500 lines
56 KiB
XML
500 lines
56 KiB
XML
<document id="DAE266D59C9824BA106F047B02A5450C" ID-CLB-Dataset="29875" ID-DOI="10.11646/zootaxa.4434.3.3" ID-GBIF-Dataset="3ebb3cb7-3b8f-4c4d-8858-d37185292c28" ID-ISSN="1175-5326" ID-Zenodo-Dep="1292189" ID-ZooBank="8B95D692-E8F7-402B-8485-3D3AE20111E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529330577762" checkinUser="plazi" docAuthor="Seeman, Owen D., Minor, Maria, Baker, Michelle R. & Walter, David Evans" docDate="2018" docId="0398D71B524A73515FFAFD313C4AE94D" docLanguage="en" docName="zootaxa.4434.3.3.pdf.imf" docOrigin="Zootaxa 4434 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heatherella osleri Seeman & Minor & Baker & Walter 2018, n. sp." docType="treatment" docUuid="66DFC33A-FA01-4994-8C2C-42B4E7BF1A30" docUuidSource="ZooBank" docVersion="8" lastPageNumber="450" masterDocId="FFA1AF63524973585F6DFFFA3F4BE851" masterDocTitle="A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia" masterLastPageNumber="465" masterPageNumber="441" pageNumber="444" updateTime="1698549773811" updateUser="ExternalLinkService">
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<mods:mods id="12DFDF73CBB5F4E423AD2B02C6979DBC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AFE79E5E9DED16E2034DE5DC2A44B2B0">
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<mods:title id="5B0E313A8394EF866806ED0DB8F5B51F">A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia</mods:title>
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<mods:name id="A6B1DB8E17740171A219FEF0EF17975D" type="personal">
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<mods:namePart id="622E50A49658401E38B9F0A0413DC399">Seeman, Owen D.</mods:namePart>
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<mods:roleTerm id="ABAF2F74DF1E45D7462B76B752D9F24A">Author</mods:roleTerm>
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<mods:namePart id="79828529E9056CF08F9D19FEA13298BB">Minor, Maria</mods:namePart>
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<mods:name id="18CA4832F91AC773FBA4F8398C1C5C04" type="personal">
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<mods:namePart id="01E5C677BBA0A828EB1BC01294A4FABF">Baker, Michelle R.</mods:namePart>
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<mods:name id="B337FCCDEC74C04C4A0D2288C95994C0" type="personal">
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<mods:roleTerm id="5877DDEC13C7DF35CE87F30F12926177">Author</mods:roleTerm>
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<mods:namePart id="E6D88AE0158EE9DDDDCCFDE7A304E511">Walter, David Evans</mods:namePart>
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<mods:typeOfResource id="0ACF9ABD4168595A3B5B333EBC968753">text</mods:typeOfResource>
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<mods:title id="E8A71739D4D387C09148DEAAA840A920">Zootaxa</mods:title>
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<mods:part id="197F861099C55EB6338A3B6E066855F9">
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<mods:date id="A896E3373ACD91B22D6136D02A48F16F">2018</mods:date>
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<mods:number id="439B67BEE2468DD075EEB2ADD29EE02F">2018-06-18</mods:number>
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<mods:number id="BC6C8CEC82CD05EE0F63C264D6E0E958">4434</mods:number>
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<mods:classification id="D7DD7F84FD97ED164FD8EC80119DD572">journal article</mods:classification>
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<treatment id="0398D71B524A73515FFAFD313C4AE94D" ID-DOI="http://doi.org/10.5281/zenodo.5966633" ID-GBIF-Taxon="145968987" ID-Zenodo-Dep="5966633" LSID="urn:lsid:zoobank.org:act:66DFC33A-FA01-4994-8C2C-42B4E7BF1A30" httpUri="http://treatment.plazi.org/id/0398D71B524A73515FFAFD313C4AE94D" lastPageId="9" lastPageNumber="450" pageId="3" pageNumber="444">
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<subSubSection id="C32B3586524A735B5FFAFD313E08EB56" pageId="3" pageNumber="444" type="nomenclature">
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<paragraph id="8B8E660D524A735B5FFAFD313C1DEAB7" blockId="3.[151,854,715,775]" box="[151,854,715,742]" pageId="3" pageNumber="444">
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<heading id="D0C6D161524A735B5FFAFD313C1DEAB7" bold="true" box="[151,854,715,742]" fontSize="11" level="1" pageId="3" pageNumber="444" reason="1">
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<emphasis id="B945BA1F524A735B5FFAFD313C1DEAB7" bold="true" box="[151,854,715,742]" pageId="3" pageNumber="444">
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<taxonomicName id="4C311D8E524A735B5FFAFD313E3BEAB4" authority="Seeman & Minor & Baker & Walter, 2018" authorityName="Seeman & Minor & Baker & Walter" authorityYear="2018" box="[151,368,715,741]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="osleri" status="sp. nov.">
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<emphasis id="B945BA1F524A735B5FFAFD313E3BEAB4" bold="true" box="[151,368,715,741]" italics="true" pageId="3" pageNumber="444">Heatherella osleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2760764524A735B5E1AFD363EF1EAB7" box="[375,442,716,742]" pageId="3" pageNumber="444" rank="species">n. sp.</taxonomicNameLabel>
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Seeman, Minor, Baker & Walter
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8E660D524A735B5FFAFD143E08EB56" blockId="3.[151,854,715,775]" box="[151,323,750,775]" pageId="3" pageNumber="444">
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(
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<figureCitation id="130A7A88524A735B5FF2FD143E5AEB56" box="[159,273,750,775]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURES 3–8" captionStart-3="FIGURES 9–11" captionStart-4="FIGURES 12–14" captionStartId-0="4.[151,250,1999,2021]" captionStartId-1="5.[151,250,1997,2019]" captionStartId-2="6.[151,264,1953,1975]" captionStartId-3="7.[151,264,1236,1258]" captionStartId-4="8.[151,264,1439,1461]" captionTargetBox-0="[158,1436,230,1948]" captionTargetBox-1="[201,1387,277,1878]" captionTargetBox-2="[252,1329,209,1919]" captionTargetBox-3="[174,1415,202,1206]" captionTargetBox-4="[213,1369,199,1413]" captionTargetId-0="figure@4.[151,1436,227,1948]" captionTargetId-1="figure@5.[151,1436,265,1888]" captionTargetId-2="figure@6.[246,1341,193,1932]" captionTargetId-3="figure@7.[164,1424,193,1214]" captionTargetId-4="figure@8.[212,1376,185,1424]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 1. Heatherella osleri n. sp. female holotype, venter." captionText-1="FIGURE 2. Heatherella osleri n. sp. female holotype, dorsum." captionText-2="FIGURES 3–8. Heatherella osleri n. sp. (3) female, ventral gnathosoma (seta h3 eXpressed asymmetrically in holotype); (4) female, gnathotectum; (5) male, gnathotectum; (6) female, chelicera; (7) male, chelicera; (8) female, leg I." captionText-3="FIGURES 9–11. Heatherella osleri n. sp. female holotype, posterior views: (9) leg II; (10) leg III; (11) leg IV." captionText-4="FIGURES 12–14. Heatherella osleri n. sp. male (12) venter; (13) dorsum (marginal shields not shown laterally; torn anteriorly from marginal shield); (14) leg II, anterior view." httpUri-0="https://zenodo.org/record/1292191/files/figure.png" httpUri-1="https://zenodo.org/record/1292193/files/figure.png" httpUri-2="https://zenodo.org/record/1292195/files/figure.png" httpUri-3="https://zenodo.org/record/1292197/files/figure.png" httpUri-4="https://zenodo.org/record/1292199/files/figure.png" pageId="3" pageNumber="444">Figs 1–14</figureCitation>
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,
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<figureCitation id="130A7A88524A735B5E70FD143E71EB56" box="[285,314,750,775]" captionStart="FIGURES 34–37" captionStartId="22.[151,264,1832,1854]" captionTargetBox="[326,1233,232,1771]" captionTargetId="figure@22.[309,1273,196,1784]" captionTargetPageId="22" captionText="FIGURES 34–37. (34) Heatherella acanthocharis walter, 1997, dorsomarginal protuberances, pustules and pores of paratype female (P-s5 pustule; P-s6 pore; P-S1 protuberance), dorsal view of flattened specimen, right side; (35) Aheatherella mira n. sp., dorsomarginal protuberances, pustules and pores of paratype female (P-s5 pustule; P-s6 protuberance with closely associated seta), dorsal view of ventrolateral fragment with broken r-setae, left side; (36) Heatherella osleri n. sp., peritreme of paratype female (ventral view, right peritreme of mite); (37) Aheatherella mira n. sp. peritreme of paratype female (ventral view, left peritreme of mite; sclerotised eXopodal portion of eXopodal-peritrematal shield shaded)." httpUri="https://zenodo.org/record/1292223/files/figure.png" pageId="3" pageNumber="444">36</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B3586524A735B5FFAFCCC3C6AEC3E" pageId="3" pageNumber="444" type="diagnosis">
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<paragraph id="8B8E660D524A735B5FFAFCCC3BF9EC77" blockId="3.[151,1437,822,1999]" pageId="3" pageNumber="444">
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<emphasis id="B945BA1F524A735B5FFAFCCC3E41EB1E" bold="true" box="[151,266,822,847]" pageId="3" pageNumber="444">Diagnosis</emphasis>
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: With the characteristics of the genus. Adult: sternal shield (bearing
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<emphasis id="B945BA1F524A735B5B33FCCD3B34EB1E" box="[1118,1151,823,847]" italics="true" pageId="3" pageNumber="444">st2</emphasis>
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) subhexagonal, without endopodal elements, poorly differentiated anteriorly; ventral setae all short, bulbous and barbed; dorsal and marginal setae with ciliate crests; dorsal shield heavily ornamented; idiosomal margin with ten pairs of idiosomatic protuberances (protuberances
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<emphasis id="B945BA1F524A735B5E87FC5E3D6FEBEA" box="[490,548,931,955]" italics="true" pageId="3" pageNumber="444">P - s4</emphasis>
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and
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<emphasis id="B945BA1F524A735B5D30FC5E3DDAEBEA" box="[605,657,930,955]" italics="true" pageId="3" pageNumber="444">P -s6</emphasis>
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present), distinct pustule associated with seta
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<emphasis id="B945BA1F524A735B5BF1FC593BFFEBEB" box="[1180,1204,931,954]" italics="true" pageId="3" pageNumber="444">s5</emphasis>
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absent,
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<emphasis id="B945BA1F524A735B5A7EFC5E3A0DEBEB" box="[1299,1350,931,955]" italics="true" pageId="3" pageNumber="444">P -s5</emphasis>
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similar to other protuberances; dorsum with seta
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<emphasis id="B945BA1F524A735B5D0EFC3D3D30EB8E" box="[611,635,967,991]" italics="true" pageId="3" pageNumber="444">z2</emphasis>
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; seta
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<emphasis id="B945BA1F524A735B5DD1FC3C3D9EEB8E" box="[700,725,966,991]" italics="true" pageId="3" pageNumber="444">z6</emphasis>
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anterior to
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<emphasis id="B945BA1F524A735B5C38FC3C3C24EB8E" box="[853,879,966,991]" italics="true" pageId="3" pageNumber="444">s6</emphasis>
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, seta
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<emphasis id="B945BA1F524A735B5CC3FC3D3C80EB8E" box="[942,971,967,991]" italics="true" pageId="3" pageNumber="444">Z1</emphasis>
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anterior to
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<emphasis id="B945BA1F524A735B5B21FC3D3B23EB8E" box="[1100,1128,967,991]" italics="true" pageId="3" pageNumber="444">S1</emphasis>
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, seta
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<emphasis id="B945BA1F524A735B5BCAFC3D3B8BEB8F" box="[1191,1216,967,990]" italics="true" pageId="3" pageNumber="444">z5</emphasis>
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placed obliquely to seta
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<emphasis id="B945BA1F524A735B5FA3FC113FAFEC53" box="[206,228,1003,1026]" italics="true" pageId="3" pageNumber="444">j5</emphasis>
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; marginal
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<emphasis id="B945BA1F524A735B5E08FC163EC1EC52" box="[357,394,1003,1027]" italics="true" pageId="3" pageNumber="444">r -R</emphasis>
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series setae not duplicated, 25–28 setae on each side, sometimes asymmetrical. Female: female with
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<emphasis id="B945BA1F524A735B5E4EFBF53E0FEC76" box="[291,324,1039,1063]" italics="true" pageId="3" pageNumber="444">
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st
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<quantity id="4CC9CBE8524A735B5E5BFBF53E0FEC76" box="[310,324,1039,1063]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="444" unit="in" value="3.0">3</quantity>
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</emphasis>
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in soft cuticle and
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<emphasis id="B945BA1F524A735B5D74FBF53D71EC76" box="[537,570,1039,1063]" italics="true" pageId="3" pageNumber="444">st4</emphasis>
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on small shields. Male: seta
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<emphasis id="B945BA1F524A735B5C13FBF53CEDEC76" box="[894,934,1039,1063]" italics="true" pageId="3" pageNumber="444">av1</emphasis>
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on genu II a blunt spur.
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</paragraph>
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<paragraph id="8B8E660D524A735B5FAAFBC83C6AEC3E" blockId="3.[151,1437,822,1999]" pageId="3" pageNumber="444">
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<emphasis id="B945BA1F524A735B5FAAFBC83E81EC1A" bold="true" box="[199,458,1074,1099]" pageId="3" pageNumber="444">Idiosomal dimensions</emphasis>
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: Females (n= 3) 360–375 long x 255–265 wide (
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<typeStatus id="548AD8AF524A735B5C9CFBC83B1EEC1A" box="[1009,1109,1074,1099]" pageId="3" pageNumber="444" type="holotype">holotype</typeStatus>
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360 x 265, small lateral split on one side); males (n= 9) 345–370 long x 220–255 wide.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B3586524A735F5FAAFB803BD4EFE3" lastPageId="7" lastPageNumber="448" pageId="3" pageNumber="444" type="description">
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<paragraph id="8B8E660D524A735B5FAAFB803DDCEE4E" blockId="3.[151,1437,822,1999]" pageId="3" pageNumber="444">
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<emphasis id="B945BA1F524A735B5FAAFB803E6EECC2" bold="true" box="[199,293,1146,1171]" pageId="3" pageNumber="444">Female.</emphasis>
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<emphasis id="B945BA1F524A735B5E40FB863E3EECC2" box="[301,373,1148,1171]" italics="true" pageId="3" pageNumber="444">Venter</emphasis>
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(
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<figureCitation id="130A7A88524A735B5EE8FB803E83ECC2" box="[389,456,1146,1171]" captionStart="FIGURE 1" captionStartId="4.[151,250,1999,2021]" captionTargetBox="[158,1436,230,1948]" captionTargetId="figure@4.[151,1436,227,1948]" captionTargetPageId="4" captionText="FIGURE 1. Heatherella osleri n. sp. female holotype, venter." httpUri="https://zenodo.org/record/1292191/files/figure.png" pageId="3" pageNumber="444">Fig. 1</figureCitation>
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). Tritosternum bifurcate; sternal region with a pair of small presternal platelets;
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<emphasis id="B945BA1F524A735B5A30FB813A35ECC2" box="[1373,1406,1147,1171]" italics="true" pageId="3" pageNumber="444">
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st
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<quantity id="4CC9CBE8524A735B5A1FFB813A35ECC2" box="[1394,1406,1147,1171]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="444" unit="in" value="1.0">1</quantity>
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</emphasis>
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in plicate cuticle,
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<emphasis id="B945BA1F524A735B5E29FB653E2DECE6" box="[324,358,1183,1207]" italics="true" pageId="3" pageNumber="444">st2</emphasis>
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on subhexagonal, lineate-microtuberculate shield with pair of large pores in posterior corners;
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<emphasis id="B945BA1F524A735B5FFAFB393FF3EC8A" box="[151,184,1219,1243]" italics="true" pageId="3" pageNumber="444">
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st
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<quantity id="4CC9CBE8524A735B5FC7FB393FF3EC8A" box="[170,184,1219,1243]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="444" unit="in" value="3.0">3</quantity>
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</emphasis>
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in soft cuticle;
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<emphasis id="B945BA1F524A735B5E1CFB393ED9EC8A" box="[369,402,1219,1243]" italics="true" pageId="3" pageNumber="444">st4</emphasis>
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on small, subtriangular shields; all sternal setae short (
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<emphasis id="B945BA1F524A735B5B79FB393B7EEC8A" box="[1044,1077,1219,1243]" italics="true" pageId="3" pageNumber="444">st1</emphasis>
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4–7,
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<emphasis id="B945BA1F524A735B5B17FB393BD1EC8A" box="[1146,1178,1219,1243]" italics="true" pageId="3" pageNumber="444">
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||
st
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<date id="FF8F40CD524A735B5BE0FB393BD1EC8A" box="[1165,1178,1219,1243]" pageId="3" pageNumber="444" value="2010-09-02">2</date>
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</emphasis>
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9–10,
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<emphasis id="B945BA1F524A735B5B80FB393A45EC8A" box="[1261,1294,1219,1243]" italics="true" pageId="3" pageNumber="444">st3</emphasis>
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9,
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<emphasis id="B945BA1F524A735B5A5AFB393A13EC8A" box="[1335,1368,1219,1243]" italics="true" pageId="3" pageNumber="444">st4</emphasis>
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8–9). Genital shield broadly subtriangular, flaring posteriorly, rounded anteriorly, irregularly reticulate with short (8–9), simple
|
||
<emphasis id="B945BA1F524A735B5F86FAF13E40ED73" box="[235,267,1291,1314]" italics="true" pageId="3" pageNumber="444">st5</emphasis>
|
||
; shield length 84–86, maximum width 90–98. Opisthogaster (
|
||
<figureCitation id="130A7A88524A735B5CB8FAF03B53ED72" box="[981,1048,1290,1315]" captionStart="FIGURE 1" captionStartId="4.[151,250,1999,2021]" captionTargetBox="[158,1436,230,1948]" captionTargetId="figure@4.[151,1436,227,1948]" captionTargetPageId="4" captionText="FIGURE 1. Heatherella osleri n. sp. female holotype, venter." httpUri="https://zenodo.org/record/1292191/files/figure.png" pageId="3" pageNumber="444">Fig. 1</figureCitation>
|
||
) with seven pairs of short (7–14) setae (
|
||
<emphasis id="B945BA1F524A735B5F8DFAD53E6CED16" box="[224,295,1327,1351]" italics="true" pageId="3" pageNumber="444">JV1–2</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5E5BFAD53E29ED16" box="[310,354,1327,1351]" italics="true" pageId="3" pageNumber="444">JV4</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5E1DFAD53EF1ED16" box="[368,442,1327,1351]" italics="true" pageId="3" pageNumber="444">ZV1–4</emphasis>
|
||
), bulbous basally, tapering to fine point and ciliate on one side, lengthening posteriorly; two pairs of gland openings with raised rims posterolaterad coxae IV; one pair laterad para-anal setae; one pair laterad
|
||
<emphasis id="B945BA1F524A735B5F81FA8D3E51EDDE" box="[236,282,1399,1423]" italics="true" pageId="3" pageNumber="444">ZV1</emphasis>
|
||
; one pair anteriad
|
||
<emphasis id="B945BA1F524A735B5E99FA8D3D68EDDE" box="[500,547,1399,1423]" italics="true" pageId="3" pageNumber="444">ZV4</emphasis>
|
||
. Setae
|
||
<emphasis id="B945BA1F524A735B5D15FA8D3DEFEDDF" box="[632,676,1399,1422]" italics="true" pageId="3" pageNumber="444">JV5</emphasis>
|
||
caudal, posterior to final
|
||
<emphasis id="B945BA1F524A735B5CA0FA8D3C95EDDF" box="[973,990,1399,1422]" italics="true" pageId="3" pageNumber="444">R</emphasis>
|
||
seta and beneath most posterior spout. Anal region raised, with pair short (~5) simple setae on anterior corners anal opening (
|
||
<emphasis id="B945BA1F524A735B5BE2FA613BF1EDE2" box="[1167,1210,1435,1459]" italics="true" pageId="3" pageNumber="444">JV3</emphasis>
|
||
) and para-anal and postanal setae similar to ventral setae. Peritrematal shield obsolete, lateral region spiculate-microtuberculate, microtubercles becoming subreticulate across anterior ventral region and evanescent posteriorly; peritremes absent, stigma in raised tuberculate process (
|
||
<figureCitation id="130A7A88524A735B5D5BF9FC3DC1EE4E" box="[566,650,1542,1567]" captionStart="FIGURES 34–37" captionStartId="22.[151,264,1832,1854]" captionTargetBox="[326,1233,232,1771]" captionTargetId="figure@22.[309,1273,196,1784]" captionTargetPageId="22" captionText="FIGURES 34–37. (34) Heatherella acanthocharis walter, 1997, dorsomarginal protuberances, pustules and pores of paratype female (P-s5 pustule; P-s6 pore; P-S1 protuberance), dorsal view of flattened specimen, right side; (35) Aheatherella mira n. sp., dorsomarginal protuberances, pustules and pores of paratype female (P-s5 pustule; P-s6 protuberance with closely associated seta), dorsal view of ventrolateral fragment with broken r-setae, left side; (36) Heatherella osleri n. sp., peritreme of paratype female (ventral view, right peritreme of mite); (37) Aheatherella mira n. sp. peritreme of paratype female (ventral view, left peritreme of mite; sclerotised eXopodal portion of eXopodal-peritrematal shield shaded)." httpUri="https://zenodo.org/record/1292223/files/figure.png" pageId="3" pageNumber="444">Fig. 36</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E660D524A735B5FAAF9D63E74EF9E" blockId="3.[151,1437,822,1999]" pageId="3" pageNumber="444">
|
||
<emphasis id="B945BA1F524A735B5FAAF9D63E6AEE12" box="[199,289,1580,1603]" italics="true" pageId="3" pageNumber="444">Dorsum</emphasis>
|
||
(
|
||
<figureCitation id="130A7A88524A735B5E5CF9D03E3CEE12" box="[305,375,1578,1603]" captionStart="FIGURE 2" captionStartId="5.[151,250,1997,2019]" captionTargetBox="[201,1387,277,1878]" captionTargetId="figure@5.[151,1436,265,1888]" captionTargetPageId="5" captionText="FIGURE 2. Heatherella osleri n. sp. female holotype, dorsum." httpUri="https://zenodo.org/record/1292193/files/figure.png" pageId="3" pageNumber="444">Fig. 2</figureCitation>
|
||
). Dorsal shields comprising single median podonotal-mesonotal shield, latinotal shields, eight pre-pygidial platelets (often concealed), three pygidial shields (one medial, two lateral), all surrounded by dorsomarginal portion of opisthogastric-peritrematal-marginal shield; anteriorly, podonotal shield extending ventrally and fused with marginal shield. Median dorsal shield irregularly areolate, bearing eight pairs of setae (
|
||
<emphasis id="B945BA1F524A735B5A73F96D3A1BEEFE" box="[1310,1360,1686,1711]" italics="true" pageId="3" pageNumber="444">j4–6</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5A33F96D3ADEEEFE" box="[1374,1429,1687,1711]" italics="true" pageId="3" pageNumber="444">J1–4</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5FFAF9413FFBEE83" box="[151,176,1723,1746]" italics="true" pageId="3" pageNumber="444">z5</emphasis>
|
||
; not including setae on vertex, i.e., ventrally placed
|
||
<emphasis id="B945BA1F524A735B5C78F9413C60EE82" box="[789,811,1723,1747]" italics="true" pageId="3" pageNumber="444">j1</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5C51F9413C1FEE82" box="[828,852,1723,1747]" italics="true" pageId="3" pageNumber="444">z1</emphasis>
|
||
and dorsally placed
|
||
<emphasis id="B945BA1F524A735B5B25F9413B31EE82" box="[1096,1146,1723,1747]" italics="true" pageId="3" pageNumber="444">j2–3</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5BE6F9413BE8EE82" box="[1163,1187,1723,1747]" italics="true" pageId="3" pageNumber="444">z2</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5BD9F9413B85EE82" box="[1204,1230,1723,1747]" italics="true" pageId="3" pageNumber="444">s1</emphasis>
|
||
); latinotal shields fused with podonotal shield anteriorly, each with four pairs of setae (
|
||
<emphasis id="B945BA1F524A735B5CF5F9243CFAEEA6" box="[920,945,1758,1783]" italics="true" pageId="3" pageNumber="444">z6</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5CD2F9253CB2EEA6" box="[959,1017,1759,1783]" italics="true" pageId="3" pageNumber="444">Z1–4</emphasis>
|
||
); pre-pygidial platelets nude; median pygidial shield with two setae (
|
||
<emphasis id="B945BA1F524A735B5E93F8F93D52EF4B" box="[510,537,1795,1818]" italics="true" pageId="3" pageNumber="444">J5</emphasis>
|
||
), lateral pygidial shield each with one seta (
|
||
<emphasis id="B945BA1F524A735B5B7BF8F93B7FEF4A" box="[1046,1076,1795,1819]" italics="true" pageId="3" pageNumber="444">Z4</emphasis>
|
||
). All dorsal and marginal setae thickened, barbed, with ciliate crests, setal length 17–23. Seta
|
||
<emphasis id="B945BA1F524A735B5C27F8DC3C28EF6E" box="[842,867,1830,1855]" italics="true" pageId="3" pageNumber="444">z6</emphasis>
|
||
anterior to
|
||
<emphasis id="B945BA1F524A735B5C8EF8DC3CB7EF6E" box="[995,1020,1830,1855]" italics="true" pageId="3" pageNumber="444">s6</emphasis>
|
||
, seta
|
||
<emphasis id="B945BA1F524A735B5B56F8DD3B12EF6E" box="[1083,1113,1831,1855]" italics="true" pageId="3" pageNumber="444">Z1</emphasis>
|
||
anterior to
|
||
<emphasis id="B945BA1F524A735B5BB5F8DD3BBEEF6E" box="[1240,1269,1831,1855]" italics="true" pageId="3" pageNumber="444">S1</emphasis>
|
||
, seta
|
||
<emphasis id="B945BA1F524A735B5A59F8DD3A06EF6F" box="[1332,1357,1831,1854]" italics="true" pageId="3" pageNumber="444">z5</emphasis>
|
||
placed obliquely to seta
|
||
<emphasis id="B945BA1F524A735B5E34F8B13E24EF33" box="[345,367,1867,1890]" italics="true" pageId="3" pageNumber="444">j5</emphasis>
|
||
. Dorsal margin of shield bearing ten pairs of gland-bearing protuberances mostly aligned with
|
||
<emphasis id="B945BA1F524A735B5FFAF8953FF2EFD7" box="[151,185,1903,1926]" italics="true" pageId="3" pageNumber="444">s -S</emphasis>
|
||
series setae (
|
||
<emphasis id="B945BA1F524A735B5E3EF8953E27EFD6" box="[339,364,1903,1927]" italics="true" pageId="3" pageNumber="444">s1</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5E16F8953EDFEFD6" box="[379,404,1903,1927]" italics="true" pageId="3" pageNumber="444">s2</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5ECEF8953E93EFD6" box="[419,472,1902,1927]" italics="true" pageId="3" pageNumber="444">s4–6</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5E8AF8953D54EFD7" box="[487,543,1903,1927]" italics="true" pageId="3" pageNumber="444">S1–5</emphasis>
|
||
) but also
|
||
<emphasis id="B945BA1F524A735B5DFFF8953DE3EFD6" box="[658,680,1903,1927]" italics="true" pageId="3" pageNumber="444">j2</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524A735B5DDAF8953DA7EFD6" box="[695,748,1903,1927]" italics="true" pageId="3" pageNumber="444">z2–4</emphasis>
|
||
; protuberance
|
||
<emphasis id="B945BA1F524A735B5CFBF8953C81EFD7" box="[918,970,1903,1926]" italics="true" pageId="3" pageNumber="444">P -s5</emphasis>
|
||
not pustule-like, similar in form to other protuberances (
|
||
<tableCitation id="C6B353B6524A735B5E2BF8683ED4EFFA" box="[326,415,1938,1963]" captionStart="TABLE 1" captionStartId="1.[151,239,799,822]" captionText="TABLE 1. Modified pores in the Heatherellidae. Most are within spout-like protuberances but P-s5 can be a raised pustule, P-s4 can be an enlarged pore (possibly a small pustule) and P-s6 is sometimes a large pore similar in form to several other unmodified idiosomal pores (see Fig. 15). Variable states are in bold." pageId="3" pageNumber="444">Table 1</tableCitation>
|
||
). Marginal setae (
|
||
<emphasis id="B945BA1F524A735B5D17F86E3DD5EFFA" box="[634,670,1939,1963]" italics="true" pageId="3" pageNumber="444">r -R</emphasis>
|
||
series) hypertrichous, with 25–28 setae on each side (sometimes asymmetrical).
|
||
</paragraph>
|
||
<caption id="DF4E3685524D735C5FFAF8353C47EFB4" box="[151,780,1999,2021]" httpUri="https://zenodo.org/record/1292191/files/figure.png" pageId="4" pageNumber="445" startId="4.[151,250,1999,2021]" targetBox="[158,1436,230,1948]" targetPageId="4">
|
||
<paragraph id="8B8E660D524D735C5FFAF8353C47EFB4" blockId="4.[151,780,1999,2021]" box="[151,780,1999,2021]" pageId="4" pageNumber="445">
|
||
<emphasis id="B945BA1F524D735C5FFAF8353E58EFB4" bold="true" box="[151,275,1999,2021]" pageId="4" pageNumber="445">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C311D8E524D735C5E74F8353E9AEFB4" authority="Seeman & Minor & Baker & Walter, 2018" authorityName="Seeman & Minor & Baker & Walter" authorityYear="2018" box="[281,465,1999,2021]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="osleri" status="sp. nov.">
|
||
<emphasis id="B945BA1F524D735C5E74F8353E9AEFB4" box="[281,465,1999,2021]" italics="true" pageId="4" pageNumber="445">Heatherella osleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945BA1F524D735C5EBAF8353D5BEFB4" bold="true" box="[471,528,1999,2021]" pageId="4" pageNumber="445">
|
||
<taxonomicNameLabel id="A2760764524D735C5EBAF8353D5BEFB4" box="[471,528,1999,2021]" pageId="4" pageNumber="445" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female holotype, venter.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E3685524C735D5FFAF8373C51EFB2" box="[151,794,1997,2019]" httpUri="https://zenodo.org/record/1292193/files/figure.png" pageId="5" pageNumber="446" startId="5.[151,250,1997,2019]" targetBox="[201,1387,277,1878]" targetPageId="5">
|
||
<paragraph id="8B8E660D524C735D5FFAF8373C51EFB2" blockId="5.[151,794,1997,2019]" box="[151,794,1997,2019]" pageId="5" pageNumber="446">
|
||
<emphasis id="B945BA1F524C735D5FFAF8373E59EFB2" bold="true" box="[151,274,1997,2019]" pageId="5" pageNumber="446">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C311D8E524C735D5E74F8373E9AEFB2" authority="Seeman & Minor & Baker & Walter, 2018" authorityName="Seeman & Minor & Baker & Walter" authorityYear="2018" box="[281,465,1997,2019]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="osleri" status="sp. nov.">
|
||
<emphasis id="B945BA1F524C735D5E74F8373E9AEFB2" box="[281,465,1997,2019]" italics="true" pageId="5" pageNumber="446">Heatherella osleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945BA1F524C735D5EBAF8373D5BEFB2" bold="true" box="[471,528,1997,2019]" pageId="5" pageNumber="446">
|
||
<taxonomicNameLabel id="A2760764524C735D5EBAF8373D5BEFB2" box="[471,528,1997,2019]" pageId="5" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female holotype, dorsum.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E3685524F735E5FFAF85B3B81EF87" httpUri="https://zenodo.org/record/1292195/files/figure.png" pageId="6" pageNumber="447" startId="6.[151,264,1953,1975]" targetBox="[252,1329,209,1919]" targetPageId="6">
|
||
<paragraph id="8B8E660D524F735E5FFAF85B3B81EF87" blockId="6.[151,1436,1953,2006]" pageId="6" pageNumber="447">
|
||
<emphasis id="B945BA1F524F735E5FFAF85B3E70EFE6" bold="true" box="[151,315,1953,1975]" pageId="6" pageNumber="447">FIGURES 3–8.</emphasis>
|
||
<taxonomicName id="4C311D8E524F735E5E2EF85B3EB7EFE6" authority="Seeman & Minor & Baker & Walter, 2018" authorityName="Seeman & Minor & Baker & Walter" authorityYear="2018" box="[323,508,1953,1975]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="447" phylum="Arthropoda" rank="species" species="osleri" status="sp. nov.">
|
||
<emphasis id="B945BA1F524F735E5E2EF85B3EB7EFE6" box="[323,508,1953,1975]" italics="true" pageId="6" pageNumber="447">Heatherella osleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945BA1F524F735E5D6EF85B3D75EFE6" bold="true" box="[515,574,1953,1975]" pageId="6" pageNumber="447">
|
||
<taxonomicNameLabel id="A2760764524F735E5D6EF85B3D75EFE6" box="[515,574,1953,1975]" pageId="6" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(3) female, ventral
|
||
<taxonomicName id="4C311D8E524F735E5C67F85B3CCFEFE6" authorityName="Krefft" authorityYear="1871" box="[778,900,1953,1975]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="6" pageNumber="460" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
|
||
(seta
|
||
<emphasis id="B945BA1F524F735E5CA9F85B3C95EFE6" box="[964,990,1953,1975]" italics="true" pageId="6" pageNumber="447">h3</emphasis>
|
||
eXpressed asymmetrically in holotype); (4) female, gnathotectum; (5) male, gnathotectum; (6) female, chelicera; (7) male, chelicera; (8) female, leg I.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E3685524E735F5FFAFB2E3BB2ECBB" box="[151,1273,1236,1258]" httpUri="https://zenodo.org/record/1292197/files/figure.png" pageId="7" pageNumber="448" startId="7.[151,264,1236,1258]" targetBox="[174,1415,202,1206]" targetPageId="7">
|
||
<paragraph id="8B8E660D524E735F5FFAFB2E3BB2ECBB" blockId="7.[151,1273,1236,1258]" box="[151,1273,1236,1258]" pageId="7" pageNumber="448">
|
||
<emphasis id="B945BA1F524E735F5FFAFB2E3E0EECBB" bold="true" box="[151,325,1236,1258]" pageId="7" pageNumber="448">FIGURES 9–11.</emphasis>
|
||
<taxonomicName id="4C311D8E524E735F5E21FB2E3D4FECBB" authority="Seeman & Minor & Baker & Walter, 2018" authorityName="Seeman & Minor & Baker & Walter" authorityYear="2018" box="[332,516,1236,1258]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="448" phylum="Arthropoda" rank="species" species="osleri" status="sp. nov.">
|
||
<emphasis id="B945BA1F524E735F5E21FB2E3D4FECBB" box="[332,516,1236,1258]" italics="true" pageId="7" pageNumber="448">Heatherella osleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945BA1F524E735F5D67FB2E3D08ECBB" bold="true" box="[522,579,1236,1258]" pageId="7" pageNumber="448">
|
||
<taxonomicNameLabel id="A2760764524E735F5D67FB2E3D08ECBB" box="[522,579,1236,1258]" pageId="7" pageNumber="448" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female holotype, posterior views: (9) leg II; (10) leg III; (11) leg IV.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E660D524E735F5FAAFAEB3B44EE77" blockId="7.[151,1436,1297,1970]" pageId="7" pageNumber="448">
|
||
<taxonomicName id="4C311D8E524E735F5FAAFAEB3E1DED7B" authorityName="Krefft" authorityYear="1871" box="[199,342,1297,1322]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="7" pageNumber="448" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B945BA1F524E735F5FAAFAEB3E1DED7B" box="[199,342,1297,1322]" italics="true" pageId="7" pageNumber="448">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130A7A88524E735F5E09FAEB3E81ED78" box="[356,458,1297,1322]" captionStart="FIGURES 3–8" captionStartId="6.[151,264,1953,1975]" captionTargetBox="[252,1329,209,1919]" captionTargetId="figure@6.[246,1341,193,1932]" captionTargetPageId="6" captionText="FIGURES 3–8. Heatherella osleri n. sp. (3) female, ventral gnathosoma (seta h3 eXpressed asymmetrically in holotype); (4) female, gnathotectum; (5) male, gnathotectum; (6) female, chelicera; (7) male, chelicera; (8) female, leg I." httpUri="https://zenodo.org/record/1292195/files/figure.png" pageId="7" pageNumber="448">Figs 3–7</figureCitation>
|
||
). Subcapitulum with strong row of denticles between palpcoxal setae; and three shorter, less distinct rows between setae
|
||
<emphasis id="B945BA1F524E735F5D7BFACE3D78ED1C" box="[534,563,1332,1357]" italics="true" pageId="7" pageNumber="448">h3</emphasis>
|
||
and posterior to palpcoxal setae; all but most posterior row extend across deutosternum as fine denticles; deutosternal gutter narrow, with two additional rows of fine denticles anterior to palpcoxal setae; all subcapitular setae simple, acuminate, length
|
||
<emphasis id="B945BA1F524E735F5CF8FA873CFAEDC7" box="[917,945,1405,1430]" italics="true" pageId="7" pageNumber="448">
|
||
h
|
||
<date id="FF8F40CD524E735F5CC8FA873CFAEDC7" box="[933,945,1405,1430]" pageId="7" pageNumber="448" value="2011-01-12">1</date>
|
||
</emphasis>
|
||
11–12,
|
||
<emphasis id="B945BA1F524E735F5B7BFA873B78EDC7" box="[1046,1075,1405,1430]" italics="true" pageId="7" pageNumber="448">
|
||
h
|
||
<date id="FF8F40CD524E735F5B48FA873B78EDC7" box="[1061,1075,1405,1430]" pageId="7" pageNumber="448" value="2010-09-02">2</date>
|
||
</emphasis>
|
||
9–10,
|
||
<emphasis id="B945BA1F524E735F5BE7FA873BEDEDC7" box="[1162,1190,1405,1430]" italics="true" pageId="7" pageNumber="448">
|
||
h
|
||
<date id="FF8F40CD524E735F5BF5FA873BEDEDC7" box="[1176,1190,1405,1430]" pageId="7" pageNumber="448" value="2011-03-12">3</date>
|
||
</emphasis>
|
||
11–12,
|
||
<emphasis id="B945BA1F524E735F5A66FA843A6DEDC4" box="[1291,1318,1406,1429]" italics="true" pageId="7" pageNumber="448">pc</emphasis>
|
||
8–10 (
|
||
<emphasis id="B945BA1F524E735F5AEDFA873AD7EDC7" box="[1408,1436,1405,1430]" italics="true" pageId="7" pageNumber="448">h3</emphasis>
|
||
expressed asymmetrically in
|
||
<typeStatus id="548AD8AF524E735F5EB1FA5B3D08EDEB" box="[476,579,1441,1466]" pageId="7" pageNumber="448" type="holotype">holotype</typeStatus>
|
||
); corniculi simple. Chelicerae (~75 long), second article (55–65 long) with 3 subdistal teeth around pilus dentilis, deep notch receives fixed digit; moveable digit (25–28) tridentate; cheliceral seta (~12 long) thick, somewhat flattened, acuminate, inserted on paraxial side of chelicera. Tectum coarsely denticulate, with median prong (sometimes bifid) and 2–3 pairs of lateral teeth.
|
||
</paragraph>
|
||
<paragraph id="8B8E660D524E735F5FAAF9C83EB6EE3F" blockId="7.[151,1436,1297,1970]" pageId="7" pageNumber="448">
|
||
<emphasis id="B945BA1F524E735F5FAAF9C83FB7EE18" box="[199,252,1586,1609]" italics="true" pageId="7" pageNumber="448">Legs</emphasis>
|
||
(
|
||
<figureCitation id="130A7A88524E735F5E61F9CB3ECBEE18" box="[268,384,1585,1610]" captionStart-0="FIGURES 3–8" captionStart-1="FIGURES 9–11" captionStartId-0="6.[151,264,1953,1975]" captionStartId-1="7.[151,264,1236,1258]" captionTargetBox-0="[252,1329,209,1919]" captionTargetBox-1="[174,1415,202,1206]" captionTargetId-0="figure@6.[246,1341,193,1932]" captionTargetId-1="figure@7.[164,1424,193,1214]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 3–8. Heatherella osleri n. sp. (3) female, ventral gnathosoma (seta h3 eXpressed asymmetrically in holotype); (4) female, gnathotectum; (5) male, gnathotectum; (6) female, chelicera; (7) male, chelicera; (8) female, leg I." captionText-1="FIGURES 9–11. Heatherella osleri n. sp. female holotype, posterior views: (9) leg II; (10) leg III; (11) leg IV." httpUri-0="https://zenodo.org/record/1292195/files/figure.png" httpUri-1="https://zenodo.org/record/1292197/files/figure.png" pageId="7" pageNumber="448">Figs 8–11</figureCitation>
|
||
). Leg setae generally unmodified; dorsal femoral setae on leg I barbed (
|
||
<figureCitation id="130A7A88524E735F5BDAF9CB3BB7EE18" box="[1207,1276,1585,1610]" captionStart="FIGURES 3–8" captionStartId="6.[151,264,1953,1975]" captionTargetBox="[252,1329,209,1919]" captionTargetId="figure@6.[246,1341,193,1932]" captionTargetPageId="6" captionText="FIGURES 3–8. Heatherella osleri n. sp. (3) female, ventral gnathosoma (seta h3 eXpressed asymmetrically in holotype); (4) female, gnathotectum; (5) male, gnathotectum; (6) female, chelicera; (7) male, chelicera; (8) female, leg I." httpUri="https://zenodo.org/record/1292195/files/figure.png" pageId="7" pageNumber="448">Fig. 8</figureCitation>
|
||
); femoral and genual setae subequal in length.
|
||
</paragraph>
|
||
<paragraph id="8B8E660D524E735F5FAAF9833BD4EFE3" blockId="7.[151,1436,1297,1970]" pageId="7" pageNumber="448">
|
||
<emphasis id="B945BA1F524E735F5FAAF9833E40EEC3" bold="true" box="[199,267,1657,1682]" pageId="7" pageNumber="448">Male.</emphasis>
|
||
Similar to female except as follows.
|
||
<emphasis id="B945BA1F524E735F5DA1F9803C5FEEC0" box="[716,788,1658,1681]" italics="true" pageId="7" pageNumber="448">Venter</emphasis>
|
||
(
|
||
<figureCitation id="130A7A88524E735F5C4AF9833C34EEC0" box="[807,895,1657,1682]" captionStart="FIGURES 12–14" captionStartId="8.[151,264,1439,1461]" captionTargetBox="[213,1369,199,1413]" captionTargetId="figure@8.[212,1376,185,1424]" captionTargetPageId="8" captionText="FIGURES 12–14. Heatherella osleri n. sp. male (12) venter; (13) dorsum (marginal shields not shown laterally; torn anteriorly from marginal shield); (14) leg II, anterior view." httpUri="https://zenodo.org/record/1292199/files/figure.png" pageId="7" pageNumber="448">Fig. 12</figureCitation>
|
||
). Shield bearing
|
||
<emphasis id="B945BA1F524E735F5B26F9833BC3EEC0" box="[1099,1160,1657,1681]" italics="true" pageId="7" pageNumber="448">st2–4</emphasis>
|
||
and subcircular genital opening covered by a pair of platelets;
|
||
<emphasis id="B945BA1F524E735F5D1FF9643DD9EEE4" box="[626,658,1694,1717]" italics="true" pageId="7" pageNumber="448">st5</emphasis>
|
||
on ventral, subtriangular extension of opisthogastric-peritrematalmarginal shield (and so eight pairs of ventral setae appearing to be present).
|
||
<emphasis id="B945BA1F524E735F5B4FF9383B37EE88" box="[1058,1148,1730,1753]" italics="true" pageId="7" pageNumber="448">Dorsum</emphasis>
|
||
(
|
||
<figureCitation id="130A7A88524E735F5BE2F93B3BADEE88" box="[1167,1254,1729,1754]" captionStart="FIGURES 12–14" captionStartId="8.[151,264,1439,1461]" captionTargetBox="[213,1369,199,1413]" captionTargetId="figure@8.[212,1376,185,1424]" captionTargetPageId="8" captionText="FIGURES 12–14. Heatherella osleri n. sp. male (12) venter; (13) dorsum (marginal shields not shown laterally; torn anteriorly from marginal shield); (14) leg II, anterior view." httpUri="https://zenodo.org/record/1292199/files/figure.png" pageId="7" pageNumber="448">Fig. 13</figureCitation>
|
||
). Dorsally with podonotal and mesonotal shields; podonotal shield fused with lateral shields, bearing five pairs of setae (
|
||
<emphasis id="B945BA1F524E735F5A59F91F3A2DEEAF" box="[1332,1382,1765,1790]" italics="true" pageId="7" pageNumber="448">j4–6</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524E735F5A18F91C3FEEEF73" italics="true" pageId="7" pageNumber="448">z5– 6</emphasis>
|
||
) excluding setae at vertex (
|
||
<emphasis id="B945BA1F524E735F5E8BF8F33D53EF70" box="[486,536,1801,1825]" italics="true" pageId="7" pageNumber="448">j1–3</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524E735F5D45F8F33D16EF70" box="[552,605,1801,1825]" italics="true" pageId="7" pageNumber="448">z1–3</emphasis>
|
||
,
|
||
<emphasis id="B945BA1F524E735F5D00F8F33DCCEF70" box="[621,647,1801,1825]" italics="true" pageId="7" pageNumber="448">s1</emphasis>
|
||
); mesonotal shield mostly free from lateral mesonotal shields (fused anteriorly), mesonotal shield bearing four pairs of setae (
|
||
<emphasis id="B945BA1F524E735F5C40F8D73C28EF14" box="[813,867,1837,1861]" italics="true" pageId="7" pageNumber="448">J1–4</emphasis>
|
||
), lateral mesonotal shields bearing three pairs of setae (
|
||
<emphasis id="B945BA1F524E735F5F8DF8AB3E52EF38" box="[224,281,1873,1897]" italics="true" pageId="7" pageNumber="448">Z1–3</emphasis>
|
||
); eight pre-pygidial platelets and pygidial shields as in female; protuberances and
|
||
<emphasis id="B945BA1F524E735F5BD3F8A83BABEF38" box="[1214,1248,1874,1897]" italics="true" pageId="7" pageNumber="448">s -S</emphasis>
|
||
and
|
||
<emphasis id="B945BA1F524E735F5A75F8A83A76EF38" box="[1304,1341,1874,1897]" italics="true" pageId="7" pageNumber="448">r -R</emphasis>
|
||
setae as in female.
|
||
<taxonomicName id="4C311D8E524E735F5E63F88F3ED6EFDF" authorityName="Krefft" authorityYear="1871" box="[270,413,1909,1934]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="7" pageNumber="448" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B945BA1F524E735F5E63F88F3ED6EFDF" box="[270,413,1909,1934]" italics="true" pageId="7" pageNumber="448">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130A7A88524E735F5EC1F88F3D5AEFDC" box="[428,529,1909,1934]" captionStart="FIGURES 3–8" captionStartId="6.[151,264,1953,1975]" captionTargetBox="[252,1329,209,1919]" captionTargetId="figure@6.[246,1341,193,1932]" captionTargetPageId="6" captionText="FIGURES 3–8. Heatherella osleri n. sp. (3) female, ventral gnathosoma (seta h3 eXpressed asymmetrically in holotype); (4) female, gnathotectum; (5) male, gnathotectum; (6) female, chelicera; (7) male, chelicera; (8) female, leg I." httpUri="https://zenodo.org/record/1292195/files/figure.png" pageId="7" pageNumber="448">Figs 5, 7</figureCitation>
|
||
). Similar to female; teeth on gnathotectum less developed.
|
||
<emphasis id="B945BA1F524E735F5BC7F88C3B94EFDC" box="[1194,1247,1910,1933]" italics="true" pageId="7" pageNumber="448">Legs</emphasis>
|
||
(
|
||
<figureCitation id="130A7A88524E735F5B82F88F3A09EFDC" box="[1263,1346,1909,1934]" captionStart="FIGURES 12–14" captionStartId="8.[151,264,1439,1461]" captionTargetBox="[213,1369,199,1413]" captionTargetId="figure@8.[212,1376,185,1424]" captionTargetPageId="8" captionText="FIGURES 12–14. Heatherella osleri n. sp. male (12) venter; (13) dorsum (marginal shields not shown laterally; torn anteriorly from marginal shield); (14) leg II, anterior view." httpUri="https://zenodo.org/record/1292199/files/figure.png" pageId="7" pageNumber="448">Fig. 14</figureCitation>
|
||
). Leg II with setae
|
||
<emphasis id="B945BA1F524E735F5E63F8633E7CEFE0" box="[270,311,1945,1969]" italics="true" pageId="7" pageNumber="448">av1</emphasis>
|
||
on femur and genu a blunt spur (8–
|
||
<date id="FF8F40CD524E735F5DA5F8633DA0EFE0" box="[712,747,1945,1969]" pageId="7" pageNumber="448">9 x</date>
|
||
5 wide, 5–
|
||
<date id="FF8F40CD524E735F5C05F8633CC1EFE0" box="[872,906,1945,1970]" pageId="7" pageNumber="448">6 x</date>
|
||
2–3 wide, respectively).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4E3685524173505FFAFA653DCBED82" httpUri="https://zenodo.org/record/1292199/files/figure.png" pageId="8" pageNumber="449" startId="8.[151,264,1439,1461]" targetBox="[213,1369,199,1413]" targetPageId="8">
|
||
<paragraph id="8B8E660D524173505FFAFA653DCBED82" blockId="8.[151,1435,1439,1491]" pageId="8" pageNumber="449">
|
||
<emphasis id="B945BA1F524173505FFAFA653E18EDE4" bold="true" box="[151,339,1439,1461]" pageId="8" pageNumber="449">FIGURES 12–14.</emphasis>
|
||
<taxonomicName id="4C311D8E524173505E34FA653D5BEDE4" authority="Seeman & Minor & Baker & Walter, 2018" authorityName="Seeman & Minor & Baker & Walter" authorityYear="2018" box="[345,528,1439,1461]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="449" phylum="Arthropoda" rank="species" species="osleri" status="sp. nov.">
|
||
<emphasis id="B945BA1F524173505E34FA653D5BEDE4" box="[345,528,1439,1461]" italics="true" pageId="8" pageNumber="449">Heatherella osleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945BA1F524173505D7BFA653D04EDE4" bold="true" box="[534,591,1439,1461]" pageId="8" pageNumber="449">
|
||
<taxonomicNameLabel id="A2760764524173505D7BFA653D04EDE4" box="[534,591,1439,1461]" pageId="8" pageNumber="449" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
male (12) venter; (13) dorsum (marginal shields not shown laterally; torn anteriorly from marginal shield); (14) leg II, anterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32B3586524173505FAAF9FA3C52EEF8" pageId="8" pageNumber="449" type="materials_examined">
|
||
<paragraph id="8B8E660D524173505FAAF9FA3C52EEF8" blockId="8.[151,1436,1536,2029]" pageId="8" pageNumber="449">
|
||
<emphasis id="B945BA1F524173505FAAF9FA3EE0EE48" bold="true" box="[199,427,1536,1561]" pageId="8" pageNumber="449">Material examined</emphasis>
|
||
. Holotype female, 2 female, 9 male paratypes, Australia, New South Wales, Lansdowne State Forest, 31°45′S 152°37′E, Graham Osler,
|
||
<date id="FF8F40CD524173505DC0F9DF3C33EE6C" box="[685,888,1572,1597]" pageId="8" pageNumber="449" value="1996-02-22">22 February 1996</date>
|
||
, Tullgren extraction of top
|
||
<quantity id="4CC9CBE8524173505BC6F9DE3BA8EE6D" box="[1195,1251,1572,1597]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="449" unit="cm" value="5.0">5 cm</quantity>
|
||
of soil in mixed hardwood forest with
|
||
<taxonomicName id="4C311D8E524173505EFDF9B23D38EE30" box="[400,627,1608,1633]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="449" phylum="Tracheophyta" rank="species" species="pilularis">
|
||
<emphasis id="B945BA1F524173505EFDF9B23D38EE30" box="[400,627,1608,1633]" italics="true" pageId="8" pageNumber="449">Eucalyptus pilularis</emphasis>
|
||
</taxonomicName>
|
||
&
|
||
<taxonomicName id="4C311D8E524173505DF5F9B33C67EE31" authorityName="Deane & Maiden" authorityYear="1896" box="[664,812,1609,1632]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="449" phylum="Tracheophyta" rank="species" species="propinqua">
|
||
<emphasis id="B945BA1F524173505DF5F9B33DE2EE31" box="[664,681,1609,1632]" italics="true" pageId="8" pageNumber="449">E</emphasis>
|
||
.
|
||
<emphasis id="B945BA1F524173505DD5F9B33C67EE31" box="[696,812,1609,1632]" italics="true" pageId="8" pageNumber="449">propinqua</emphasis>
|
||
</taxonomicName>
|
||
. Holotype, 1 female and 7 male paratypes deposited in QM; 1 female (ANIC 51-006365), 1 male paratype deposited in ANIC (ANIC 51-006366); 1 male paratype deposited in each of CNC and NZAC (NZAC02016473).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B3586524173505FAAF94E3EB3EEA0" pageId="8" pageNumber="449" type="etymology">
|
||
<paragraph id="8B8E660D524173505FAAF94E3EB3EEA0" blockId="8.[151,1436,1536,2029]" pageId="8" pageNumber="449">
|
||
<emphasis id="B945BA1F524173505FAAF94E3E03EE9C" bold="true" box="[199,328,1716,1741]" pageId="8" pageNumber="449">Etymology</emphasis>
|
||
: The new species is named in honour of the young acarologist who collected these specimens many years ago, Graham Osler.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B3586524173515FAAF9063C4AE94D" lastPageId="9" lastPageNumber="450" pageId="8" pageNumber="449" type="discussion">
|
||
<paragraph id="8B8E660D524173505FAAF9063CF4EFBD" blockId="8.[151,1436,1536,2029]" pageId="8" pageNumber="449">
|
||
<emphasis id="B945BA1F524173505FAAF9063E04EF44" bold="true" box="[199,335,1788,1813]" pageId="8" pageNumber="449">Comments.</emphasis>
|
||
This species is found in wet sclerophyll forests; more details on the mite’s ecology may be found in
|
||
<bibRefCitation id="EFA01BFC524173505FD9F8DA3EF1EF68" box="[180,442,1824,1849]" pageId="8" pageNumber="449" refId="ref9334" refString="Osler, G. H. R. & Beattie, A. J. (2001) Contribution of oribatid and mesostigmatid soil mites in ecologically based estimates of global species richness. Austral Ecology, 26, 70 - 79." type="journal article">Osler & Beattie (2001)</bibRefCitation>
|
||
. This species is obviously different from the two species from
|
||
<collectingRegion id="49F5A8EF524173505B10F8DA3A4AEF68" box="[1149,1281,1824,1849]" country="Australia" name="Queensland" pageId="8" pageNumber="449">Queensland</collectingRegion>
|
||
in having the dorsal setae with ciliate crests (simple setae in other
|
||
<taxonomicName id="4C311D8E524173505D84F8BE3C25EF0C" authorityName="walter" authorityYear="1997" box="[745,878,1860,1885]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945BA1F524173505D84F8BE3C25EF0C" box="[745,878,1860,1885]" italics="true" pageId="8" pageNumber="449">Heatherella</emphasis>
|
||
</taxonomicName>
|
||
), seta
|
||
<emphasis id="B945BA1F524173505CD5F8BF3C9AEF0C" box="[952,977,1861,1885]" italics="true" pageId="8" pageNumber="449">z2</emphasis>
|
||
present (absent in other
|
||
<taxonomicName id="4C311D8E524173505B85F8BE3ADBEF0C" authorityName="Walter" authorityYear="1997" box="[1256,1424,1860,1885]" class="Arachnida" family="Heatherellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="449" phylum="Arthropoda" rank="family">Heatherellidae</taxonomicName>
|
||
), and ten pairs of lateral protuberances (8–9 pairs in other
|
||
<taxonomicName id="4C311D8E524173505C5CF8923CFDEFD0" authorityName="walter" authorityYear="1997" box="[817,950,1896,1921]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945BA1F524173505C5CF8923CFDEFD0" box="[817,950,1896,1921]" italics="true" pageId="8" pageNumber="449">Heatherella</emphasis>
|
||
</taxonomicName>
|
||
including
|
||
<emphasis id="B945BA1F524173505B59F8933B2CEFD1" box="[1076,1127,1897,1920]" italics="true" pageId="8" pageNumber="449">P -s5</emphasis>
|
||
; see
|
||
<tableCitation id="C6B353B6524173505BCBF8923BB0EFD0" box="[1190,1275,1896,1921]" captionStart="TABLE 1" captionStartId="1.[151,239,799,822]" captionText="TABLE 1. Modified pores in the Heatherellidae. Most are within spout-like protuberances but P-s5 can be a raised pustule, P-s4 can be an enlarged pore (possibly a small pustule) and P-s6 is sometimes a large pore similar in form to several other unmodified idiosomal pores (see Fig. 15). Variable states are in bold." pageId="8" pageNumber="449">Table 1</tableCitation>
|
||
).
|
||
<taxonomicName id="4C311D8E524173505A7AF8923F9CEFF4" authorityName="Seeman & Minor & Baker & Walter" authorityYear="2018" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="449" phylum="Arthropoda" rank="species" species="osleri">
|
||
<emphasis id="B945BA1F524173505A7AF8923F9CEFF4" italics="true" pageId="8" pageNumber="449">Heatherella osleri</emphasis>
|
||
</taxonomicName>
|
||
seems more closely related to
|
||
<taxonomicName id="4C311D8E524173505D41F8773D9BEFF4" authorityName="Walter" authorityYear="1997" box="[556,720,1932,1957]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="449" phylum="Arthropoda" rank="species" species="callimaulos">
|
||
<emphasis id="B945BA1F524173505D41F8773D0BEFF5" box="[556,576,1933,1956]" italics="true" pageId="8" pageNumber="449">H</emphasis>
|
||
.
|
||
<emphasis id="B945BA1F524173505D23F8763D9BEFF4" box="[590,720,1932,1957]" italics="true" pageId="8" pageNumber="449">callimaulos</emphasis>
|
||
</taxonomicName>
|
||
in sharing the position of the dorsal setae (i.e. seta
|
||
<emphasis id="B945BA1F524173505A64F8763A69EFF4" box="[1289,1314,1932,1957]" italics="true" pageId="8" pageNumber="449">z6</emphasis>
|
||
anterior to
|
||
<emphasis id="B945BA1F524173505FFAF84A3FFBEF98" box="[151,176,1968,1993]" italics="true" pageId="8" pageNumber="449">s6</emphasis>
|
||
, seta
|
||
<emphasis id="B945BA1F524173505F9AF84B3E5EEF98" box="[247,277,1969,1993]" italics="true" pageId="8" pageNumber="449">Z1</emphasis>
|
||
anterior to
|
||
<emphasis id="B945BA1F524173505ECFF84B3EF5EF98" box="[418,446,1969,1993]" italics="true" pageId="8" pageNumber="449">S1</emphasis>
|
||
, seta
|
||
<emphasis id="B945BA1F524173505D6BF84B3D54EF99" box="[518,543,1969,1992]" italics="true" pageId="8" pageNumber="449">z5</emphasis>
|
||
placed obliquely to seta
|
||
<emphasis id="B945BA1F524173505C25F84B3C15EF99" box="[840,862,1969,1992]" italics="true" pageId="8" pageNumber="449">j5</emphasis>
|
||
), having marginal protuberance
|
||
<emphasis id="B945BA1F524173505BB7F84B3A46EF98" box="[1242,1293,1969,1993]" italics="true" pageId="8" pageNumber="449">P -s4</emphasis>
|
||
present and having males with seta
|
||
<emphasis id="B945BA1F524173505EF0F82F3E8DEFBC" box="[413,454,2005,2029]" italics="true" pageId="8" pageNumber="449">av1</emphasis>
|
||
on genu II modified into a spur-like process.
|
||
</paragraph>
|
||
<paragraph id="8B8E660D524073515FAAFF6D3C4AE94D" blockId="9.[151,1436,151,285]" pageId="9" pageNumber="450">
|
||
The presence of ten pairs of protuberances, the maximum number known so far in
|
||
<taxonomicName id="4C311D8E524073515B1BFF6D3A57E8E1" authorityName="Walter" authorityYear="1997" box="[1142,1308,151,176]" class="Arachnida" family="Heatherellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="450" phylum="Arthropoda" rank="family">Heatherellidae</taxonomicName>
|
||
, suggests a trend towards an increase in protuberance number. The presence of protuberance
|
||
<emphasis id="B945BA1F524073515B4CFF473B1FE885" box="[1057,1108,188,212]" italics="true" pageId="9" pageNumber="450">
|
||
P -s
|
||
<date id="FF8F40CD524073515B2BFF463B1FE885" box="[1094,1108,188,212]" pageId="9" pageNumber="450">4</date>
|
||
</emphasis>
|
||
may be a synapomorphy, but protuberance
|
||
<emphasis id="B945BA1F524073515E5CFF1B3E2FE8A6" box="[305,356,224,248]" italics="true" pageId="9" pageNumber="450">P -s5</emphasis>
|
||
is an autapomorphy, and we interpret protuberance
|
||
<emphasis id="B945BA1F524073515CDCFF1B3CAEE8A9" box="[945,997,223,248]" italics="true" pageId="9" pageNumber="450">P -s6</emphasis>
|
||
as a homoplasy as it is present only in
|
||
<taxonomicName id="4C311D8E524073515FFAFEFF3FB2E94C" authorityName="Seeman & Minor & Baker & Walter" authorityYear="2018" box="[151,249,260,285]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="450" phylum="Arthropoda" rank="species" species="osleri">
|
||
<emphasis id="B945BA1F524073515FFAFEFF3FE0E94D" box="[151,171,261,284]" italics="true" pageId="9" pageNumber="450">H</emphasis>
|
||
.
|
||
<emphasis id="B945BA1F524073515FD4FEFE3FB2E94C" box="[185,249,260,285]" italics="true" pageId="9" pageNumber="450">osleri</emphasis>
|
||
</taxonomicName>
|
||
and the new genus described below (
|
||
<tableCitation id="C6B353B6524073515DF1FEFE3DA4E94D" box="[668,751,260,285]" captionStart="TABLE 1" captionStartId="1.[151,239,799,822]" captionText="TABLE 1. Modified pores in the Heatherellidae. Most are within spout-like protuberances but P-s5 can be a raised pustule, P-s4 can be an enlarged pore (possibly a small pustule) and P-s6 is sometimes a large pore similar in form to several other unmodified idiosomal pores (see Fig. 15). Variable states are in bold." pageId="9" pageNumber="450">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |