344 lines
44 KiB
XML
344 lines
44 KiB
XML
<document id="7D014B3AA0EE8EB799840B3D2902FC91" ID-DOI="10.37520/fi.2021.022" ID-ISSN="2533-4069" ID-Zenodo-Dep="7167583" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665260545767" checkinUser="felipe" docAuthor="Wheeler, Elisabeth A. & Manchester, Steven R." docDate="2021" docId="039787B7FF96FF99FBECDE6AB5AEFC0C" docLanguage="en" docName="FossilImprint.77.2.299-329.pdf" docOrigin="Fossil Imprint 77 (2)" docSource="http://dx.doi.org/10.37520/fi.2021.022" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Fagus dodgei Wheeler & Manchester 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="308" masterDocId="FFAEFFCFFF91FF90FFFED964B118FFA2" masterDocTitle="A Diverse Assemblage Of Late Eocene Woods From Oregon, Western Usa" masterLastPageNumber="329" masterPageNumber="299" pageNumber="306" updateTime="1699373241204" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="7DCBF91CD615A62340369BA6358BD23D">A Diverse Assemblage Of Late Eocene Woods From Oregon, Western Usa</mods:title>
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<mods:namePart id="7159BE776A1D9EA56BED97376BAFA757">Wheeler, Elisabeth A.</mods:namePart>
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<mods:namePart id="267AACB0BF72B76CB1F25144743E86E4">Manchester, Steven R.</mods:namePart>
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<mods:typeOfResource id="90ACDE40717761F26DD9D4379533C19F">text</mods:typeOfResource>
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<mods:title id="5C083D0DB1D56F9B83CEC4DC8915959D">Fossil Imprint</mods:title>
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<mods:part id="EAB1AD936BD7B60B86132BEBD2F5C0C8">
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<mods:date id="C135571A614E691B796A957B4244C467">2021</mods:date>
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<mods:number id="F487EAFEE09D7804ECEF9B8D6C0BDA1A">2022-02-18</mods:number>
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<mods:number id="E92CF63FB09F9B20D98E499851F3DB2A">77</mods:number>
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<mods:url id="EA2F600DB7E230945F0FBEF8DCC480D6">http://dx.doi.org/10.37520/fi.2021.022</mods:url>
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<mods:identifier id="6C88F3328129A206774E115A263BF857" type="DOI">10.37520/fi.2021.022</mods:identifier>
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<subSubSection id="C324652AFF96FF97FBECDE6AB5CCF8E4" pageId="7" pageNumber="306" type="nomenclature">
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<paragraph id="8B8136A1FF96FF97FBECDE6AB41BF884" blockId="7.[1042,1283,1806,1830]" box="[1042,1283,1806,1830]" pageId="7" pageNumber="306">
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<heading id="D0C981CDFF96FF97FBECDE6AB41BF884" box="[1042,1283,1806,1830]" centered="true" fontSize="8" level="2" pageId="7" pageNumber="306" reason="2">
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<emphasis id="B94AEAB3FF96FF97FBECDE6AB41BF884" bold="true" box="[1042,1283,1806,1830]" pageId="7" pageNumber="306">
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<taxonomicName id="4C3E4D22FF96FF97FBECDE6AB5BFF884" authority="Wheeler & Manchester, 2021" authorityName="Wheeler & Manchester" authorityYear="2021" box="[1042,1191,1806,1830]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="306" phylum="Tracheophyta" rank="species" species="dodgei" status="sp. nov.">
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<emphasis id="B94AEAB3FF96FF97FBECDE6AB5BFF884" bold="true" box="[1042,1191,1806,1830]" italics="true" pageId="7" pageNumber="306">Fagus dodgei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27957C8FF96FF97FB53DE6AB41BF884" box="[1197,1283,1806,1830]" pageId="7" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8136A1FF96FF97FBBEDE4AB5CCF8E4" blockId="7.[1088,1236,1838,1862]" box="[1088,1236,1838,1862]" pageId="7" pageNumber="306">
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<figureCitation id="13052A24FF96FF97FBBEDE4AB5CCF8E4" box="[1088,1236,1838,1862]" captionStart="Text-fig" captionStartId="8.[125,206,1353,1374]" captionTargetBox="[125,1440,151,1334]" captionTargetId="figure-275@8.[125,1440,150,1841]" captionTargetPageId="8" captionText="Text-fig. 4. Fagus dodgei sp. nov., a, b, d–h: UF 279-34468; c: UF 279-30165. a: Wood diffuse-porous to semi-ring-porous with distinct latewoodzone with narrower vessels; vessels solitary andin short multiples;diffuse, diffuse-in-aggregatesaxial parenchyma visible in latewood, TS. b: Growth ring boundary, TS. c: Opposite intervessel pitting, TLS. d: Scalariform perforation plate with fewer than 10 bars, RLS. e: Simple perforation plates (PP), RLS. f: Vessel-ray parenchyma pitting with reduced borders and frequently oval in outline, RLS. g: Rays 1–4(–5)-seriate with variable numbers of marginal rows, TLS. h: Rays of two distinct sizes, widest rays>10-seriate, TLS. Scale bars: 200 µm in a, h; 100 µm in b, e, g; 50 µm in d, f." figureDoi="http://doi.org/10.5281/zenodo.7167591" httpUri="https://zenodo.org/record/7167591/files/figure.png" pageId="7" pageNumber="306">Text-fig. 4a–h</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C324652AFF96FF99FC8FDE3AB5AEFC0C" lastPageId="9" lastPageNumber="308" pageId="7" pageNumber="306" type="description">
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<paragraph id="8B8136A1FF96FF97FC8FDE3AB4A9F837" blockId="7.[843,1482,1885,1941]" pageId="7" pageNumber="306">
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H o l o t y p e. Designated here. UF 279-34468 (Textfig. 4a, b, d–h). Minimum estimated axis diameter
|
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.
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</paragraph>
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<paragraph id="8B8136A1FF96FF97FC8FDEC9B285F847" blockId="7.[843,1482,1964,2021]" pageId="7" pageNumber="306">A d d i t i o n a l m a t e r i a l. UF 279-30165 (Textfig. 4c).</paragraph>
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<paragraph id="8B8136A1FF96FF97FC8FDE98B4DFF7B1" blockId="7.[843,1479,2044,2100]" box="[881,1479,2044,2068]" pageId="7" pageNumber="306">P l a n t F o s s i l N a m e s R e g i s t r y N u m b e r.</paragraph>
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<paragraph id="8B8136A1FF96FF97FCB5D178B580F796" blockId="7.[843,1479,2044,2100]" box="[843,1176,2076,2100]" pageId="7" pageNumber="306">PFN 002679 (for new species).</paragraph>
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<caption id="DF416629FF99FF98FF83DC2DB2E6FA52" ID-DOI="http://doi.org/10.5281/zenodo.7167591" ID-Zenodo-Dep="7167591" httpUri="https://zenodo.org/record/7167591/files/figure.png" pageId="8" pageNumber="307" startId="8.[125,206,1353,1374]" targetBox="[125,1440,151,1334]" targetPageId="8">
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<paragraph id="8B8136A1FF99FF98FF83DC2DB2E6FA52" blockId="8.[125,1440,1353,1520]" pageId="8" pageNumber="307">
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Text-fig. 4.
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<taxonomicName id="4C3E4D22FF99FF98FF0ADC2DB064FAFC" authority="Wheeler & Manchester, 2021" authorityName="Wheeler & Manchester" authorityYear="2021" box="[244,380,1353,1374]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="307" phylum="Tracheophyta" rank="species" species="dodgei" status="sp. nov.">
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<emphasis id="B94AEAB3FF99FF98FF0ADC2DB064FAFC" bold="true" box="[244,380,1353,1374]" italics="true" pageId="8" pageNumber="307">Fagus dodgei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27957C8FF99FF98FE7ADC2DB0CCFAFC" box="[388,468,1353,1374]" pageId="8" pageNumber="307" rank="species">sp. nov.</taxonomicNameLabel>
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, a, b, d–h: UF 279-34468; c: UF 279-30165. a:
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<taxonomicName id="4C3E4D22FF99FF98FC3EDC2DB2E4FAFC" box="[960,1020,1353,1374]" family="Wood" pageId="8" pageNumber="324" rank="family">Wood</taxonomicName>
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diffuse-porous to semi-ring-porous with distinct latewood zone with narrower vessels; vessels solitary and in short multiples; diffuse, diffuse-in-aggregates axial parenchyma visible in latewood, TS. b: Growth ring boundary, TS. c: Opposite intervessel pitting, TLS. d: Scalariform perforation plate with fewer than 10 bars, RLS. e: Simple perforation plates (PP), RLS. f: Vessel-ray parenchyma pitting with reduced borders and frequently oval in outline, RLS. g: Rays 1–4(–5)-seriate with variable numbers of marginal rows, TLS. h: Rays of two distinct sizes, widest rays>10-seriate, TLS. Scale bars: 200 µm in a, h; 100 µm in b, e, g; 50 µm in d, f.
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</paragraph>
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</caption>
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<paragraph id="8B8136A1FF99FF98FF6EDF4DB3A3F9C0" blockId="8.[106,745,1576,1634]" pageId="8" pageNumber="307">
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<materialsCitation id="3B563CFCFF99FF98FF6EDF4DB3F1F9E2" box="[144,745,1576,1601]" collectionCode="R" location="Paleobotany Collections" pageId="8" pageNumber="307" specimenCount="1" stateProvince="Florida">
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<collectionCode id="ED2FAE64FF99FF98FF6EDF4DB1BAF9E2" box="[144,162,1577,1600]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="307">R</collectionCode>
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e p o s i t o r y.
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<location id="8EE1607AFF99FF98FEA2DF4CB362F9E2" LSID="urn:lsid:plazi:treatment:039787B7FF96FF99FBECDE6AB5AEFC0C:8EE1607AFF99FF98FEA2DF4CB362F9E2" box="[348,634,1576,1600]" name="Paleobotany Collections" pageId="8" pageNumber="307" stateProvince="Florida">Paleobotany Collections</location>
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,
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<collectingRegion id="49FAF843FF99FF98FD62DF4CB3F1F9E2" box="[668,745,1576,1600]" country="United States of America" name="Florida" pageId="8" pageNumber="307">Florida</collectingRegion>
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</materialsCitation>
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<materialsCitation id="3B563CFCFF99FF98FF94DF2FB3AFF9C0" box="[106,695,1610,1634]" collectionCode="R" country="United States of America" location="Museum of Natural History" pageId="8" pageNumber="307" specimenCount="1" stateProvince="Florida">
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<location id="8EE1607AFF99FF98FF94DF2FB08AF9C0" LSID="urn:lsid:plazi:treatment:039787B7FF96FF99FBECDE6AB5AEFC0C:8EE1607AFF99FF98FF94DF2FB08AF9C0" box="[106,402,1610,1634]" country="United States of America" name="Museum of Natural History" pageId="8" pageNumber="307" stateProvince="Florida">Museum of Natural History</location>
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, Gainesville,
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<collectingRegion id="49FAF843FF99FF98FDDADF2EB36CF9C0" box="[548,628,1610,1634]" country="United States of America" name="Florida" pageId="8" pageNumber="307">Florida</collectingRegion>
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,
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<collectingCountry id="F3297631FF99FF98FD80DF2EB3AFF9C0" box="[638,695,1610,1634]" name="United States of America" pageId="8" pageNumber="307">USA</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8136A1FF99FF98FF6EDF18B0F5F975" blockId="8.[106,745,1659,1751]" pageId="8" pageNumber="307">
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E t y m o l o g y. Named for William W. Dodge IV, who made sure paleoxylotomy in
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<collectingRegion id="49FAF843FF99FF98FE56DFF9B353F917" box="[424,587,1693,1717]" country="United States of America" name="North Carolina" pageId="8" pageNumber="307">North Carolina</collectingRegion>
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was alive and well during the Covid-19 pandemic.
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</paragraph>
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<paragraph id="8B8136A1FF99FF98FF6EDF94B3F1F8A5" blockId="8.[106,745,1775,1833]" box="[144,745,1775,1800]" pageId="8" pageNumber="307">
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T y p e l o c a l i t y. UF 279. About
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<quantity id="4CC69B44FF99FF98FDE1DF94B340F8A5" box="[543,600,1775,1800]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="307" unit="km" value="3.0">3 km</quantity>
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east of Post,
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</paragraph>
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<paragraph id="8B8136A1FF99FF98FF94DE75B0BFF88B" blockId="8.[106,745,1775,1833]" box="[106,423,1809,1833]" pageId="8" pageNumber="307">
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Crook County,
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<collectingRegion id="49FAF843FF99FF98FEF3DE75B078F88B" box="[269,352,1809,1833]" country="United States of America" name="Oregon" pageId="8" pageNumber="307">Oregon</collectingRegion>
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,
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<collectingCountry id="F3297631FF99FF98FE95DE75B0BCF88B" box="[363,420,1809,1833]" name="United States of America" pageId="8" pageNumber="307">USA</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B8136A1FF99FF98FF6EDE27B3F1F8F8" blockId="8.[106,745,1858,1915]" box="[144,745,1858,1883]" pageId="8" pageNumber="307">T y p e s t r a t u m a n d a g e. John Day Formation,</paragraph>
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<paragraph id="8B8136A1FF99FF98FF94DE00B1D8F8D9" blockId="8.[106,745,1858,1915]" box="[106,192,1892,1915]" pageId="8" pageNumber="307">Eocene.</paragraph>
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<paragraph id="8B8136A1FF99FF98FF6EDEF1B52FF821" blockId="8.[106,745,1941,2100]" lastBlockId="8.[820,1460,1576,1923]" pageId="8" pageNumber="307">
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D i a g n o s i s. Growth rings distinct. Diffuse-porous to semi-ring-porous. Vessels mostly solitary, but also in occasional radial, oblique, or tangential pairs; tangential diameters average <100 µm; perforations simple and scalariform; intervessel pits opposite, transitional to scalariform; pits to ray parenchyma oval to horizontally elongate, with reduced borders. Non-septate fibers with distinctly bordered pits on radial walls. Axial parenchyma diffuse and diffuse-in-aggregates as short uniseriate bands. Rays uniseriate and multiseriate, tending to be of two sizes, wider multiseriate rays>10-seriate; wide rays irregularly spaced; average multiseriate ray height>
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||
<quantity id="4CC69B44FF99FF98FB1CDF8EB406F8A0" box="[1250,1310,1770,1794]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="307" unit="mm" value="1.0">1 mm</quantity>
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; homocellular to heterocellular, body composed of procumbent cells with marginal rows of square and upright cells. Solitary rhomboidal crystals occasional in procumbent ray cells. Storied structure absent.
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||
</paragraph>
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||
<paragraph id="8B8136A1FF99FF98FCA4DEFFB4ABF796" blockId="8.[820,1459,1947,2100]" pageId="8" pageNumber="307">
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D e s c r i p t i o n. Growth rings distinct, marked by radially narrow fibers, and differences between latewood and earlywood vessels of subsequent rings (latewood vessels are narrower and not as crowded as earlywood vessels) (Textfig. 4a, b). Diffuse-porous to semi-ring-porous (
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||
<figureCitation id="13052A24FF99FF98FAD6D178B4BEF796" box="[1320,1446,2076,2100]" captionStart="Text-fig" captionStartId="8.[125,206,1353,1374]" captionTargetBox="[125,1440,151,1334]" captionTargetId="figure-275@8.[125,1440,150,1841]" captionTargetPageId="8" captionText="Text-fig. 4. Fagus dodgei sp. nov., a, b, d–h: UF 279-34468; c: UF 279-30165. a: Wood diffuse-porous to semi-ring-porous with distinct latewoodzone with narrower vessels; vessels solitary andin short multiples;diffuse, diffuse-in-aggregatesaxial parenchyma visible in latewood, TS. b: Growth ring boundary, TS. c: Opposite intervessel pitting, TLS. d: Scalariform perforation plate with fewer than 10 bars, RLS. e: Simple perforation plates (PP), RLS. f: Vessel-ray parenchyma pitting with reduced borders and frequently oval in outline, RLS. g: Rays 1–4(–5)-seriate with variable numbers of marginal rows, TLS. h: Rays of two distinct sizes, widest rays>10-seriate, TLS. Scale bars: 200 µm in a, h; 100 µm in b, e, g; 50 µm in d, f." figureDoi="http://doi.org/10.5281/zenodo.7167591" httpUri="https://zenodo.org/record/7167591/files/figure.png" pageId="8" pageNumber="307">Text-fig. 4a</figureCitation>
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||
).
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||
</paragraph>
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||
<paragraph id="8B8136A1FF98FF99FF58D9F3B31EFE13" blockId="9.[129,768,151,882]" pageId="9" pageNumber="308">
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||
Vessels mostly solitary, but also occasional radial, oblique and tangential pairs; average earlywood tangential diameter 55 (11), range 39–79 µm; vessel frequency in first half of growth ring ca. 90/mm
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||
<superScript id="7C4B9BE9FF98FF99FDEFD992B301FEA6" attach="left" box="[529,537,246,260]" fontSize="6" pageId="9" pageNumber="308">2</superScript>
|
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; perforations simple and scalariform with 4-24 bars, mostly 8–12 bars (Textfig. 4d, e); intervessel pits opposite (5–7 µm), transitional to scalariform (
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<figureCitation id="13052A24FF98FF99FEDAD83CB0BCFED2" box="[292,420,344,368]" captionStart="Text-fig" captionStartId="8.[125,206,1353,1374]" captionTargetBox="[125,1440,151,1334]" captionTargetId="figure-275@8.[125,1440,150,1841]" captionTargetPageId="8" captionText="Text-fig. 4. Fagus dodgei sp. nov., a, b, d–h: UF 279-34468; c: UF 279-30165. a: Wood diffuse-porous to semi-ring-porous with distinct latewoodzone with narrower vessels; vessels solitary andin short multiples;diffuse, diffuse-in-aggregatesaxial parenchyma visible in latewood, TS. b: Growth ring boundary, TS. c: Opposite intervessel pitting, TLS. d: Scalariform perforation plate with fewer than 10 bars, RLS. e: Simple perforation plates (PP), RLS. f: Vessel-ray parenchyma pitting with reduced borders and frequently oval in outline, RLS. g: Rays 1–4(–5)-seriate with variable numbers of marginal rows, TLS. h: Rays of two distinct sizes, widest rays>10-seriate, TLS. Scale bars: 200 µm in a, h; 100 µm in b, e, g; 50 µm in d, f." figureDoi="http://doi.org/10.5281/zenodo.7167591" httpUri="https://zenodo.org/record/7167591/files/figure.png" pageId="9" pageNumber="308">Text-fig. 4c</figureCitation>
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); pits to ray parenchyma oval to horizontally elongate, with reduced borders (
|
||
<figureCitation id="13052A24FF98FF99FD8CD81CB3EBFE32" box="[626,755,376,400]" captionStart="Text-fig" captionStartId="8.[125,206,1353,1374]" captionTargetBox="[125,1440,151,1334]" captionTargetId="figure-275@8.[125,1440,150,1841]" captionTargetPageId="8" captionText="Text-fig. 4. Fagus dodgei sp. nov., a, b, d–h: UF 279-34468; c: UF 279-30165. a: Wood diffuse-porous to semi-ring-porous with distinct latewoodzone with narrower vessels; vessels solitary andin short multiples;diffuse, diffuse-in-aggregatesaxial parenchyma visible in latewood, TS. b: Growth ring boundary, TS. c: Opposite intervessel pitting, TLS. d: Scalariform perforation plate with fewer than 10 bars, RLS. e: Simple perforation plates (PP), RLS. f: Vessel-ray parenchyma pitting with reduced borders and frequently oval in outline, RLS. g: Rays 1–4(–5)-seriate with variable numbers of marginal rows, TLS. h: Rays of two distinct sizes, widest rays>10-seriate, TLS. Scale bars: 200 µm in a, h; 100 µm in b, e, g; 50 µm in d, f." figureDoi="http://doi.org/10.5281/zenodo.7167591" httpUri="https://zenodo.org/record/7167591/files/figure.png" pageId="9" pageNumber="308">Text-fig. 4f</figureCitation>
|
||
); vessel element lengths 450–520 µm.
|
||
</paragraph>
|
||
<paragraph id="8B8136A1FF98FF99FF58D8DDB1D8FE53" blockId="9.[129,768,151,882]" pageId="9" pageNumber="308">Non-septate fibers with distinctly bordered pits on radial walls.</paragraph>
|
||
<paragraph id="8B8136A1FF98FF99FF58D89DB30BFDF3" blockId="9.[129,768,151,882]" pageId="9" pageNumber="308">
|
||
Axial parenchyma apotracheal, diffuse as isolated cells and diffuse-in-aggregates as short uniseriate bands, more easily seen in latewood (
|
||
<figureCitation id="13052A24FF98FF99FE78DB5DB31EFDF3" box="[390,518,569,593]" captionStart="Text-fig" captionStartId="8.[125,206,1353,1374]" captionTargetBox="[125,1440,151,1334]" captionTargetId="figure-275@8.[125,1440,150,1841]" captionTargetPageId="8" captionText="Text-fig. 4. Fagus dodgei sp. nov., a, b, d–h: UF 279-34468; c: UF 279-30165. a: Wood diffuse-porous to semi-ring-porous with distinct latewoodzone with narrower vessels; vessels solitary andin short multiples;diffuse, diffuse-in-aggregatesaxial parenchyma visible in latewood, TS. b: Growth ring boundary, TS. c: Opposite intervessel pitting, TLS. d: Scalariform perforation plate with fewer than 10 bars, RLS. e: Simple perforation plates (PP), RLS. f: Vessel-ray parenchyma pitting with reduced borders and frequently oval in outline, RLS. g: Rays 1–4(–5)-seriate with variable numbers of marginal rows, TLS. h: Rays of two distinct sizes, widest rays>10-seriate, TLS. Scale bars: 200 µm in a, h; 100 µm in b, e, g; 50 µm in d, f." figureDoi="http://doi.org/10.5281/zenodo.7167591" httpUri="https://zenodo.org/record/7167591/files/figure.png" pageId="9" pageNumber="308">Text-fig. 4a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136A1FF98FF99FF58DB3DB218FC90" blockId="9.[129,768,151,882]" pageId="9" pageNumber="308">
|
||
Rays uniseriate and multiseriate, tending to be of two sizes, with rays 1–5(–6)-seriate common, and wider multiseriate rays to 18–20-seriate, with few intermediate widths; wide rays irregularly spaced (
|
||
<figureCitation id="13052A24FF98FF99FE83DBDDB30EFD73" box="[381,534,697,721]" captionStart="Text-fig" captionStartId="8.[125,206,1353,1374]" captionTargetBox="[125,1440,151,1334]" captionTargetId="figure-275@8.[125,1440,150,1841]" captionTargetPageId="8" captionText="Text-fig. 4. Fagus dodgei sp. nov., a, b, d–h: UF 279-34468; c: UF 279-30165. a: Wood diffuse-porous to semi-ring-porous with distinct latewoodzone with narrower vessels; vessels solitary andin short multiples;diffuse, diffuse-in-aggregatesaxial parenchyma visible in latewood, TS. b: Growth ring boundary, TS. c: Opposite intervessel pitting, TLS. d: Scalariform perforation plate with fewer than 10 bars, RLS. e: Simple perforation plates (PP), RLS. f: Vessel-ray parenchyma pitting with reduced borders and frequently oval in outline, RLS. g: Rays 1–4(–5)-seriate with variable numbers of marginal rows, TLS. h: Rays of two distinct sizes, widest rays>10-seriate, TLS. Scale bars: 200 µm in a, h; 100 µm in b, e, g; 50 µm in d, f." figureDoi="http://doi.org/10.5281/zenodo.7167591" httpUri="https://zenodo.org/record/7167591/files/figure.png" pageId="9" pageNumber="308">
|
||
Text-fig.
|
||
<quantity id="4CC69B44FF98FF99FE21DBDDB0E4FD73" box="[479,508,697,721]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="9" pageNumber="308" unit="g" value="4.0">4g</quantity>
|
||
, h
|
||
</figureCitation>
|
||
); average multiseriate ray height
|
||
<quantity id="4CC69B44FF98FF99FF08DBBDB041FD53" box="[246,345,729,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" pageId="9" pageNumber="308" unit="mm" value="1.56">1.56 mm</quantity>
|
||
, range
|
||
<quantity id="4CC69B44FF98FF99FE57DBBDB351FD53" box="[425,585,729,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.7149999999999999" metricValueMax="2.69" metricValueMin="0.74" pageId="9" pageNumber="308" unit="mm" value="1.7149999999999999" valueMax="2.69" valueMin="0.74">0.74–2.69 mm</quantity>
|
||
; homocellular to heterocellular, body composed of procumbent cells usually with 1–4 (up to 8) marginal rows of square and upright cells.
|
||
</paragraph>
|
||
<paragraph id="8B8136A1FF98FF99FF58DA5EB0F3FCD0" blockId="9.[129,768,151,882]" pageId="9" pageNumber="308">Solitary rhomboidal crystals occasional in procumbent ray cells. Storied structure absent.</paragraph>
|
||
<paragraph id="8B8136A1FF98FF99FF58DAEDB0EAFA80" blockId="9.[129,768,905,1860]" pageId="9" pageNumber="308">
|
||
C o m p a r i s o n s w i t h e x t a n t w o o d s. The following combination of features indicates affinities with
|
||
<taxonomicName id="4C3E4D22FF98FF99FF44DAAEB067FC43" box="[186,383,969,993]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FF44DAAEB1E5FC43" box="[186,253,970,993]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
(Fagaceae)
|
||
</taxonomicName>
|
||
: distinct growth rings (1p), simple and scalariform perforation plates (13p 14p), opposite intervessel pitting (21p), vessel-ray parenchyma pits with reduced borders (30a), narrow numerous vessels (42a 43a 46a 47a), non-septate fibers with distinctly bordered pits on radial walls (62p 66p), diffuse and diffuse-in-aggregates axial parenchyma (76p 77p), wide rays>10 cells wide and>
|
||
<quantity id="4CC69B44FF98FF99FF6DDDCEB1D4FB60" box="[147,204,1194,1218]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="308" unit="mm" value="1.0">1 mm</quantity>
|
||
high (99p 102p). To the best of our knowledge, there is not a reliable means of distinguishing present-day
|
||
<taxonomicName id="4C3E4D22FF98FF99FD43DDAFB1D6FAA0" pageId="9" pageNumber="308">
|
||
<emphasis id="B94AEAB3FF98FF99FD43DDAFB218FB40" box="[701,768,1227,1250]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
based on wood anatomy alone (
|
||
<bibRefCitation id="EFAF4B50FF98FF99FDDCDD8EB1A1FA80" author="Brazier, J. D. & Franklin, G. L." pageId="9" pageNumber="308" pagination="1 - 96" refId="ref22957" refString="Brazier, J. D., Franklin, G. L. (1961): Identification of hardwoods. A microscope key. - Forest Products Research Bulletin, 46: 1 - 96." type="journal article" year="1961">Brazier and Franklin 1961</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAF4B50FF98FF99FF3DDC6EB0FFFA80" author="Wheeler, E. A. & Dillhoff, T. A." box="[195,487,1290,1314]" pageId="9" pageNumber="308" pagination="1 - 101" refId="ref28301" refString="Wheeler, E. A., Dillhoff, T. A. (2009): The Middle Miocene Wood Flora of Vantage, Washington, USA. - IAWA Journal, Supplement 7: 1 - 101." type="journal article" year="2009">Wheeler and Dillhoff 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136A1FF98FF99FF58DC4EB031F8E6" blockId="9.[129,768,905,1860]" pageId="9" pageNumber="308">
|
||
Woods of the
|
||
<taxonomicName id="4C3E4D22FF98FF99FEC2DC4EB0A2FAE0" box="[316,442,1322,1346]" class="Magnoliopsida" family="Platanaceae" kingdom="Plantae" order="Proteales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="family">Platanaceae</taxonomicName>
|
||
are common in the Paleogene of the western
|
||
<collectingCountry id="F3297631FF98FF99FEC8DC2FB077FAC1" box="[310,367,1355,1379]" name="United States of America" pageId="9" pageNumber="308">USA</collectingCountry>
|
||
. and superficially resemble
|
||
<taxonomicName id="4C3E4D22FF98FF99FD48DC2FB3E1FAC0" box="[694,761,1355,1378]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FD48DC2FB3E1FAC0" box="[694,761,1355,1378]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
. However, there are features useful in distinguishing them (e.g.,
|
||
<bibRefCitation id="EFAF4B50FF98FF99FF3FDCEFB0E2FA01" author="Panshin, A. J. & DeZeeuw, C." box="[193,506,1419,1443]" pageId="9" pageNumber="308" refId="ref26488" refString="Panshin, A. J., DeZeeuw, C. (1980): Textbook of Wood Technology. 4 th ed. - McGraw-Hill, New York, 722 pp." type="book" year="1980">Panshin and DeZeeuw 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAF4B50FF98FF99FDF7DCEFB3BFFA01" author="Hoadley, B." box="[521,679,1419,1443]" pageId="9" pageNumber="308" refId="ref24597" refString="Hoadley, B. (1990): Identifying wood. Accurate results with simple tools. - Taunton Press, Newtown, Connecticut, 233 pp." type="book" year="1990">Hoadley 1990</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C3E4D22FF98FF99FD43DCE8B218FA01" box="[701,768,1420,1443]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FD43DCE8B218FA01" box="[701,768,1420,1443]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
consistently has vessel-ray parenchyma pits with reduced borders that are somewhat horizontally elongate, while
|
||
<taxonomicName id="4C3E4D22FF98FF99FF7FDC8FB1F8F9A1" box="[129,224,1515,1539]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FF7FDC8FB1F8F9A1" box="[129,224,1515,1539]" italics="true" pageId="9" pageNumber="308">Platanus</emphasis>
|
||
</taxonomicName>
|
||
L. has vessel-ray parenchyma pits similar to intervessel pits. Vessel multiples are rare in both, but
|
||
<taxonomicName id="4C3E4D22FF98FF99FD5EDF6FB3E7F981" box="[672,767,1547,1571]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FD5EDF6FB3E7F981" box="[672,767,1547,1571]" italics="true" pageId="9" pageNumber="308">Platanus</emphasis>
|
||
</taxonomicName>
|
||
has some tangential multiples and a tendency to vessels being tangentially arranged; consequently, intervessel pits are more common in
|
||
<taxonomicName id="4C3E4D22FF98FF99FE8BDF0FB0CCF921" box="[373,468,1643,1667]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FE8BDF0FB0CCF921" box="[373,468,1643,1667]" italics="true" pageId="9" pageNumber="308">Platanus</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C3E4D22FF98FF99FDC6DF08B363F921" box="[568,635,1644,1667]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FDC6DF08B363F921" box="[568,635,1644,1667]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
. Intervessel pits are consistently opposite in
|
||
<emphasis id="B94AEAB3FF98FF99FE03DFE8B37DF906" box="[509,613,1676,1700]" italics="true" pageId="9" pageNumber="308">
|
||
<taxonomicName id="4C3E4D22FF98FF99FE03DFE8B346F906" box="[509,606,1676,1700]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">Platanus</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
while
|
||
<taxonomicName id="4C3E4D22FF98FF99FD43DFE8B218F901" box="[701,768,1676,1699]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FD43DFE8B218F901" box="[701,768,1676,1699]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
usually has transitional (opposite- scalariform) intervessel pitting. Spacing of and size of the wide rays is more uniform in
|
||
<taxonomicName id="4C3E4D22FF98FF99FF06DF88B04FF8A6" box="[248,343,1772,1796]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FF06DF88B04FF8A6" box="[248,343,1772,1796]" italics="true" pageId="9" pageNumber="308">Platanus</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C3E4D22FF98FF99FE52DF89B0F7F8A6" box="[428,495,1773,1796]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FE52DF89B0F7F8A6" box="[428,495,1773,1796]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
. All these features can be reviewed in the InsideWood image collection (InsideWood 2004-onwards).
|
||
</paragraph>
|
||
<paragraph id="8B8136A1FF98FF99FF58DE38B525FF6D" blockId="9.[128,768,1883,2100]" lastBlockId="9.[843,1482,151,528]" pageId="9" pageNumber="308">
|
||
C o m p a r i s o n s w i t h f o s s i l w o o d s.
|
||
<taxonomicName id="4C3E4D22FF98FF99FD4ADE38B3EFF8D1" box="[692,759,1884,1907]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FD4ADE38B3EFF8D1" box="[692,759,1884,1907]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
- like woods have been described as species of
|
||
<taxonomicName id="4C3E4D22FF98FF99FD48DE18B3E1F831" box="[694,761,1916,1939]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FD48DE18B3E1F831" box="[694,761,1916,1939]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E4D22FF98FF99FF7EDEF8B051F810" authority="STOPES" authorityName="STOPES" box="[128,329,1947,1972]" genus="Fagoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="308" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FF7EDEF8B1EAF816" box="[128,242,1948,1972]" italics="true" pageId="9" pageNumber="308">Fagoxylon</emphasis>
|
||
STOPES
|
||
</taxonomicName>
|
||
et fUJII, and
|
||
<taxonomicName id="4C3E4D22FF98FF99FE0BDEF8B1DAF871" authority="UNGER" authorityName="UNGER" genus="Fegonium" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="308" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FE0BDEF8B379F811" box="[501,609,1948,1971]" italics="true" pageId="9" pageNumber="308">Fegonium</emphasis>
|
||
UNGER (
|
||
<bibRefCitation id="EFAF4B50FF98FF99FD33DEFFB1A3F871" author="Suss, H." pageId="9" pageNumber="308" pagination="161 - 183" refId="ref27625" refString="Suss, H. (1986): Untersuchungen uber fossile Buchenholzer. Beitrage zu einer Monographie der Gattung Fagoxylon Stopes & Fujii. - Feddes Repertorium, 97 (3 - 4): 161 - 183. https: // doi. org / 10.1002 / fedr. 4910970310" type="journal article" year="1986">Süss 1986</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Descriptions of most lack details that would permit confirming their relationships with
|
||
<taxonomicName id="4C3E4D22FF98FF99FE07DEB8B324F851" box="[505,572,2012,2035]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FE07DEB8B324F851" box="[505,572,2012,2035]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
. There are at least ten
|
||
<taxonomicName id="4C3E4D22FF98FF99FF53DE98B1E8F7B1" box="[173,240,2044,2067]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FF53DE98B1E8F7B1" box="[173,240,2044,2067]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
-like woods reported from the Mio-Pliocene of Europe (
|
||
<bibRefCitation id="EFAF4B50FF98FF99FF24D178B0B3F796" author="Gregory, M. & Poole, I. & Wheeler, E. A." box="[218,427,2076,2100]" pageId="9" pageNumber="308" pagination="1 - 230" refId="ref24259" refString="Gregory, M., Poole, I., Wheeler, E. A. (2009): Fossil dicot wood names. - IAWA Journal, Supplement 6: 1 - 230." type="journal article" year="2009">Gregory et al. 2009</bibRefCitation>
|
||
), but their descriptions are brief, so it is not possible to do detailed comparisons of them with this late Eocene wood.
|
||
</paragraph>
|
||
<paragraph id="8B8136A1FF98FF99FC8FD9B3B5D1FDAD" blockId="9.[843,1482,151,528]" pageId="9" pageNumber="308">
|
||
There are differences between
|
||
<taxonomicName id="4C3E4D22FF98FF99FB4FD9BCB45AFF4D" authorityName="Wheeler & Manchester" authorityYear="2021" box="[1201,1346,215,239]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="species" species="dodgei">
|
||
<emphasis id="B94AEAB3FF98FF99FB4FD9BCB45AFF4D" box="[1201,1346,215,239]" italics="true" pageId="9" pageNumber="308">Fagus dodgei</emphasis>
|
||
</taxonomicName>
|
||
and the midMiocene
|
||
<taxonomicName id="4C3E4D22FF98FF99FC50D99CB5FBFEAF" authority="E. A. WHEELER" authorityName="E. A. WHEELER" box="[942,1251,248,272]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="species" species="manosii">
|
||
<emphasis id="B94AEAB3FF98FF99FC50D99CB553FEAD" box="[942,1099,248,271]" italics="true" pageId="9" pageNumber="308">Fagus manosii</emphasis>
|
||
E.A.WHEELER
|
||
</taxonomicName>
|
||
et T.A.DILLHOff from
|
||
<collectingRegion id="49FAF843FF98FF99FCB5D873B510FE92" box="[843,1032,279,304]" country="United States of America" name="Washington" pageId="9" pageNumber="308">Washington State</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3297631FF98FF99FBECD87CB55EFE92" box="[1042,1094,280,304]" name="United States of America" pageId="9" pageNumber="308">USA</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFAF4B50FF98FF99FBADD873B46EFE92" author="Wheeler, E. A. & Dillhoff, T. A." box="[1107,1398,279,304]" pageId="9" pageNumber="308" pagination="1 - 101" refId="ref28301" refString="Wheeler, E. A., Dillhoff, T. A. (2009): The Middle Miocene Wood Flora of Vantage, Washington, USA. - IAWA Journal, Supplement 7: 1 - 101." type="journal article" year="2009">Wheeler and Dillhoff 2009</bibRefCitation>
|
||
), which had more scalariform intervessel pits and larger opposite intervessel pits, narrower and shorter rays (to 16-seriate and
|
||
<quantity id="4CC69B44FF98FF99FCB5D81CB287FE2D" box="[843,927,376,400]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="308" unit="mm" value="1.7">1.7 mm</quantity>
|
||
high), less axial parenchyma, and lacked crystals. Thus, although there is no reliable way to distinguish between extant species of
|
||
<taxonomicName id="4C3E4D22FF98FF99FBFED8DCB55BFE6D" box="[1024,1091,440,463]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FBFED8DCB55BFE6D" box="[1024,1091,440,463]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
, we are choosing to recognize
|
||
<taxonomicName id="4C3E4D22FF98FF99FA79D8DCB28CFE4D" authorityName="Wheeler & Manchester" authorityYear="2021" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="species" species="dodgei">
|
||
<emphasis id="B94AEAB3FF98FF99FA79D8DCB28CFE4D" italics="true" pageId="9" pageNumber="308">Fagus dodgei</emphasis>
|
||
</taxonomicName>
|
||
as a new species to emphasize that it differs from the mid-Miocene Vantage
|
||
<taxonomicName id="4C3E4D22FF98FF99FBC3D89CB598FDAD" box="[1085,1152,504,527]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FBC3D89CB598FDAD" box="[1085,1152,504,527]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
wood.
|
||
</paragraph>
|
||
<paragraph id="8B8136A1FF98FF99FC8FDB43B2F4FC9D" blockId="9.[843,1482,551,831]" pageId="9" pageNumber="308">
|
||
R e m a r k s.
|
||
<bibRefCitation id="EFAF4B50FF98FF99FBF7DB43B43DFD9D" author="Denk, T. & Grimm, G. W." box="[1033,1317,551,575]" pageId="9" pageNumber="308" pagination="83 - 100" refId="ref23724" refString="Denk, T., Grimm, G. W. (2009): The biogeographic history of beech trees. - Review of Palaeobotany and Palynology, 158: 83 - 100. https: // doi. org / 10.1016 / j. revpalbo. 2009.08.007" type="journal article" year="2009">Denk and Grimm (2009)</bibRefCitation>
|
||
proposed that
|
||
<taxonomicName id="4C3E4D22FF98FF99FCB5DB23B296FDFC" box="[843,910,583,606]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FCB5DB23B296FDFC" box="[843,910,583,606]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
evolved in the Northern Pacific Region. Fossil species of
|
||
<taxonomicName id="4C3E4D22FF98FF99FC38DB03B511FDDC" box="[966,1033,615,638]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FC38DB03B511FDDC" box="[966,1033,615,638]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
, based on fruits and leaves, are known from the Paleogene of the Pacific Northwest. The middle Eocene
|
||
<taxonomicName id="4C3E4D22FF98FF99FC61DBC3B549FD1D" box="[927,1105,679,703]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="species" species="langevinii">
|
||
<emphasis id="B94AEAB3FF98FF99FC61DBC3B549FD1D" box="[927,1105,679,703]" italics="true" pageId="9" pageNumber="308">Fagus langevinii</emphasis>
|
||
</taxonomicName>
|
||
<collectingRegion id="49FAF843FF98FF99FBA8DBC3B5C4FD1F" box="[1110,1244,679,702]" country="United Kingdom" name="Manchester" pageId="9" pageNumber="308">MANCHESTER</collectingRegion>
|
||
et R.M.DILLHOff from McAbee,
|
||
<collectingRegion id="49FAF843FF98FF99FC40DBA3B59AFD7D" box="[958,1154,711,735]" country="Canada" name="British Columbia" pageId="9" pageNumber="308">British Columbia</collectingRegion>
|
||
, and Republic,
|
||
<collectingRegion id="49FAF843FF98FF99FABADBA3B4DEFD7D" box="[1348,1478,711,735]" country="United States of America" name="Washington" pageId="9" pageNumber="308">Washington</collectingRegion>
|
||
, differs from the early Oligocene
|
||
<taxonomicName id="4C3E4D22FF98FF99FB5EDB8CB4D1FD5F" authority="R. W. CHANEY" authorityName="R. W. CHANEY" box="[1184,1481,743,767]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="species" species="pacifica">
|
||
<emphasis id="B94AEAB3FF98FF99FB5EDB8CB423FD5D" box="[1184,1339,743,767]" italics="true" pageId="9" pageNumber="308">Fagus pacifica</emphasis>
|
||
R.W.CHANEY
|
||
</taxonomicName>
|
||
from the Bridge Creek flora of
|
||
<collectingRegion id="49FAF843FF98FF99FB48DA63B41EFCBD" box="[1206,1286,775,799]" country="United States of America" name="Oregon" pageId="9" pageNumber="308">Oregon</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFAF4B50FF98FF99FAE7DA63B2F9FC9D" author="Manchester, S. R. & Dillhoff, R. M." pageId="9" pageNumber="308" pagination="1509 - 1517" refId="ref25747" refString="Manchester, S. R., Dillhoff, R. M. (2004): Fagus (Fagaceae) fruits, foliage, and pollen from the Middle Eocene of Pacific Northwestern North America. - Canadian Journal of Botany, 82: 1509 - 1517. https: // doi. org / 10.1139 / b 04 - 112" type="journal article" year="2004">
|
||
<collectingRegion id="49FAF843FF98FF99FAE7DA63B48FFCBD" box="[1305,1431,775,799]" country="United Kingdom" name="Manchester" pageId="9" pageNumber="308">Manchester</collectingRegion>
|
||
and Dillhoff 2004
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136A1FF98FF99FC8FDA32B5AEFC0C" blockId="9.[843,1482,854,942]" pageId="9" pageNumber="308">
|
||
C o - o c c u r r i n g f r u i t s. A single silicified
|
||
<taxonomicName id="4C3E4D22FF98FF99FCB5DA13B296FC2C" box="[843,910,887,910]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="308" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B94AEAB3FF98FF99FCB5DA13B296FC2C" box="[843,910,887,910]" italics="true" pageId="9" pageNumber="308">Fagus</emphasis>
|
||
</taxonomicName>
|
||
nut was recovered from the same locality, UF 279 (
|
||
<bibRefCitation id="EFAF4B50FF98FF99FCACDAF2B5B3FC0C" author="Manchester, S. R. & McIntosh, W. C." box="[850,1195,918,942]" pageId="9" pageNumber="308" pagination="7 - 17" refId="ref25878" refString="Manchester, S. R., McIntosh, W. C. (2007): Late Eocene silicified fruits and seeds from the John Day Formation near Post, Oregon. - PaleoBios, 27 (1): 7 - 17." type="journal article" year="2007">
|
||
<collectingRegion id="49FAF843FF98FF99FCACDAF2B2C8FC0C" box="[850,976,918,942]" country="United Kingdom" name="Manchester" pageId="9" pageNumber="308">Manchester</collectingRegion>
|
||
and McIntosh 2007
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |