blockId="10.[151,1437,1428,2028]" pageId="10" pageNumber="11">All the infraorbital and anterior infraorbital bones are also ornamented with tubercles covered with a very thin layer of ganoin, which are denser towards the ventral and posteroventral border of the circumborbital ring.</paragraph>
|
||
<paragraph id="8B8F7571FFED0205FF10FA576E1AFB06" blockId="10.[151,1437,1428,2028]" pageId="10" pageNumber="11">
|
||
The most anterior bone of the infraorbital series is usually interpreted as the antorbital. In
|
||
<taxonomicName id="4C300EF2FFED0205FB1DFA576FB3F892" box="[1224,1436,1500,1524]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFED0205FB1DFA576FB3F892" box="[1224,1436,1500,1524]" italics="true" pageId="10" pageNumber="11">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
this most anterior element resembles the small, almost tubular antorbital in “
|
||
<taxonomicName id="4C300EF2FFED0205FC3BF98B6E74FB7E" box="[1006,1115,1536,1560]" class="Actinopterygii" family="Semionotidae" genus="Lepidotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944A963FFED0205FC3BF98B6E74FB7E" box="[1006,1115,1536,1560]" italics="true" pageId="10" pageNumber="11">Lepidotes</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="4C300EF2FFED0205FBBBF98A6E9EFB7E" box="[1134,1201,1537,1560]" class="Actinopterygii" family="Semionotidae" genus="Lepidotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="minor">
|
||
<emphasis id="B944A963FFED0205FBBBF98A6E9EFB7E" box="[1134,1201,1537,1560]" italics="true" pageId="10" pageNumber="11">minor</emphasis>
|
||
</taxonomicName>
|
||
illustrated by
|
||
<bibRefCitation id="EFA10880FFED0205FA87F98A6B5AFB5A" author="Patterson" pageId="10" pageNumber="24" refString="Patterson, C. (1975) The braincase of pholidophorid and leptolepid fishes, with a review of the actinopterygian braincase. Philosophical Transactions of the Royal Society of London, 269, 275 - 579." type="journal article" year="1975">Patterson (1975: fig. 135)</bibRefCitation>
|
||
(
|
||
<figureCitation id="130B69F4FFED0205FE51F9AF6BC5FB5A" box="[388,490,1572,1596]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="10" pageNumber="11">Figs 6–7</figureCitation>
|
||
). An irregularly shaped ossification preserved anterior to the antorbital and dorsal to the anterior end of the first anterior infraorbital might represent a displaced rostral.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFED0205FF10F9E76832FA72" blockId="10.[151,1437,1428,2028]" pageId="10" pageNumber="11">
|
||
<emphasis id="B944A963FFED0205FF10F9E76BB4FBE3" bold="true" box="[197,411,1644,1669]" pageId="10" pageNumber="11">Suborbital bones.</emphasis>
|
||
There are three suborbital bones in
|
||
<taxonomicName id="4C300EF2FFED0205FC92F9E76E32FBE2" box="[839,1053,1644,1668]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFED0205FC92F9E76E32FBE2" box="[839,1053,1644,1668]" italics="true" pageId="10" pageNumber="11">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
, and they are also densely ornamented with tubercles covered with a thin layer of ganoin. They are all of similar size and aligned in a single row completely covering the area between the circumborbital bones and the preoperculum, but do not extend anteriorly, ventral to the orbit. The most dorsal suborbital is expanded posteriorly and completely separates the preoperculum from the dermopterotic (
|
||
<figureCitation id="130B69F4FFED0205FE7CF9776820FA72" box="[425,527,1788,1812]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionStartId-2="10.[151,249,1361,1384]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetBox-2="[151,1436,193,1339]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetId-2="figure@10.[151,1436,193,1340]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-2="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" httpUri-2="https://zenodo.org/record/205975/files/figure.png" pageId="10" pageNumber="11">Figs 6–8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFED0204FF10F8946F1FFE72" blockId="10.[151,1437,1428,2028]" lastBlockId="11.[151,1437,151,788]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
<emphasis id="B944A963FFED0205FF10F8946BBCFA5E" bold="true" box="[197,403,1823,1848]" pageId="10" pageNumber="11">Opercular bones.</emphasis>
|
||
The preoperculum is a crescent-shape bone, very narrow and gently curved, narrowing both dorsally and anteriorly (
|
||
<figureCitation id="130B69F4FFED0205FE7CF8CF683FFA3A" box="[425,528,1860,1884]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionStartId-2="10.[151,249,1361,1384]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetBox-2="[151,1436,193,1339]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetId-2="figure@10.[151,1436,193,1340]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-2="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" httpUri-2="https://zenodo.org/record/205975/files/figure.png" pageId="10" pageNumber="11">Figs 6–8</figureCitation>
|
||
). Anteriorly, the preoperculum extends beyond the anterior margin of the orbit. Dorsally, the preoperculum does not reach the dermopterotic; its dorsal tip being widely separated from that bone by the most dorsal suborbital. The operculum is large and subrectangular, a little narrower dorsally than ventrally. It is twice as deep as long in the
|
||
<typeStatus id="548BCBD3FFED0205FE21F83B6879FAAE" box="[500,598,1968,1992]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
and in IGM 6720 (
|
||
<figureCitation id="130B69F4FFED0205FCE7F83B69B7FAAE" box="[818,920,1968,1992]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionStartId-2="10.[151,249,1361,1384]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetBox-2="[151,1436,193,1339]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetId-2="figure@10.[151,1436,193,1340]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-2="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" httpUri-2="https://zenodo.org/record/205975/files/figure.png" pageId="10" pageNumber="11">Figs 6–8</figureCitation>
|
||
), but only 1,6 times deeper than long in IGM 6717 (
|
||
<figureCitation id="130B69F4FFED0205FF0AF85F6B0DFA8A" box="[223,290,2004,2028]" captionStart="FIGURE 3" captionStartId="6.[151,249,1801,1824]" captionTargetBox="[151,1436,193,1780]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tlayuamichin itztli gen. et sp. nov. A, Specimen IGM 6717 preserved in right lateroventral view. B, specimen IGM 6718 preserved in right lateral view. Scale bars equal 70 mm." httpUri="https://zenodo.org/record/205969/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
|
||
) and, thus, the shape of this bone is somewhat variable within the species. There are three irregularly shaped opercular ossifications in IGM 6719: a large dorsal operculum, an even larger ventral operculum, and a small rounded ossification at the anteroventral end of the ventral operculum (
|
||
<figureCitation id="130B69F4FFEC0204FBF1FF376E43FDB2" box="[1060,1132,188,212]" captionStart="FIGURE 9" captionStartId="11.[151,249,1832,1855]" captionTargetBox="[151,1436,841,1811]" captionTargetId="figure@11.[151,1436,841,1811]" captionTargetPageId="11" captionText="FIGURE 9. Tlayuamichin itztli gen. et sp. nov., details of the skull in IGM 6719. A, posterior part of the skull showing multiple opercular ossifications and the peculiar shape of the suboperculum in this specimen. B, detailed photograph of the dermopterotic bone; the arrow points to the peculiar anteroventral process of this bone in the new taxon. C, same photograph as in B, but in this case the arrows point to the pores of the preopercular sensory canal in the preoperculum (below), suborbital (middle) and dermopterotic (above). Scale bars equal = 10 mm." httpUri="https://zenodo.org/record/205976/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
|
||
). The three bones occupy together an area about 1,5 times deeper than long, but the dorsal operculum is about 0,6 times narrower than the ventral operculum. The suboperculum has a very high ascending process, which is about 70% of the maximal length of the bone (completely exposed only in the
|
||
<typeStatus id="548BCBD3FFEC0204FD30FEAC6964FC59" box="[741,843,295,319]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
;
|
||
<figureCitation id="130B69F4FFEC0204FC82FEAC6991FC59" box="[855,958,295,319]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="11" pageNumber="12">Figs 6–7</figureCitation>
|
||
). The ascending process forms an interior angle of about 90° with the dorsal border of the suboperculum. In IGM 6716, 6717 and 6720 the suboperculum is deepest anteriorly, where it reaches about 30% of the depth of the operculum, and tapers posteriorly with curved dorsal and ventral borders (
|
||
<figureCitation id="130B69F4FFEC0204FE03FE1F680EFCCA" box="[470,545,404,428]" captionStart="FIGURE 3" captionStartId="6.[151,249,1801,1824]" captionTargetBox="[151,1436,193,1780]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tlayuamichin itztli gen. et sp. nov. A, Specimen IGM 6717 preserved in right lateroventral view. B, specimen IGM 6718 preserved in right lateral view. Scale bars equal 70 mm." httpUri="https://zenodo.org/record/205969/files/figure.png" pageId="11" pageNumber="12">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFEC0204FDE4FE1F6871FCCA" box="[561,606,404,428]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionStartId-2="10.[151,249,1361,1384]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetBox-2="[151,1436,193,1339]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetId-2="figure@10.[151,1436,193,1340]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-2="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" httpUri-2="https://zenodo.org/record/205975/files/figure.png" pageId="11" pageNumber="12">6–8</figureCitation>
|
||
). In IGM 6719 the suboperculum is almost equally deep throughout its length and forms a sinuous articulation with the ventral operculum (
|
||
<figureCitation id="130B69F4FFEC0204FC46FE3C69F8FCA9" box="[915,983,439,463]" captionStart="FIGURE 9" captionStartId="11.[151,249,1832,1855]" captionTargetBox="[151,1436,841,1811]" captionTargetId="figure@11.[151,1436,841,1811]" captionTargetPageId="11" captionText="FIGURE 9. Tlayuamichin itztli gen. et sp. nov., details of the skull in IGM 6719. A, posterior part of the skull showing multiple opercular ossifications and the peculiar shape of the suboperculum in this specimen. B, detailed photograph of the dermopterotic bone; the arrow points to the peculiar anteroventral process of this bone in the new taxon. C, same photograph as in B, but in this case the arrows point to the pores of the preopercular sensory canal in the preoperculum (below), suborbital (middle) and dermopterotic (above). Scale bars equal = 10 mm." httpUri="https://zenodo.org/record/205976/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
|
||
). The ascending process of the suboperculum in this fish is hidden by the somewhat displaced preoperculum. Anteriorly, the suboperculum articulates with the interoperculum through an almost vertical suture. The interoperculum extends anteriorly up to the level of the anterior end of the preoperculum, thus extending medioventral to the whole ventral margin of that bone. The ventral border of the interoperculum is gently convex. All the opercular bones are ornamented with numerous tubercles covered with ganoin, except for the preoperculum where the tubercles are only very few and scattered. It is worthy to note that the peculiarities of the opercular bones in IGM 6719 described above are considered as abnormalities. IGM 6719 presents multiple ossification of the operculum and probably also de preoperculum and an abnormal shape of the suboperculum. Although it is important to report such cases, in the lack of ontogenetic information it is impossible to know the causes or processes that might led to such deviations in this fish.
|
||
</paragraph>
|
||
<caption id="DF4F25F9FFEC0204FF42F8A368C3FADF" httpUri="https://zenodo.org/record/205976/files/figure.png" pageId="11" pageNumber="12" targetBox="[151,1436,841,1811]" targetPageId="11">
|
||
<paragraph id="8B8F7571FFEC0204FF42F8A368C3FADF" blockId="11.[151,1437,1832,1977]" pageId="11" pageNumber="12">
|
||
<emphasis id="B944A963FFEC0204FF42F8A36B3DFA58" bold="true" box="[151,274,1832,1855]" pageId="11" pageNumber="12">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C300EF2FFEC0204FECCF8A26BF7FA59" box="[281,472,1833,1855]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFEC0204FECCF8A26BF7FA59" box="[281,472,1833,1855]" italics="true" pageId="11" pageNumber="12">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFEC0204FE0BF8A26854FA58" bold="true" box="[478,635,1833,1854]" pageId="11" pageNumber="12">gen. et sp. nov.</emphasis>
|
||
, details of the skull in IGM 6719. A, posterior part of the skull showing multiple opercular ossifications and the peculiar shape of the suboperculum in this specimen. B, detailed photograph of the dermopterotic bone; the arrow points to the peculiar anteroventral process of this bone in the new taxon. C, same photograph as in B, but in this case the arrows point to the pores of the preopercular sensory canal in the preoperculum (below), suborbital (middle) and dermopterotic (above). Scale bars equal = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F25F9FFEB0203FF42FBB969B4F901" httpUri="https://zenodo.org/record/205977/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1436,193,1053]" targetPageId="12">
|
||
<paragraph id="8B8F7571FFEB0203FF42FBB969B4F901" blockId="12.[151,1436,1074,1128]" pageId="12" pageNumber="13">
|
||
<emphasis id="B944A963FFEB0203FF42FBB96B0FF92E" bold="true" box="[151,288,1074,1097]" pageId="12" pageNumber="13">FIGURE 10.</emphasis>
|
||
<taxonomicName id="4C300EF2FFEB0203FEF3FBB86BC9F92F" box="[294,486,1075,1097]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFEB0203FEF3FBB86BC9F92F" box="[294,486,1075,1097]" italics="true" pageId="12" pageNumber="13">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFEB0203FE38FBB868A3F92E" bold="true" box="[493,652,1075,1096]" pageId="12" pageNumber="13">gen. et sp. nov.</emphasis>
|
||
, ethmoidal region of the skull in IGM 6716 (holotype) showing the paraethmoids as indicated in Figure 7 (labeled as “pet”). Scale bar equals = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F25F9FFEB0203FF42F8196916FAAE" httpUri="https://zenodo.org/record/205978/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1431,1181,1915]" targetPageId="12">
|
||
<paragraph id="8B8F7571FFEB0203FF42F8196916FAAE" blockId="12.[151,1436,1938,1992]" pageId="12" pageNumber="13">
|
||
<emphasis id="B944A963FFEB0203FF42F8196B31FACE" bold="true" box="[151,286,1938,1961]" pageId="12" pageNumber="13">FIGURE 11.</emphasis>
|
||
<taxonomicName id="4C300EF2FFEB0203FEF1F8186BCDFACF" box="[292,482,1939,1961]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFEB0203FEF1F8186BCDFACF" box="[292,482,1939,1961]" italics="true" pageId="12" pageNumber="13">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFEB0203FE3DF81868AAFACE" bold="true" box="[488,645,1939,1960]" pageId="12" pageNumber="13">gen. et sp. nov.</emphasis>
|
||
, jaws in IGM 6719. A, photograph; B, line drawing. Note that the retroarticular bone is a little displaced posteriorly. Scale bar equals = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F7571FFEA0202FF12FF136E86FE3A" blockId="13.[151,1437,151,1976]" pageId="13" pageNumber="14">
|
||
<emphasis id="B944A963FFEA0202FF12FF136B26FDD6" bold="true" box="[199,265,152,176]" pageId="13" pageNumber="14">Jaws.</emphasis>
|
||
The jaws are best preserved in IGM 6719 (
|
||
<figureCitation id="130B69F4FFEA0202FCD1FF1C6977FDC9" box="[772,856,151,175]" captionStart="FIGURE 11" captionStartId="12.[151,249,1938,1961]" captionTargetBox="[151,1431,1181,1915]" captionTargetId="figure@12.[151,1436,1177,1917]" captionTargetPageId="12" captionText="FIGURE 11. Tlayuamichin itztli gen. et sp. nov., jaws in IGM 6719. A, photograph; B, line drawing. Note that the retroarticular bone is a little displaced posteriorly. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205978/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
|
||
), but they are also relatively well preserved in the
|
||
<typeStatus id="548BCBD3FFEA0202FF42FF376AD6FDB2" box="[151,249,188,212]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
and IGM 6717 (
|
||
<figureCitation id="130B69F4FFEA0202FE66FF376BD4FDB2" box="[435,507,188,212]" captionStart="FIGURE 3" captionStartId="6.[151,249,1801,1824]" captionTargetBox="[151,1436,193,1780]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tlayuamichin itztli gen. et sp. nov. A, Specimen IGM 6717 preserved in right lateroventral view. B, specimen IGM 6718 preserved in right lateral view. Scale bars equal 70 mm." httpUri="https://zenodo.org/record/205969/files/figure.png" pageId="13" pageNumber="14">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFEA0202FDD2FF37681BFDB2" box="[519,564,188,212]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="13" pageNumber="14">6–7</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFEA0202FDEAFF376871FDB2" box="[575,606,188,212]" captionStart="FIGURE 12" captionStartId="14.[153,251,1156,1179]" captionTargetBox="[155,1436,177,1140]" captionTargetId="figure@14.[155,1440,177,1141]" captionTargetPageId="14" captionText="FIGURE 12. Tlayuamichin itztli gen. et sp. nov., detailed photograph of the anterior part of the skull preserved in ventral view in IGM 6717. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205979/files/figure.png" pageId="13" pageNumber="14">12</figureCitation>
|
||
). The lower jaw joints the quadrate at the level of the anterior orbital margin. It is a triangular structure, about 2.5 times longer than deep. The posterior and dorsal borders of the lower jaw are convex, the ventral border bents ventrally at the alveolar portion, and the alveolar border is straight, inclined anteriorly. The dentary symphysis is straight, vertical, and is contained about 2.5 times in the maximum depth of the coronoid process and more than 5 times in the total length of the lower jaw. The lower jaw is only exposed in lateral view in the available specimens, and the dentary, angular, surangular and retroarticular bones are recognized. The dentary is large and elongate, with a short posteroventral process, which is only about 15 % of the total length of the lower jaw. The posterior border of the dentary, which articulates with the angular, forms an obtuse angle with the posteroventral process. Small tubercles ornament the external lateral surface of the anterior portion of the dentary below the alveolar border. There is a single row of dentary teeth, which have short pedicles and rounded crowns. Of the right side of the skull, two coronoid bones, anterior and posterior, are exposed in IGM 6716 (
|
||
<figureCitation id="130B69F4FFEA0202FF4AFDCC6B29FF39" box="[159,262,583,607]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="13" pageNumber="14">Figs 6–7</figureCitation>
|
||
). Only the posterior of these coronoid bones is exposed on the left side of this specimen. The coronoids bear only one raw of teeth, which are identical to the marginal teeth of the dentary. The angular articulates with the dentary between the coronoid process and the posteroventral process, with the retroarticular posteroventrally, and with the surangular anterodorsally (
|
||
<figureCitation id="130B69F4FFEA0202FDFEFD3F6851FFAA" box="[555,638,692,716]" captionStart="FIGURE 11" captionStartId="12.[151,249,1938,1961]" captionTargetBox="[151,1431,1181,1915]" captionTargetId="figure@12.[151,1436,1177,1917]" captionTargetPageId="12" captionText="FIGURE 11. Tlayuamichin itztli gen. et sp. nov., jaws in IGM 6719. A, photograph; B, line drawing. Note that the retroarticular bone is a little displaced posteriorly. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205978/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
|
||
). The angular is excavated for the articulation of the quadrate posteriorly. This articular facet is concave and well distinct, but not deeply excavated.
|
||
<collectingCountry id="F32735E1FFEA0202FBFBFD536E62FF96" box="[1070,1101,728,752]" name="American Samoa" pageId="13" pageNumber="14">As</collectingCountry>
|
||
usual in semionotiforms, the surangular places on top of the lower jaw articulating dorsal to the angular and dentary, and it is dorsally convex. The retroarticular is relatively large and well ossified, forming the posteroventral end of the lower jaw. It articulates with the posteroventral process of the dentary anteriorly, and with the angular anterodorsally.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFEA0202FF10FCEC69B3F81E" blockId="13.[151,1437,151,1976]" pageId="13" pageNumber="14">
|
||
The upper jaw is also shown in IGM 6716, 6717 and 6719 (
|
||
<figureCitation id="130B69F4FFEA0202FCB8FCEC69FCFE19" box="[877,979,871,895]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="13" pageNumber="14">Figs 6–7</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFEA0202FC35FCEC69D2FE19" box="[992,1021,871,895]" captionStart="FIGURE 11" captionStartId="12.[151,249,1938,1961]" captionTargetBox="[151,1431,1181,1915]" captionTargetId="figure@12.[151,1436,1177,1917]" captionTargetPageId="12" captionText="FIGURE 11. Tlayuamichin itztli gen. et sp. nov., jaws in IGM 6719. A, photograph; B, line drawing. Note that the retroarticular bone is a little displaced posteriorly. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205978/files/figure.png" pageId="13" pageNumber="14">11</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFEA0202FBDFFCEC6E06FE19" box="[1034,1065,871,895]" captionStart="FIGURE 12" captionStartId="14.[153,251,1156,1179]" captionTargetBox="[155,1436,177,1140]" captionTargetId="figure@14.[155,1440,177,1141]" captionTargetPageId="14" captionText="FIGURE 12. Tlayuamichin itztli gen. et sp. nov., detailed photograph of the anterior part of the skull preserved in ventral view in IGM 6717. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205979/files/figure.png" pageId="13" pageNumber="14">12</figureCitation>
|
||
). The premaxillae are badly preserved; in IGM 6716, only the nasal processes of both premaxillae are exposed. In IGM 6717 the two premaxillae are preserved in ventral view (
|
||
<figureCitation id="130B69F4FFEA0202FE38FC24686FFEA1" box="[493,576,943,967]" captionStart="FIGURE 12" captionStartId="14.[153,251,1156,1179]" captionTargetBox="[155,1436,177,1140]" captionTargetId="figure@14.[155,1440,177,1141]" captionTargetPageId="14" captionText="FIGURE 12. Tlayuamichin itztli gen. et sp. nov., detailed photograph of the anterior part of the skull preserved in ventral view in IGM 6717. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205979/files/figure.png" pageId="13" pageNumber="14">Fig. 12</figureCitation>
|
||
). The alveolar portion of the premaxilla is narrow. The exact number of premaxillary teeth is unknown; three alveoli are preserved in the right premaxilla and four in the left premaxilla, but in both cases the bone is broken and more alveoli might have been present. According to the size of the alveoli, the premaxillary teeth are notably smaller than the vomerine or dermopalatine teeth and even fairly smaller than the dentary teeth. Therefore, the premaxillary teeth are the smallest in the jaws and palate. The maxilla is best preserved in IGM 6719 and is an edentulous flat bone. The articular process is strong, rod-like and long (broken in IGM 6719,
|
||
<figureCitation id="130B69F4FFEA0202FECCFB036B45F9C6" box="[281,362,1160,1184]" captionStart="FIGURE 11" captionStartId="12.[151,249,1938,1961]" captionTargetBox="[151,1431,1181,1915]" captionTargetId="figure@12.[151,1436,1177,1917]" captionTargetPageId="12" captionText="FIGURE 11. Tlayuamichin itztli gen. et sp. nov., jaws in IGM 6719. A, photograph; B, line drawing. Note that the retroarticular bone is a little displaced posteriorly. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205978/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
|
||
, but complete in IGM 6717,
|
||
<figureCitation id="130B69F4FFEA0202FD79FB0368D2F9C6" box="[684,765,1160,1184]" captionStart="FIGURE 12" captionStartId="14.[153,251,1156,1179]" captionTargetBox="[155,1436,177,1140]" captionTargetId="figure@14.[155,1440,177,1141]" captionTargetPageId="14" captionText="FIGURE 12. Tlayuamichin itztli gen. et sp. nov., detailed photograph of the anterior part of the skull preserved in ventral view in IGM 6717. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205979/files/figure.png" pageId="13" pageNumber="14">Fig. 12</figureCitation>
|
||
). Immediately posterior to this process, the maxilla expands abruptly dorsally forming a subrectangular plate with rounded corners, which is about 1.2 times longer than deep (
|
||
<figureCitation id="130B69F4FFEA0202FF4AFB5B6ADDF98E" box="[159,242,1232,1256]" captionStart="FIGURE 11" captionStartId="12.[151,249,1938,1961]" captionTargetBox="[151,1431,1181,1915]" captionTargetId="figure@12.[151,1436,1177,1917]" captionTargetPageId="12" captionText="FIGURE 11. Tlayuamichin itztli gen. et sp. nov., jaws in IGM 6719. A, photograph; B, line drawing. Note that the retroarticular bone is a little displaced posteriorly. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205978/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
|
||
). The posterior border of the maxilla is almost well preserved in IGM 6719 only. However, the border is broken and, thus, it was reconstructed in
|
||
<figureCitation id="130B69F4FFEA0202FDB4FB7F68E4F86A" box="[609,715,1268,1292]" captionStart="FIGURE 11" captionStartId="12.[151,249,1938,1961]" captionTargetBox="[151,1431,1181,1915]" captionTargetId="figure@12.[151,1436,1177,1917]" captionTargetPageId="12" captionText="FIGURE 11. Tlayuamichin itztli gen. et sp. nov., jaws in IGM 6719. A, photograph; B, line drawing. Note that the retroarticular bone is a little displaced posteriorly. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205978/files/figure.png" pageId="13" pageNumber="14">Figure 11</figureCitation>
|
||
B according to the imprint left on the dentary. The posterior bor- der of the maxilla is therefore almost straight and immediately below the broken piece there is a sharp indentation close to the posteroventral corner of the bone, which is not due to any kind of fracture or brake. A long and slender supramaxilla lies along the whole dorsal border of the maxillar plate.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFEA0202FF10FA0F6863FADE" blockId="13.[151,1437,151,1976]" pageId="13" pageNumber="14">
|
||
<emphasis id="B944A963FFEA0202FF10FA0F6830F8FA" bold="true" box="[197,543,1411,1436]" pageId="13" pageNumber="14">Palatoquadrate ossifications.</emphasis>
|
||
In
|
||
<taxonomicName id="4C300EF2FFEA0202FD9CFA0F6932F8FA" box="[585,797,1412,1436]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFEA0202FD9CFA0F6932F8FA" box="[585,797,1412,1436]" italics="true" pageId="13" pageNumber="14">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
the cheek is naked and the palatoquadrate ossifications are best exposed and well preserved in the
|
||
<typeStatus id="548BCBD3FFEA0202FDACFA2368F4F8A6" box="[633,731,1448,1472]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
(IGM 6716), which shows large part of the quadrate, ectopterygoid, metapterygoid, and only some parts of the entopterygoid (
|
||
<figureCitation id="130B69F4FFEA0202FCBCFA4769E0F882" box="[873,975,1484,1508]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="13" pageNumber="14">Figs 6–7</figureCitation>
|
||
). The quadrate is fan shaped with convex dorsal border and the articular process directed ventrally. The articular process of the quadrate is small and short. Only the most anterior portion of the quadratojugal is visible in IGM 6716 attached to the ventral border of the quadrate and far back far from the articular facet for the lower jaw. The ectopterygoid articulates with the whole anterior border of the quadrate; its shape is crescent, with gently convex dorsal and concave ventral borders, and it is about two times longer than the quadrate. The anterior portion of the ectopterygoid is broken in the
|
||
<typeStatus id="548BCBD3FFEA0202FADFF90B6F43FBFE" box="[1290,1388,1664,1688]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
and no teeth are preserved in this area. The two dermopalatines are well exposed in IGM 6717 (
|
||
<figureCitation id="130B69F4FFEA0202FB42F92F6EC4FBDA" box="[1175,1259,1700,1724]" captionStart="FIGURE 12" captionStartId="14.[153,251,1156,1179]" captionTargetBox="[155,1436,177,1140]" captionTargetId="figure@14.[155,1440,177,1141]" captionTargetPageId="14" captionText="FIGURE 12. Tlayuamichin itztli gen. et sp. nov., detailed photograph of the anterior part of the skull preserved in ventral view in IGM 6717. Scale bar equals = 10 mm." httpUri="https://zenodo.org/record/205979/files/figure.png" pageId="13" pageNumber="14">Fig. 12</figureCitation>
|
||
). The bones are broad, rounded, and bear 7–9 teeth irregularly ordered. IGM 6717 also shows the paired vomers, which are still articulated. Each vomer bears 4–5 irregularly arranged teeth. These palatine and vomerine teeth are similar to those on the jaws, also with relatively short pedicles and rounded crowns, though somewhat larger than them. The entopterygoid is also exposed in IGM 6717 and it is completely edentulous. Only the posterior portion of the entopterygoid is exposed in IGM 6716 suturing with the ectopterygoid and metapterygoid, but not with the quadrate (
|
||
<figureCitation id="130B69F4FFEA0202FA9DF8D36A88FAF2" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="13" pageNumber="14">Figs 6– 7</figureCitation>
|
||
). The metapterygoid is very broad, meeting the dorsal border of the entopterygoid anteriorly and the ectopterygoid and quadrate ventrally (
|
||
<figureCitation id="130B69F4FFEA0202FE0CF82B6810FADE" box="[473,575,1952,1976]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="13" pageNumber="14">Figs 6–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF4F25F9FFE90201FF4CFB0F681AF9DF" httpUri="https://zenodo.org/record/205979/files/figure.png" pageId="14" pageNumber="15" targetBox="[155,1436,177,1140]" targetPageId="14">
|
||
<paragraph id="8B8F7571FFE90201FF4CFB0F681AF9DF" blockId="14.[151,1437,1156,2036]" pageId="14" pageNumber="15">
|
||
<emphasis id="B944A963FFE90201FF4CFB0F6B0FF9FF" bold="true" box="[153,288,1156,1179]" pageId="14" pageNumber="15">FIGURE 12.</emphasis>
|
||
<taxonomicName id="4C300EF2FFE90201FEF3FB0F6BCAF9FC" box="[294,485,1156,1178]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFE90201FEF3FB0F6BCAF9FC" box="[294,485,1156,1178]" italics="true" pageId="14" pageNumber="15">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFE90201FE3EFB0E68A8F9FC" bold="true" box="[491,647,1157,1178]" pageId="14" pageNumber="15">gen. et sp. nov.</emphasis>
|
||
, detailed photograph of the anterior part of the skull preserved in ventral view in IGM 6717. Scale bar equals = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F7571FFE90201FF10FB5E6858FB61" blockId="14.[151,1437,1156,2036]" pageId="14" pageNumber="15">
|
||
<emphasis id="B944A963FFE90201FF10FB5E6854F988" bold="true" box="[197,635,1237,1262]" pageId="14" pageNumber="15">Hyoid arch and branchiostegal rays.</emphasis>
|
||
The hyoid arch is partially shown in IGM 6716, 6717, and 6719 (
|
||
<figureCitation id="130B69F4FFE90201FABFFB5D6A8BF877" captionStart="FIGURE 3" captionStartId="6.[151,249,1801,1824]" captionTargetBox="[151,1436,193,1780]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tlayuamichin itztli gen. et sp. nov. A, Specimen IGM 6717 preserved in right lateroventral view. B, specimen IGM 6718 preserved in right lateral view. Scale bars equal 70 mm." httpUri="https://zenodo.org/record/205969/files/figure.png" pageId="14" pageNumber="15">Figs 3</figureCitation>
|
||
A, 4, 6–7, 12). The hyomandibula is not shown in any of the available specimens. The anterior ceratohyal is relatively large and slightly waisted. Its posterior border is about two times deeper the anterior one. The constriction or waist divides the anterior ceratohyal in two portions, the posterior portion being twice as long as the anterior portion. The longitudinal axis and the anterior and posterior borders of the anterior ceratohyal are massively ossified, whereas the rest of the bone is laminar. The posterior ceratohyal is partially shown in the specimens mentioned above. The anterior and posterior ceratohyals articulate through cartilage and a superficial medial irregular suture. No hypohyal or basihyal is preserved, not even in IGM 6717, which preserves the skull in ventral view and, thus, these bones probably remain cartilaginous.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE90201FF10F9996F4CFBDE" blockId="14.[151,1437,1156,2036]" pageId="14" pageNumber="15">
|
||
The branchiostegal rays are preserved partially articulated in IGM 6716 and 6719 (
|
||
<figureCitation id="130B69F4FFE90201FBADF9996EECFB4C" box="[1144,1219,1554,1578]" captionStart="FIGURE 4" captionStartId="7.[151,249,938,961]" captionTargetBox="[151,1436,193,916]" captionTargetId="figure@7.[151,1436,193,917]" captionTargetPageId="7" captionText="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." httpUri="https://zenodo.org/record/205970/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFE90201FB05F9996ED0FB4C" box="[1232,1279,1554,1578]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="14" pageNumber="15">6–7</figureCitation>
|
||
). Considering the two specimens, the following arrangement is interpreted: five slender branchiostegal rays articulate with the anterior ceratohyal, and two larger, plate-like branchiostegals articulate with the posterior ceratohyal. The most posterior branchiostegal ray is overlapped by the suboperculum and has approximately the same length as the ventral margin of that bone, and about three times the length of the most anterior and smallest branchiostegal ray.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE90201FF12F9496B5EFA0E" blockId="14.[151,1437,1156,2036]" pageId="14" pageNumber="15">
|
||
<emphasis id="B944A963FFE90201FF12F949683FFBBD" bold="true" box="[199,528,1730,1755]" pageId="14" pageNumber="15">Sensory canals of the head.</emphasis>
|
||
The trajectories of the cephalic sensory canals are best preserved in IGM 6716 and 6719 (
|
||
<figureCitation id="130B69F4FFE90201FEC5F96D6B74FB98" box="[272,347,1766,1790]" captionStart="FIGURE 4" captionStartId="7.[151,249,938,961]" captionTargetBox="[151,1436,193,916]" captionTargetId="figure@7.[151,1436,193,917]" captionTargetPageId="7" captionText="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." httpUri="https://zenodo.org/record/205970/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFE90201FEBDF96D6BBAFB98" box="[360,405,1766,1790]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="14" pageNumber="15">6–7</figureCitation>
|
||
). The supraorbital sensory canal is represented by a groove that is perforated from time to time, throughout the central portion of the nasals, the lateral portion of the frontals and the central portion of the anterior halves of the parietals. In the parietals, the anterior, middle and posterior pit lines are also represented by perforated grooves.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE90201FF42F8F96BBEFA92" blockId="14.[151,1437,1156,2036]" pageId="14" pageNumber="15">The temporal sensory canal is represented by some relatively large pores on the anterior and ventral margins of the dermopterotic, and on the lateral margin of the most lateral extrascapular. In the extrascapulars, the supraoccipital commissure is also represented by pores, but it is not possible to know whether it was continuous from one to the other side of the skull.</paragraph>
|
||
<paragraph id="8B8F7571FFE80200FF12FF1C6B09FF69" blockId="15.[151,1437,151,2017]" pageId="15" pageNumber="16">
|
||
The supraorbital bones are all pierced by foramina that are interpreted as the orbital sensory canal as described in López-Arbarello and Sferco (in press). The orbital canal meets the temporal and infraorbital canals in the dermosphenotic. The trajectory of the infraorbital sensory canal is represented by grooves in several infraorbital bones. In the small infraorbital bones placed at the posterior border of the orbit, the sensory canal probably traversed their thickened anterior margin. More anteriorly, the grooves representing the canal are shown in the ventral half of the infraorbital bones. From the posteroventral corner of the orbit forwards, the grooves are gradually closer to the ventral border of these bones. Additionally, a series of relatively large pores at the ventral margin of the infraorbital bones most probably represent the exits of branches of the infraorbital sensory canal. In the antorbital the infraorbital canal is only represented by two pores placed at its ventral margin.
|
||
<collectingCountry id="F32735E1FFE80200FC0AFE3969D1FCAC" box="[991,1022,434,458]" name="American Samoa" pageId="15" pageNumber="16">As</collectingCountry>
|
||
mentioned above, a small bone traversed by a groove and placed anterior to the antorbital probably represents a rostral traversed by the ethmoidal commissure.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE80200FF12FD906900FE60" blockId="15.[151,1437,151,2017]" pageId="15" pageNumber="16">
|
||
The preopercular sensory canal apparently traversed the thickened anterior margin of the preopercular bone. At least 8 pores distributed throughout the central portion of the preoperculum in IGM 6716 indicate the exits of branches from this sensory canal. In the most dorsal suborbital there is a pore at the ventral margin, placed directly opposite to the thickened anterior margin of the preoperculum, and followed by a superficial slight groove that ends opposite to a pore at the ventral margin of the dermopterotic (
|
||
<figureCitation id="130B69F4FFE80200FC96FD2369AAFFA6" box="[835,901,680,704]" captionStart="FIGURE 9" captionStartId="11.[151,249,1832,1855]" captionTargetBox="[151,1436,841,1811]" captionTargetId="figure@11.[151,1436,841,1811]" captionTargetPageId="11" captionText="FIGURE 9. Tlayuamichin itztli gen. et sp. nov., details of the skull in IGM 6719. A, posterior part of the skull showing multiple opercular ossifications and the peculiar shape of the suboperculum in this specimen. B, detailed photograph of the dermopterotic bone; the arrow points to the peculiar anteroventral process of this bone in the new taxon. C, same photograph as in B, but in this case the arrows point to the pores of the preopercular sensory canal in the preoperculum (below), suborbital (middle) and dermopterotic (above). Scale bars equal = 10 mm." httpUri="https://zenodo.org/record/205976/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
|
||
C). These structures indicate that the preopercular sensory canal traversed the posterior portion of the most dorsal suborbital, a bone that, as explained above, completely separates the preoperculum from the dermopterotic.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE80200FF12FC9A6F1AFEF5" blockId="15.[151,1437,151,2017]" pageId="15" pageNumber="16">Anteriorly, the preopercular sensory canal continues as the mandibular sensory canal through the angular and the dentary. In these bones, the mandibular canal is represented by several fenestrae and a groove running close to the ventral border of the lower jaw. Additional pores in one or the other specimen are not constantly present and might represent branches of the main canal or might have a different origin (e.g. innervation or nutrition).</paragraph>
|
||
<paragraph id="8B8F7571FFE80200FF12FC156BEDF9CA" blockId="15.[151,1437,151,2017]" pageId="15" pageNumber="16">
|
||
<emphasis id="B944A963FFE80200FF12FC156B51FED1" bold="true" box="[199,382,926,951]" pageId="15" pageNumber="16">Pectoral girdle.</emphasis>
|
||
The dermal bones of the pectoral girdle are best preserved in the
|
||
<typeStatus id="548BCBD3FFE80200FB80FC156E98FED0" box="[1109,1207,926,950]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
(IGM 6716;
|
||
<figureCitation id="130B69F4FFE80200FA9CFC156A88FEBF" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="15" pageNumber="16">Figs 6– 7</figureCitation>
|
||
), in which the left girdle is complete and articulated, including a posttemporal bone, a supracleithrum, a cleithrum, probably a serrated organ and four postcleithra.
|
||
<collectingCountry id="F32735E1FFE80200FC97FC6E694EFE9B" box="[834,865,997,1021]" name="American Samoa" pageId="15" pageNumber="16">As</collectingCountry>
|
||
usual in semionotiforms the posttemporal has an approximately triangular outline, narrowing dorsally. The bone is densely ornamented with ridges and/or tubercles covered with a thin layer of ganoin, and ends dorsally at the level of the dorsal border of the median extrascapular bone when three extrascapular bones are present. The anteroventral process of the posttemporal is short and broad, laterally compressed. The posteroventral border of the posttemporal is convex and articulates with a concave surface in the supracleithrum.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE80200FF12FB336BF9F88F" blockId="15.[151,1437,151,2017]" pageId="15" pageNumber="16">
|
||
The supracleithrum is dorsoventrally elongate and the exposed portion is relatively narrow. The supracleithrum is overlapped by the posttemporal dorsally and the operculum anteriorly, and it ventrally overlaps the cleithrum and the most dorsal postcleithrum. The posterior border of the supracleithrum is straight through most of its length, until meeting the most dorsal postcleithrum, where it bents anteroventrally, so that the bone tapers in that direction. Only dorsal to the postcleithrum, the posterior margin of the supracleithrum is ornamented. In the
|
||
<typeStatus id="548BCBD3FFE80200FB27FACF6F7BF83A" box="[1266,1364,1348,1372]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
(IGM 6716) this area is garnished with a series of parallel and short ridges, which become gradually more tenuous in dorso-ventral direction (
|
||
<figureCitation id="130B69F4FFE80200FE73FA006822F8C5" box="[422,525,1419,1443]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="15" pageNumber="16">Figs 6–7</figureCitation>
|
||
). In the dorsal portion of the bone, these ridges project as spines beyond the bor- der of the bone. However, the same area is ornamented with tubercles in IGM 6719 (
|
||
<figureCitation id="130B69F4FFE80200FB99FA256EA0F8A0" box="[1100,1167,1454,1478]" captionStart="FIGURE 9" captionStartId="11.[151,249,1832,1855]" captionTargetBox="[151,1436,841,1811]" captionTargetId="figure@11.[151,1436,841,1811]" captionTargetPageId="11" captionText="FIGURE 9. Tlayuamichin itztli gen. et sp. nov., details of the skull in IGM 6719. A, posterior part of the skull showing multiple opercular ossifications and the peculiar shape of the suboperculum in this specimen. B, detailed photograph of the dermopterotic bone; the arrow points to the peculiar anteroventral process of this bone in the new taxon. C, same photograph as in B, but in this case the arrows point to the pores of the preopercular sensory canal in the preoperculum (below), suborbital (middle) and dermopterotic (above). Scale bars equal = 10 mm." httpUri="https://zenodo.org/record/205976/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
|
||
) and with tubercles and ridges in IGM 6720 (
|
||
<figureCitation id="130B69F4FFE80200FE51FA5A6BE7F88F" box="[388,456,1489,1513]" captionStart="FIGURE 8" captionStartId="10.[151,249,1361,1384]" captionTargetBox="[151,1436,193,1339]" captionTargetId="figure@10.[151,1436,193,1340]" captionTargetPageId="10" captionText="FIGURE 8. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6720. Scale bar equals = 45 mm." httpUri="https://zenodo.org/record/205975/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFE80200FF12FA7E6EEEFA87" blockId="15.[151,1437,151,2017]" pageId="15" pageNumber="16">
|
||
In lateral view the cleithrum has a crescent shape, slightly deeper than long and anteriorly reaches a level around the posterior rim of the orbit. The cleithrum has up to seven sinuous and variably interrupted series of denticles along the edge between the branchial and lateral surfaces. The lateral surface of the cleithrum is notably narrow and its free border is smooth. The four postcleithra are aligned following the curved made by the posteroventral border of the cleithrum. Differing from the other dermal components of the pectoral girdle, the exposed portion of the postcleithra is covered with a layer of ganoin. The first, most dorsal postcleithrum is dorsoventrally elongate, narrowing dorsally. It is the deepest among the postcleithra, but it is narrower than the adjacent postcleithrum. This second postcleithrum is approximately as deep as long and has a very gently concave anterior border, and a convex posterior border. The two following postcleithra are much smaller than the elements described before. Both are elongate and their shape accommodate to the shape of the posterior border of the cleithrum.
|
||
<collectingCountry id="F32735E1FFE80200FAB6F8B26FADFA37" box="[1379,1410,1849,1873]" name="American Samoa" pageId="15" pageNumber="16">As</collectingCountry>
|
||
is the case of the supracleithrum, the two most dorsal postcleithra are variably ornamented with tubercles and/or ridges, which are sometimes projected in spines. Ventral and medial to the second postcleithrum there is a modified scale with three strong spines projecting backwards (
|
||
<figureCitation id="130B69F4FFE80200FD3CF82E6960FADB" box="[745,847,1957,1981]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="15" pageNumber="16">Figs 6–7</figureCitation>
|
||
). To facilitate comparisons (see discussion) we refer to this scale, which is well exposed in the
|
||
<typeStatus id="548BCBD3FFE80200FDBBF84268FFFA87" box="[622,720,1993,2017]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
(IGM 6716), as the modified pectoral scale.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFF7021FFF12FF1C6E77FD91" blockId="16.[151,1437,151,1114]" pageId="16" pageNumber="17">
|
||
Of the endochondral components of the pectoral girdle, only five radials are exposed in the
|
||
<typeStatus id="548BCBD3FFF7021FFB00FF1C6F18FDC9" box="[1237,1335,151,175]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="130B69F4FFF7021FFA93FF1C6A88FDB2" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="16" pageNumber="17">Figs 6– 7</figureCitation>
|
||
). Three of these radials are only partially exposed because they are preserved in articulation. The other two radials are disarticulated and displaced ventral to the skull and anterior to the pectoral fin.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFF7021FFF12FE88684FFF39" blockId="16.[151,1437,151,1114]" pageId="16" pageNumber="17">
|
||
<emphasis id="B944A963FFF7021FFF12FE886B7EFC7A" bold="true" box="[199,337,259,284]" pageId="16" pageNumber="17">Paired fins.</emphasis>
|
||
The pectoral fin is best and almost completely preserved (only the distal fin rays are missing) in the
|
||
<typeStatus id="548BCBD3FFF7021FFF14FEAC6B0CFC59" box="[193,291,295,319]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
(IGM 6716;
|
||
<figureCitation id="130B69F4FFF7021FFE6EFEAC680EFC59" box="[443,545,295,319]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,249,1202,1225]" captionStartId-1="9.[151,249,1517,1540]" captionTargetBox-0="[278,1307,179,1187]" captionTargetBox-1="[188,1398,625,1463]" captionTargetId-0="figure@8.[278,1308,179,1188]" captionTargetId-1="figure@9.[151,1436,589,1496]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." captionText-1="FIGURE 7. Tlayuamichin itztli gen. et sp. nov. Line drawing of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri-0="https://zenodo.org/record/205972/files/figure.png" httpUri-1="https://zenodo.org/record/205973/files/figure.png" pageId="16" pageNumber="17">Figs 6–7</figureCitation>
|
||
), comprising nine lepidotrichia. The proximal third to one half of each lepidotrichium is unsegmented, contrasting strongly with the distal portion, which is finely segmented and intensively branched. The first lepidotrichium does not branch and its terminal segment is elongate, covered with ganoin and completes the series of fringing fulcra, which are laying on this first ray. The proximal portion of the first pectoral ray forms a lateral ridge, which is also covered with ganoin. Scattered remains of a few small broken spines are preserved on the proximal portion of the second to fourth pectoral rays. From the second ray onwards, the lepidotrichia are asymmetrically branched, the medial portion being always more intensively branched than the lateral one. Also, the branching is gradually more intense towards the medial rays. Two short and paired basal fulcra are preceding the series of short fringing fulcra.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFF7021FFF12FDE76B24FE51" blockId="16.[151,1437,151,1114]" pageId="16" pageNumber="17">
|
||
The pelvic fins are also best, though only partially preserved in the
|
||
<typeStatus id="548BCBD3FFF7021FFC67FDE76E3BFFE2" box="[946,1044,620,644]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
(IGM 6716;
|
||
<figureCitation id="130B69F4FFF7021FFB73FDE76EC6FFE2" box="[1190,1257,620,644]" captionStart="FIGURE 2" captionStartId="4.[151,249,1014,1037]" captionTargetBox="[151,1436,427,993]" captionTargetId="figure@4.[151,1436,427,993]" captionTargetPageId="4" captionText="FIGURE 2. Tlayuamichin itztli gen. et sp. nov. Holotype (IGM 6716) preserved in left lateral view. Standard length of the specimen = 403 mm. Scale bar equals 70 mm." httpUri="https://zenodo.org/record/205968/files/figure.png" pageId="16" pageNumber="17">Fig. 2</figureCitation>
|
||
), where the pelvic fin is located about 0.52 of the standard length, right behind the seventh vertical row of scales. The total number of rays is unknown, but at least two lepidotrichia are preserved. Resembling the case of the pectoral fin, the proximal segment of the first pelvic fin ray forms a lateral ridge, which is covered with a thin layer of ganoin. A series of fulcra garnish the pelvic fin, including one unpaired basal fulcrum, one paired basal fulcra, and at least eight fringing fulcra.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFF7021FFF12FCC868D8F93C" blockId="16.[151,1437,151,1114]" pageId="16" pageNumber="17">
|
||
<emphasis id="B944A963FFF7021FFF12FCC86B5EFE3A" bold="true" box="[199,369,835,860]" pageId="16" pageNumber="17">Unpaired fins.</emphasis>
|
||
The unpaired fins are best preserved in IGM 6718 (
|
||
<figureCitation id="130B69F4FFF7021FFC61FCCF69D5FE3A" box="[948,1018,836,860]" captionStart="FIGURE 3" captionStartId="6.[151,249,1801,1824]" captionTargetBox="[151,1436,193,1780]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tlayuamichin itztli gen. et sp. nov. A, Specimen IGM 6717 preserved in right lateroventral view. B, specimen IGM 6718 preserved in right lateral view. Scale bars equal 70 mm." httpUri="https://zenodo.org/record/205969/files/figure.png" pageId="16" pageNumber="17">Figs 3</figureCitation>
|
||
B, 13–14). The dorsal fin is short and originates right back of the level of the pelvic-fin insertion. The dorsal fin is incompletely preserved in IGM 6716 and 6720 (
|
||
<figureCitation id="130B69F4FFF7021FFED8FC046B79FEC1" box="[269,342,911,935]" captionStart="FIGURE 2" captionStartId="4.[151,249,1014,1037]" captionTargetBox="[151,1436,427,993]" captionTargetId="figure@4.[151,1436,427,993]" captionTargetPageId="4" captionText="FIGURE 2. Tlayuamichin itztli gen. et sp. nov. Holotype (IGM 6716) preserved in left lateral view. Standard length of the specimen = 403 mm. Scale bar equals 70 mm." httpUri="https://zenodo.org/record/205968/files/figure.png" pageId="16" pageNumber="17">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFF7021FFEB7FC046B5EFEC1" box="[354,369,911,935]" captionStart="FIGURE 5" captionStartId="7.[151,249,1636,1659]" captionTargetBox="[151,1436,1010,1614]" captionTargetId="figure@7.[151,1436,1010,1615]" captionTargetPageId="7" captionText="FIGURE 5. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6720 preserved in right lateral view. Scale bar equals = 50 mm." httpUri="https://zenodo.org/record/205971/files/figure.png" pageId="16" pageNumber="17">5</figureCitation>
|
||
), but both specimens show that this fin originates at the 23rd vertical row of scales, at about 0.6 of the standard length. There are 13 lepidotrichia in the dorsal fin of IGM 6718 (
|
||
<figureCitation id="130B69F4FFF7021FFBE9FC396EA0FEAC" box="[1084,1167,946,970]" captionStart="FIGURE 13" captionStartId="16.[151,249,1619,1642]" captionTargetBox="[151,1436,1154,1598]" captionTargetId="figure@16.[151,1436,1154,1598]" captionTargetPageId="16" captionText="FIGURE 13. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of: A, the dorsal fin in IGM 6718; B, the anal fin in IGM 6718. Scale bars equal = 20 mm." httpUri="https://zenodo.org/record/205980/files/figure.png" pageId="16" pageNumber="17">Fig. 13</figureCitation>
|
||
A), the height of which decreases progressively posteriorly and, thus, the shape of the fin was probably triangular. All dorsal fin rays, including the first one, branch at least two times. There are five large paired basal fulcra, and at least 10 large fringing fulcra. Among these fulcra, the two posterior paired basal fulcra and two anterior fringing fulcra are the largest, being almost 1.5 times higher than the adjacent fulcra.
|
||
</paragraph>
|
||
<caption id="DF4F25F9FFF7021FFF42F9D86BCBFBEE" httpUri="https://zenodo.org/record/205980/files/figure.png" pageId="16" pageNumber="17" targetBox="[151,1436,1154,1598]" targetPageId="16">
|
||
<paragraph id="8B8F7571FFF7021FFF42F9D86BCBFBEE" blockId="16.[151,1436,1619,1672]" pageId="16" pageNumber="17">
|
||
<emphasis id="B944A963FFF7021FFF42F9D86B30FB0E" bold="true" box="[151,287,1619,1642]" pageId="16" pageNumber="17">FIGURE 13.</emphasis>
|
||
<taxonomicName id="4C300EF2FFF7021FFEF1F9D86BCCFB0F" box="[292,483,1619,1641]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFF7021FFEF1F9D86BCCFB0F" box="[292,483,1619,1641]" italics="true" pageId="16" pageNumber="17">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFF7021FFE3CF9DF68AAFB0F" bold="true" box="[489,645,1620,1641]" pageId="16" pageNumber="17">gen. et sp. nov.</emphasis>
|
||
Detailed photograph of: A, the dorsal fin in IGM 6718; B, the anal fin in IGM 6718. Scale bars equal = 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F7571FFF7021FFF10F9246B38FAC4" blockId="16.[151,1436,1711,2026]" pageId="16" pageNumber="17">
|
||
The anal fin is also incompletely preserved in both IGM 6716 and 6720 (
|
||
<figureCitation id="130B69F4FFF7021FFC28F9246E69FBA1" box="[1021,1094,1711,1735]" captionStart="FIGURE 2" captionStartId="4.[151,249,1014,1037]" captionTargetBox="[151,1436,427,993]" captionTargetId="figure@4.[151,1436,427,993]" captionTargetPageId="4" captionText="FIGURE 2. Tlayuamichin itztli gen. et sp. nov. Holotype (IGM 6716) preserved in left lateral view. Standard length of the specimen = 403 mm. Scale bar equals 70 mm." httpUri="https://zenodo.org/record/205968/files/figure.png" pageId="16" pageNumber="17">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="130B69F4FFF7021FFB86F9246E4DFBA1" box="[1107,1122,1711,1735]" captionStart="FIGURE 5" captionStartId="7.[151,249,1636,1659]" captionTargetBox="[151,1436,1010,1614]" captionTargetId="figure@7.[151,1436,1010,1615]" captionTargetPageId="7" captionText="FIGURE 5. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6720 preserved in right lateral view. Scale bar equals = 50 mm." httpUri="https://zenodo.org/record/205971/files/figure.png" pageId="16" pageNumber="17">5</figureCitation>
|
||
), but these specimens show its origin at the 18th vertical row of scales, at 0.72–0.74 of the standard length. The anal fin is almost complete and well preserved in IGM 6718 (
|
||
<figureCitation id="130B69F4FFF7021FFE39F9716810FA74" box="[492,575,1786,1810]" captionStart="FIGURE 13" captionStartId="16.[151,249,1619,1642]" captionTargetBox="[151,1436,1154,1598]" captionTargetId="figure@16.[151,1436,1154,1598]" captionTargetPageId="16" captionText="FIGURE 13. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of: A, the dorsal fin in IGM 6718; B, the anal fin in IGM 6718. Scale bars equal = 20 mm." httpUri="https://zenodo.org/record/205980/files/figure.png" pageId="16" pageNumber="17">Fig. 13</figureCitation>
|
||
B), comprising seven lepidotrichia, the two distal thirds of which are finely segmented and branched, including the first ray that branches at least once. The proximal half of the first lepidotrichium is covered with lateral patches of ganoin. The anal fin is garnished with three paired basal fulcra, and at least 19 small fringing fulcra. There is a single large preanal scute, which is about three times larger than the adjacent scales.
|
||
</paragraph>
|
||
<paragraph id="8B8F7571FFF7021EFF10F8256B05FCB6" blockId="16.[151,1436,1711,2026]" lastBlockId="17.[151,1436,151,464]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
|
||
The caudal fin is also very well preserved in IGM 6718 (
|
||
<figureCitation id="130B69F4FFF7021FFC89F825699EFAA0" box="[860,945,1966,1990]" captionStart="FIGURE 14" captionStartId="17.[151,249,1236,1259]" captionTargetBox="[375,1211,505,1215]" captionTargetId="figure@17.[375,1211,505,1215]" captionTargetPageId="17" captionText="FIGURE 14. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the caudal fin in IGM 6718. Scale bar equals = 20 mm." httpUri="https://zenodo.org/record/205981/files/figure.png" pageId="16" pageNumber="17">Fig. 14</figureCitation>
|
||
), but the distal portions of the fin rays are missing, so that the shape of the fin is unknown. Both in this specimen and in IGM 6720, there are nine and eight lepidotrichia forming the dorsal and ventral caudal lobes respectively.
|
||
<collectingCountry id="F32735E1FFF6021EFC60FF1369FBFDD6" box="[949,980,152,176]" name="American Samoa" pageId="17" pageNumber="18">As</collectingCountry>
|
||
shown in IGM 6716, all caudal fin rays are laterally covered with patches of ganoin. The caudal fin rays are intensively branched, with the exception of the two marginal rays, and finely segmented like the lepidotrichia in the other unpaired fins. In this case however, the proximal unsegmented portion is short and almost totally covered with scales.
|
||
<collectingCountry id="F32735E1FFF6021EFBC6FE8F6E1DFC7A" box="[1043,1074,260,284]" name="American Samoa" pageId="17" pageNumber="18">As</collectingCountry>
|
||
far as preserved, the dorsal and ventral marginal rays are unbranched. The most posterior flank scales are relatively longer and oval. There are at least six unpaired dorsal caudal fulcra covering the body lobe of the tail, followed by three paired basal fulcra and at least 13 fringing fulcra laying on the dorsal marginal ray. Ventrally, the caudal fin is preceded by one short unpaired basal fulcrum and five paired basal fulcra, and there are at least 22 small fringing fulcra on the ventral marginal ray.
|
||
</paragraph>
|
||
<caption id="DF4F25F9FFF6021EFF42FB5F6AEAF86C" httpUri="https://zenodo.org/record/205981/files/figure.png" pageId="17" pageNumber="18" targetBox="[375,1211,505,1215]" targetPageId="17">
|
||
<paragraph id="8B8F7571FFF6021EFF42FB5F6AEAF86C" blockId="17.[151,1436,1236,1290]" pageId="17" pageNumber="18">
|
||
<emphasis id="B944A963FFF6021EFF42FB5F6B0EF98C" bold="true" box="[151,289,1236,1259]" pageId="17" pageNumber="18">FIGURE 14.</emphasis>
|
||
<taxonomicName id="4C300EF2FFF6021EFEFDFB5E6BC6F98D" box="[296,489,1237,1259]" class="Actinopterygii" family="Semionotidae" genus="Tlayuamichin" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amiiformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="itztli">
|
||
<emphasis id="B944A963FFF6021EFEFDFB5E6BC6F98D" box="[296,489,1237,1259]" italics="true" pageId="17" pageNumber="18">Tlayuamichin itztli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944A963FFF6021EFE24FB5E68BCF98C" bold="true" box="[497,659,1237,1258]" pageId="17" pageNumber="18">gen. et sp. nov.</emphasis>
|
||
Detailed photograph of the caudal fin in IGM 6718. Scale bar equals = 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F7571FFF6021EFF12FABB6BCDFA27" blockId="17.[151,1437,1328,2037]" pageId="17" pageNumber="18">
|
||
<emphasis id="B944A963FFF6021EFF12FABB6B70F82F" bold="true" box="[199,351,1328,1353]" pageId="17" pageNumber="18">Squamation.</emphasis>
|
||
The body is covered with large and thick rhomboid scales. The squamation pattern is best preserved in the
|
||
<typeStatus id="548BCBD3FFF6021EFEF8FADE6BA0F80B" box="[301,399,1365,1389]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
|
||
(IGM 6716;
|
||
<figureCitation id="130B69F4FFF6021EFDF6FADE6848F80B" box="[547,615,1365,1389]" captionStart="FIGURE 2" captionStartId="4.[151,249,1014,1037]" captionTargetBox="[151,1436,427,993]" captionTargetId="figure@4.[151,1436,427,993]" captionTargetPageId="4" captionText="FIGURE 2. Tlayuamichin itztli gen. et sp. nov. Holotype (IGM 6716) preserved in left lateral view. Standard length of the specimen = 403 mm. Scale bar equals 70 mm." httpUri="https://zenodo.org/record/205968/files/figure.png" pageId="17" pageNumber="18">Fig. 2</figureCitation>
|
||
), in which are 34 vertical rows of scales from the operculum to the hinge line, and a complete ridge of scales along the dorsal midline between the skull and the dorsal fin. The scales are not ornamented and have smooth surfaces. Most of the scales on the flank in IGM 6716 form one or two spines at their posteroventral angle (
|
||
<figureCitation id="130B69F4FFF6021EFE5FFA4A6BE1F8BF" box="[394,462,1473,1497]" captionStart="FIGURE 6" captionStartId="8.[151,249,1202,1225]" captionTargetBox="[278,1307,179,1187]" captionTargetId="figure@8.[278,1308,179,1188]" captionTargetPageId="8" captionText="FIGURE 6. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of the skull in IGM 6716 (holotype). Scale bar equals = 50 mm." httpUri="https://zenodo.org/record/205972/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
|
||
). However, such spine is only insinuated in the scales of IGM 6719 (
|
||
<figureCitation id="130B69F4FFF6021EFB06FA4A6F39F8BF" box="[1235,1302,1473,1497]" captionStart="FIGURE 4" captionStartId="7.[151,249,938,961]" captionTargetBox="[151,1436,193,916]" captionTargetId="figure@7.[151,1436,193,917]" captionTargetPageId="7" captionText="FIGURE 4. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6719 preserved in right lateral view. Scale bar equals 50 mm." httpUri="https://zenodo.org/record/205970/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
|
||
) and generally absent in IGM 6720 (
|
||
<figureCitation id="130B69F4FFF6021EFE69FA6E682FF89B" box="[444,512,1509,1533]" captionStart="FIGURE 5" captionStartId="7.[151,249,1636,1659]" captionTargetBox="[151,1436,1010,1614]" captionTargetId="figure@7.[151,1436,1010,1615]" captionTargetPageId="7" captionText="FIGURE 5. Tlayuamichin itztli gen. et sp. nov. Specimen IGM 6720 preserved in right lateral view. Scale bar equals = 50 mm." httpUri="https://zenodo.org/record/205971/files/figure.png" pageId="17" pageNumber="18">Fig. 5</figureCitation>
|
||
), which shows that the feature is highly variable within the species. On the flank, the scales are generally rectangular, approximately as deep as long, only somewhat deeper than long immediately behind the operculum. The dorsal ridge scales are conspicuous, slightly longer than those adjacent scales and bears a short posterior spine directed backward (
|
||
<figureCitation id="130B69F4FFF6021EFDABF9DA68FFFB0F" box="[638,720,1617,1641]" captionStart="FIGURE 15" captionStartId="18.[151,249,807,830]" captionTargetBox="[152,1435,181,786]" captionTargetId="figure@18.[152,1437,181,786]" captionTargetPageId="18" captionText="FIGURE 15. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of: A, the dorsal ridge scales in IGM 6719; B, several partially articulated scales in IGM 6717 showing the well-developed articular, anterodorsal and anteroventral processes. Scale bars equal = 10 mm." httpUri="https://zenodo.org/record/205982/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
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A). In the caudal peduncle, the scales are about 2 times longer than deep and have smooth borders in all the specimens. There are seven inverted rows of scales forming the body lobe of the tail. Among them, the scales of the marginal row are distinctively elongate and not diamond shape as the other scales in the caudal region. Six scales are preserved in this marginal row, but there is a fragment of a seventh scale and one scales is evidently missing both ventrally and dorsally and, thus, nine is the total number of scales in this row. Nevertheless, this marginal row does not reach the tip of the body lobe, which is formed by the two following rows of scales.
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</paragraph>
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<paragraph id="8B8F7571FFF6021EFF10F8C66FA0FA93" blockId="17.[151,1437,1328,2037]" pageId="17" pageNumber="18">
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The articular processes of the scales are only shown in the middle flank region of IGM 6717. In this area, the scales show well-developed articular, anterodorsal and anteroventral processes (anterior pegs) (
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<figureCitation id="130B69F4FFF6021EFB38F8FA6F6CFAEF" box="[1261,1347,1905,1929]" captionStart="FIGURE 15" captionStartId="18.[151,249,807,830]" captionTargetBox="[152,1435,181,786]" captionTargetId="figure@18.[152,1437,181,786]" captionTargetPageId="18" captionText="FIGURE 15. Tlayuamichin itztli gen. et sp. nov. Detailed photograph of: A, the dorsal ridge scales in IGM 6719; B, several partially articulated scales in IGM 6717 showing the well-developed articular, anterodorsal and anteroventral processes. Scale bars equal = 10 mm." httpUri="https://zenodo.org/record/205982/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
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B). The articular process that serves in the peg-and-socket articulation (dorsal peg) is not very high and places right at the base of the anterodorsal process. The anterodorsal and anteroventral processes serve in the longitudinal articulation and are oriented in diverging directions, with the anterodorsal process being larger than the anteroventral process.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |