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<document id="83D2FAB4E911F5287F9A14B392F81A00" ID-DOI="10.5281/zenodo.183532" ID-GBIF-Dataset="90941c20-aece-4da0-b4bf-19e800996840" ID-ISSN="1175-5326" ID-Zenodo-Dep="183532" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459920883928" checkinUser="plazi" docAuthor="Short, Andrew Edward Z." docDate="2008" docId="03993C2AFFAD320BB8ACF918FC605DB9" docLanguage="en" docName="zt01855p055.pdf" docOrigin="Zootaxa 1855" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tritonus crenulatus Short, 2008, sp.n." docType="treatment" docVersion="7" lastPageNumber="49" masterDocId="FFA04452FFAA3203B83BFF8DFFC45B06" masterDocTitle="Revision of the Afrotropical genus Tritonus Mulsant (Coleoptera: Hydrophilidae)" masterLastPageNumber="55" masterPageNumber="41" pageNumber="48" updateTime="1698229214647" updateUser="plazi">
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<mods:titleInfo id="B70FE20847E46292F4B0E46BE7A6487D">
<mods:title id="6BB3B60B9D2EB09D56DDC7CECC9CF0B3">Revision of the Afrotropical genus Tritonus Mulsant (Coleoptera: Hydrophilidae)</mods:title>
</mods:titleInfo>
<mods:name id="5D2D429F36BC5187E7D48CB239264977" type="personal">
<mods:role id="EA82A8555E042B31C16A50D8C0AD9D08">
<mods:roleTerm id="4ED85224570CE1325FF5D63AD41CB9CA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="36FD719A38A486765D380075DC500384">Short, Andrew Edward Z.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1A8BDB358D5ADD7200103D06977F8659">text</mods:typeOfResource>
<mods:relatedItem id="4856A35E075A76C0B0336E223C4A3779" type="host">
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<mods:title id="EB6862456433397CA6F53D18B989E2E3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="918FE23B782ACACF615258E1B8CCDBB9">
<mods:date id="F270829C73ECF21B1537674E339B6CEA">2008</mods:date>
<mods:detail id="DEC9C9DF40C95E3ED29B6061B5E66F04" type="volume">
<mods:number id="049B2A3586CF57E07F5A9E0A9C4F1B99">1855</mods:number>
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<mods:classification id="73D5B5CBD8037C232CEDECAFD28AC993">journal article</mods:classification>
<mods:identifier id="12876F9FD8871F9005445F2ED8B386A1" type="DOI">10.5281/zenodo.183532</mods:identifier>
<mods:identifier id="557A2B208FFBE907456481E4BCB6AE7C" type="GBIF-Dataset">90941c20-aece-4da0-b4bf-19e800996840</mods:identifier>
<mods:identifier id="FE4ADB35F34245AB314C9BE0E77B9A36" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="994F42ED29DC81B8E66D85EFFE8934A7" type="Zenodo-Dep">183532</mods:identifier>
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<treatment id="03993C2AFFAD320BB8ACF918FC605DB9" ID-DOI="http://doi.org/10.5281/zenodo.6232623" ID-GBIF-Taxon="119373676" ID-Zenodo-Dep="6232623" LSID="urn:lsid:plazi:treatment:03993C2AFFAD320BB8ACF918FC605DB9" httpUri="http://treatment.plazi.org/id/03993C2AFFAD320BB8ACF918FC605DB9" lastPageId="8" lastPageNumber="49" pageId="7" pageNumber="48">
<subSubSection id="C32ADEB7FFAD3204B8ACF918FEF05DD4" pageId="7" pageNumber="48" type="nomenclature">
<paragraph id="8B8F8D3CFFAD3204B8ACF918FE0C5DA9" blockId="7.[151,456,1685,1746]" box="[151,456,1685,1711]" pageId="7" pageNumber="48">
<heading id="D0C73A50FFAD3204B8ACF918FE0C5DA9" bold="true" box="[151,456,1685,1711]" fontSize="11" level="1" pageId="7" pageNumber="48" reason="1">
<emphasis id="B944512EFFAD3204B8ACF918FE0C5DA9" bold="true" box="[151,456,1685,1711]" pageId="7" pageNumber="48">
<taxonomicName id="4C30F6BFFFAD3204B8ACF918FE425DA9" ID-CoL="7D87S" box="[151,390,1685,1711]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="crenulatus" status="sp. nov.">
<emphasis id="B944512EFFAD3204B8ACF918FE425DA9" bold="true" box="[151,390,1685,1711]" italics="true" pageId="7" pageNumber="48">Tritonus crenulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277EC55FFAD3204B9B6F918FE0C5DA9" box="[397,456,1685,1711]" pageId="7" pageNumber="48" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F8D3CFFAD3204B8ACF935FEF05DD4" blockId="7.[151,456,1685,1746]" box="[151,308,1720,1746]" pageId="7" pageNumber="48">
(
<figureCitation id="130B91B9FFAD3204B8A4F935FEC65DD4" box="[159,258,1720,1746]" captionStart="FIGURES 11 12" captionStartId="10.[151,269,1197,1221]" captionTargetBox="[151,1435,634,1171]" captionTargetId="figure@10.[151,1436,634,1171]" captionTargetPageId="10" captionText="FIGURES 11 12. Abdomen of Tritonus spp.; 11) T. riambavy; 12) T. crenulatus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/183538/files/figure.png" pageId="7" pageNumber="48">Figs. 12</figureCitation>
,
<figureCitation id="130B91B9FFAD3204B935F935FEE95DD4" box="[270,301,1720,1746]" captionStart="FIGURES 15 21" captionStartId="11.[151,269,1696,1720]" captionTargetBox="[242,1353,216,1672]" captionTargetId="figure@11.[234,1353,194,1672]" captionTargetPageId="11" captionText="FIGURES 15 21. Aedeagus of Tritonus spp., dorsal view; 15) T. riana; 16) T. steineri; 17) T. riambavy; 18) T. cribratus; 19) T. madagascarensis; 20) T. crenulatus; 21) T. complanatus. Scale bar = 0.45 mm." httpUri="https://zenodo.org/record/183540/files/figure.png" pageId="7" pageNumber="48">20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32ADEB7FFAD3204B8ACF88FFE3E5C6B" pageId="7" pageNumber="48" type="materials_examined">
<paragraph id="8B8F8D3CFFAD3204B8ACF88FFE3E5C6B" blockId="7.[151,1436,1794,2021]" pageId="7" pageNumber="48">
<emphasis id="B944512EFFAD3204B8ACF88FFE0D5C1A" bold="true" box="[151,457,1794,1820]" pageId="7" pageNumber="48">
<typeStatus id="548B339EFFAD3204B8ACF88FFF105C1A" box="[151,212,1794,1820]" pageId="7" pageNumber="48">Type</typeStatus>
Material.
<typeStatus id="548B339EFFAD3204B96CF88FFE0D5C1A" box="[343,457,1794,1820]" pageId="7" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
(male): “
<collectingCountry id="F327CDACFFAD3204BA0DF88EFCCD5C1B" box="[566,777,1795,1821]" name="Madagascar" pageId="7" pageNumber="48">MADAGASCAR</collectingCountry>
/ Antananarivo/ Manakambahiny/
<date id="FF8EABFCFFAD3204BCA7F88EFAAF5C1B" box="[1180,1387,1795,1821]" pageId="7" pageNumber="48" value="1990-02-15">15 February 1990</date>
/ W. E. Steiner”, “In seepage over/ exposed granite/ cliff face”. (
<collectionCode id="ED2115F9FFAD3204BB61F8A6FC7E5C43" box="[858,954,1835,1861]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="48">NMNH</collectionCode>
).
<emphasis id="B944512EFFAD3204BBF4F8A7FB495C42" bold="true" box="[975,1165,1834,1860]" pageId="7" pageNumber="48">
<typeStatus id="548B339EFFAD3204BBF4F8A7FB885C42" box="[975,1100,1834,1860]" pageId="7" pageNumber="48" type="paratype">Paratypes</typeStatus>
(16).
</emphasis>
same data as
<typeStatus id="548B339EFFAD3204BD0EF8A6FA5F5C43" box="[1333,1435,1835,1861]" pageId="7" pageNumber="48" type="holotype">holotype</typeStatus>
(16,
<collectionCode id="ED2115F9FFAD3204B8F7F8DEFEED5C6B" box="[204,297,1875,1901]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="48">NMNH</collectionCode>
,
<collectionCode id="ED2115F9FFAD3204B90EF8DEFE4F5C6B" box="[309,395,1875,1901]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="48">NMPC</collectionCode>
,
<collectionCode id="ED2115F9FFAD3204B9ACF8DEFE2A5C6B" box="[407,494,1875,1901]" pageId="7" pageNumber="48">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C32ADEB7FFAD3204B8FEF8F7FE775CE3" pageId="7" pageNumber="48" type="diagnosis">
<paragraph id="8B8F8D3CFFAD3204B8FEF8F7FE775CE3" blockId="7.[151,1436,1794,2021]" pageId="7" pageNumber="48">
<emphasis id="B944512EFFAD3204B8FEF8F7FE815C92" bold="true" box="[197,325,1914,1940]" pageId="7" pageNumber="48">Diagnosis.</emphasis>
The coarsely crenulate anterior margins of abdominal ventrites 2 and 3 will serve to distinguish this species from all others. It is very similar to the slightly smaller
<taxonomicName id="4C30F6BFFFAD3204BC39F82EFB985CBA" box="[1026,1116,1955,1980]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="riana">
<emphasis id="B944512EFFAD3204BC39F82EFB985CBA" box="[1026,1116,1955,1980]" italics="true" pageId="7" pageNumber="48">T. riana</emphasis>
</taxonomicName>
, but this species has much less coarse crenulations.
</paragraph>
</subSubSection>
<caption id="DF4FDDB4FFA2320BB8ACFCAAFB9C5867" httpUri="https://zenodo.org/record/183536/files/figure.png" pageId="8" pageNumber="49" targetBox="[151,1435,178,783]" targetPageId="8">
<paragraph id="8B8F8D3CFFA2320BB8ACFCAAFB9C5867" blockId="8.[151,1435,807,865]" pageId="8" pageNumber="49">
<emphasis id="B944512EFFA2320BB8ACFCAAFE875839" bold="true" box="[151,323,807,831]" pageId="8" pageNumber="49">FIGURES 68.</emphasis>
SEMs of thoracic sterna of
<taxonomicName id="4C30F6BFFFA2320BBA4AFCA5FD0E5839" box="[625,714,808,831]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA2320BBA4AFCA5FD0E5839" box="[625,714,808,831]" italics="true" pageId="8" pageNumber="49">Tritonus</emphasis>
</taxonomicName>
species; 6) meso- and metaventrite of
<taxonomicName id="4C30F6BFFFA2320BBC52FCA5FAC85839" box="[1129,1292,808,831]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="complanatus">
<emphasis id="B944512EFFA2320BBC52FCA5FAC85839" box="[1129,1292,808,831]" italics="true" pageId="8" pageNumber="49">T. complanatus</emphasis>
</taxonomicName>
, scale bar= 1 mm; 7) prosternum of
<taxonomicName id="4C30F6BFFFA2320BB9B3FCC7FDEE5867" box="[392,554,842,865]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="complanatus">
<emphasis id="B944512EFFA2320BB9B3FCC7FDEE5867" box="[392,554,842,865]" italics="true" pageId="8" pageNumber="49">T. complanatus</emphasis>
</taxonomicName>
, 8) prosternum of
<taxonomicName id="4C30F6BFFFA2320BBAC9FCC7FCAB5867" box="[754,879,842,865]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="riambavy">
<emphasis id="B944512EFFA2320BBAC9FCC7FCAB5867" box="[754,879,842,865]" italics="true" pageId="8" pageNumber="49">T. riambavy</emphasis>
</taxonomicName>
. Scale bars= 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C32ADEB7FFA2320BB8FEFC19FD015D49" pageId="8" pageNumber="49" type="description">
<paragraph id="8B8F8D3CFFA2320BB8FEFC19FD015D49" blockId="8.[151,1437,916,1727]" pageId="8" pageNumber="49">
<emphasis id="B944512EFFA2320BB8FEFC19FE9958A8" bold="true" box="[197,349,916,942]" pageId="8" pageNumber="49">Description.</emphasis>
Total length
<quantity id="4CC820D9FFA2320BB9C2FC18FD4F58A9" box="[505,651,917,943]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="7.6" metricValueMin="7.1" pageId="8" pageNumber="49" unit="mm" value="7.35" valueMax="7.6" valueMin="7.1">7.17.6 mm</quantity>
. Color: Dorsum black with general shagreen. Pronotum with a few dark black patches around margins. Elytra with indistinct green iridescent patches along lateral margins.
<emphasis id="B944512EFFA2320BB8ACFC52FF2558FF" bold="true" box="[151,225,991,1017]" pageId="8" pageNumber="49">Head.</emphasis>
Ground punctation fine but distinct; irregular in size. Systematic punctures distinct; 3× as large as largest surrounding ground punctures; on clypeus bearing short decumbent setae; those on frons anterior to each eye relatively long and erect. Antennae with club distinctly shorter in length than antennomeres 26 combined. Maxillary palpi short, apical segment ca. 1.2× length of penultimate segment; penultimate segment and segment 2 subequal in length. Mentum with anterior margin slightly convex; anterior half with fine to very fine, transverse microsculpture; entire mentum with fine, irregularly scattered punctures. Anterior margin of head moderately wide and broadly rounded.
<emphasis id="B944512EFFA2320BBA9AFB32FCC15FDF" bold="true" box="[673,773,1215,1241]" pageId="8" pageNumber="49">Thorax.</emphasis>
Ground punctation on pronotum and elytra fine, slightly finer than ground punctation on head. Pronotal and elytral systematic punctures large and distinct; bearing short setae. Prosternum slightly tectiform with a sharp, narrow and glabrous median carina; not distinctly elevated or dilated at anterior margin. Mesoventrite elevated medially into a carina, with anterior half slightly more elevated than posterior half; surface of carina narrowly glabrous. Metasternum with large elevated glabrous region that half as wide as long. Metafemora glabrous and shiny on basal third; apical two thirds dull, with moderately impressed microsculpture in the form of fine transverse wavy lines.
<emphasis id="B944512EFFA2320BBCB7FA12FACA5EBF" bold="true" box="[1164,1294,1439,1465]" pageId="8" pageNumber="49">Abdomen.</emphasis>
Posteromedial glabrous region of ventrite 1 moderately small, slightly more than half as wide as glabrous region on ventrite 2. Glabrous region of ventrite hemi-ovoid to subtriangular in shape; about one third the width of the ventrite. Anterior margin of ventrite 2 coarsely and deeply crenulate; margins of ventrites 1, 3 and 4 weakly but distinctly crenulate. Aedeagus as in
<figureCitation id="130B91B9FFA2320BBA51F9B8FD055D49" box="[618,705,1589,1615]" captionStart="FIGURES 15 21" captionStartId="11.[151,269,1696,1720]" captionTargetBox="[242,1353,216,1672]" captionTargetId="figure@11.[234,1353,194,1672]" captionTargetPageId="11" captionText="FIGURES 15 21. Aedeagus of Tritonus spp., dorsal view; 15) T. riana; 16) T. steineri; 17) T. riambavy; 18) T. cribratus; 19) T. madagascarensis; 20) T. crenulatus; 21) T. complanatus. Scale bar = 0.45 mm." httpUri="https://zenodo.org/record/183540/files/figure.png" pageId="8" pageNumber="49">Fig. 20</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32ADEB7FFA2320BB8FEF9D7FEBC5D9F" pageId="8" pageNumber="49" type="etymology">
<paragraph id="8B8F8D3CFFA2320BB8FEF9D7FEBC5D9F" blockId="8.[151,1437,916,1727]" pageId="8" pageNumber="49">
<emphasis id="B944512EFFA2320BB8FEF9D7FE965D72" bold="true" box="[197,338,1626,1652]" pageId="8" pageNumber="49">Etymology.</emphasis>
Named for the coarsely crenulate margins of abdominal ventrites 2 and 3, which are distinctive of this species.
</paragraph>
</subSubSection>
<subSubSection id="C32ADEB7FFA2320BB8FEF929FC605DB9" box="[197,932,1700,1727]" pageId="8" pageNumber="49" type="distribution">
<paragraph id="8B8F8D3CFFA2320BB8FEF929FC605DB9" blockId="8.[151,1437,916,1727]" box="[197,932,1700,1727]" pageId="8" pageNumber="49">
<emphasis id="B944512EFFA2320BB8FEF929FEA25DB8" bold="true" box="[197,358,1700,1726]" pageId="8" pageNumber="49">Distribution.</emphasis>
<collectingCountry id="F327CDACFFA2320BB956F928FE395DB9" box="[365,509,1701,1727]" name="Madagascar" pageId="8" pageNumber="49">Madagascar</collectingCountry>
. Known only from the
<typeStatus id="548B339EFFA2320BBB37F928FCFA5DB9" box="[780,830,1701,1727]" pageId="8" pageNumber="49">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>