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<document id="B4EC516BDEC0BE507F629EA02A89F104" ID-DOI="10.1206/0003-0090.467.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="12171703" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718827875798" checkinUser="felipe" docAuthor="Criscione-Vastano, Julia &amp; Grimaldi, David A." docDate="2024" docId="0398F72AFF8DFFF587C3F521A557BF7E" docLanguage="en" docName="BulAmeMusNatHis.2024.467.1-137.pdf" docOrigin="Bulletin of the American Museum of Natural History 2024 (467)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-2024/issue-467/0003-0090.467.1.1/Remarkable-Diversity-of-Beetles-Coleoptera-in-the-Late-Triassic-Norian/10.1206/0003-0090.467.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Holcoptera solitensis Thomson 2017" docType="treatment" docVersion="3" lastPageNumber="24" masterDocId="FFA18F52FF9AFFED8512F11AA730BB08" masterDocTitle="Remarkable Diversity Of Beetles (Coleoptera) In The Late Triassic (Norian) “ Solite Deposit ” Of Virginia And North Carolina" masterLastPageNumber="141" masterPageNumber="1" pageNumber="23" updateTime="1718834141660" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EE047001F54EB5EBCC023E29028CDFCA">Remarkable Diversity Of Beetles (Coleoptera) In The Late Triassic (Norian) “ Solite Deposit ” Of Virginia And North Carolina</mods:title>
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<mods:namePart id="40F6FC9AC74D264AC7C0D45FE3546381">Criscione-Vastano, Julia</mods:namePart>
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<mods:namePart id="FDD23A28AE9D58771837E6A5C6DC5A7E">Grimaldi, David A.</mods:namePart>
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<mods:date id="66BA8AAB628DEB1AA25281A1A105E334">2024</mods:date>
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<mods:number id="6081DABDA4A09DD11EE77216FF560993">2024-05-31</mods:number>
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<mods:url id="87664EB9808554D40421700D9F2E1EC7">https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-2024/issue-467/0003-0090.467.1.1/Remarkable-Diversity-of-Beetles-Coleoptera-in-the-Late-Triassic-Norian/10.1206/0003-0090.467.1.1.full</mods:url>
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<mods:identifier id="C2CE1290F1D42D1B34267ACF4008180B" type="DOI">10.1206/0003-0090.467.1.1</mods:identifier>
<mods:identifier id="CB8E97CFB317CC0D21730C7949AD861F" type="ISSN">0003-0090</mods:identifier>
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<treatment id="0398F72AFF8DFFF587C3F521A557BF7E" ID-DOI="http://doi.org/10.5281/zenodo.12173733" ID-Zenodo-Dep="12173733" LSID="urn:lsid:plazi:treatment:0398F72AFF8DFFF587C3F521A557BF7E" httpUri="http://treatment.plazi.org/id/0398F72AFF8DFFF587C3F521A557BF7E" lastPageId="24" lastPageNumber="24" pageId="23" pageNumber="23">
<subSubSection id="C32B15B7FF8DFFFA87C3F521A3A7BF5B" box="[721,1175,1083,1107]" pageId="23" pageNumber="23" type="nomenclature">
<paragraph id="8B8E463CFF8DFFFA87C3F521A3A7BF5B" blockId="23.[721,1175,1083,1107]" box="[721,1175,1083,1107]" pageId="23" pageNumber="23">
<heading id="D0C6F150FF8DFFFA87C3F521A3A7BF5B" box="[721,1175,1083,1107]" centered="true" fontSize="9" level="2" pageId="23" pageNumber="23" reason="2">
<taxonomicName id="4C313DBFFF8DFFFA87C3F521A3A7BF5B" authority="Thomson et al., 2017" authorityName="Thomson" authorityYear="2017" box="[721,1175,1083,1107]" class="Insecta" family="Coptoclavidae" genus="Holcoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="23" phylum="Arthropoda" rank="species" species="solitensis">
<emphasis id="B9459A2EFF8DFFFA87C3F521A49ABF5B" box="[721,938,1083,1107]" italics="true" pageId="23" pageNumber="23">Holcoptera solitensis</emphasis>
<bibRefCitation id="EFA03BCDFF8DFFFA86A0F521A3A7BF5B" author="Thomson, U. &amp; A. J. Ross &amp; P. Davidson" box="[946,1175,1083,1107]" pageId="23" pageNumber="23" pagination="659 - 674" refId="ref42846" refString="Thomson, U., A. J. Ross, and P. Davidson. 2017. Mesozoic Holcoptera (Coleoptera: Coptoclavidae) from England and the United States. Proceedings of the Geologists' Association 128 (4): 659 - 674." type="journal article" year="2017">Thomson et al., 2017</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B15B7FF8DFFFA865CF575A31EBDC2" pageId="23" pageNumber="23" type="description">
<paragraph id="8B8E463CFF8DFFFA865CF575A329BF8F" blockId="23.[846,1049,1135,1159]" box="[846,1049,1135,1159]" pageId="23" pageNumber="23">
<heading id="D0C6F150FF8DFFFA865CF575A329BF8F" box="[846,1049,1135,1159]" centered="true" fontSize="9" level="2" pageId="23" pageNumber="23" reason="2">
<figureCitation id="130A5AB9FF8DFFFA865CF575A4C5BF8F" box="[846,1013,1135,1159]" captionStart="FIG" captionStartId="75.[108,150,1674,1695]" captionTargetBox="[108,1212,224,1648]" captionTargetId="figure-10@75.[108,1212,224,1648]" captionTargetPageId="75" captionText="FIG. 37. Holcoptera solitensis. A, B. VMNH 97986. C. VMNH 96762 with partial head. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12171792" httpUri="https://zenodo.org/record/12171792/files/figure.png" pageId="23" pageNumber="23">Figures 37A–C</figureCitation>
,
<figureCitation id="130A5AB9FF8DFFFA86EDF575A329BF8F" box="[1023,1049,1135,1159]" captionStart="FIG" captionStartId="76.[109,151,1677,1698]" captionTargetBox="[106,1210,224,1648]" captionTargetId="figure-10@76.[106,1210,224,1648]" captionTargetPageId="76" captionText="FIG. 38. Holcoptera sp., AMNH 04-96. Note width of second innermost longitudinal bands compared to those of VMNH 97986 and 96762 (fig. 37). Specimen possibly belonging to a separate holcopteran species. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12171794" httpUri="https://zenodo.org/record/12171794/files/figure.png" pageId="23" pageNumber="23">38</figureCitation>
</heading>
</paragraph>
<paragraph id="8B8E463CFF8DFFFA87DEF5BBA31EBDC2" blockId="23.[684,1213,1185,1738]" pageId="23" pageNumber="23">
DESCRIPTION: Small beetle (
<quantity id="4CC9EBD9FF8DFFFA811BF5BBA352BFB0" box="[1033,1122,1185,1209]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="23" pageNumber="23" unit="mm" value="4.7">4.7 mm</quantity>
long as preserved,
<quantity id="4CC9EBD9FF8DFFFA860CF5D8A45FBFD1" box="[798,879,1218,1242]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="23" pageNumber="23" unit="mm" value="2.8">2.8 mm</quantity>
wide) preserved in dorsal view; preservation fair; no counterpart. Peculiar specimen with body margins poorly defined, head missing. Elytra and some underlying structures preserved. Anterior portion of specimen with two symmetrical, raised circular structures laterally. Elytra with distinctive color pattern consisting of four longitudinal bands: two outer bands connecting posteriorly to form U-shape, innermost band thinner than others and extending to a color patch along the anterior margin where the orientation becomes perpendicular to the other bands. Outer elytral margins not preserved. Abdominal segments visible beneath elytra, four subequal tergites discernible; additional faint bands laterally, possibly outer margins of abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C32B15B7FF82FFF5859EF1F8A55FBAC8" pageId="24" pageNumber="24" type="materials_examined">
<paragraph id="8B8E463CFF82FFF5859EF1F8A55FBAC8" blockId="24.[108,637,226,1142]" pageId="24" pageNumber="24">
MATERIAL: Illustration and description based on
<collectionCode id="ED20DEF9FF82FFF58583F019A7D5BA12" box="[145,229,259,282]" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="24" pageNumber="24">VMNH</collectionCode>
97986. Twenty-five additional specimens are known:
<collectionCode id="ED20DEF9FF82FFF5844CF03EA684BA34" box="[350,436,292,316]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="24">AMNH</collectionCode>
04-96,
<collectionCode id="ED20DEF9FF82FFF58737F03EA54BBA33" box="[549,635,292,315]" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="24" pageNumber="24">VMNH</collectionCode>
49735/49736, 51498, 51511, 51839, 54135, 54136, 96728, 96729, 96731, 96736, 96762, 96765, 96766, 96768, 96769, 96782, 96784, 96787, 96789, 96791, 97234, 97243, 97247, and 97251.
</paragraph>
</subSubSection>
<subSubSection id="C32B15B7FF82FFF5859EF0D3A557BF7E" pageId="24" pageNumber="24" type="discussion">
<paragraph id="8B8E463CFF82FFF5859EF0D3A6A1B824" blockId="24.[108,637,226,1142]" pageId="24" pageNumber="24">
COMMENTS: There are 26 known specimens in the Solite material sharing this distinctive elytral pattern. Seven of these consist of more than a single elytron, and
<specimenCount id="9D378DB5FF82FFF585EFF337A6A6B94C" box="[253,406,556,580]" count="1" pageId="24" pageNumber="24" type="generic">one specimen</specimenCount>
(VMNH 96762, fig. 37C) includes the head, though it is not attached to the body. The illustrated specimen (VMNH 97986) appears shorter than the others and is possibly longitudinally distorted. Additionally,
<specimenCount id="9D378DB5FF82FFF584F4F3ABA54CB9C0" box="[486,636,688,712]" count="1" pageId="24" pageNumber="24" type="generic">one specimen</specimenCount>
(AMNH 04-96, fig. 38) differs in the thickness of the second innermost longitudinal band and might represent a separate species.
</paragraph>
<paragraph id="8B8E463CFF82FFF5859EF22FA557BF7E" blockId="24.[108,637,226,1142]" pageId="24" pageNumber="24">
This morphotype was previously described as
<taxonomicName id="4C313DBFFF82FFF5857EF24CA67DB866" box="[108,333,854,878]" class="Insecta" family="Coptoclavidae" genus="Holcoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="solitensis">
<emphasis id="B9459A2EFF82FFF5857EF24CA67DB866" box="[108,333,854,878]" italics="true" pageId="24" pageNumber="24">Holcoptera solitensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFA03BCDFF82FFF5846FF24CA54CB866" author="Thomson, U. &amp; A. J. Ross &amp; P. Davidson" box="[381,636,854,878]" pageId="24" pageNumber="24" pagination="659 - 674" refId="ref42846" refString="Thomson, U., A. J. Ross, and P. Davidson. 2017. Mesozoic Holcoptera (Coleoptera: Coptoclavidae) from England and the United States. Proceedings of the Geologists' Association 128 (4): 659 - 674." type="journal article" year="2017">Thomson et al. (2017)</bibRefCitation>
based on two isolated elytra. It should be noted that the original description states that
<taxonomicName id="4C313DBFFF82FFF58739F283A797B8D9" class="Insecta" family="Coptoclavidae" genus="Holcoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="solitensis">
<emphasis id="B9459A2EFF82FFF58739F283A797B8D9" italics="true" pageId="24" pageNumber="24">H. solitensis</emphasis>
</taxonomicName>
specimens “are the only ones seen where none of the [elytral color] bands connect, either to a color patch at the base or distally” (
<bibRefCitation id="EFA03BCDFF82FFF58701F2E1A618BF3C" author="Thomson, U. &amp; A. J. Ross &amp; P. Davidson" pageId="24" pageNumber="24" pagination="659 - 674" refId="ref42846" refString="Thomson, U., A. J. Ross, and P. Davidson. 2017. Mesozoic Holcoptera (Coleoptera: Coptoclavidae) from England and the United States. Proceedings of the Geologists' Association 128 (4): 659 - 674." type="journal article" year="2017">Thomson et al., 2017: 671</bibRefCitation>
). The two outer bands of the Solite specimens do, in fact, connect distally, contradicting the species’ original description.
</paragraph>
</subSubSection>
</treatment>
</document>
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