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<document id="1F6A4BBF945DA29D760F1AA94D5F3A1C" ID-CLB-Dataset="21703" ID-DOI="10.24349/acarologia/20204357" ID-GBIF-Dataset="621e183b-789b-4d5e-bc43-64000219e1a8" ID-ISSN="2107-7207" ID-Zenodo-Dep="3890497" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591394849704" checkinUser="admin" docAuthor="Ermilov, Sergey G. &amp; Starý, Josef" docDate="2020" docId="039887E7B545FF8ABD9A8503FC8229FA" docLanguage="en" docName="Acarologia.60.1.64-74.pdf.imf" docOrigin="Acarologia 60 (1)" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.6:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="6" docTitle="Galumna (Galumna) sandormahunkai Ermilov &amp; Starý 2020, n. sp." docType="treatment" docVersion="10" lastPageNumber="74" masterDocId="FFA1FF9FB547FF8FBC718312FFA72D58" masterDocTitle="New and interesting species of the generaGalumna andPergalumna (Acari, Oribatida, Galumnidae) from the Montagne dAmbre National Park, Madagascar" masterLastPageNumber="74" masterPageNumber="64" pageNumber="64" updateTime="1698842185176" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0E2117C9C0B59FE2DC735783778D9B58">New and interesting species of the generaGalumna andPergalumna (Acari, Oribatida, Galumnidae) from the Montagne dAmbre National Park, Madagascar</mods:title>
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<mods:name id="E2DF849D00F0C99F553AE9D6985E502E" type="personal">
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<mods:namePart id="9F1C6C1D668E5D2935304179175ED7BC">Ermilov, Sergey G.</mods:namePart>
<mods:affiliation id="138F863EB8B1F0EB14A46D3CAEB8798B">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
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<mods:namePart id="1F0822925B2ECCDC6A1F1735712D8FE9">Starý, Josef</mods:namePart>
<mods:affiliation id="E17F57ED52D612160918410F03B38F5D">Biology Centre v. v. i., Czech Academy of Sciences, Institute of Soil Biology, České Budejovice, Czech</mods:affiliation>
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<mods:title id="0BD528F9ED92E8CD65D1293C36EF9479">Acarologia</mods:title>
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<mods:date id="313994FAE12422F4BA83F915528B9018">2020</mods:date>
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<treatment id="039887E7B545FF8ABD9A8503FC8229FA" ID-DOI="http://doi.org/10.5281/zenodo.5579922" ID-GBIF-Taxon="164501351" ID-Zenodo-Dep="5579922" LSID="urn:lsid:zoobank.org:act:98E9F2F2-5384-4E4D-BADE-C6A461B64755" httpUri="http://treatment.plazi.org/id/039887E7B545FF8ABD9A8503FC8229FA" lastPageId="5" pageId="2">
<subSubSection id="C32B657AB545FF8DBD9A8503FBE52B75" box="[491,1090,1553,1581]" pageId="2" type="nomenclature">
<paragraph id="8B8E36F1B545FF8DBD9A8503FBE52B75" blockId="2.[491,1090,1553,1581]" box="[491,1090,1553,1581]" pageId="2">
<heading id="D0C6819DB545FF8DBD9A8503FBE52B75" box="[491,1090,1553,1581]" fontSize="12" level="2" pageId="2" reason="2">
<emphasis id="B945EAE3B545FF8DBD9A8503FBE52B75" bold="true" box="[491,1090,1553,1581]" pageId="2">
<taxonomicName id="4C314D72B545FF8DBD9A8503FC572B75" ID-CoL="8T9VR" authority="Ermilov &amp; Starý, 2020" authorityName="Ermilov &amp; Starý" authorityYear="2020" box="[491,1008,1553,1581]" class="Arachnida" family="Galumnidae" genus="Galumna" kingdom="Animalia" order="Sarcoptiformes" pageId="2" phylum="Arthropoda" rank="species" species="sandormahunkai" status="sp. nov." subGenus="Galumna">
<emphasis id="B945EAE3B545FF8DBD9A8503FDC02B75" bold="true" box="[491,615,1553,1581]" italics="true" pageId="2">Galumna</emphasis>
(
<emphasis id="B945EAE3B545FF8DBE088503FD522B75" bold="true" box="[633,757,1553,1581]" italics="true" pageId="2">Galumna</emphasis>
)
<emphasis id="B945EAE3B545FF8DBF768503FC572B75" bold="true" box="[775,1008,1553,1581]" italics="true" pageId="2">sandormahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765798B545FF8DBF898503FBE52B75" box="[1016,1090,1553,1581]" pageId="2" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C32B657AB545FF8DBD9A8559FD0C2BDA" pageId="2" type="description">
<paragraph id="8B8E36F1B545FF8DBD9A8559FC4F2B06" blockId="2.[491,1522,1611,2045]" box="[491,1000,1611,1630]" pageId="2">
Zoobank:
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<paragraph id="8B8E36F1B545FF8DBE628578FD0C2BDA" blockId="2.[491,1522,1611,2045]" box="[531,683,1642,1666]" pageId="2">
(
<figureCitation id="130A2A74B545FF8DBE6A8578FD042BDA" box="[539,675,1642,1666]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[515,582,1939,1960]" captionStartId-1="4.[515,582,1472,1493]" captionStartId-2="6.[515,582,1666,1687]" captionTargetBox-0="[527,1473,971,1875]" captionTargetBox-1="[539,1474,239,1394]" captionTargetBox-2="[598,1440,221,1600]" captionTargetId-0="figure@3.[508,1492,961,2352]" captionTargetId-1="figure@4.[456,1495,185,1654]" captionTargetId-2="figure@6.[503,1487,221,1612]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="Figure 1 Galumna sandormahunkai n. sp., adult: a dorsal view; b ventral view (gnathosoma and legs omitted). Scale bar 100 μm." captionText-1="Figure 2 Galumna sandormahunkai n. sp., adult: a anterior part of body, lateral view (gnathosoma and legs omitted); b posterior part of body, lateral view; c anterior part of prodorsum, frontal view; d subcapitulum, ventral view; e palp, right, antiaxial view; f chelicera, left, paraxial view. Scale bar 100 μm (a, b), scale bar 50 μm (c, d, f), scale bar 20 μm (e)." captionText-2="Figure 3 Galumna sandormahunkai n. sp., adult: a leg I, without trochanter, right, antiaxial view; b leg IV, left, antiaxial view. Scale bar 50 μm." figureDoi-0="http://doi.org/10.5281/zenodo.3890499" figureDoi-1="http://doi.org/10.5281/zenodo.3890501" figureDoi-2="http://doi.org/10.5281/zenodo.3890503" httpUri-0="https://zenodo.org/record/3890499/files/figure.png" httpUri-1="https://zenodo.org/record/3890501/files/figure.png" httpUri-2="https://zenodo.org/record/3890503/files/figure.png" pageId="2">Figures 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32B657AB545FF8DBE62859FFA932ACF" pageId="2" type="diagnosis">
<paragraph id="8B8E36F1B545FF8DBE62859FFA932ACF" blockId="2.[491,1522,1611,2045]" pageId="2">
<emphasis id="B945EAE3B545FF8DBE62859FFD2F2BFD" bold="true" box="[531,648,1677,1701]" pageId="2">Diagnosis</emphasis>
— Body size 780996 × 564780. Rostrum protruding, narrowly rounded. Lamellar and sublamellar lines divergent distally,
<emphasis id="B945EAE3B545FF8DB87B85A2FBBE2B9F" box="[1034,1049,1712,1735]" italics="true" pageId="2">L</emphasis>
directed to lateral sides of prodorsum,
<emphasis id="B945EAE3B545FF8DBD9A85C0FE5F2BB2" box="[491,504,1746,1770]" italics="true" pageId="2">S</emphasis>
curving backwards. Rostral and lamellar setae of medium size, setiform, slightly barbed. Interlamellar setae minute. Bothridial setae long, narrowly lanceolate, barbed. Dorsosejugal porose areas present. Dorsosejugal suture complete. Four pairs of small porose areas,
<emphasis id="B945EAE3B545FF8DB9F08405FA382A77" box="[1409,1439,1815,1839]" italics="true" pageId="2">Aa</emphasis>
oval, others rounded. Median pore absent. Epimeral setal formula 1-0-1-2. Epimeral and anogenital setae short, setiform, roughened. Circumpedal carinae short. Postanal porose area narrowly elongate oval. Solenidion on tibiae IV inserted in anterior part of the segment.
</paragraph>
</subSubSection>
<subSubSection id="C32B657AB545FF8ABE6284B3FDF02F4A" lastPageId="5" pageId="2" type="description">
<paragraph id="8B8E36F1B545FF8DBE6284B3FAC82AA5" blockId="2.[491,1522,1611,2045]" pageId="2">
<emphasis id="B945EAE3B545FF8DBE6284B3FD312AE1" bold="true" box="[531,662,1953,1977]" pageId="2">Description</emphasis>
— Measurements Large species. Body length 996 (
<typeStatus id="548A8853B545FF8DB89B84B2FAEE2AE0" box="[1258,1353,1952,1976]" pageId="2" type="holotype">holotype</typeStatus>
, female), 780 996 (
<specimenCount id="9D37FD78B545FF8DBE6984D1FD072A84" box="[536,672,1987,2012]" pageId="2" type="generic" typeStatus="paratypes">13 paratypes</specimenCount>
,
<specimenCount id="9D37FD78B545FF8DBEDB84D1FCB92A83" box="[682,798,1987,2011]" pageId="2" type="female">10 females</specimenCount>
and three males); notogaster width 763 (
<typeStatus id="548A8853B545FF8DB89184D1FAE72A83" box="[1248,1344,1987,2011]" pageId="2" type="holotype">holotype</typeStatus>
), 564780 (
<specimenCount id="9D37FD78B545FF8DB9C384D1FDF12AA5" pageId="2" type="generic" typeStatus="paratypes">13 paratypes</specimenCount>
). Females larger than males: 946996 × 747780 versus 780846 × 564614.
</paragraph>
<paragraph id="8B8E36F1B544FF8CBE6283F4FA2B2C1A" blockId="3.[491,1542,229,908]" pageId="3">
Integument Body color light brown to brown. Body surface densely microgranulate, granules (less than 1) poorly developed (visible only at high magnification
<date id="FF8F1031B544FF8CB94B821AFA262C78" box="[1338,1409,264,288]" pageId="3" value="1000-10">x1000</date>
). Antiaxial sides of all leg femora and trochanters III, IV with rounded and elongated tubercles.
</paragraph>
<paragraph id="8B8E36F1B544FF8CBE62825FFA2D2F0E" blockId="3.[491,1542,229,908]" pageId="3">
Prodorsum (
<figureCitation id="130A2A74B544FF8CBEE5825FFD552C3D" box="[660,754,333,357]" captionStart="Figure 1" captionStartId="3.[515,582,1939,1960]" captionTargetBox="[527,1473,971,1875]" captionTargetId="figure@3.[508,1492,961,2352]" captionTargetPageId="3" captionText="Figure 1 Galumna sandormahunkai n. sp., adult: a dorsal view; b ventral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3890499" httpUri="https://zenodo.org/record/3890499/files/figure.png" pageId="3">Figs 1a</figureCitation>
,
<figureCitation id="130A2A74B544FF8CBF70825FFCE22C3D" box="[769,837,333,357]" captionStart="Figure 2" captionStartId="4.[515,582,1472,1493]" captionTargetBox="[539,1474,239,1394]" captionTargetId="figure@4.[456,1495,185,1654]" captionTargetPageId="4" captionText="Figure 2 Galumna sandormahunkai n. sp., adult: a anterior part of body, lateral view (gnathosoma and legs omitted); b posterior part of body, lateral view; c anterior part of prodorsum, frontal view; d subcapitulum, ventral view; e palp, right, antiaxial view; f chelicera, left, paraxial view. Scale bar 100 μm (a, b), scale bar 50 μm (c, d, f), scale bar 20 μm (e)." figureDoi="http://doi.org/10.5281/zenodo.3890501" httpUri="https://zenodo.org/record/3890501/files/figure.png" pageId="3">2a, 2c</figureCitation>
) Rostrum strongly protruding (visible in frontal view), narrowly rounded. Lamellar and sublamellar lines slightly divergent distally,
<emphasis id="B945EAE3B544FF8CB9498262FAE02CDF" box="[1336,1351,368,391]" italics="true" pageId="3">L</emphasis>
thickened, directed to lateral sides of prodorsum,
<emphasis id="B945EAE3B544FF8CBFFF8280FC3C2CF2" box="[910,923,402,426]" italics="true" pageId="3">S</emphasis>
thin, curving backwards. Lateral structures
<emphasis id="B945EAE3B544FF8CB90E8281FA362CF2" box="[1407,1425,403,426]" italics="true" pageId="3">N</emphasis>
and ridges
<emphasis id="B945EAE3B544FF8CBE4582A7FDE32C94" box="[564,580,437,460]" italics="true" pageId="3">E</emphasis>
and
<emphasis id="B945EAE3B544FF8CBE0882A7FD2F2C94" box="[633,648,437,460]" italics="true" pageId="3">T</emphasis>
well developed. Rostral (4953) and lamellar (4953) setae setiform, slightly barbed. Interlamellar setae very short (46), setiform, thin, smooth. Bothridial setae (159168) with long stalk and short, narrowly lanceolate, barbed head. Exobothridial setae represented by alveoli. Dorsosejugal porose areas (2432 × 46) elongate oval, transversely oriented, located posterior or posterolateral to
<emphasis id="B945EAE3B544FF8CBF51812DFC922F0E" box="[800,821,575,598]" italics="true" pageId="3">in</emphasis>
. Dorsophragmata slightly elongated longitudinally.
</paragraph>
<paragraph id="8B8E36F1B544FF8CBE628173FDC82E1E" blockId="3.[491,1542,229,908]" pageId="3">
Notogaster (
<figureCitation id="130A2A74B544FF8CBEEA8172FD512F20" box="[667,758,608,632]" captionStart="Figure 1" captionStartId="3.[515,582,1939,1960]" captionTargetBox="[527,1473,971,1875]" captionTargetId="figure@3.[508,1492,961,2352]" captionTargetPageId="3" captionText="Figure 1 Galumna sandormahunkai n. sp., adult: a dorsal view; b ventral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3890499" httpUri="https://zenodo.org/record/3890499/files/figure.png" pageId="3">Figs 1a</figureCitation>
,
<figureCitation id="130A2A74B544FF8CBF708172FCE02F20" box="[769,839,608,632]" captionStart="Figure 2" captionStartId="4.[515,582,1472,1493]" captionTargetBox="[539,1474,239,1394]" captionTargetId="figure@4.[456,1495,185,1654]" captionTargetPageId="4" captionText="Figure 2 Galumna sandormahunkai n. sp., adult: a anterior part of body, lateral view (gnathosoma and legs omitted); b posterior part of body, lateral view; c anterior part of prodorsum, frontal view; d subcapitulum, ventral view; e palp, right, antiaxial view; f chelicera, left, paraxial view. Scale bar 100 μm (a, b), scale bar 50 μm (c, d, f), scale bar 20 μm (e)." figureDoi="http://doi.org/10.5281/zenodo.3890501" httpUri="https://zenodo.org/record/3890501/files/figure.png" pageId="3">2a, 2b</figureCitation>
) Dorsosejugal suture complete. With 10 pairs of setal alveoli or vestigial setae (up to 1) and four pairs of small porose areas having distinct borders,
<emphasis id="B945EAE3B544FF8CB9D38191FA672FC3" box="[1442,1472,643,667]" italics="true" pageId="3">Aa</emphasis>
(2226 × 1012) oval,
<emphasis id="B945EAE3B544FF8CBE9981B7FCA22FE5" box="[744,773,677,701]" italics="true" pageId="3">A1</emphasis>
,
<emphasis id="B945EAE3B544FF8CBF6481B7FC952FE5" box="[789,818,677,701]" italics="true" pageId="3">A2</emphasis>
and
<emphasis id="B945EAE3B544FF8CBF1B81B7FC202FE5" box="[874,903,677,701]" italics="true" pageId="3">A3</emphasis>
(1020) rounded. Porose areas
<emphasis id="B945EAE3B544FF8CB89881B7FAA02FE5" box="[1257,1287,677,701]" italics="true" pageId="3">Aa</emphasis>
located close to pteromorphal hinges, anteriorly to
<emphasis id="B945EAE3B544FF8CBF2F81D5FCD42F87" box="[862,883,711,735]" italics="true" pageId="3">la</emphasis>
. Median pore absent in females and males. Opisthonotal gland openings and all lyrifissures distinct,
<emphasis id="B945EAE3B544FF8CBFCF81F8FC442E5A" box="[958,995,746,770]" italics="true" pageId="3">gla</emphasis>
located anterolateral to
<emphasis id="B945EAE3B544FF8CB89581F8FAA62E5A" box="[1252,1281,746,770]" italics="true" pageId="3">A1</emphasis>
and removed from them,
<emphasis id="B945EAE3B544FF8CBE5E801FFDED2E7C" box="[559,586,781,804]" italics="true" pageId="3">im</emphasis>
anterior to
<emphasis id="B945EAE3B544FF8CBEB7801EFD432E7C" box="[710,740,780,804]" italics="true" pageId="3">A1</emphasis>
and slightly removed from them,
<emphasis id="B945EAE3B544FF8CB823801FFBC02E7C" box="[1106,1127,781,804]" italics="true" pageId="3">ip</emphasis>
lateral to
<emphasis id="B945EAE3B544FF8CB8A2801FFB472E7C" box="[1235,1248,781,804]" italics="true" pageId="3">p</emphasis>
<subScript id="17B534B4B544FF8CB891800AFB4E2E71" attach="left" box="[1248,1257,792,809]" fontSize="7" pageId="3">1</subScript>
,
<emphasis id="B945EAE3B544FF8CB888801EFAA92E7C" box="[1273,1294,780,804]" italics="true" pageId="3">ih</emphasis>
anterior to
<emphasis id="B945EAE3B544FF8CB9FB801FFA302E7C" box="[1418,1431,781,804]" italics="true" pageId="3">p</emphasis>
<subScript id="17B534B4B544FF8CB9E9800AFA392E71" attach="left" box="[1432,1438,792,809]" fontSize="7" pageId="3">3</subScript>
,
<emphasis id="B945EAE3B544FF8CB9C1801FFA682E7C" box="[1456,1487,781,804]" italics="true" pageId="3">ips</emphasis>
lateral to
<emphasis id="B945EAE3B544FF8CBE218022FDFA2E1F" box="[592,605,816,839]" italics="true" pageId="3">p</emphasis>
<subScript id="17B534B4B544FF8CBE2C8029FDC42E14" attach="left" box="[605,611,827,844]" fontSize="7" pageId="3">3</subScript>
.
</paragraph>
<paragraph id="8B8E36F1B544FF8BBE628043FC2F2BB2" blockId="3.[491,1542,229,908]" lastBlockId="4.[491,1528,1677,2045]" lastPageId="4" pageId="3">
Gnathosoma (
<figureCitation id="130A2A74B544FF8CBED68043FD522E31" box="[679,757,849,873]" captionStart="Figure 2" captionStartId="4.[515,582,1472,1493]" captionTargetBox="[539,1474,239,1394]" captionTargetId="figure@4.[456,1495,185,1654]" captionTargetPageId="4" captionText="Figure 2 Galumna sandormahunkai n. sp., adult: a anterior part of body, lateral view (gnathosoma and legs omitted); b posterior part of body, lateral view; c anterior part of prodorsum, frontal view; d subcapitulum, ventral view; e palp, right, antiaxial view; f chelicera, left, paraxial view. Scale bar 100 μm (a, b), scale bar 50 μm (c, d, f), scale bar 20 μm (e)." figureDoi="http://doi.org/10.5281/zenodo.3890501" httpUri="https://zenodo.org/record/3890501/files/figure.png" pageId="3">Figs 2</figureCitation>
d-f) Subcapitulum size 196205 × 180192. Subcapitular setae (
<emphasis id="B945EAE3B544FF8CB9CB8040FA602E31" box="[1466,1479,850,873]" italics="true" pageId="3">a</emphasis>
, 32;
<emphasis id="B945EAE3B544FF8CBE648067FD8F2ED4" box="[533,552,885,908]" italics="true" pageId="3">m</emphasis>
, 32;
<emphasis id="B945EAE3B544FF8CBE108066FDC92ED4" box="[609,622,884,908]" italics="true" pageId="3">h</emphasis>
, 24) setiform, slightly barbed;
<emphasis id="B945EAE3B544FF8CBFCC8067FC6D2ED4" box="[957,970,885,908]" italics="true" pageId="3">a</emphasis>
thickest,
<emphasis id="B945EAE3B544FF8CB8428066FBE72ED4" box="[1075,1088,884,908]" italics="true" pageId="3">h</emphasis>
thinnest. Adoral setae (24) setiform, barbed. Length of palps 139143. Postpalpal setae (8) spiniform, smooth. Length of chelicerae 237241. Cheliceral setae (
<emphasis id="B945EAE3B543FF8BBF6985BDFC992B9F" box="[792,830,1711,1735]" italics="true" pageId="4">cha</emphasis>
, 8286;
<emphasis id="B945EAE3B543FF8BBFD185BDFC612B9F" box="[928,966,1711,1735]" italics="true" pageId="4">chb</emphasis>
, 4549) setiform, barbed. Trägårdhs organ of chelicerae long, elongate triangular.
</paragraph>
<caption id="DF4E6679B544FF8CBE728481FCD12AE7" ID-DOI="http://doi.org/10.5281/zenodo.3890499" ID-Zenodo-Dep="3890499" httpUri="https://zenodo.org/record/3890499/files/figure.png" pageId="3" targetBox="[527,1473,971,1875]" targetPageId="3">
<paragraph id="8B8E36F1B544FF8CBE728481FCD12AE7" pageId="3">
<emphasis id="B945EAE3B544FF8CBE728481FDFD2AF0" bold="true" box="[515,602,1939,1960]" pageId="3">Figure 1</emphasis>
<taxonomicName id="4C314D72B544FF8CBE188481FCCD2AF1" authority="Ermilov &amp; Starý, 2020" authorityName="Ermilov &amp; Starý" authorityYear="2020" box="[617,874,1939,1961]" class="Arachnida" family="Galumnidae" genus="Galumna" kingdom="Animalia" order="Sarcoptiformes" pageId="3" phylum="Arthropoda" rank="species" species="sandormahunkai" status="sp. nov.">
<emphasis id="B945EAE3B544FF8CBE188481FCCD2AF1" box="[617,874,1939,1961]" italics="true" pageId="3">Galumna sandormahunkai</emphasis>
</taxonomicName>
<emphasis id="B945EAE3B544FF8CBF008481FC0A2AF0" bold="true" box="[881,941,1939,1960]" pageId="3">
<taxonomicNameLabel id="A2765798B544FF8CBF008481FC0A2AF0" box="[881,941,1939,1960]" pageId="3" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult: a dorsal view; b ventral view (gnathosoma and legs omitted). Scale bar 100 μm.
</paragraph>
</caption>
<caption id="DF4E6679B543FF8BBE7286D2FBD02B41" ID-DOI="http://doi.org/10.5281/zenodo.3890501" ID-Zenodo-Dep="3890501" httpUri="https://zenodo.org/record/3890501/files/figure.png" pageId="4" targetBox="[539,1474,239,1394]" targetPageId="4">
<paragraph id="8B8E36F1B543FF8BBE7286D2FBD02B41" pageId="4">
<emphasis id="B945EAE3B543FF8BBE7286D2FDFF288D" bold="true" box="[515,600,1472,1493]" pageId="4">Figure 2</emphasis>
<taxonomicName id="4C314D72B543FF8BBE1086ADFCC7288D" authority="Ermilov &amp; Starý, 2020" authorityName="Ermilov &amp; Starý" authorityYear="2020" box="[609,864,1471,1493]" class="Arachnida" family="Galumnidae" genus="Galumna" kingdom="Animalia" order="Sarcoptiformes" pageId="4" phylum="Arthropoda" rank="species" species="sandormahunkai" status="sp. nov.">
<emphasis id="B945EAE3B543FF8BBE1086ADFCC7288D" box="[609,864,1471,1493]" italics="true" pageId="4">Galumna sandormahunkai</emphasis>
</taxonomicName>
<emphasis id="B945EAE3B543FF8BBF1486D2FC3A288D" bold="true" box="[869,925,1472,1493]" pageId="4">
<taxonomicNameLabel id="A2765798B543FF8BBF1486D2FC3A288D" box="[869,925,1472,1493]" pageId="4" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult: a anterior part of body, lateral view (gnathosoma and legs omitted); b posterior part of body, lateral view; c anterior part of prodorsum, frontal view; d subcapitulum, ventral view; e palp, right, antiaxial view; f chelicera, left, paraxial view. Scale bar 100 μm (a, b), scale bar 50 μm (c, d, f), scale bar 20 μm (e).
</paragraph>
</caption>
<paragraph id="8B8E36F1B543FF8BBE6285E6FA682A2B" blockId="4.[491,1528,1677,2045]" pageId="4">
Epimeral and lateral podosomal regions (
<figureCitation id="130A2A74B543FF8BBFA785E6FB9F2A54" box="[982,1080,1780,1804]" captionStart="Figure 1" captionStartId="3.[515,582,1939,1960]" captionTargetBox="[527,1473,971,1875]" captionTargetId="figure@3.[508,1492,961,2352]" captionTargetPageId="3" captionText="Figure 1 Galumna sandormahunkai n. sp., adult: a dorsal view; b ventral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3890499" httpUri="https://zenodo.org/record/3890499/files/figure.png" pageId="4">Figs 1b</figureCitation>
,
<figureCitation id="130A2A74B543FF8BB83685E6FBC62A54" box="[1095,1121,1780,1804]" captionStart="Figure 2" captionStartId="4.[515,582,1472,1493]" captionTargetBox="[539,1474,239,1394]" captionTargetId="figure@4.[456,1495,185,1654]" captionTargetPageId="4" captionText="Figure 2 Galumna sandormahunkai n. sp., adult: a anterior part of body, lateral view (gnathosoma and legs omitted); b posterior part of body, lateral view; c anterior part of prodorsum, frontal view; d subcapitulum, ventral view; e palp, right, antiaxial view; f chelicera, left, paraxial view. Scale bar 100 μm (a, b), scale bar 50 μm (c, d, f), scale bar 20 μm (e)." figureDoi="http://doi.org/10.5281/zenodo.3890501" httpUri="https://zenodo.org/record/3890501/files/figure.png" pageId="4">2a</figureCitation>
) Anterior margin of epimere I smooth. Epimeral setal formula 1-0-1-2. Epimeral setae setiform, thin, roughened,
<emphasis id="B945EAE3B543FF8BB9138405FAD92A77" box="[1378,1406,1815,1839]" italics="true" pageId="4">3b</emphasis>
(3032) longer than
<emphasis id="B945EAE3B543FF8BBE1A842BFD212A09" box="[619,646,1849,1873]" italics="true" pageId="4">1b</emphasis>
,
<emphasis id="B945EAE3B543FF8BBEE5842BFD082A09" box="[660,687,1849,1873]" italics="true" pageId="4">4a</emphasis>
and
<emphasis id="B945EAE3B543FF8BBE92842BFCA72A09" box="[739,768,1849,1873]" italics="true" pageId="4">4b</emphasis>
(1820). Pedotecta I broadly rounded, pedotecta II quadringular in ventral view. Discidia triangular. Circumpedal carinae short, thin, directed to acetabula IV.
</paragraph>
<paragraph id="8B8E36F1B543FF8ABE62846CFCD42C1B" blockId="4.[491,1528,1677,2045]" lastBlockId="5.[491,1562,229,1186]" lastPageId="5" pageId="4">
Anogenital region (
<figureCitation id="130A2A74B543FF8BBE82846CFCF32ACE" box="[755,852,1918,1942]" captionStart="Figure 1" captionStartId="3.[515,582,1939,1960]" captionTargetBox="[527,1473,971,1875]" captionTargetId="figure@3.[508,1492,961,2352]" captionTargetPageId="3" captionText="Figure 1 Galumna sandormahunkai n. sp., adult: a dorsal view; b ventral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3890499" httpUri="https://zenodo.org/record/3890499/files/figure.png" pageId="4">Figs 1b</figureCitation>
,
<figureCitation id="130A2A74B543FF8BBF10846CFC272ACE" box="[865,896,1918,1942]" captionStart="Figure 2" captionStartId="4.[515,582,1472,1493]" captionTargetBox="[539,1474,239,1394]" captionTargetId="figure@4.[456,1495,185,1654]" captionTargetPageId="4" captionText="Figure 2 Galumna sandormahunkai n. sp., adult: a anterior part of body, lateral view (gnathosoma and legs omitted); b posterior part of body, lateral view; c anterior part of prodorsum, frontal view; d subcapitulum, ventral view; e palp, right, antiaxial view; f chelicera, left, paraxial view. Scale bar 100 μm (a, b), scale bar 50 μm (c, d, f), scale bar 20 μm (e)." figureDoi="http://doi.org/10.5281/zenodo.3890501" httpUri="https://zenodo.org/record/3890501/files/figure.png" pageId="4">2b</figureCitation>
) Six pairs of genital setae
<emphasis id="B945EAE3B543FF8BB8E9846DFB012ACE" box="[1176,1190,1919,1942]" italics="true" pageId="4">g</emphasis>
<subScript id="17B534B4B543FF8BB8D78498FB082AC3" attach="both" box="[1190,1199,1930,1947]" fontSize="7" pageId="4">1</subScript>
, (
<emphasis id="B945EAE3B543FF8BB8CF846DFB6B2ACE" box="[1214,1228,1919,1942]" italics="true" pageId="4">g</emphasis>
<subScript id="17B534B4B543FF8BB8BD8498FB722AC3" attach="left" box="[1228,1237,1930,1947]" fontSize="7" pageId="4">2</subScript>
, 2628;
<emphasis id="B945EAE3B543FF8BB947846DFAE32ACE" box="[1334,1348,1919,1942]" italics="true" pageId="4">g</emphasis>
<subScript id="17B534B4B543FF8BB9358498FAED2AC3" attach="left" box="[1348,1354,1930,1947]" fontSize="7" pageId="4">3</subScript>
<emphasis id="B945EAE3B543FF8BB92D846DFACD2ACE" box="[1372,1386,1919,1942]" italics="true" pageId="4">g</emphasis>
<subScript id="17B534B4B543FF8BB91B8498FAD42AC3" attach="left" box="[1386,1395,1930,1947]" fontSize="7" pageId="4">6</subScript>
, 1820), one pair of aggenital (1820), two pairs of anal (1820) and three pairs of adanal (1820) setae setiform, thin, roughened. Anterior edge of genital plates with two setae. Aggenital setae located between genital and anal apertures, nearer to genital aperture. Adanal lyrifissures located close and parallel to anal plates. Adanal setae
<emphasis id="B945EAE3B542FF8AB84D83F7FBF02DA5" box="[1084,1111,229,253]" italics="true" pageId="5">ad</emphasis>
<subScript id="17B534B4B542FF8AB82983E0FBC62C5B" attach="left" box="[1112,1121,242,259]" fontSize="7" pageId="5">1</subScript>
and
<emphasis id="B945EAE3B542FF8AB8E883F7FB132DA5" box="[1177,1204,229,253]" italics="true" pageId="5">ad</emphasis>
<subScript id="17B534B4B542FF8AB8C483E0FB192C5B" attach="left" box="[1205,1214,242,259]" fontSize="7" pageId="5">2</subScript>
posterior,
<emphasis id="B945EAE3B542FF8AB94583F7FAE82DA5" box="[1332,1359,229,253]" italics="true" pageId="5">ad</emphasis>
<subScript id="17B534B4B542FF8AB92183E0FAF12C5B" attach="left" box="[1360,1366,242,259]" fontSize="7" pageId="5">3</subScript>
lateral to anal aperture. Distance
<emphasis id="B945EAE3B542FF8ABE9B821AFCA22C78" box="[746,773,264,288]" italics="true" pageId="5">ad</emphasis>
<subScript id="17B534B4B542FF8ABF778206FCA82C7D" attach="left" box="[774,783,276,293]" fontSize="7" pageId="5">1</subScript>
<emphasis id="B945EAE3B542FF8ABF6C821AFC9F2C78" box="[797,824,264,288]" italics="true" pageId="5">ad</emphasis>
<subScript id="17B534B4B542FF8ABF488206FCE52C7D" attach="left" box="[825,834,276,293]" fontSize="7" pageId="5">2</subScript>
slightly shorter than
<emphasis id="B945EAE3B542FF8AB856821AFBE52C78" box="[1063,1090,264,288]" italics="true" pageId="5">ad</emphasis>
<subScript id="17B534B4B542FF8AB8338206FBEC2C7D" attach="left" box="[1090,1099,276,293]" fontSize="7" pageId="5">2</subScript>
<emphasis id="B945EAE3B542FF8AB82B821AFBD22C78" box="[1114,1141,264,288]" italics="true" pageId="5">ad</emphasis>
<subScript id="17B534B4B542FF8AB8078206FBDB2C7D" attach="left" box="[1142,1148,276,293]" fontSize="7" pageId="5">3</subScript>
. Unpaired postanal porose area narrowly elongate oval (4149 × 46).
</paragraph>
<paragraph id="8B8E36F1B542FF8ABE62825CFDF02F4A" blockId="5.[491,1562,229,1186]" pageId="5">
Legs (
<figureCitation id="130A2A74B542FF8ABE13825FFD792C3D" box="[610,734,333,357]" captionStart="Figure 3" captionStartId="6.[515,582,1666,1687]" captionTargetBox="[598,1440,221,1600]" captionTargetId="figure@6.[503,1487,221,1612]" captionTargetPageId="6" captionText="Figure 3 Galumna sandormahunkai n. sp., adult: a leg I, without trochanter, right, antiaxial view; b leg IV, left, antiaxial view. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.3890503" httpUri="https://zenodo.org/record/3890503/files/figure.png" pageId="5">Figs 3a, 3b</figureCitation>
) Median claw distinctly thicker than lateral claws, all slightly barbed on dorsal side. Porose area on all femora and on trochanters III, IV well visible. Formulae of leg setation and solenidia I (
<date id="FF8F1031B542FF8ABE818280FCFF2CF2" box="[752,856,402,426]" pageId="5" value="1920-03-01" valueMax="1920-04-04" valueMin="1920-03-01">1-4-3-4-20</date>
) [1-2-2], II (
<date id="FF8F1031B542FF8ABF908280FBEE2CF2" box="[993,1097,402,426]" pageId="5" value="2015-03-01" valueMax="2015-04-04" valueMin="2015-03-01">1-4-3-4-15</date>
) [1-1-2], III (
<date id="FF8F1031B542FF8AB8AC8280FAEB2CF2" box="[1245,1356,402,426]" pageId="5" value="2015-01-01" valueMax="2015-03-02" valueMin="2015-01-01">1-2-1-3-15</date>
) [1-1-0], IV (
<date id="FF8F1031B542FF8ABE6782A7FD222C95" box="[534,645,437,461]" pageId="5" value="2012-02-01" valueMax="2012-03-02" valueMin="2012-02-01">1-2-2-3-12</date>
) [0-1-0]; homology of setae and solenidia indicated in
<tableCitation id="C6B3034AB542FF8AB8BE82A7FA852C95" box="[1231,1314,437,461]" captionStart="Table 1" captionStartId="5.[119,182,1646,1667]" captionTargetBox="[132,1425,1747,1949]" captionTargetPageId="5" captionText="Table 1 Leg setation and solenidia of adultGalumna sandormahunkai n. sp. andPergalumna janosbaloghin. sp." httpUri="http://table.plazi.org/id/DF4E6679B542FF8ABC06857CFB652BDB" pageId="5" tableUuid="DF4E6679B542FF8ABC06857CFB652BDB">Table 1</tableCitation>
. Famulus on tarsi I inserted between solenidia ω
<subScript id="17B534B4B542FF8ABF5F82F1FC902CAC" attach="left" box="[814,823,483,500]" fontSize="7" pageId="5">1</subScript>
and ω
<subScript id="17B534B4B542FF8ABF0F82F1FC202CAC" attach="left" box="[894,903,483,500]" fontSize="7" pageId="5">2</subScript>
. Solenidion on tibiae IV inserted in anterior part of the segment.
</paragraph>
</subSubSection>
<subSubSection id="C32B657AB542FF8ABE62810FFCEA2E7E" pageId="5" type="materials_examined">
<paragraph id="8B8E36F1B542FF8ABE62810FFAE22FC4" blockId="5.[491,1562,229,1186]" pageId="5">
<materialsCitation id="3B593CACB542FF8ABE62810FFAE62FC4" ID-GBIF-Occurrence="2634683306" collectedFrom="sifting of leaf litter sample under big unidentified tree" collectingDate="2014-01-13" collectingMethod="Winkler apparatus extraction" country="Madagascar" latitude="-12.524445" location="National Park" longLatPrecision="21" longitude="49.164448" municipality="Montagne" pageId="5" specimenCount="962" specimenCount-female="11" specimenCount-male="951" typeStatus="holotype">
<emphasis id="B945EAE3B542FF8ABE62810FFD562F6D" bold="true" box="[531,753,541,565]" pageId="5">Material examined</emphasis>
<typeStatus id="548A8853B542FF8ABF62810EFCCA2F6C" box="[787,877,540,564]" pageId="5" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D37FD78B542FF8ABF0A810EFC602F6C" box="[891,967,540,564]" pageId="5" type="female">female</specimenCount>
) and 13
<typeStatus id="548A8853B542FF8AB852810FFB2E2F6D" box="[1059,1161,541,565]" pageId="5" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D37FD78B542FF8AB8E9810EFAAA2F6C" box="[1176,1293,540,564]" pageId="5" type="female">10 females</specimenCount>
and three
<specimenCount id="9D37FD78B542FF8AB9F7810EFA6E2F6C" box="[1414,1481,540,564]" pageId="5" type="male">males</specimenCount>
): North
<collectingCountry id="F3267661B542FF8ABE42812DFD642F0F" box="[563,707,575,599]" name="Madagascar" pageId="5" value="Madagascar">Madagascar</collectingCountry>
,
<collectingMunicipality id="6BEAAC8BB542FF8ABEA18152FCE42F00" box="[720,835,576,600]" pageId="5">Montagne</collectingMunicipality>
dAmbre
<location id="8EEE602AB542FF8ABFCA812DFBF72F0F" LSID="urn:lsid:plazi:treatment:039887E7B545FF8ABD9A8503FC8229FA:8EEE602AB542FF8ABFCA812DFBF72F0F" box="[955,1104,575,599]" country="Madagascar" latitude="-12.524445" longLatPrecision="21" longitude="49.164448" municipality="Montagne" name="National Park" pageId="5">National Park</location>
, circuit
<location id="8EEE602AB542FF8AB8D8812DFA882F0F" LSID="urn:lsid:plazi:treatment:039887E7B545FF8ABD9A8503FC8229FA:8EEE602AB542FF8AB8D8812DFA882F0F" box="[1193,1327,575,599]" country="Madagascar" latitude="-12.524445" longLatPrecision="21" longitude="49.164448" municipality="Montagne" name="Ampijoroana" pageId="5">Ampijoroana</location>
, evergreen rain forest,
<geoCoordinate id="EE055036B542FF8ABE418173FD012F21" box="[560,678,609,633]" degrees="12" direction="south" minutes="31" orientation="latitude" pageId="5" precision="15" seconds="28" value="-12.524445">12°3128”S</geoCoordinate>
,
<geoCoordinate id="EE055036B542FF8ABEDE8173FC832F21" box="[687,804,609,633]" degrees="49" direction="east" minutes="09" orientation="longitude" pageId="5" precision="15" seconds="52" value="49.164448">49°0952”E</geoCoordinate>
,
<quantity id="4CC99B14B542FF8ABF5F8173FCCB2F21" box="[814,876,609,633]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="5" unit="m" value="950.0">
<specimenCount id="9D37FD78B542FF8ABF5F8173FCCB2F21" box="[814,876,609,633]" pageId="5" type="male">950 m</specimenCount>
</quantity>
a.s.l.,
<collectedFrom id="382D750CB542FF8ABFC18173FDB22FC4" pageId="5">sifting of leaf litter sample under big unidentified tree</collectedFrom>
,
<collectingMethod id="52704EE6B542FF8ABE508196FCF12FC4" box="[545,854,644,668]" pageId="5">Winkler apparatus extraction</collectingMethod>
,
<date id="FF8F1031B542FF8ABF118196FC1A2FC4" box="[864,957,644,668]" pageId="5" value="2014-01-13">
<collectingDate id="EFCBE9D9B542FF8ABF118196FC1A2FC4" box="[864,957,644,668]" pageId="5" value="2014-01-13">13.I.2014</collectingDate>
</date>
(R. Ravebolun and L. Rabotenoson)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8E36F1B542FF8ABE6281B5FCEA2E7E" blockId="5.[491,1562,229,1186]" pageId="5">
<materialsCitation id="3B593CACB542FF8ABE6281B5FC0B2FB9" ID-GBIF-Occurrence="2634683304" country="Germany" location="Germany" pageId="5" specimenCount="3" typeStatus="holotype">
<emphasis id="B945EAE3B542FF8ABE6281B5FD6F2FE7" bold="true" box="[531,712,679,703]" pageId="5">
<typeStatus id="548A8853B542FF8ABE6281B5FDEB2FE7" box="[531,588,679,703]" pageId="5">Type</typeStatus>
deposition
</emphasis>
— The
<typeStatus id="548A8853B542FF8ABF5781B4FC272FE6" box="[806,896,678,702]" pageId="5" type="holotype">holotype</typeStatus>
and two
<typeStatus id="548A8853B542FF8ABF9081B5FBE02FE7" box="[993,1095,679,703]" pageId="5" type="paratype">paratypes</typeStatus>
are deposited in the collection of the Senckenberg Institute, Görlitz,
<collectingCountry id="F3267661B542FF8ABF3681DBFC0B2FB9" box="[839,940,713,737]" name="Germany" pageId="5" value="Germany">Germany</collectingCountry>
</materialsCitation>
.
<materialsCitation id="3B593CACB542FF8ABFC981DBFB2A2E5C" ID-GBIF-Occurrence="2634683303" country="Russia" location="Tyumen State" pageId="5" specimenCount="1" stateProvince="Tyumen State" typeStatus="paratype">
Eleven
<typeStatus id="548A8853B542FF8AB87681D8FBCA2FBA" box="[1031,1133,714,738]" pageId="5" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="49F5F813B542FF8ABD9A81FEFDD92E5C" box="[491,638,748,772]" country="Russia" name="Tyumen'" pageId="5">Tyumen State</collectingRegion>
University Museum of Zoology,
<collectingRegion id="49F5F813B542FF8ABFAF81FEFB9F2E5C" box="[990,1080,748,772]" country="Russia" name="Tyumen'" pageId="5">Tyumen</collectingRegion>
,
<collectingCountry id="F3267661B542FF8AB83281FEFB2A2E5C" box="[1091,1165,748,772]" name="Russia" pageId="5" value="Russia">Russia</collectingCountry>
</materialsCitation>
. All specimens are preserved in ethanol with drop of glycerol.
</paragraph>
</subSubSection>
<subSubSection id="C32B657AB542FF8ABE628023FA962E33" pageId="5" type="etymology">
<paragraph id="8B8E36F1B542FF8ABE628023FA962E33" blockId="5.[491,1562,229,1186]" pageId="5">
<emphasis id="B945EAE3B542FF8ABE628023FD372E11" bold="true" box="[531,656,817,841]" pageId="5">Etymology</emphasis>
— The new species is named after late Prof. Dr. S. Mahunka, the Hungarian acarologist, for his extensive contributions to our knowledge of oribatid mites.
</paragraph>
</subSubSection>
<subSubSection id="C32B657AB542FF8ABE628064FC8229FA" pageId="5" type="discussion">
<paragraph id="8B8E36F1B542FF8ABE628064FC8229FA" blockId="5.[491,1562,229,1186]" pageId="5">
<emphasis id="B945EAE3B542FF8ABE628064FD252ED6" bold="true" box="[531,642,886,910]" pageId="5">Remarks</emphasis>
— In the presence of long rostral and lamellar setae, short interlamellar setae, long bothridial setae with developed head, complete dorsosejugal suture, and four pairs of rounded notogastral porose areas,
<taxonomicName id="4C314D72B542FF8ABF2A80A9FBDF2E8B" authority="Ermilov &amp; Starý, 2020" authorityName="Ermilov &amp; Starý" authorityYear="2020" box="[859,1144,955,979]" class="Arachnida" family="Galumnidae" genus="Galumna" kingdom="Animalia" order="Sarcoptiformes" pageId="5" phylum="Arthropoda" rank="species" species="sandormahunkai" status="sp. nov.">
<emphasis id="B945EAE3B542FF8ABF2A80A9FBDF2E8B" box="[859,1144,955,979]" italics="true" pageId="5">Galumna sandormahunkai</emphasis>
</taxonomicName>
<emphasis id="B945EAE3B542FF8AB80E80A9FB1B2E8B" bold="true" box="[1151,1212,955,979]" pageId="5">
<taxonomicNameLabel id="A2765798B542FF8AB80E80A9FB1B2E8B" box="[1151,1212,955,979]" pageId="5" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is morphologically most similar to
<taxonomicName id="4C314D72B542FF8ABE2980CFFB7F2EAD" authority="Ermilov, Hugo-Coetzee and Theron, 2018" authorityName="Ermilov, Hugo-Coetzee and Theron" authorityYear="2018" box="[600,1240,989,1013]" class="Arachnida" family="Galumnidae" genus="Galumna" kingdom="Animalia" order="Sarcoptiformes" pageId="5" phylum="Arthropoda" rank="species" species="sphagni">
<emphasis id="B945EAE3B542FF8ABE2980CFFCBF2EAD" box="[600,792,989,1013]" italics="true" pageId="5">Galumna sphagni</emphasis>
<bibRefCitation id="EFA04B00B542FF8ABF6D80CFFB7F2EAD" author="Ermilov S. G. &amp; Hugo-Coetzee E. A. &amp; Theron P. D." box="[796,1240,989,1013]" journalOrPublisher="Zool. Zh." pageId="5" pagination="515 - 527" part="97" refId="ref5548" refString="Ermilov S. G., Hugo-Coetzee E. A., Theron P. D. 2018. To the knowledge of oribatid mites of the subgenus Galumna (Galumna) Heyden 1826 (Acari, Oribatida, Galumnidae) in South Africa, with a key to species known from the Ethiopian region. Zool. Zh., 97 (5): 515 - 527. doi: 10.7868 / S 0044513418050033" title="To the knowledge of oribatid mites of the subgenus Galumna (Galumna) Heyden 1826 (Acari, Oribatida, Galumnidae) in South Africa, with a key to species known from the Ethiopian region" type="journal article" year="2018">Ermilov, Hugo-Coetzee and Theron, 2018</bibRefCitation>
</taxonomicName>
from South Africa, but differs from the letter by the larger body size (780996 × 564780 versus 415431 × 315332), the presence of strongly protruding rostrum (versus not protruding), lanceolate, pointed apically bothridial setae (versus unilaterally dilated, rounded apically), the direction of lamellar lines to lateral sides of the prodorsum (versus anterior part of the ventral plate), and the absence of median pore (versus present).
</paragraph>
</subSubSection>
</treatment>
</document>