299 lines
36 KiB
XML
299 lines
36 KiB
XML
<document id="5B8DF5E56061B2E1FCBC4093FC038589" ID-DOI="10.11646/zootaxa.4819.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4396744" ID-ZooBank="80258C53-F94D-46CB-9E86-AF0C3226DF9E" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1609199984737" checkinUser="plazi" docAuthor="Polizei, Thiago T. S., Barclay, Maxwell V. L. & Bispo, Pitágoras C." docDate="2020" docId="039887D2FF85AF7E2A8A3C04FCBEF8D7" docLanguage="en" docName="zootaxa.4819.1.2.pdf" docOrigin="Zootaxa 4819 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexacylloepus maierae Polizei & Barclay & Bispo 2020, sp. nov." docType="treatment" docUuid="3325B8D0-1CB4-4DDE-8010-9E357098387D" docUuidSource="ZooBank" docVersion="1" lastPageNumber="26" masterDocId="FFA1FFAAFF97AF6A2A1D3A09FFC5FF9A" masterDocTitle="Hexacylloepus Hinton, 1940 rises again; 18 new species of a neglected genus (Coleoptera: Byrrhoidea: Elmidae)" masterLastPageNumber="48" masterPageNumber="6" pageNumber="24" updateTime="1730286293773" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:titleInfo id="65912D43B04B050A1CA34CB3B67B4747">
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<mods:title id="38D5BA3892BFA0CB5880771A51B3B530">Hexacylloepus Hinton, 1940 rises again; 18 new species of a neglected genus (Coleoptera: Byrrhoidea: Elmidae)</mods:title>
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<mods:name id="D48EBBBEF82328926EEE1F3D63258F6D" type="personal">
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<mods:roleTerm id="FA21537B4940773728A712F2E97EDF37">Author</mods:roleTerm>
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<mods:namePart id="68FA219B6AAAA930D0C70456F2FE613C">Polizei, Thiago T. S.</mods:namePart>
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<mods:nameIdentifier id="03944B963BE3D2137039AF2D2E14F9CE" type="ORCID">0000-0003-3746-8865</mods:nameIdentifier>
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<mods:affiliation id="4B3BA20619A36F0D14458DFE7257FCF1">Laboratório de Coleoptera (LaC) - Museu de Zoologia da Universidade de São Paulo (MZSP). São Paulo, São Paulo State, Brazil. thiagopolizei @ hotmail. com; https: // orcid. org / 0000 - 0003 - 3746 - 8865 & Programa de Pós-Graduação em Entomologia-Departamento de Biologia - Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto (FFCL-RP) - Universidade de São Paulo (USP). Ribeirão Preto, São Paulo State, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="E54DB9F5CE94ABD9AF714D31D70089E3" type="email">thiagopolizei@hotmail.com</mods:nameIdentifier>
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<mods:roleTerm id="E695CB3B37B59527C6380539CE698742">Author</mods:roleTerm>
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<mods:namePart id="A9A64222DC3A8EC53244CC8A7BEA388A">Barclay, Maxwell V. L.</mods:namePart>
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<mods:affiliation id="E60307BDE17FA0C8E0519D9735A43DC9">Department of Life Sciences, Natural History Museum. London SW 7 5 BD, United Kingdom.</mods:affiliation>
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</mods:name>
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<mods:name id="304AE649069A7A3D42E2CCE73E46A0A7" type="personal">
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<mods:roleTerm id="D484AD13E8EA981D2A657F6E4DB800A1">Author</mods:roleTerm>
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<mods:namePart id="A392B70197B3FD185EA554E76F2668C4">Bispo, Pitágoras C.</mods:namePart>
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<mods:affiliation id="67CEB3F79928D690E59A02D0B8909731">Programa de Pós-Graduação em Entomologia-Departamento de Biologia - Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto (FFCL-RP) - Universidade de São Paulo (USP). Ribeirão Preto, São Paulo State, Brazil. & Laboratório de Biologia Aquática (LABIA). Departamento de Ciências Biológicas - Faculdade de Ciências e Letras de Assis (FCL) - Universidade Estadual Paulista “ Júlio de Mesquita Filho ” (Unesp). Assis, São Paulo State, Brazil.</mods:affiliation>
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<mods:title id="B21606D350163D2CA070C403C1298BC0">Zootaxa</mods:title>
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<mods:part id="5318ABB9F0465CDF5391AB9CA2EAD48C">
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<mods:date id="B168573FB8BFDE93E7CB22C1AF2F450D">2020</mods:date>
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<mods:number id="C2C32CC4DC9D0087B505896921EE9C01">2020-07-22</mods:number>
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<mods:detail id="1918086867E7D53431D2A1A03A44AD24" type="volume">
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<mods:number id="BDF2849B3D9EA8A005385BD74FBC1089">4819</mods:number>
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<mods:classification id="919C0B1131387B490468CBE47D52EB83">journal article</mods:classification>
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<mods:identifier id="236A88BE2B8EA4F3223CCD8C15EB0FF9" type="DOI">10.11646/zootaxa.4819.1.2</mods:identifier>
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<mods:identifier id="334ED8829703B3491BC3DF5530F6DEB6" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039887D2FF85AF7E2A8A3C04FCBEF8D7" LSID="urn:lsid:zoobank.org:act:3325B8D0-1CB4-4DDE-8010-9E357098387D" httpUri="http://treatment.plazi.org/id/039887D2FF85AF7E2A8A3C04FCBEF8D7" lastPageId="20" lastPageNumber="26" pageId="18" pageNumber="24">
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<subSubSection id="C32B654FFF85AF782A8A3C04FDD5F9BD" box="[151,528,1549,1575]" pageId="18" pageNumber="24" type="nomenclature">
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<paragraph id="8B8E36C4FF85AF782A8A3C04FDD5F9BD" blockId="18.[151,528,1549,1611]" box="[151,528,1549,1575]" pageId="18" pageNumber="24">
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<taxonomicName id="4C314D47FF85AF782A8A3C04FE6EF9BD" authority="Polizei & Barclay & Bispo, 2020" authorityName="Polizei & Barclay & Bispo" authorityYear="2020" box="[151,427,1549,1575]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="24" phylum="Arthropoda" rank="species" species="maierae" status="sp. nov.">
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<emphasis id="B945EAD6FF85AF782A8A3C04FE6EF9BD" box="[151,427,1549,1575]" italics="true" pageId="18" pageNumber="24">Hexacylloepus maierae</emphasis>
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</taxonomicName>
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<emphasis id="B945EAD6FF85AF782BAE3C04FDD5F9BD" bold="true" box="[435,528,1549,1575]" pageId="18" pageNumber="24">
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<taxonomicNameLabel id="A27657ADFF85AF782BAE3C04FDD5F9BD" box="[435,528,1549,1575]" pageId="18" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B654FFF85AF782A8A3C38FEE3F9D1" box="[151,294,1585,1611]" pageId="18" pageNumber="24" type="description">
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<paragraph id="8B8E36C4FF85AF782A8A3C38FEE3F9D1" blockId="18.[151,528,1549,1611]" box="[151,294,1585,1611]" pageId="18" pageNumber="24">
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(
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<figureCitation id="130A2A41FF85AF782ABD3C38FF27F9D1" box="[160,226,1585,1611]" captionStart="FIGURE 9" captionStartId="19.[151,250,1917,1941]" captionTargetBox="[189,1362,382,1885]" captionTargetId="figure-115@19.[183,1404,381,1893]" captionTargetPageId="19" captionText="FIGURE 9. Hexacylloepus maierae sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396764" httpUri="https://zenodo.org/record/4396764/files/figure.png" pageId="18" pageNumber="24">Fig. 9</figureCitation>
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A–C)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B654FFF85AF782A8A3C7DFD4AF914" box="[151,655,1652,1678]" pageId="18" pageNumber="24" type="materials_examined">
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<paragraph id="8B8E36C4FF85AF782A8A3C7DFD4AF914" blockId="18.[151,1437,1652,2039]" box="[151,655,1652,1678]" pageId="18" pageNumber="24">
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<emphasis id="B945EAD6FF85AF782A8A3C7DFEFCF914" bold="true" box="[151,313,1652,1678]" pageId="18" pageNumber="24">
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<typeStatus id="548A8866FF85AF782A8A3C7DFF17F914" box="[151,210,1652,1678]" pageId="18" pageNumber="24">Type</typeStatus>
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locality:
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</emphasis>
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<collectingCountry id="F3267654FF85AF782B5D3C7DFE6BF914" box="[320,430,1652,1678]" name="Suriname" pageId="18" pageNumber="24">Suriname</collectingCountry>
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,
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<collectingRegion id="49F5F826FF85AF782BA73C7DFD49F914" box="[442,652,1652,1678]" country="Suriname" name="Sipaliwini" pageId="18" pageNumber="24">Sipaliwini District</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B654FFF85AF782ADA3C91FE1DF834" pageId="18" pageNumber="24" type="diagnosis">
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<paragraph id="8B8E36C4FF85AF782ADA3C91FE1DF834" blockId="18.[151,1437,1652,2039]" pageId="18" pageNumber="24">
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<emphasis id="B945EAD6FF85AF782ADA3C91FEFFF928" bold="true" box="[199,314,1688,1714]" pageId="18" pageNumber="24">Diagnosis</emphasis>
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: Antennae with thin setae at apex of each antennomere. Frontoclypeal suture straight. Pronotum with a long, wide, deep impression on disc, reaching the anterior and posterior margins. Prosternum with a pair of reddish-brown spots. Hypomera with a belt of tomentum reaching the anterior and lateral margins. Epipleura with tomentum. Tibiae each with a pair of spurs at apex, protibiae each with a row of short spines on distal half; meso- and metatibiae each with a groove on distal half, with two rows: one of short spines and the other of long setae. Abdomen with the disc of the first ventrite without tomentum. Male genitalia sharp; phallobase twice as long as wide; parameres reaching 2/3 of the penis length; penis with anterior quarter constricted, and slightly tapering toward the apex, apex sharp.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B654FFF85AF7E2ADA3DB1FF08FAAB" lastPageId="20" lastPageNumber="26" pageId="18" pageNumber="24" type="description">
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<paragraph id="8B8E36C4FF85AF782ADA3DB1FC8CF86D" blockId="18.[151,1437,1652,2039]" pageId="18" pageNumber="24">
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<emphasis id="B945EAD6FF85AF782ADA3DB1FE9EF848" bold="true" box="[199,347,1976,2002]" pageId="18" pageNumber="24">Description:</emphasis>
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Male. Body elongated, subparallel. Yellow coloured, with the antenna, mouthparts and legs reddish-brown. Total length:
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<quantity id="4CC99B21FF85AF782BA43DD5FDE5F86D" box="[441,544,2012,2039]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" pageId="18" pageNumber="24" unit="mm" value="1.64">1.64 mm</quantity>
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. Greatest width:
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<quantity id="4CC99B21FF85AF7828C33DD5FC80F86D" box="[734,837,2012,2039]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="18" pageNumber="24" unit="mm" value="0.65">0.65 mm</quantity>
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.
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</paragraph>
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<paragraph id="8B8E36C4FF84AF792ADA3A9EFC18FEFF" blockId="19.[151,1437,151,357]" pageId="19" pageNumber="25">
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<emphasis id="B945EAD6FF84AF792ADA3A9EFEC8FF2B" box="[199,269,151,177]" italics="true" pageId="19" pageNumber="25">Head:</emphasis>
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surface with micropunctures. Antenna inserted between the anterior margin of the eyes and the frontoclypeal suture, filiform, with 11 antennomeres, each one with a few setae at apex. Frontoclypeal suture straight. Clypeus 3x as wide as long; anterior margin straight and posterior margin sinuous; surface similar to that of head. Labrum twice as wide as long; antero-lateral angles rounded; surface alutaceous with long, thin setae on lateral margin. Maxillary palpus 4-segmented; terminal palpomere truncated apically. Labial palpus 3-segmented. Mentum and submentum squared. Gula rectangular. Genae covered with tomentum.
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</paragraph>
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<caption id="DF4E664CFF84AF792A8A3D74FCF0F820" ID-DOI="http://doi.org/10.5281/zenodo.4396764" ID-Zenodo-Dep="4396764" httpUri="https://zenodo.org/record/4396764/files/figure.png" pageId="19" pageNumber="25" startId="19.[151,250,1917,1941]" targetBox="[189,1362,382,1885]" targetPageId="19">
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<paragraph id="8B8E36C4FF84AF792A8A3D74FCF0F820" blockId="19.[151,1436,1917,1978]" pageId="19" pageNumber="25">
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<emphasis id="B945EAD6FF84AF792A8A3D74FED1F80F" bold="true" box="[151,276,1917,1941]" pageId="19" pageNumber="25">FIGURE 9.</emphasis>
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<taxonomicName id="4C314D47FF84AF792B063D74FDCFF80F" authority="Polizei & Barclay & Bispo, 2020" authorityName="Polizei & Barclay & Bispo" authorityYear="2020" box="[283,522,1917,1941]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="25" phylum="Arthropoda" rank="species" species="maierae" status="sp. nov.">
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<emphasis id="B945EAD6FF84AF792B063D74FDCFF80F" box="[283,522,1917,1941]" italics="true" pageId="19" pageNumber="25">Hexacylloepus maierae</emphasis>
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</taxonomicName>
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<emphasis id="B945EAD6FF84AF79280F3D77FDA6F80C" bold="true" box="[530,611,1918,1942]" pageId="19" pageNumber="25">
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<taxonomicNameLabel id="A27657ADFF84AF79280F3D77FDA6F80C" box="[530,611,1918,1942]" pageId="19" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm.
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</paragraph>
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</caption>
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<paragraph id="8B8E36C4FF83AF7E2ADA3A9EFDFEFE87" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
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<emphasis id="B945EAD6FF83AF7E2ADA3A9EFEE4FF2B" box="[199,289,151,177]" italics="true" pageId="20" pageNumber="26">Thorax:</emphasis>
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Pronotum as long as wide, arcuate anteriorly; sides sinuate and serrate, posterior margin smooth with two prescutellar foveae (
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<figureCitation id="130A2A41FF83AF7E2BB03AB2FDCDFF4F" box="[429,520,187,213]" captionStart="FIGURE 9" captionStartId="19.[151,250,1917,1941]" captionTargetBox="[189,1362,382,1885]" captionTargetId="figure-115@19.[183,1404,381,1893]" captionTargetPageId="19" captionText="FIGURE 9. Hexacylloepus maierae sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396764" httpUri="https://zenodo.org/record/4396764/files/figure.png" pageId="20" pageNumber="26">Fig. 9A</figureCitation>
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). Surface with micropunctures and short setae. Sublateral carinae complete, sinuous and raised (
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<figureCitation id="130A2A41FF83AF7E2B553AD6FE67FF63" box="[328,418,223,249]" captionStart="FIGURE 9" captionStartId="19.[151,250,1917,1941]" captionTargetBox="[189,1362,382,1885]" captionTargetId="figure-115@19.[183,1404,381,1893]" captionTargetPageId="19" captionText="FIGURE 9. Hexacylloepus maierae sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396764" httpUri="https://zenodo.org/record/4396764/files/figure.png" pageId="20" pageNumber="26">Fig. 9A</figureCitation>
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). Longitudinal impression long, wide and deep on disc, reaching the anterior and posterior margin, and wider on middle portion.
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</paragraph>
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<paragraph id="8B8E36C4FF83AF7E2ADA3B2EFDF1FE4B" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
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Elytra twice as long as wide; anterior margin arcuate, lateral margin moderately arcuate and serrate; rounded apex (
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<figureCitation id="130A2A41FF83AF7E2AC73B42FEF6FEFF" box="[218,307,331,357]" captionStart="FIGURE 9" captionStartId="19.[151,250,1917,1941]" captionTargetBox="[189,1362,382,1885]" captionTargetId="figure-115@19.[183,1404,381,1893]" captionTargetPageId="19" captionText="FIGURE 9. Hexacylloepus maierae sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396764" httpUri="https://zenodo.org/record/4396764/files/figure.png" pageId="20" pageNumber="26">Fig. 9A</figureCitation>
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). Humeri prominent. Surface with rows of punctures. Elytral suture moderately elevated. Carina short and elevated on interval IV, extending from base to 1/6 of the elytral length; sublateral carinae on intervals VI and VIII, extending from base to 3/4 of the elytral length. Epipleura covered with tomentum (
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<figureCitation id="130A2A41FF83AF7E2EBB3B9AFAC4FE37" box="[1190,1281,403,429]" captionStart="FIGURE 9" captionStartId="19.[151,250,1917,1941]" captionTargetBox="[189,1362,382,1885]" captionTargetId="figure-115@19.[183,1404,381,1893]" captionTargetPageId="19" captionText="FIGURE 9. Hexacylloepus maierae sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396764" httpUri="https://zenodo.org/record/4396764/files/figure.png" pageId="20" pageNumber="26">Fig. 9B</figureCitation>
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). Hind wings macropterous. Scutellum shield-like.
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</paragraph>
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<paragraph id="8B8E36C4FF83AF7E2ADA3BD2FEA3FD1F" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
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Prosternum broad, surface with micropunctures, anterior margin straight and lateral margin covered with tomentum and with a reddish-brown coloured spot. Prosternal process long, and beyond the procoxae; lateral margin flat; rounded apex; disc shallow. Notosternal suture slightly sinuous. Hypomera posteriorly wider than anteriorly, lateral margin with serrate surface with micropunctures, with a belt of tomentum reaching the anterior and lateral margins (
|
||
<figureCitation id="130A2A41FF83AF7E2B1D3862FE9CFD1F" box="[256,345,619,645]" captionStart="FIGURE 9" captionStartId="19.[151,250,1917,1941]" captionTargetBox="[189,1362,382,1885]" captionTargetId="figure-115@19.[183,1404,381,1893]" captionTargetPageId="19" captionText="FIGURE 9. Hexacylloepus maierae sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396764" httpUri="https://zenodo.org/record/4396764/files/figure.png" pageId="20" pageNumber="26">Fig. 9B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA3886FDE4FD6B" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">Mesoventrite with a shallow groove to receive the prosternal process. Mesepimeron and mesepisternum covered with tomentum. Metaventrite broad with a longitudinal medial depression; surface with micropunctures and with sides covered with tomentum.</paragraph>
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA38F2FAD1FC3F" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
|
||
Legs (except tarsi) covered with punctures and tomentum (
|
||
<figureCitation id="130A2A41FF83AF7E296538F2FC11FC8F" box="[888,980,763,789]" captionStart="FIGURE 9" captionStartId="19.[151,250,1917,1941]" captionTargetBox="[189,1362,382,1885]" captionTargetId="figure-115@19.[183,1404,381,1893]" captionTargetPageId="19" captionText="FIGURE 9. Hexacylloepus maierae sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396764" httpUri="https://zenodo.org/record/4396764/files/figure.png" pageId="20" pageNumber="26">Fig. 9B</figureCitation>
|
||
). Coxae rounded; trochanter elongated; femora 4x as long as wide, 1/3 of the length of the elytra; tibiae long and thin, 8x longer than wide, 1/3 of the length of the elytra and with a pair of spurs at apex. Protibiae each with a row of short spines on distal half. Meso- and metatibiae each with a groove on distal half and with two rows: one with short spines and other with long setae. Tarsi elongate, with setae; apical tarsomere with the length of the basal four combined. Claws long and thin.
|
||
</paragraph>
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA39A6FCEEFBAF" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
|
||
<emphasis id="B945EAD6FF83AF7E2ADA39A6FEF4FC53" box="[199,305,943,969]" italics="true" pageId="20" pageNumber="26">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="130A2A41FF83AF7E2B5E39A6FE5AFC53" box="[323,415,943,969]" captionStart="FIGURE 9" captionStartId="19.[151,250,1917,1941]" captionTargetBox="[189,1362,382,1885]" captionTargetId="figure-115@19.[183,1404,381,1893]" captionTargetPageId="19" captionText="FIGURE 9. Hexacylloepus maierae sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396764" httpUri="https://zenodo.org/record/4396764/files/figure.png" pageId="20" pageNumber="26">Fig. 9B</figureCitation>
|
||
): five ventrites; as long as wide. Anterior and posterior margins straight on the first and second ventrites and arcuate on the subsequent ventrites, lateral margins rounded. Surface covered with tomentum (excepted for disc of first ventrite). Disc of the first ventrite broad and flat. Fifth ventrite with lateral margins slightly produced posterolaterad and apex rounded with long setae.
|
||
</paragraph>
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA3E36FA89FB73" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
|
||
<emphasis id="B945EAD6FF83AF7E2ADA3E36FEAEFBC3" box="[199,363,1087,1113]" italics="true" pageId="20" pageNumber="26">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="130A2A41FF83AF7E2B643E36FE17FBC3" box="[377,466,1087,1113]" captionStart="FIGURE 9" captionStartId="19.[151,250,1917,1941]" captionTargetBox="[189,1362,382,1885]" captionTargetId="figure-115@19.[183,1404,381,1893]" captionTargetPageId="19" captionText="FIGURE 9. Hexacylloepus maierae sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396764" httpUri="https://zenodo.org/record/4396764/files/figure.png" pageId="20" pageNumber="26">Fig. 9C</figureCitation>
|
||
): (Total length:
|
||
<quantity id="4CC99B21FF83AF7E289C3E37FD22FBC3" box="[641,743,1086,1113]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="20" pageNumber="26" unit="mm" value="0.55">0.55 mm</quantity>
|
||
. Greatest width:
|
||
<quantity id="4CC99B21FF83AF7E29BF3E37FBCCFBC3" box="[930,1033,1086,1113]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="26" unit="mm" value="0.15">0.15 mm</quantity>
|
||
), symmetrical and acute. Phallobase twice as long as wide, moderately longer than the penis length. Parameres fused, and articulated with the phallobase, reaching 2/3 of the penis length; thin 6x as long as wide, with sharp apex and curved to internal margin. Penis with anterior quarter constricted, and slightly tapering toward the apex; apex sharp; fibula elongated, very narrow, approximately the same length as the penis; corona present, as an arrangement of tiny rod-like sclerotizations.
|
||
</paragraph>
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA3EFAFF08FAAB" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
|
||
<emphasis id="B945EAD6FF83AF7E2ADA3EFAFEEDFA97" bold="true" box="[199,296,1267,1293]" pageId="20" pageNumber="26">Female:</emphasis>
|
||
External morphology similar to male, except for absence of the groove and row of spines on metatibiae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B654FFF83AF7E2ADA3F33FD9CFACF" box="[199,601,1338,1365]" pageId="20" pageNumber="26" type="materials_examined">
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA3F33FD9CFACF" blockId="20.[151,1437,151,1869]" box="[199,601,1338,1365]" pageId="20" pageNumber="26">
|
||
<emphasis id="B945EAD6FF83AF7E2ADA3F33FE76FACE" bold="true" box="[199,435,1338,1364]" pageId="20" pageNumber="26">Material examined:</emphasis>
|
||
<specimenCount id="9D37FD4DFF83AF7E2BA73F33FD90FACF" box="[442,597,1338,1365]" pageId="20" pageNumber="26" type="generic">15 specimens</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B654FFF83AF7E2ADA3F57FD7FFAE3" box="[199,698,1374,1401]" pageId="20" pageNumber="26" type="distribution">
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA3F57FD7FFAE3" blockId="20.[151,1437,151,1869]" box="[199,698,1374,1401]" pageId="20" pageNumber="26">
|
||
<emphasis id="B945EAD6FF83AF7E2ADA3F57FEA6FAE2" bold="true" box="[199,355,1374,1400]" pageId="20" pageNumber="26">Distribution:</emphasis>
|
||
<collectingCountry id="F3267654FF83AF7E2B773F56FE1FFAE3" box="[362,474,1375,1401]" name="Suriname" pageId="20" pageNumber="26">Suriname</collectingCountry>
|
||
:
|
||
<collectingRegion id="49F5F826FF83AF7E2BF93F56FD73FAE3" box="[484,694,1375,1401]" country="Suriname" name="Sipaliwini" pageId="20" pageNumber="26">Sipaliwini District</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B654FFF83AF7E2ADA3F8AFD2EF9CB" pageId="20" pageNumber="26" type="materials_examined">
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA3F8AFF3CFA7F" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
|
||
<emphasis id="B945EAD6FF83AF7E2ADA3F8AFEC1FA07" bold="true" box="[199,260,1411,1437]" pageId="20" pageNumber="26">Male</emphasis>
|
||
<materialsCitation id="3B593C99FF83AF7E2B113F8AFF30FA7F" collectingDate="2013-08-15" collectorName="A. E. Z. Short" country="Suriname" elevation="600" location="Augustus Creek" longitude="-3.9266667" pageId="20" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Sipaliwini" typeStatus="holotype">
|
||
<emphasis id="B945EAD6FF83AF7E2B113F8AFEB9FA07" bold="true" box="[268,380,1411,1437]" pageId="20" pageNumber="26">
|
||
<typeStatus id="548A8866FF83AF7E2B113F8AFEB2FA07" box="[268,375,1411,1437]" pageId="20" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Holo- type |
|
||
<specimenCount id="9D37FD4DFF83AF7E28093F8AFDECFA07" box="[532,553,1411,1437]" pageId="20" pageNumber="26" type="male">♂</specimenCount>
|
||
|
|
||
<collectingCountry id="F3267654FF83AF7E28223F8AFD13FA07" box="[575,726,1411,1437]" name="Suriname" pageId="20" pageNumber="26">SURINAME</collectingCountry>
|
||
:
|
||
<collectingRegion id="49F5F826FF83AF7E28FC3F8AFC74FA07" box="[737,945,1411,1437]" country="Suriname" name="Sipaliwini" pageId="20" pageNumber="26">Sipaliwini District</collectingRegion>
|
||
/
|
||
<geoCoordinate id="EE055003FF83AF7E29D43F8BFB8DFA07" box="[969,1096,1410,1437]" degrees="3" direction="west" minutes="55.600" orientation="longitude" pageId="20" pageNumber="26" precision="1" value="-3.9266667">N3º55.600’</geoCoordinate>
|
||
<geoCoordinate id="EE055003FF83AF7E2E4D3F8BFB23FA07" box="[1104,1254,1410,1437]" degrees="3" direction="west" minutes="55.600" orientation="longitude" pageId="20" pageNumber="26" precision="1" value="-3.9266667">W56º11.300’</geoCoordinate>
|
||
,
|
||
<quantity id="4CC99B21FF83AF7E2EEE3F8AFAF6FA07" box="[1267,1331,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="26" unit="m" value="600.0">
|
||
<elevation id="001CD1F7FF83AF7E2EEE3F8AFAF6FA07" box="[1267,1331,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="26" unit="m" value="600.0">600m</elevation>
|
||
</quantity>
|
||
/ CSNR:
|
||
<collectorName id="26C45312FF83AF7E2A8A3FAEFEADFA5B" box="[151,360,1447,1473]" pageId="20" pageNumber="26">Tafelberg Summit</collectorName>
|
||
/ nr
|
||
<collectorName id="26C45312FF83AF7E2BB33FAEFD7CFA5B" box="[430,697,1447,1473]" pageId="20" pageNumber="26">
|
||
<location id="8EEE601FFF83AF7E2BB33FAEFDACFA5B" LSID="urn:lsid:plazi:treatment:039887D2FF85AF7E2A8A3C04FCBEF8D7:8EEE601FFF83AF7E2BB33FAEFDACFA5B" box="[430,617,1447,1473]" country="Suriname" longitude="-3.9266667" name="Augustus Creek" pageId="20" pageNumber="26" stateProvince="Sipaliwini">Augustus Creek</location>
|
||
Camp
|
||
</collectorName>
|
||
/ leg.
|
||
<collectorName id="26C45312FF83AF7E290F3FAEFC6EFA5B" box="[786,939,1447,1473]" pageId="20" pageNumber="26">A.E.Z. Short</collectorName>
|
||
;
|
||
<date id="FF8F1004FF83AF7E29A03FAFFB81FA5B" box="[957,1092,1446,1473]" pageId="20" pageNumber="26" value="2013-08-15">
|
||
<collectingDate id="EFCBE9ECFF83AF7E29A03FAFFB81FA5B" box="[957,1092,1446,1473]" pageId="20" pageNumber="26" value="2013-08-15">15.viii.2013</collectingDate>
|
||
</date>
|
||
/ UV light; SR13-0815-LT2 (NZCS)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA3FE6FD2EF9CB" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
|
||
<materialsCitation id="3B593C99FF83AF7E2ADA3FE6FD22F9CB" collectingDate="2013-08-15" collectorName="A. E. Z. Short" country="Suriname" elevation="600" location="Augustus Creek" longitude="-3.9266667" pageId="20" pageNumber="26" specimenCount="14" stateProvince="Sipaliwini" typeStatus="paratype">
|
||
<emphasis id="B945EAD6FF83AF7E2ADA3FE6FE8DF993" bold="true" box="[199,328,1519,1545]" pageId="20" pageNumber="26">
|
||
<typeStatus id="548A8866FF83AF7E2ADA3FE6FE86F993" box="[199,323,1519,1545]" pageId="20" pageNumber="26" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D37FD4DFF83AF7E2B4E3FE6FE2BF993" box="[339,494,1519,1545]" pageId="20" pageNumber="26" type="generic">14 specimens</specimenCount>
|
||
Para- type |
|
||
<collectingCountry id="F3267654FF83AF7E28953FE6FCDAF993" box="[648,799,1519,1545]" name="Suriname" pageId="20" pageNumber="26">SURINAME</collectingCountry>
|
||
:
|
||
<collectingRegion id="49F5F826FF83AF7E29303FE6FC3AF993" box="[813,1023,1519,1545]" country="Suriname" name="Sipaliwini" pageId="20" pageNumber="26">Sipaliwini District</collectingRegion>
|
||
/
|
||
<geoCoordinate id="EE055003FF83AF7E2E063FE7FB5FF993" box="[1051,1178,1518,1545]" degrees="3" direction="west" minutes="55.600" orientation="longitude" pageId="20" pageNumber="26" precision="1" value="-3.9266667">N3º55.600’</geoCoordinate>
|
||
<geoCoordinate id="EE055003FF83AF7E2EB93FE7FAF2F993" box="[1188,1335,1518,1545]" degrees="3" direction="west" minutes="55.600" orientation="longitude" pageId="20" pageNumber="26" precision="1" value="-3.9266667">W56º11.300</geoCoordinate>
|
||
’,
|
||
<quantity id="4CC99B21FF83AF7E2F573FE6FA4FF993" box="[1354,1418,1519,1545]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="26" unit="m" value="600.0">
|
||
<elevation id="001CD1F7FF83AF7E2F573FE6FA4FF993" box="[1354,1418,1519,1545]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="26" unit="m" value="600.0">600m</elevation>
|
||
</quantity>
|
||
/ CSNR:
|
||
<collectorName id="26C45312FF83AF7E2AF23C1AFE7EF9B7" box="[239,443,1555,1581]" pageId="20" pageNumber="26">Tafelberg Summit</collectorName>
|
||
/ nr
|
||
<collectorName id="26C45312FF83AF7E2BF23C1AFD2BF9B7" box="[495,750,1555,1581]" pageId="20" pageNumber="26">
|
||
<location id="8EEE601FFF83AF7E2BF23C1AFD61F9B7" LSID="urn:lsid:plazi:treatment:039887D2FF85AF7E2A8A3C04FCBEF8D7:8EEE601FFF83AF7E2BF23C1AFD61F9B7" box="[495,676,1555,1581]" country="Suriname" longitude="-3.9266667" name="Augustus Creek" pageId="20" pageNumber="26" stateProvince="Sipaliwini">Augustus Creek</location>
|
||
Camp
|
||
</collectorName>
|
||
/ leg.
|
||
<collectorName id="26C45312FF83AF7E29283C1AFC0DF9B7" box="[821,968,1555,1581]" pageId="20" pageNumber="26">A.E.Z. Short</collectorName>
|
||
;
|
||
<date id="FF8F1004FF83AF7E29C93C1BFB99F9B7" box="[980,1116,1554,1581]" pageId="20" pageNumber="26" value="2013-08-15">
|
||
<collectingDate id="EFCBE9ECFF83AF7E29C93C1BFB99F9B7" box="[980,1116,1554,1581]" pageId="20" pageNumber="26" value="2013-08-15">15.viii.2013</collectingDate>
|
||
</date>
|
||
/ UV light; SR13-0815-LT2 (4 NZCS; 5 SEMK; 5 MZSP) (MZSP 35760-35764)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B654FFF83AF7E2ADA3C52FCE1F903" pageId="20" pageNumber="26" type="etymology">
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA3C52FCE1F903" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
|
||
<emphasis id="B945EAD6FF83AF7E2ADA3C52FE97F9EF" bold="true" box="[199,338,1627,1653]" pageId="20" pageNumber="26">Etymology:</emphasis>
|
||
<taxonomicName id="4C314D47FF83AF7E2B473C52FE73F9EE" authorityName="Polizei & Barclay & Bispo" authorityYear="2020" box="[346,438,1627,1652]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="26" phylum="Arthropoda" rank="species" species="maierae">
|
||
<emphasis id="B945EAD6FF83AF7E2B473C52FE73F9EE" box="[346,438,1627,1652]" italics="true" pageId="20" pageNumber="26">maierae</emphasis>
|
||
</taxonomicName>
|
||
is a tribute to Dr. Crystal Maier, specialist in Dryopoidea and curator at the Museum of Comparative Zoology—Cambridge,
|
||
<collectingRegion id="49F5F826FF83AF7E28293C76FD1CF903" box="[564,729,1663,1689]" country="United States of America" name="Massachusetts" pageId="20" pageNumber="26">Massachusetts</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3267654FF83AF7E28F93C76FCE5F903" box="[740,800,1663,1689]" name="United States of America" pageId="20" pageNumber="26">USA</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B654FFF83AF7E2ADA3CABFCBEF8D7" pageId="20" pageNumber="26" type="discussion">
|
||
<paragraph id="8B8E36C4FF83AF7E2ADA3CABFCBEF8D7" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="26">
|
||
<emphasis id="B945EAD6FF83AF7E2ADA3CABFE76F927" bold="true" box="[199,435,1698,1725]" pageId="20" pageNumber="26">Comparative notes:</emphasis>
|
||
<taxonomicName id="4C314D47FF83AF7E2BA63CAAFD01F926" authority="Polizei & Barclay & Bispo, 2020" authorityName="Polizei & Barclay & Bispo" authorityYear="2020" box="[443,708,1699,1725]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="26" phylum="Arthropoda" rank="species" species="maierae" status="sp. nov.">
|
||
<emphasis id="B945EAD6FF83AF7E2BA63CAAFD01F926" box="[443,708,1699,1725]" italics="true" pageId="20" pageNumber="26">Hexacylloepus maierae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EAD6FF83AF7E28D63CAAFCE3F927" bold="true" box="[715,806,1699,1725]" pageId="20" pageNumber="26">
|
||
<taxonomicNameLabel id="A27657ADFF83AF7E28D63CAAFCE3F927" box="[715,806,1699,1725]" pageId="20" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles
|
||
<taxonomicName id="4C314D47FF83AF7E29B83CAAFB25F927" authority="(Grouvelle, 1898)" baseAuthorityName="Grouvelle" baseAuthorityYear="1898" box="[933,1248,1699,1725]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="26" phylum="Arthropoda" rank="species" species="smithi">
|
||
<emphasis id="B945EAD6FF83AF7E29B83CAAFBC9F927" box="[933,1036,1699,1725]" italics="true" pageId="20" pageNumber="26">H. smithi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA04B35FF83AF7E2E013CAAFB12F927" author="Grouvelle, A." box="[1052,1239,1699,1725]" pageId="20" pageNumber="26" pagination="35 - 48" refId="ref25839" refString="Grouvelle, A. (1898) Clavicornes de Grenada et de St. Vincent (Antilles) recoltes par MHH Smith, et appartenant au Musee de Cambridge. Notes from the Leyden Museum, 20 (1), 35 - 48." type="journal article" year="1898">Grouvelle, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, both having the longitudinal impression on pronotum long and wide; hypomera with a belt of tomentum reaching the anterior and lateral margins, and mesotibiae each with a groove with a row of spines. However,
|
||
<taxonomicName id="4C314D47FF83AF7E2E5F3CE2FB6CF89F" baseAuthorityName="Grouvelle" baseAuthorityYear="1898" box="[1090,1193,1771,1797]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="26" phylum="Arthropoda" rank="species" species="smithi">
|
||
<emphasis id="B945EAD6FF83AF7E2E5F3CE2FB6CF89F" box="[1090,1193,1771,1797]" italics="true" pageId="20" pageNumber="26">H. smithi</emphasis>
|
||
</taxonomicName>
|
||
has the frontoclypeal suture inconspicuous and disc of the first abdominal ventrite depressed, while
|
||
<taxonomicName id="4C314D47FF83AF7E2E133D06FB48F8B2" authority="Polizei & Barclay & Bispo, 2020" authorityName="Polizei & Barclay & Bispo" authorityYear="2020" box="[1038,1165,1807,1832]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="26" phylum="Arthropoda" rank="species" species="maierae" status="sp. nov.">
|
||
<emphasis id="B945EAD6FF83AF7E2E133D06FB48F8B2" box="[1038,1165,1807,1832]" italics="true" pageId="20" pageNumber="26">H. maierae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EAD6FF83AF7E2E883D06FB2AF8B3" bold="true" box="[1173,1263,1807,1833]" pageId="20" pageNumber="26">
|
||
<taxonomicNameLabel id="A27657ADFF83AF7E2E883D06FB2AF8B3" box="[1173,1263,1807,1833]" pageId="20" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has the frontoclypeal suture straight and disc of the first abdominal ventrite flat.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |