158 lines
18 KiB
XML
158 lines
18 KiB
XML
<document id="E39B880CCA704772356EA3BACEB4CA72" ID-CLB-Dataset="32715" ID-DOI="10.11646/zootaxa.4290.2.2" ID-GBIF-Dataset="4469947c-31f9-46e6-b963-03ae79d84554" ID-ISSN="1175-5326" ID-Zenodo-Dep="829009" ID-ZooBank="79FBA585-7662-4DE1-9BAB-37DFDA2BABBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110470243" checkinUser="plazi" docAuthor="Marshall, Stephen A." docDate="2017" docId="0399A674FFF4A02EFF0AF8BCADE48F48" docLanguage="en" docName="zootaxa.4290.2.2.pdf" docOrigin="Zootaxa 4290 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paramimegralla longicephala Marshall, 2017, new species" docType="treatment" docVersion="7" lastPageNumber="259" masterDocId="FFA0DE0CFFF8A021FF9DFF89A94D8B13" masterDocTitle="Micropezidae (Insecta, Diptera, Acalyptratae) of Madagascar and a revision of the genus Paramimegralla Hennig" masterLastPageNumber="280" masterPageNumber="244" pageNumber="256" updateTime="1698465370965" updateUser="ExternalLinkService">
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<mods:title id="C573F74F1654FBD34A9C815BB912EFC5">Micropezidae (Insecta, Diptera, Acalyptratae) of Madagascar and a revision of the genus Paramimegralla Hennig</mods:title>
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<mods:namePart id="EB54F0CAA2897EA3ADEA3F0C919D934C">Marshall, Stephen A.</mods:namePart>
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<mods:title id="9026E7563630DB03AA48AC449C79AE1F">Zootaxa</mods:title>
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<mods:part id="9A3E09F3EDD25F85B0C7DD2C62969ABE">
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<mods:date id="7EBE93107EEDDD29A608613474D9D9CD">2017</mods:date>
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<mods:number id="24D3DF63318BFF455B4D2D2BB86FC67B">4290</mods:number>
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<mods:number id="BA751C54B40406CC3A022AB5C6D60981">2</mods:number>
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<mods:identifier id="1E1C644C75EF7C8357844B242206C64A" type="CLB-Dataset">32715</mods:identifier>
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<mods:identifier id="5086A53E713D2CC224D409B3DBF1D45F" type="DOI">10.11646/zootaxa.4290.2.2</mods:identifier>
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<mods:identifier id="6FAF403BBC1D6526408C85C0BC829542" type="GBIF-Dataset">4469947c-31f9-46e6-b963-03ae79d84554</mods:identifier>
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<mods:identifier id="9F264C598C9921ABE7DD712A2D2F083B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0399A674FFF4A02EFF0AF8BCADE48F48" ID-DOI="http://doi.org/10.5281/zenodo.6033428" ID-GBIF-Taxon="132549249" ID-Zenodo-Dep="6033428" LSID="urn:lsid:plazi:treatment:0399A674FFF4A02EFF0AF8BCADE48F48" httpUri="http://treatment.plazi.org/id/0399A674FFF4A02EFF0AF8BCADE48F48" lastPageId="15" lastPageNumber="259" pageId="12" pageNumber="256">
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<subSubSection id="C32A44E9FFF4A02DFF0AF8BCA85B8C62" pageId="12" pageNumber="256" type="nomenclature">
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<paragraph id="8B8F1762FFF4A02DFF0AF8BCABDB8C5C" blockId="12.[151,662,1845,1905]" box="[151,662,1845,1871]" pageId="12" pageNumber="256">
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<heading id="D0C7A00EFFF4A02DFF0AF8BCABDB8C5C" bold="true" box="[151,662,1845,1871]" fontSize="11" level="1" pageId="12" pageNumber="256" reason="1">
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<emphasis id="B944CB70FFF4A02DFF0AF8BCABDB8C5C" bold="true" box="[151,662,1845,1871]" pageId="12" pageNumber="256">
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<taxonomicName id="4C306CE1FFF4A02DFF0AF8BCA8B28C5C" ID-CoL="9HV3Z" box="[151,511,1845,1871]" class="Insecta" family="Micropezidae" genus="Paramimegralla" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="256" phylum="Arthropoda" rank="species" species="longicephala" status="sp. nov.">
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<emphasis id="B944CB70FFF4A02DFF0AF8BCA8B28C5C" bold="true" box="[151,511,1845,1871]" italics="true" pageId="12" pageNumber="256">Paramimegralla longicephala</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A277760BFFF4A02DFD9AF8BCABDB8C5C" box="[519,662,1845,1871]" pageId="12" pageNumber="256" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8F1762FFF4A02DFF0AF8D1A85B8C62" blockId="12.[151,662,1845,1905]" box="[151,278,1880,1905]" pageId="12" pageNumber="256">
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<figureCitation id="130B0BE7FFF4A02DFF0AF8D1A9A68C62" box="[151,235,1880,1905]" captionStart="FIGURES 31 – 36" captionStartId="14.[151,264,1680,1703]" captionTargetBox="[151,1436,183,1658]" captionTargetId="figure@14.[151,1436,183,1659]" captionTargetPageId="14" captionText="FIGURES 31 – 36. Paramimegralla lOngiCephala sp. nov., Male holotype. 31. Head and anterior thorax, dorsal; 32. Holotype prior to capture, Andasibe; 33. Base of abdomen; 34. Abdominal segments 3 ‾ 6, lateral; 35. Sternite 5 with genital fork; 36. Male terminalia, left lateral." httpUri="https://zenodo.org/record/829019/files/figure.png" pageId="12" pageNumber="256">Figs 31</figureCitation>
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̄36
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A44E9FFF4A02EFF0AF829A8CD884C" lastPageId="15" lastPageNumber="259" pageId="12" pageNumber="256" type="description">
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<paragraph id="8B8F1762FFF4A02DFF0AF829AD9B8CAA" blockId="12.[151,1238,1952,1977]" box="[151,1238,1952,1977]" pageId="12" pageNumber="256">
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<emphasis id="B944CB70FFF4A02DFF0AF829A86F8CAA" bold="true" box="[151,290,1952,1977]" pageId="12" pageNumber="256">Description</emphasis>
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(male only). Length
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<quantity id="4CC8BA87FFF4A02DFD93F829AB148CAA" box="[526,601,1952,1977]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="256" unit="mm" value="15.0">15mm</quantity>
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. Ground colour dull black, mostly silvery microsetulose.
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</paragraph>
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<caption id="DF4F47EAFFF5A02CFF0AF8CCAD858CAB" httpUri="https://zenodo.org/record/829017/files/figure.png" pageId="13" pageNumber="257" targetBox="[151,1436,193,1840]" targetPageId="13">
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<paragraph id="8B8F1762FFF5A02CFF0AF8CCAD858CAB" blockId="13.[151,1436,1861,1976]" pageId="13" pageNumber="257">
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<emphasis id="B944CB70FFF5A02CFF0AF8CCABAA8C48" bold="true" box="[151,743,1861,1884]" pageId="13" pageNumber="257">
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FIGURES 24–30.
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<taxonomicName id="4C306CE1FFF5A02CFEC2F8CCABCB8C48" box="[351,646,1861,1883]" class="Insecta" family="Micropezidae" genus="Paramimegralla" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="256" phylum="Arthropoda" rank="species" species="campanula" status="sp. nov.">
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<emphasis id="B944CB70FFF5A02CFEC2F8CCABCB8C48" bold="true" box="[351,646,1861,1883]" italics="true" pageId="13" pageNumber="257">Paramimegralla Campanula</emphasis>
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</taxonomicName>
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sp. nov.,
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</emphasis>
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Male holotype. 34. Dorsolateral; 25. Head, anterolateral; 26. Male terminalia, left lateral; 27. Male terminalia, ventral; 28. Base of aedeagus, ventral; 29. Apex of abdomen with terminalia removed; 30. Sternite 5 with genital fork. Abbreviations: aa, phallapodeme; bd, basal distiphallus; c, cercus; dd, distal distiphallus; e, epandrium; ea, ejaculatory apodeme; ed, ejaculatory duct; h, hypandrium; pb, phallic bulb.
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</paragraph>
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</caption>
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<caption id="DF4F47EAFFF6A02FFF0AF919A8FE8DF7" httpUri="https://zenodo.org/record/829019/files/figure.png" pageId="14" pageNumber="258" targetBox="[151,1436,183,1658]" targetPageId="14">
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<paragraph id="8B8F1762FFF6A02FFF0AF919A8FE8DF7" blockId="14.[151,1436,1680,1764]" pageId="14" pageNumber="258">
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<emphasis id="B944CB70FFF6A02FFF0AF919ABA18DB5" bold="true" box="[151,748,1680,1703]" pageId="14" pageNumber="258">
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FIGURES 31–36.
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<taxonomicName id="4C306CE1FFF6A02FFEC1F919ABD98DB5" box="[348,660,1680,1702]" class="Insecta" family="Micropezidae" genus="Paramimegralla" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="256" phylum="Arthropoda" rank="species" species="longicephala" status="sp. nov.">
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<emphasis id="B944CB70FFF6A02FFEC1F919ABD98DB5" bold="true" box="[348,660,1680,1702]" italics="true" pageId="14" pageNumber="258">Paramimegralla lOngiCephala</emphasis>
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</taxonomicName>
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sp. nov.
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</emphasis>
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, Male holotype. 31. Head and anterior thorax, dorsal; 32. Holotype prior to capture, Andasibe; 33. Base of abdomen; 34. Abdominal segments 3‾6, lateral; 35. Sternite 5 with genital fork; 36. Male terminalia, left lateral.
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</paragraph>
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</caption>
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<paragraph id="8B8F1762FFF6A02FFF5AF882ABF08CE5" blockId="14.[151,1437,1803,2038]" pageId="14" pageNumber="258">
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Head with frons broad, raised and conspicuously rugose, frontal vitta densely setulose but frons otherwise sparsely setulose (most) to bare (epicephalon), frontal vitta broad anterior to ocelli, abruptly tapered to a thin groove immediately behind ocelli and ending far before back of head. Head greatly elongated behind ocelli, inner vertical setae at mid point between ocelli and back of head, postocellar and outer vertical setae in line along back of head; head behind eyes about one third of eye width. Two frontal setae. Palpus parallel-sided, black, golden microsetulose. Occipital suture shining black. Clypeus shining black, sparsely microsetulose, twice as long as face. Length of first flagellomere
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<date id="FF8E31A2FFF6A02FFE49F857A8BB8CE5" box="[468,502,2014,2038]" pageId="14" pageNumber="258">2X</date>
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maximum width.
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</paragraph>
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<paragraph id="8B8F1762FFF7A02EFF5AFF1EACC88A9B" blockId="15.[151,1437,151,1115]" pageId="15" pageNumber="259">
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Thorax with scutellum silvery microsetulose; scutum black, finely microsetulose with two narrow and narrowly separated longitudinal silver submedian strips. Proepisternum with short, inconspicuous ventral setae. Pleuron entirely microsetulose, mesopleuron with indistinct silvery pruinose diagonal strip extending from corner of transverse suture to mid coxa. Legs black to dark brown except as follows: fore femur brown at apex, fore tarsus white, first tarsomere with dense golden setulae ventrally. Mid and hind femora with a yellow–brown apex, hind femur also with a broad yellow–brown band near middle. Hind tarsus mostly white, apical two tarsomeres black. Hind femur flattened and distinctly sulcate on
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<typeStatus id="548BA9C0FFF7A02EFD02FEF9AB828A9B" box="[671,719,368,392]" pageId="15" pageNumber="259">type</typeStatus>
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specimen, hind tibia cylindrical. Hind trochanter dark brown.
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</paragraph>
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<paragraph id="8B8F1762FFF7A02EFF5AFE1DA8AC8909" blockId="15.[151,1437,151,1115]" pageId="15" pageNumber="259">
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Wing with broad basal, discal and preapical bands as well as an infuscated apex widely separated from the preapical band by a clear area apparently made of 3 coalesced circular spots; discal band concave on distal margin and convex on basal margin. Anal cell bare except for a very narrow microsetulose fringe, CuA2 straight, in line with dm̄cu, distal angle 35
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<emphasis id="B944CB70FFF7A02EFE52FD8BA8978908" bold="true" box="[463,474,514,539]" pageId="15" pageNumber="259">°</emphasis>
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.
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</paragraph>
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<paragraph id="8B8F1762FFF7A02EFF5AFDAFA8CD884C" blockId="15.[151,1437,151,1115]" pageId="15" pageNumber="259">
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Abdominal syntergite 1+2 humped and silvery at junction between T1 and T2; T1 evenly setose anterior to hump; pleural membrane black with a broad, almost vertical, parallel-sided white band on anterior margin of P3. Oviscape black with orange apex. Female unknown. Male with genital fork (S5) with long distally tapered and incurved arms with inner margins densely packed with short, stout setae, base between arms with two long, parallel-sided processes (unique in the genus,
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<figureCitation id="130B0BE7FFF7A02EFD34FD3FABB389DC" box="[681,766,694,719]" captionStart="FIGURES 31 – 36" captionStartId="14.[151,264,1680,1703]" captionTargetBox="[151,1436,183,1658]" captionTargetId="figure@14.[151,1436,183,1659]" captionTargetPageId="14" captionText="FIGURES 31 – 36. Paramimegralla lOngiCephala sp. nov., Male holotype. 31. Head and anterior thorax, dorsal; 32. Holotype prior to capture, Andasibe; 33. Base of abdomen; 34. Abdominal segments 3 ‾ 6, lateral; 35. Sternite 5 with genital fork; 36. Male terminalia, left lateral." httpUri="https://zenodo.org/record/829019/files/figure.png" pageId="15" pageNumber="259">Fig. 35</figureCitation>
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). Distiphallus with an exceptionally broad and short basal part ending in a dense phallic bulb; distal part (beyond phallic bulb) thin and very long, about 4 times as long as basal part, loosely coiled distally and apically thin. Blade of ejaculatory apodeme broad, much larger than sperm pump, width subequal to epandrial height, total area of ejaculatory apodeme including sperm pump subequal in length to epandrium.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A44E9FFF7A02EFF5AFCE3AD3F88B4" pageId="15" pageNumber="259" type="materials_examined">
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<paragraph id="8B8F1762FFF7A02EFF5AFCE3AD3F88B4" blockId="15.[151,1437,151,1115]" pageId="15" pageNumber="259">
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<materialsCitation id="3B581D3FFFF7A02EFF5AFCE3AD3F88B4" ID-GBIF-Occurrence="1572325768" collectionCode="CAS, MYCRO" collectorName="S. A. Marshall" country="Madagascar" location="Andasibe." pageId="15" pageNumber="259" specimenCode="MYCRO192" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B944CB70FFF7A02EFF5AFCE3A8378890" bold="true" box="[199,378,874,899]" pageId="15" pageNumber="259">
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||
<typeStatus id="548BA9C0FFF7A02EFF5AFCE3A84F8890" box="[199,258,874,899]" pageId="15" pageNumber="259">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="548BA9C0FFF7A02EFE1BFCE3A8A38890" box="[390,494,874,899]" pageId="15" pageNumber="259" type="holotype">Holotype</typeStatus>
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(
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<specimenCount id="9D36DCEBFFF7A02EFD99FCE2AB568890" box="[516,539,875,899]" pageId="15" pageNumber="259" type="male">♂</specimenCount>
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,
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<collectionCode id="ED218FA7FFF7A02EFDB1FCE2AB298890" box="[556,612,875,899]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="15" pageNumber="259">CAS</collectionCode>
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)
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<collectingCountry id="F32757F2FFF7A02EFDE5FCE3AA4F8890" box="[632,770,874,899]" name="Madagascar" pageId="15" pageNumber="259">Madagascar</collectingCountry>
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||
. Moramanga district, Tamatave, Torotorofotsy,
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<quantity id="4CC8BA87FFF7A02EFAB2FCE2AC318890" box="[1327,1404,874,899]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="259" unit="km" value="22.0">22 km</quantity>
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N
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<location id="8EEF41B9FFF7A02EFF0AFC07A84588B4" LSID="urn:lsid:plazi:treatment:0399A674FFF4A02EFF0AF8BCADE48F48:8EEF41B9FFF7A02EFF0AFC07A84588B4" box="[151,264,910,935]" country="Madagascar" name="Andasibe." pageId="15" pageNumber="259">Andasibe.</location>
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18°46.25', 48°25.93', 23̄25.2014,
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<collectorName id="26C572B4FFF7A02EFD0BFC06AA7888B4" box="[662,821,910,935]" pageId="15" pageNumber="259">S.A. Marshall</collectorName>
|
||
, sequence
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||
<specimenCode id="DB96BF19FFF7A02EFC33FC07AD0788B4" box="[942,1098,910,935]" collectionCode="MYCRO" pageId="15" pageNumber="259">
|
||
<collectionCode id="ED218FA7FFF7A02EFC33FC07AD5288B4" box="[942,1055,910,935]" pageId="15" pageNumber="259">MYCRO</collectionCode>
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||
192
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||
</specimenCode>
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̄15.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C32A44E9FFF7A02EFF5AFC3BA8C18F24" pageId="15" pageNumber="259" type="discussion">
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||
<paragraph id="8B8F1762FFF7A02EFF5AFC3BA8C18F24" blockId="15.[151,1437,151,1115]" pageId="15" pageNumber="259">
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<emphasis id="B944CB70FFF7A02EFF5AFC3BA80288D8" bold="true" box="[199,335,946,971]" pageId="15" pageNumber="259">Comments.</emphasis>
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Although this species is known only from the
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<typeStatus id="548BA9C0FFF7A02EFCF4FC3BAA8688D8" box="[873,971,946,971]" pageId="15" pageNumber="259" type="holotype">holotype</typeStatus>
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male, the unusually extended postocular part of the head, the long parallel-sided process of the base of the genital fork, and the combination of a very long distal distiphallus and a very short basal basiphallus make
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<taxonomicName id="4C306CE1FFF7A02EFCF9FC72AD548F00" box="[868,1049,1019,1043]" class="Insecta" family="Micropezidae" genus="Paramimegralla" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="longicephala">
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||
<emphasis id="B944CB70FFF7A02EFCF9FC72AA3F8F01" box="[868,882,1019,1042]" italics="true" pageId="15" pageNumber="259">P</emphasis>
|
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.
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<emphasis id="B944CB70FFF7A02EFC1AFC72AD548F00" box="[903,1049,1019,1043]" italics="true" pageId="15" pageNumber="259">longicephala</emphasis>
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</taxonomicName>
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an easily recognized and well– characterized species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A44E9FFF7A02EFF5AFBCBADE48F48" box="[199,1193,1090,1115]" pageId="15" pageNumber="259" type="etymology">
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<paragraph id="8B8F1762FFF7A02EFF5AFBCBADE48F48" blockId="15.[151,1437,151,1115]" box="[199,1193,1090,1115]" pageId="15" pageNumber="259">
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||
<emphasis id="B944CB70FFF7A02EFF5AFBCBA80B8F48" bold="true" box="[199,326,1090,1115]" pageId="15" pageNumber="259">Etymology</emphasis>
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. The specific name refers to the exceptionally elongated head of this species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |