263 lines
40 KiB
XML
263 lines
40 KiB
XML
<document id="9E94190AF589F4C1791BBCFF543547E0" ID-CLB-Dataset="8626" ID-DOI="10.5281/zenodo.4476396" ID-GBIF-Dataset="f638455f-2539-4bb0-816f-bafbc0b7c1ac" ID-ISSN="ISSN 0374-1036" ID-Zenodo-Dep="4476396" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1611663472831" checkinUser="felipe" docAuthor="Kment, Petr" docDate="2005" docId="039887C4A317F47D7CEB7D3D33AC52CD" docLanguage="en" docName="ActaEntMusNatPra.45.21-50.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl.) 45" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Mahea distanti, sp. nov." docType="treatment" docVersion="6" lastPageNumber="33" masterDocId="FFA1FFBCA31EF4717D327C42312D533A" masterDocTitle="Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles" masterLastPageNumber="50" masterPageNumber="21" pageNumber="30" updateTime="1698925359088" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FE39044A29F1E866FDAFAFC7A16A037B">Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles</mods:title>
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<mods:name id="4ED351F695878CAC6B3283FCA992CBBA" type="personal">
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<mods:namePart id="C239A184EEDA60E0949F6A414CAC280B">Kment, Petr</mods:namePart>
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<mods:affiliation id="87BD07061A904116BBE49CF334009D42">) &) Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: sigara @ post. cz &) &) Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<mods:title id="BF127B2B553A65B17B2A9CEB2869F224">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="85764B7E256B62469405702BCD176332">2005</mods:date>
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<mods:title id="E49CE259FD52F95316203810D9999A79">suppl.</mods:title>
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<mods:number id="63353E54AEF8D8684E771F395514001D">2005-12-31</mods:number>
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<mods:number id="FDB89750E1714689E133D18F0777D872">45</mods:number>
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<mods:identifier id="6B7F48D0D58FC908B0B9B32169DFC98A" type="DOI">10.5281/zenodo.4476396</mods:identifier>
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<treatment id="039887C4A317F47D7CEB7D3D33AC52CD" ID-DOI="http://doi.org/10.5281/zenodo.4476426" ID-GBIF-Taxon="177135175" ID-Zenodo-Dep="4476426" LSID="urn:lsid:plazi:treatment:039887C4A317F47D7CEB7D3D33AC52CD" httpUri="http://treatment.plazi.org/id/039887C4A317F47D7CEB7D3D33AC52CD" lastPageId="12" lastPageNumber="33" pageId="9" pageNumber="30">
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<subSubSection id="C32B6559A317F4787CEB7D3D33F752AD" box="[473,730,383,407]" pageId="9" pageNumber="30" type="nomenclature">
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<paragraph id="8B8E36D2A317F4787CEB7D3D33F752AD" blockId="9.[473,730,383,407]" box="[473,730,383,407]" pageId="9" pageNumber="30">
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<heading id="D0C681BEA317F4787CEB7D3D33F752AD" bold="true" box="[473,730,383,407]" centered="true" fontSize="8" level="2" pageId="9" pageNumber="30" reason="3">
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<emphasis id="B945EAC0A317F4787CEB7D3D33F752AD" bold="true" box="[473,730,383,407]" pageId="9" pageNumber="30">
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<taxonomicName id="4C314D51A317F4787CEB7D3D335052AD" ID-CoL="7VRC4" authority="Kment, 2005" box="[473,637,383,407]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="distanti" status="sp. nov.">
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<emphasis id="B945EAC0A317F4787CEB7D3D335052AD" bold="true" box="[473,637,383,407]" italics="true" pageId="9" pageNumber="30">Mahea distanti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27657BBA317F4787FB67D3D33F752AD" box="[644,730,383,407]" pageId="9" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B6559A317F4787F237DDD338C5288" box="[529,673,415,435]" pageId="9" pageNumber="30" type="description">
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<paragraph id="8B8E36D2A317F4787F237DDD338C5288" blockId="9.[113,853,415,496]" box="[529,673,415,435]" pageId="9" pageNumber="30">
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(
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<figureCitation id="130A2A57A317F4787F257DDD33575288" box="[535,634,415,435]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStartId-0="10.[151,190,1518,1537]" captionStartId-1="5.[113,152,1493,1512]" captionStartId-2="6.[151,190,654,673]" captionStartId-3="8.[151,190,917,936]" captionTargetBox-0="[151,1124,361,1478]" captionTargetBox-1="[114,1065,227,1462]" captionTargetBox-2="[151,1126,220,628]" captionTargetBox-3="[151,1126,220,883]" captionTargetId-0="figure-11@10.[151,1124,361,1478]" captionTargetId-1="figure-16@5.[114,1065,220,1467]" captionTargetId-2="figure-323@6.[151,1126,220,628]" captionTargetId-3="figure-211@8.[151,1126,220,883]" captionTargetPageId-0="10" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="Figs. 11-16. Mahea distanti sp. nov. 11 – antenna; 12 – pronotum, dorsal view; 13 – pronotum, posterior view; 14 – pleural margin and connexiva, dorsal view; 15 – hind leg (tarsomeres and claws missing); 16 – abdomen, ventral view." captionText-1="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." captionText-2="Figs. 6-7. Mahea sexualis Distant, 1909. Pygophore (lectotype). 6 – posterior view; 7 – ventral view. Scale bars 0.5 mm." captionText-3="Figs. 8-10. Mahea andriai (Cachan, 1952). 8 – pronotum, dorsal view; 9 – pronotum, posterior view; 10 – abdomen, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.4476412" figureDoi-1="http://doi.org/10.5281/zenodo.4476402" figureDoi-2="http://doi.org/10.5281/zenodo.4476404" figureDoi-3="http://doi.org/10.5281/zenodo.4476408" httpUri-0="https://zenodo.org/record/4476412/files/figure.png" httpUri-1="https://zenodo.org/record/4476402/files/figure.png" httpUri-2="https://zenodo.org/record/4476404/files/figure.png" httpUri-3="https://zenodo.org/record/4476408/files/figure.png" pageId="9" pageNumber="30">Figs. 11-19</figureCitation>
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, 43)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B6559A317F4787D437D80339A52D5" pageId="9" pageNumber="30" type="reference_group">
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<treatmentCitationGroup id="AB2111FCA317F4787D437D80339A52D5" pageId="9" pageNumber="30">
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<paragraph id="8B8E36D2A317F4787D437D80327852EF" blockId="9.[113,853,415,496]" box="[113,853,450,469]" pageId="9" pageNumber="30">
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<taxonomicName id="4C314D51A317F4787D437D8031DA52EF" ID-CoL="7VRC7" authorityName="Distant" authorityYear="1909" box="[113,247,450,469]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="sexualis">
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<emphasis id="B945EAC0A317F4787D437D8031DA52EF" box="[113,247,450,469]" italics="true" pageId="9" pageNumber="30">Mahea sexualis</emphasis>
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</taxonomicName>
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(misidentification):
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<treatmentCitation id="0A9010C3A317F4787C947D80336152EF" author="DISTANT W. L." box="[422,588,450,469]" page="33" pageId="9" pageNumber="30" year="1909">
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<bibRefCitation id="EFA04B23A317F4787C947D80330652EF" author="DISTANT W. L." box="[422,555,450,469]" pageId="9" pageNumber="30" pagination="29 - 47" refId="ref13166" refString="DISTANT W. L. 1909: The Percy Sladen trust expedition to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner. Volume II. No. IV. " Sealark " Rhynchota. Transactions of the Linnean Society of London, Zoology 13 (1): 29 - 47 + 1 pl." type="journal article" year="1909">DISTANT (1909)</bibRefCitation>
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: 33
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</treatmentCitation>
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, pl. 4 (record, figure of male).
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</paragraph>
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<paragraph id="8B8E36D2A317F4787D437D9E339A52D5" blockId="9.[113,853,415,496]" box="[113,695,476,496]" pageId="9" pageNumber="30">
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<taxonomicName id="4C314D51A317F4787D437D9E31DA52D5" ID-CoL="7VRC7" authorityName="Distant" authorityYear="1909" box="[113,247,476,495]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="sexualis">
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<emphasis id="B945EAC0A317F4787D437D9E31DA52D5" box="[113,247,476,495]" italics="true" pageId="9" pageNumber="30">Mahea sexualis</emphasis>
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</taxonomicName>
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(misidentification):
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<treatmentCitation id="0A9010C3A317F4787C9A7D9F336B52D5" author="KUMAR R." box="[424,582,476,496]" page="44" pageId="9" pageNumber="30" year="1974">
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<bibRefCitation id="EFA04B23A317F4787C9A7D9F330A52D5" author="KUMAR R." box="[424,551,476,496]" pageId="9" pageNumber="30" pagination="1 - 60" refId="ref13645" refString="KUMAR R. 1974: A revision of world Acanthosomatidae (Heteroptera: Pentatomoidea): Keys to and descriptions of subfamilies, tribes and genera, with designation of types. Australian Journal of Zoology, Supplementary Series 34: 1 - 60." type="book chapter" year="1974">KUMAR (1974)</bibRefCitation>
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: 44
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||
</treatmentCitation>
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(taxonomy).
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</paragraph>
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</treatmentCitationGroup>
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||
</subSubSection>
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||
<subSubSection id="C32B6559A317F4787D437E5032C95010" pageId="9" pageNumber="30" type="materials_examined">
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<paragraph id="8B8E36D2A317F4787D437E5033875110" blockId="9.[113,1089,529,810]" box="[113,682,529,554]" pageId="9" pageNumber="30">
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<emphasis id="B945EAC0A317F4787D437E5030255113" bold="true" box="[113,264,529,554]" pageId="9" pageNumber="30">
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<typeStatus id="548A8870A317F4787D437E5031855110" box="[113,168,530,554]" pageId="9" pageNumber="30">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F3267642A317F4787C3D7E5030BC5110" box="[271,401,530,554]" name="Madagascar" pageId="9" pageNumber="30">Madagascar</collectingCountry>
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, Tamatave [=
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<collectingRegion id="49F5F830A317F4787F1A7E50338D5110" box="[552,672,530,554]" country="Madagascar" name="Toamasina" pageId="9" pageNumber="30">Toamasina</collectingRegion>
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||
].
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</paragraph>
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||
<paragraph id="8B8E36D2A317F4787D437E7032C95010" blockId="9.[113,1089,529,810]" pageId="9" pageNumber="30">
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<emphasis id="B945EAC0A317F4787D437E7030345173" bold="true" box="[113,281,561,586]" pageId="9" pageNumber="30">Type material.</emphasis>
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<typeStatus id="548A8870A317F4787C107E7030B85172" box="[290,405,562,585]" pageId="9" pageNumber="30" type="holotype">HOLOTYPUS</typeStatus>
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: ♁, ‘Type / H. T. [p, white circle with red margin] //
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<taxonomicName id="4C314D51A317F4787EDA7E70351D5170" authorityName=": KUMAR" authorityYear="1974" box="[1000,1072,562,586]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="47" phylum="Arthropoda" rank="genus">Mahea</taxonomicName>
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/ sexualis / type Dist. [hw] //
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<collectingCountry id="F3267642A317F4787CA17E10333E5150" box="[403,531,594,618]" name="Madagascar" pageId="9" pageNumber="30">Madagascar</collectingCountry>
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/ Tamatave / XII. [hw] // Distant Coll. / 1911-383 [p] // SYN- / TYPE [p, white circle with blue margin] //
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<typeStatus id="548A8870A317F4787F907E30326D51B0" box="[674,832,626,650]" pageId="9" pageNumber="30" type="holotype">HOLOTYPUS</typeStatus>
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/
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<taxonomicName id="4C314D51A317F4787E667E30329951B0" authorityName=": KUMAR" authorityYear="1974" box="[852,948,626,650]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="47" phylum="Arthropoda" rank="genus">MAHEA</taxonomicName>
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/ DISTANTI / sp. nov. / det. P. KMENT 2005 [p, red label]’ (
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<collectionCode id="ED20AE17A317F4787F6C7ED033995190" box="[606,692,658,682]" pageId="9" pageNumber="30">BMNH</collectionCode>
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||
).
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<typeStatus id="548A8870A317F4787FF67ED0320A5190" box="[708,807,658,682]" pageId="9" pageNumber="30" type="holotype">Holotype</typeStatus>
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on micro pin; left antenna, right antennomere 4, left fore leg, right fore and middle tibiae and tarsi and both hind tarsi missing; pronotum ventrally detached from mesonotum, dorsally cracked on the right side, right humeral spine missing. The pygophore and male genitalia of the specimen had been dissected and stored in glycerol in a small glass microvial attached to the same pin.
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C32B6559A317F47A7D437F0230775666" lastPageId="11" lastPageNumber="32" pageId="9" pageNumber="30" type="description">
|
||
<paragraph id="8B8E36D2A317F4787D437F02304C5042" blockId="9.[113,1090,832,1593]" pageId="9" pageNumber="30">
|
||
<emphasis id="B945EAC0A317F4787D437F0231D65062" bold="true" box="[113,251,832,856]" pageId="9" pageNumber="30">Description.</emphasis>
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Male (Fig. 43). Length
|
||
<quantity id="4CC99B37A317F4787CCF7F03337E5062" box="[509,595,833,857]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="30" unit="mm" value="8.2">8.2 mm</quantity>
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||
. Body ochraceous, slightly shining, with thick coarse dark punctures.
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||
</paragraph>
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||
<paragraph id="8B8E36D2A317F4787DA27FC333D357C3" blockId="9.[113,1090,832,1593]" pageId="9" pageNumber="30">
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||
Head pale ochraceous with coarse black or brown punctures, forming more or less distinct rows (two regular rows on each mandibular plate, rest of head with more irregular pattern than in
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||
<taxonomicName id="4C314D51A317F4787DF27F83301550E3" authorityName="Distant" authorityYear="1909" box="[192,312,961,985]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="sexualis">
|
||
<emphasis id="B945EAC0A317F4787DF27F83301550E3" box="[192,312,961,985]" italics="true" pageId="9" pageNumber="30">M. sexualis</emphasis>
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||
</taxonomicName>
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||
, ‘circular’ figure on vertex not developed). Head shorter than wide across eyes (1: 1.14), its width about half of pronotal width between humeral angles (1: 2.75). Mandibular plates almost parallel basally, smoothly rounded apically, enclosing apex of clypeus. Eyes brownish; ocelli reddish, situated behind imaginary line through posterior margins of eyes, but more distant from anterior pronotal margin than in
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||
<taxonomicName id="4C314D51A317F4787FC2780332455763" authorityName="Distant" authorityYear="1909" box="[752,872,1089,1113]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="sexualis">
|
||
<emphasis id="B945EAC0A317F4787FC2780332455763" box="[752,872,1089,1113]" italics="true" pageId="9" pageNumber="30">M. sexualis</emphasis>
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||
</taxonomicName>
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||
. Each antenniferous tubercle laterally with black longitudinal spot; antennomere 1 blackish brown; antennomere 2 pale castaneous, with erect pubescence, hairs slightly longer than diameter of antennomere; antennomere 3 castaneous; lengths of antennomeres: 1 –
|
||
<quantity id="4CC99B37A317F4787FEA78E332005782" box="[728,813,1185,1209]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="30" unit="mm" value="0.4">0.4 mm</quantity>
|
||
,
|
||
<quantity id="4CC99B37A317F4787E0B78E332955782" box="[825,952,1185,1209]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.9" metricValueMin="2.0" pageId="9" pageNumber="30" unit="mm" value="2.45" valueMax="2.9" valueMin="2.0">2 – 2.9 mm</quantity>
|
||
, 3 –
|
||
<quantity id="4CC99B37A317F4787EDC78E3356D5782" box="[1006,1088,1185,1209]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="30" unit="mm" value="1.3">1.3 mm</quantity>
|
||
(ratio 1: 7.25: 3.25) (
|
||
<figureCitation id="130A2A57A317F4787C6C7883308757E3" box="[350,426,1217,1241]" captionStart="Figs" captionStartId="10.[151,190,1518,1537]" captionTargetBox="[151,1124,361,1478]" captionTargetId="figure-11@10.[151,1124,361,1478]" captionTargetPageId="10" captionText="Figs. 11-16. Mahea distanti sp. nov. 11 – antenna; 12 – pronotum, dorsal view; 13 – pronotum, posterior view; 14 – pleural margin and connexiva, dorsal view; 15 – hind leg (tarsomeres and claws missing); 16 – abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476412" httpUri="https://zenodo.org/record/4476412/files/figure.png" pageId="9" pageNumber="30">Fig. 11</figureCitation>
|
||
). Head ventrally ochraceous with small brown punctures; rostrum ochraceous, apex blackened, reaching base of abdomen.
|
||
</paragraph>
|
||
<paragraph id="8B8E36D2A317F4787DA2794331D35502" blockId="9.[113,1090,832,1593]" pageId="9" pageNumber="30">
|
||
Pronotum ochraceous with uniform brown punctures, median carina not developed, humeral angles and spines reddish; anterior margin strongly concave, head more projected into pronotum than in
|
||
<taxonomicName id="4C314D51A317F4787C1E7903308E5663" authorityName="Distant" authorityYear="1909" box="[300,419,1345,1369]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="sexualis">
|
||
<emphasis id="B945EAC0A317F4787C1E7903308E5663" box="[300,419,1345,1369]" italics="true" pageId="9" pageNumber="30">M. sexualis</emphasis>
|
||
</taxonomicName>
|
||
; antero-lateral margins rounded, divergent laterad; each humeral angle conically produced into spine, directed laterad and slightly upwards (
|
||
<figureCitation id="130A2A57A317F4787E837923351E5643" box="[945,1075,1377,1401]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="9" pageNumber="30">Figs. 12, 13</figureCitation>
|
||
). Scutellum triangular, its punctures finer than those on pronotum; anterior part only slightly arched, castaneous, only antero-lateral angles ochraceous; lateral margins slightly depressed medially; posterior part flat, ochraceous, with inconspicuous dark punctures; apex black, triangular. Hemelytra ochraceous with brown punctures; suture between corium and membrane brown; membrane with brownish tinge, translucent, slightly surpassing postero-lateral angles of sternum 7.
|
||
</paragraph>
|
||
<paragraph id="8B8E36D2A314F47A7D847C9D338E50E1" blockId="10.[151,1127,223,344]" lastBlockId="11.[113,1089,932,1596]" lastPageId="11" lastPageNumber="32" pageId="10" pageNumber="31">
|
||
Thorax ventrally ochraceous with coarse brown punctures; brown colour of punctures locally coalescent, forming small to large irregular spots on median parts of pleura; acetabula, peritreme and lateral margins of pleura ochraceous. Metapleura strongly swollen, produced laterad, its posterior margin introverted, both well visible in dorsal view (
|
||
<figureCitation id="130A2A57A314F47B7EBE7D7D32F4526D" box="[908,985,319,343]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="10" pageNumber="31">Fig. 14</figureCitation>
|
||
). Legs ochraceous, hind femora darkened apically, tarsi brown. Hind femora and tibiae slightly curved inwards, with granulation on inner margins (
|
||
<figureCitation id="130A2A57A315F47A7F7A7F8633B850E1" box="[584,661,963,988]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="11" pageNumber="32">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF4E665AA314F47B7DA579AC31EB550D" ID-DOI="http://doi.org/10.5281/zenodo.4476412" ID-Zenodo-Dep="4476412" httpUri="https://zenodo.org/record/4476412/files/figure.png" pageId="10" pageNumber="31" startId="10.[151,190,1518,1537]" targetBox="[151,1124,361,1478]" targetPageId="10">
|
||
<paragraph id="8B8E36D2A314F47B7DA579AC31EB550D" blockId="10.[151,1127,1518,1591]" pageId="10" pageNumber="31">
|
||
Figs. 11-16.
|
||
<taxonomicName id="4C314D51A314F47B7C3179AC30AE553B" box="[259,387,1518,1537]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="distanti" status="sp. nov.">
|
||
<emphasis id="B945EAC0A314F47B7C3179AC30AE553B" box="[259,387,1518,1537]" italics="true" pageId="10" pageNumber="31">Mahea distanti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27657BBA314F47B7CBB79AD30E15538" box="[393,460,1519,1538]" pageId="10" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
11 – antenna; 12 – pronotum, dorsal view; 13 – pronotum, posterior view; 14 – pleural margin and connexiva, dorsal view; 15 – hind leg (tarsomeres and claws missing); 16 – abdomen, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E665AA315F47A7D437F043348504E" ID-DOI="http://doi.org/10.5281/zenodo.4476414" ID-Zenodo-Dep="4476414" httpUri="https://zenodo.org/record/4476414/files/figure.png" pageId="11" pageNumber="32" startId="11.[113,152,838,857]" targetBox="[113,1088,220,800]" targetPageId="11">
|
||
<paragraph id="8B8E36D2A315F47A7D437F043348504E" blockId="11.[113,1089,838,884]" pageId="11" pageNumber="32">
|
||
Figs. 17-19.
|
||
<taxonomicName id="4C314D51A315F47A7DEE7F0430765063" box="[220,347,838,857]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="species" species="distanti" status="sp. nov.">
|
||
<emphasis id="B945EAC0A315F47A7DEE7F0430765063" box="[220,347,838,857]" italics="true" pageId="11" pageNumber="32">Mahea distanti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27657BBA315F47A7C537F0430895063" box="[353,420,838,857]" pageId="11" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
17 – pygophore, ventral view; 18 – right paramere, dorsal view; 19 – aedeagus (basal articulatory apparatus omitted). Scale bars 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E36D2A315F47A7DA27FA633CA57E1" blockId="11.[113,1089,932,1596]" pageId="11" pageNumber="32">
|
||
Abdomen ochraceous with thick brown punctures; sterna 3-4 broadly, sternum 5 narrowly reddish brown medially, sternum 6 with sparse brown punctures, sternum 7 with sparse brown punctures and large blackish brown spots on each side near anterior margin and before postero-lateral angles; median carina well developed on sterna 3-6, gradually disappearing on sternum 7, ochraceous, acutely produced anteriad as abdominal spine between metacoxae (
|
||
<figureCitation id="130A2A57A315F47A7924782631A357A1" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="5.[113,152,1493,1512]" captionStartId-1="6.[151,190,654,673]" captionTargetBox-0="[114,1065,227,1462]" captionTargetBox-1="[151,1126,220,628]" captionTargetId-0="figure-16@5.[114,1065,220,1467]" captionTargetId-1="figure-323@6.[151,1126,220,628]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." captionText-1="Figs. 6-7. Mahea sexualis Distant, 1909. Pygophore (lectotype). 6 – posterior view; 7 – ventral view. Scale bars 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4476402" figureDoi-1="http://doi.org/10.5281/zenodo.4476404" httpUri-0="https://zenodo.org/record/4476402/files/figure.png" httpUri-1="https://zenodo.org/record/4476404/files/figure.png" pageId="11" pageNumber="32">Fig. 16</figureCitation>
|
||
); connexiva ochraceous, spines on postero-lateral angles of sterna 3-4 very small, on sternum 5 small, on sternum 6 long, narrow, directed posteriad, and on sternum 7 produced, widely triangular with parallel outer margins (
|
||
<figureCitation id="130A2A57A315F47A7F68788633F757E1" box="[602,730,1219,1244]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="5.[113,152,1493,1512]" captionStartId-1="6.[151,190,654,673]" captionTargetBox-0="[114,1065,227,1462]" captionTargetBox-1="[151,1126,220,628]" captionTargetId-0="figure-16@5.[114,1065,220,1467]" captionTargetId-1="figure-323@6.[151,1126,220,628]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." captionText-1="Figs. 6-7. Mahea sexualis Distant, 1909. Pygophore (lectotype). 6 – posterior view; 7 – ventral view. Scale bars 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4476402" figureDoi-1="http://doi.org/10.5281/zenodo.4476404" httpUri-0="https://zenodo.org/record/4476402/files/figure.png" httpUri-1="https://zenodo.org/record/4476404/files/figure.png" pageId="11" pageNumber="32">Figs. 14, 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E36D2A315F47A7DA278A633535606" blockId="11.[113,1089,932,1596]" pageId="11" pageNumber="32">
|
||
Pygophore brownish; dorso-ventrally flattened (
|
||
<figureCitation id="130A2A57A315F47A7FB878A633FA57C6" box="[650,727,1252,1276]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="5.[113,152,1493,1512]" captionStartId-1="6.[151,190,654,673]" captionTargetBox-0="[114,1065,227,1462]" captionTargetBox-1="[151,1126,220,628]" captionTargetId-0="figure-16@5.[114,1065,220,1467]" captionTargetId-1="figure-323@6.[151,1126,220,628]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." captionText-1="Figs. 6-7. Mahea sexualis Distant, 1909. Pygophore (lectotype). 6 – posterior view; 7 – ventral view. Scale bars 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4476402" figureDoi-1="http://doi.org/10.5281/zenodo.4476404" httpUri-0="https://zenodo.org/record/4476402/files/figure.png" httpUri-1="https://zenodo.org/record/4476404/files/figure.png" pageId="11" pageNumber="32">Fig. 17</figureCitation>
|
||
); hypophysis of paramere roughly semicircular, apically rounded, not pointed, inner margin deeply incised (
|
||
<figureCitation id="130A2A57A315F47A7E4C794632E15626" box="[894,972,1284,1308]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStartId-0="5.[113,152,1493,1512]" captionStartId-1="6.[151,190,654,673]" captionStartId-2="8.[151,190,917,936]" captionTargetBox-0="[114,1065,227,1462]" captionTargetBox-1="[151,1126,220,628]" captionTargetBox-2="[151,1126,220,883]" captionTargetId-0="figure-16@5.[114,1065,220,1467]" captionTargetId-1="figure-323@6.[151,1126,220,628]" captionTargetId-2="figure-211@8.[151,1126,220,883]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionText-0="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." captionText-1="Figs. 6-7. Mahea sexualis Distant, 1909. Pygophore (lectotype). 6 – posterior view; 7 – ventral view. Scale bars 0.5 mm." captionText-2="Figs. 8-10. Mahea andriai (Cachan, 1952). 8 – pronotum, dorsal view; 9 – pronotum, posterior view; 10 – abdomen, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.4476402" figureDoi-1="http://doi.org/10.5281/zenodo.4476404" figureDoi-2="http://doi.org/10.5281/zenodo.4476408" httpUri-0="https://zenodo.org/record/4476402/files/figure.png" httpUri-1="https://zenodo.org/record/4476404/files/figure.png" httpUri-2="https://zenodo.org/record/4476408/files/figure.png" pageId="11" pageNumber="32">Fig. 18</figureCitation>
|
||
); vesica of aedeagus deeply sinuated, without loop (
|
||
<figureCitation id="130A2A57A315F47A7F107966335D5606" box="[546,624,1316,1340]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStartId-0="5.[113,152,1493,1512]" captionStartId-1="6.[151,190,654,673]" captionStartId-2="8.[151,190,917,936]" captionTargetBox-0="[114,1065,227,1462]" captionTargetBox-1="[151,1126,220,628]" captionTargetBox-2="[151,1126,220,883]" captionTargetId-0="figure-16@5.[114,1065,220,1467]" captionTargetId-1="figure-323@6.[151,1126,220,628]" captionTargetId-2="figure-211@8.[151,1126,220,883]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionText-0="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." captionText-1="Figs. 6-7. Mahea sexualis Distant, 1909. Pygophore (lectotype). 6 – posterior view; 7 – ventral view. Scale bars 0.5 mm." captionText-2="Figs. 8-10. Mahea andriai (Cachan, 1952). 8 – pronotum, dorsal view; 9 – pronotum, posterior view; 10 – abdomen, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.4476402" figureDoi-1="http://doi.org/10.5281/zenodo.4476404" figureDoi-2="http://doi.org/10.5281/zenodo.4476408" httpUri-0="https://zenodo.org/record/4476402/files/figure.png" httpUri-1="https://zenodo.org/record/4476404/files/figure.png" httpUri-2="https://zenodo.org/record/4476408/files/figure.png" pageId="11" pageNumber="32">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E36D2A315F47A7DA2790630775666" blockId="11.[113,1089,932,1596]" box="[144,346,1348,1372]" pageId="11" pageNumber="32">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6559A315F47D7D437921306F526D" lastPageId="12" lastPageNumber="33" pageId="11" pageNumber="32" type="diagnosis">
|
||
<paragraph id="8B8E36D2A315F47D7D437921306F526D" blockId="11.[113,1089,932,1596]" lastBlockId="12.[151,1127,223,503]" lastPageId="12" lastPageNumber="33" pageId="11" pageNumber="32">
|
||
<emphasis id="B945EAC0A315F47A7D43792130445641" bold="true" box="[113,361,1379,1403]" pageId="11" pageNumber="32">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C314D51A315F47A7C5D792633205646" box="[367,525,1380,1404]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="species" species="distanti" status="sp. nov.">
|
||
<emphasis id="B945EAC0A315F47A7C5D792633205646" box="[367,525,1380,1404]" italics="true" pageId="11" pageNumber="32">Mahea distanti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27657BBA315F47A7F21792633485641" box="[531,613,1380,1403]" pageId="11" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from both
|
||
<taxonomicName id="4C314D51A315F47A7E16792632B15646" authorityName="Distant" authorityYear="1909" box="[804,924,1380,1404]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="species" species="sexualis">
|
||
<emphasis id="B945EAC0A315F47A7E16792632B15646" box="[804,924,1380,1404]" italics="true" pageId="11" pageNumber="32">M. sexualis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C314D51A315F47A7EFC7926356C5646" box="[974,1089,1380,1404]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="species" species="durrelli" status="sp. nov.">
|
||
<emphasis id="B945EAC0A315F47A7EFC7926356C5646" box="[974,1089,1380,1404]" italics="true" pageId="11" pageNumber="32">M. durrelli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27657BBA315F47A7D4379C631E856A1" box="[113,197,1412,1435]" pageId="11" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having the lateral pronotal angles conically produced and gradually tapering to a spinose apex. From
|
||
<taxonomicName id="4C314D51A315F47A7C6579E630E15686" box="[343,460,1444,1468]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="species" species="parvula">
|
||
<emphasis id="B945EAC0A315F47A7C6579E630E15686" box="[343,460,1444,1468]" italics="true" pageId="11" pageNumber="32">M. parvula</emphasis>
|
||
</taxonomicName>
|
||
, it differs by the larger body, more prominent lateral pronotal angles, and relatively narrower head. From
|
||
<taxonomicName id="4C314D51A315F47A7F53798633FF56E6" baseAuthorityName="Cachan" baseAuthorityYear="1952" box="[609,722,1476,1500]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="species" species="andriai">
|
||
<emphasis id="B945EAC0A315F47A7F53798633FF56E6" box="[609,722,1476,1500]" italics="true" pageId="11" pageNumber="32">M. andriai</emphasis>
|
||
</taxonomicName>
|
||
, it can be separated by the following characters: Body more vividly coloured, covered with dark punctures; basic colour ochraceous, posterior part of pronotum with reddish tones; anterior part of scutellum reddish brown, antero-lateral angles and posterior part contrasting pale; apex of scutellum black, narrowly rounded. Dark punctures on head, sternum and abdominal venter coalescent, forming conspicuous rows (usually impressed and darkened). Hairs of antennomere 2 as long as or slightly longer than its diameter. Thorax conspicuously swollen. Lateral connexival spines as in
|
||
<figureCitation id="130A2A57A312F47D7DA57D7D3013526D" box="[151,318,319,343]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="5.[113,152,1493,1512]" captionStartId-1="6.[151,190,654,673]" captionTargetBox-0="[114,1065,227,1462]" captionTargetBox-1="[151,1126,220,628]" captionTargetId-0="figure-16@5.[114,1065,220,1467]" captionTargetId-1="figure-323@6.[151,1126,220,628]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." captionText-1="Figs. 6-7. Mahea sexualis Distant, 1909. Pygophore (lectotype). 6 – posterior view; 7 – ventral view. Scale bars 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4476402" figureDoi-1="http://doi.org/10.5281/zenodo.4476404" httpUri-0="https://zenodo.org/record/4476402/files/figure.png" httpUri-1="https://zenodo.org/record/4476404/files/figure.png" pageId="12" pageNumber="33">Figs. 14 and 16</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6559A312F47D7DA57D1D3055528D" pageId="12" pageNumber="33" type="etymology">
|
||
<paragraph id="8B8E36D2A312F47D7DA57D1D351B52AD" blockId="12.[151,1127,223,503]" pageId="12" pageNumber="33">
|
||
<emphasis id="B945EAC0A312F47D7DA57D1D303A524D" bold="true" box="[151,279,351,375]" pageId="12" pageNumber="33">Etymology.</emphasis>
|
||
This species is named in honour of William Lucas Distant (1845-1922), wellknown English entomologist, who described many heteropteran taxa, including
|
||
<taxonomicName id="4C314D51A312F47D7ED47D3D350252AD" authorityName=": KUMAR" authorityYear="1974" box="[998,1071,383,407]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAC0A312F47D7ED47D3D350252AD" box="[998,1071,383,407]" italics="true" pageId="12" pageNumber="33">Mahea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8E36D2A312F47D7DA57DDD3055528D" blockId="12.[151,1127,223,503]" box="[151,376,415,439]" pageId="12" pageNumber="33">
|
||
<emphasis id="B945EAC0A312F47D7DA57DDD302F528D" bold="true" box="[151,258,415,439]" pageId="12" pageNumber="33">Bionomy.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6559A312F47D7DA57DFD328752ED" box="[151,938,447,471]" pageId="12" pageNumber="33" type="distribution">
|
||
<paragraph id="8B8E36D2A312F47D7DA57DFD328752ED" blockId="12.[151,1127,223,503]" box="[151,938,447,471]" pageId="12" pageNumber="33">
|
||
<emphasis id="B945EAC0A312F47D7DA57DFD300A52ED" bold="true" box="[151,295,447,471]" pageId="12" pageNumber="33">Distribution.</emphasis>
|
||
East
|
||
<collectingCountry id="F3267642A312F47D7C517DFD30CE52ED" box="[355,483,447,471]" name="Madagascar" pageId="12" pageNumber="33">Madagascar</collectingCountry>
|
||
(
|
||
<collectingRegion id="49F5F830A312F47D7CC37DFD334752ED" box="[497,618,447,471]" country="Madagascar" name="Toamasina" pageId="12" pageNumber="33">Toamasina</collectingRegion>
|
||
). Only the
|
||
<typeStatus id="548A8870A312F47D7FEC7DFD321652ED" box="[734,827,447,471]" pageId="12" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
is known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6559A312F47D7DA57D9D33AC52CD" box="[151,641,479,503]" pageId="12" pageNumber="33" type="discussion">
|
||
<paragraph id="8B8E36D2A312F47D7DA57D9D33AC52CD" blockId="12.[151,1127,223,503]" box="[151,641,479,503]" pageId="12" pageNumber="33">
|
||
<emphasis id="B945EAC0A312F47D7DA57D9D303552CD" bold="true" box="[151,280,479,503]" pageId="12" pageNumber="33">Comments.</emphasis>
|
||
See comments under
|
||
<taxonomicName id="4C314D51A312F47D7F307DA2335752CD" authorityName="Distant" authorityYear="1909" box="[514,634,479,503]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="sexualis">
|
||
<emphasis id="B945EAC0A312F47D7F307DA2335752CD" box="[514,634,479,503]" italics="true" pageId="12" pageNumber="33">M. sexualis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |