treatments-xml/data/03/97/87/03978783FF8DFFF2FC82FD60AFD29B3D.xml
2024-06-21 12:22:17 +02:00

540 lines
57 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="629715E775CBD137D96C189A22E38A99" ID-DOI="10.15553/c2016v711a7" ID-GBIF-Dataset="508bebc5-39aa-4f9c-b544-171e992dbbea" ID-ISSN="2235-3658" ID-Zenodo-Dep="161516" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476779283805" checkinUser="plazi" docAuthor="Wendy L. Applequist" docDate="2016" docId="03978783FF8DFFF2FC82FD60AFD29B3D" docLanguage="en" docName="Candollea.71.1.33-60.pdf" docOrigin="Candollea 71 (1)" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Homalium (sect. Eumyriantheia) Warb. 1893" docType="treatment" docVersion="8" lastPageNumber="36" masterDocId="FFAEFFFBFF8CFFF1FFA6FF9DAA7C9D57" masterDocTitle="Revision of the Malagasy species of Homalium sect. Eumyriantheia Warb. (Salicaceae)" masterLastPageNumber="60" masterPageNumber="33" pageNumber="34" updateTime="1698390116944" updateUser="plazi">
<mods:mods id="CACCAFF8BE5325AE7318F45E651684B4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C4D18994D0933409E2395EDD2FB687D3">
<mods:title id="AAA12B10F28B44CE10B66D06723E93E0">Revision of the Malagasy species of Homalium sect. Eumyriantheia Warb. (Salicaceae)</mods:title>
</mods:titleInfo>
<mods:name id="AD6926E67175318450F013652EDECF71" type="personal">
<mods:role id="160AD291B4BB8E15AC8B211DEF1B86AA">
<mods:roleTerm id="7C43FCF50F113471CA0B0CEDBE5A65E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E34463877C0A63F549B4F6AD334177A7">Wendy L. Applequist</mods:namePart>
</mods:name>
<mods:typeOfResource id="0796DB9737A6944F6B3177C08402FFD7">text</mods:typeOfResource>
<mods:relatedItem id="EFA0A18684A591F2A6ECDD623D1AE85D" type="host">
<mods:titleInfo id="FD3703CB05AC784EA72C2FD1F5141573">
<mods:title id="53FAABF9C48AA454B64EF9AC4346A26A">Candollea</mods:title>
</mods:titleInfo>
<mods:part id="65731C3B6217C80448B71B1D52B1B9A9">
<mods:date id="D93993FDF08EDE0BC263149AA56E7516">2016</mods:date>
<mods:detail id="D48372ADA955DAB3F9141D50004CF64C" type="pubDate">
<mods:number id="D9F15BC306929DA63D90098A0195B03C">2016-03-14</mods:number>
</mods:detail>
<mods:detail id="1788D38CC802D640E7807DAD9914AE14" type="volume">
<mods:number id="FE09C86911EF427F78C92A8D72D67DA4">71</mods:number>
</mods:detail>
<mods:detail id="6A2D80652446EEC66E79D83B69474586" type="issue">
<mods:number id="7D7FAF691F12E2FEBEA6D4973E9579BF">1</mods:number>
</mods:detail>
<mods:extent id="81C80E71E8F2F7D2D3D4F335308593BC" unit="page">
<mods:start id="D742A87C934D65088FC2AC7375282CBC">33</mods:start>
<mods:end id="FB223621874AF84080AD11C0520EDC19">60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="29174D085DE2C98DFD733D784AF2450F">journal article</mods:classification>
<mods:identifier id="F7C9DCAB8487B1BC4E239CE0D7F9D1DA" type="DOI">10.15553/c2016v711a7</mods:identifier>
<mods:identifier id="95976E17B73ABC3D5FD6F3062EF5B456" type="GBIF-Dataset">508bebc5-39aa-4f9c-b544-171e992dbbea</mods:identifier>
<mods:identifier id="F7F5E0A2E8094CEEC3F33BB8EE5CE689" type="ISSN">2235-3658</mods:identifier>
<mods:identifier id="6544E9C66353B860E42DA5C0E3CF0FA0" type="Zenodo-Dep">161516</mods:identifier>
</mods:mods>
<treatment id="03978783FF8DFFF2FC82FD60AFD29B3D" ID-DOI="http://doi.org/10.5281/zenodo.6300556" ID-GBIF-Taxon="177200348" ID-Zenodo-Dep="6300556" LSID="urn:lsid:plazi:treatment:03978783FF8DFFF2FC82FD60AFD29B3D" httpUri="http://treatment.plazi.org/id/03978783FF8DFFF2FC82FD60AFD29B3D" lastPageId="3" lastPageNumber="36" pageId="1" pageNumber="34">
<subSubSection id="C324651EFF8DFFF0FC82FD60AEDB9E61" pageId="1" pageNumber="34" type="nomenclature">
<paragraph id="8B813695FF8DFFF0FC82FD60AEDB9E61" blockId="1.[804,1457,719,1099]" pageId="1" pageNumber="34">
<treatmentCitation id="0A9F1084FF8DFFF0FC82FD60AEDB9E61" pageId="1" pageNumber="34">
<taxonomicName id="4C3E4D16FF8DFFF0FC82FD60AEB19E42" authority="Warb." authorityName="Warb." authorityYear="1893" box="[804,1229,765,789]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="section" section="Eumyriantheia">
<emphasis id="B94AEA87FF8DFFF0FC82FD60A9E69E42" bold="true" box="[804,922,765,789]" italics="true" pageId="1" pageNumber="34">Homalium</emphasis>
sect.
<emphasis id="B94AEA87FF8DFFF0FC7CFD60AEFE9E42" bold="true" box="[986,1154,765,789]" italics="true" pageId="1" pageNumber="34">Eumyriantheia</emphasis>
Warb.
</taxonomicName>
in Engler &amp; Prantl, Nat. Pflanzenfam. III(6a): 36. 1893.
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF8DFFF0FCECFCD9AFCD9957" pageId="1" pageNumber="34" type="reference_group">
<paragraph id="8B813695FF8DFFF0FCECFCD9AFCD9957" blockId="1.[804,1457,719,1099]" pageId="1" pageNumber="34">
Ξ
<treatmentCitation id="0A9F1084FF8DFFF0FCD6FCD5A9D19ED7" pageId="1" pageNumber="34">
<taxonomicName id="4C3E4D16FF8DFFF0FCD6FCD5AE169E37" authority="Thouars" authorityName="Thouars" box="[880,1130,840,864]" class="Magnoliopsida" family="Salicaceae" genus="Myriantheia" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8DFFF0FCD6FCD5A9829E37" box="[880,1022,840,864]" italics="true" pageId="1" pageNumber="34">Myriantheia</emphasis>
Thouars
</taxonomicName>
, Gen. Nov. Madagasc.: 21. 1806
</treatmentCitation>
. Ξ
<taxonomicName id="4C3E4D16FF8DFFF0FC7CFCF5AE2D9ED7" ID-CoL="4Y6K" authority="Warb." authorityName="Warb." authorityYear="1893" box="[986,1105,872,896]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8DFFF0FC7CFCF5AE2D9ED7" box="[986,1105,872,896]" italics="true" pageId="1" pageNumber="34">Homalium</emphasis>
</taxonomicName>
[unranked as §]
<treatmentCitation id="0A9F1084FF8DFFF0FA84FCF5AE759E97" pageId="1" pageNumber="34">
<taxonomicName id="4C3E4D16FF8DFFF0FA84FCF5AE4C9EF7" authority="(Thouars) Baill." authorityName="Baill." baseAuthorityName="Thouars" class="Magnoliopsida" family="Salicaceae" genus="Myriantheia" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8DFFF0FA84FCF5AFCC9ED7" box="[1314,1456,872,896]" italics="true" pageId="1" pageNumber="34">Myriantheia</emphasis>
(Thouars) Baill.
</taxonomicName>
in Bull. Mens. Soc. Linn. Paris 1: 576. 1886
</treatmentCitation>
[as
<taxonomicName id="4C3E4D16FF8DFFF0FB98FC35AEBF9E97" authority="Thouars" authorityName="Thouars" box="[1086,1219,936,960]" class="Magnoliopsida" family="Salicaceae" genus="Myrianthea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8DFFF0FB98FC35AEBF9E97" box="[1086,1219,936,960]" italics="true" pageId="1" pageNumber="34">Myrianthea</emphasis>
</taxonomicName>
]. Ξ
<treatmentCitation id="0A9F1084FF8DFFF0FB5BFC35AE909957" pageId="1" pageNumber="34">
<taxonomicName id="4C3E4D16FF8DFFF0FB5BFC35AEAF9EB7" authority="(Thouars) Kuntze" authorityName="Kuntze" baseAuthorityName="Thouars" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="section" section="Myriantheia">
<emphasis id="B94AEA87FF8DFFF0FB5BFC35AF0F9E97" box="[1277,1395,936,960]" italics="true" pageId="1" pageNumber="34">Homalium</emphasis>
sect.
<emphasis id="B94AEA87FF8DFFF0FCD6FC55A9869EB7" box="[880,1018,968,992]" italics="true" pageId="1" pageNumber="34">Myriantheia</emphasis>
(Thouars) Kuntze
</taxonomicName>
in Post &amp; Kuntze, Lex. Gen. Phan., Prop.: 285. 1903
</treatmentCitation>
[as
<emphasis id="B94AEA87FF8DFFF0FABDFC75AFDC9957" box="[1307,1440,1000,1024]" italics="true" pageId="1" pageNumber="34">Myriantheja</emphasis>
].
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF8DFFF0FCECFB8EAF13991C" pageId="1" pageNumber="34" type="type_taxon">
<paragraph id="8B813695FF8DFFF0FCECFB8EAF13991C" blockId="1.[804,1457,719,1099]" pageId="1" pageNumber="34">
<emphasis id="B94AEA87FF8DFFF0FCECFB8EA9E6997C" bold="true" box="[842,922,1043,1067]" pageId="1" pageNumber="34">
<typeStatus id="54858837FF8DFFF0FCECFB8EA9F3997C" box="[842,911,1043,1067]" pageId="1" pageNumber="34">Typus</typeStatus>
:
</emphasis>
<taxonomicName id="4C3E4D16FF8DFFF0FC07FB8EAF7D997C" ID-CoL="3MCMV" authority="(Tul.) Baill." authorityName="(Tul.) Baill." box="[929,1281,1043,1067]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="species" species="laxiflorum">
<emphasis id="B94AEA87FF8DFFF0FC07FB8EAE02997C" box="[929,1150,1043,1067]" italics="true" pageId="1" pageNumber="34">Homalium laxiflorum</emphasis>
(Tul.) Baill.
</taxonomicName>
<taxonomicName id="4C3E4D16FF8DFFF0FA8BFB8EA9AD991C" ID-CoL="6RRTV" authority="Tul." authorityName="Tul." class="Magnoliopsida" family="Salicaceae" genus="Myriantheia" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="species" species="laxiflora">
<emphasis id="B94AEA87FF8DFFF0FA8BFB8EA9E2991C" italics="true" pageId="1" pageNumber="34">Myriantheia laxiflora</emphasis>
Tul.
</taxonomicName>
) (designated by
<treatmentCitation id="0A9F1084FF8DFFF0FB26FBAEAF13991C" box="[1152,1391,1075,1099]" pageId="1" pageNumber="34">
<bibRefCitation id="EFAF4B64FF8DFFF0FB26FBAEAF1F991C" author="Sleumer, H." box="[1152,1379,1075,1099]" journalOrPublisher="Bull. Jard. Natl. Belg." pageId="1" pageNumber="34" pagination="239 - 328" part="43" refId="ref29189" refString="Sleumer, H. (1973). Revision du genre Homalium Jacq. (Flacourtiacees) en Afrique (y compris Madagascar et les Mascareignes). Bull. Jard. Natl. Belg. 43: 239 - 328." title="Revision du genre Homalium Jacq. (Flacourtiacees) en Afrique (y compris Madagascar et les Mascareignes)" type="journal article" year="1973">Sleumer, 1973: 315</bibRefCitation>
).
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF8DFFF0FCECFBE3AE7E9A22" pageId="1" pageNumber="34" type="description">
<paragraph id="8B813695FF8DFFF0FCECFBE3AE7E9A22" blockId="1.[804,1455,1149,1909]" pageId="1" pageNumber="34">
<emphasis id="B94AEA87FF8DFFF0FCECFBE3A9AE99C2" box="[842,978,1150,1173]" italics="true" pageId="1" pageNumber="34">Trees. Leaves</emphasis>
with stipules usually caducous (rarely in part persistent); secondary veins brochidodromous, little or not prominent; texture fairly thick to coriaceous.
<emphasis id="B94AEA87FF8DFFF0FA83FB20AFD39982" box="[1317,1455,1213,1237]" italics="true" pageId="1" pageNumber="34">Inflorescences</emphasis>
racemose or paniculate, lateral (in part pseudoterminal or clustered at apices); flowers pedicellate (sessile), often with bracteoles borne on pedicels, dehiscing immediately or slightly below flower shortly after anthesis.
<emphasis id="B94AEA87FF8DFFF0FB0DFAA0AE829802" box="[1195,1278,1341,1365]" italics="true" pageId="1" pageNumber="34">Flowers</emphasis>
with sepals and petals numbering 4 to 5 (seldom 6), not markedly accrescent nor highly reduced; sepals not (or inconsistently) reflexed; petals obovate to oblanceolate (in
<taxonomicName id="4C3E4D16FF8DFFF0FB38FA03AF5E98E2" box="[1182,1314,1438,1461]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="species" species="trigynum">
<emphasis id="B94AEA87FF8DFFF0FB38FA03AF5E98E2" box="[1182,1314,1438,1461]" italics="true" pageId="1" pageNumber="34">H. trigynum</emphasis>
</taxonomicName>
to spatulate, with sepals then ovate), not ciliate (except through pubescence of entire adaxial surface); calyx cup usually more or less funnelform (rarely shortly so, or in
<taxonomicName id="4C3E4D16FF8DFFF0FBD4FA63AE8D9B42" box="[1138,1265,1534,1557]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="species" species="trigynum">
<emphasis id="B94AEA87FF8DFFF0FBD4FA63AE8D9B42" box="[1138,1265,1534,1557]" italics="true" pageId="1" pageNumber="34">H. trigynum</emphasis>
</taxonomicName>
cylindrical with a broad rounded base); stamens inserted just below base of, but not adnate to petals in groups of 3 (aberrantly 4; in
<taxonomicName id="4C3E4D16FF8DFFF0FA9AF9A3AFD29B02" box="[1340,1454,1597,1621]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="species" species="boinense">
<emphasis id="B94AEA87FF8DFFF0FA9AF9A3AFD29B02" box="[1340,1454,1597,1621]" italics="true" pageId="1" pageNumber="34">H. boinense</emphasis>
</taxonomicName>
consistently 5) with two in each fascicle attached at the sides of the neighboring sepal glands; anthers
<quantity id="4CC69B70FF8DFFF0FB46F9E0AF1E9BC2" box="[1248,1378,1661,1685]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="34" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.2-0.5 mm</quantity>
, dorsifixed below midpoint and often somewhat versatile, dehiscing by longitudinal sutures, united by a broad connective so that the thecae are parallel or almost parallel with their adjoining lateral surfaces appressed; styles fused at base.
<emphasis id="B94AEA87FF8DFFF0FA87F963AF1D9A42" box="[1313,1377,1790,1813]" italics="true" pageId="1" pageNumber="34">Fruits</emphasis>
, where known, usually containing 1 to 2 maturing seeds (possibly several-seeded in one species); inner surface of ovary glabrous (sparsely pubescent).
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF8EFFF3FF17FE83ABF79AC2" pageId="2" pageNumber="35" type="discussion">
<paragraph id="8B813695FF8EFFF3FF17FE83A80D9FC2" blockId="2.[139,790,285,1941]" pageId="2" pageNumber="35">
<emphasis id="B94AEA87FF8EFFF3FF17FE83AB789C62" box="[177,260,286,309]" italics="true" pageId="2" pageNumber="35">Notes. </emphasis>
<bibRefCitation id="EFAF4B64FF8EFFF3FEADFE80ABBD9C61" author="Sleumer, H." box="[267,449,285,310]" journalOrPublisher="Bull. Jard. Natl. Belg." pageId="2" pageNumber="35" pagination="239 - 328" part="43" refId="ref29189" refString="Sleumer, H. (1973). Revision du genre Homalium Jacq. (Flacourtiacees) en Afrique (y compris Madagascar et les Mascareignes). Bull. Jard. Natl. Belg. 43: 239 - 328." title="Revision du genre Homalium Jacq. (Flacourtiacees) en Afrique (y compris Madagascar et les Mascareignes)" type="journal article" year="1973">Sleumer (1973)</bibRefCitation>
wrongly corrected the name of sect.
<emphasis id="B94AEA87FF8EFFF3FF64FEA0AB1D9C02" box="[194,353,317,341]" italics="true" pageId="2" pageNumber="35">Eumyriantheia</emphasis>
to
<taxonomicName id="4C3E4D16FF8EFFF3FE20FEA0A8779C02" box="[390,523,317,341]" class="Magnoliopsida" family="Salicaceae" genus="Myriantheia" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FE20FEA0A8779C02" box="[390,523,317,341]" italics="true" pageId="2" pageNumber="35">Myriantheia</emphasis>
</taxonomicName>
and treated it as having been based upon the generic name
<taxonomicName id="4C3E4D16FF8EFFF3FD87FEC0AB7D9CC1" authority="Thouars. Under Art." authorityName="Thouars. Under Art." class="Magnoliopsida" family="Salicaceae" genus="Myriantheia" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FD87FEC0A8D69C22" box="[545,682,349,373]" italics="true" pageId="2" pageNumber="35">Myriantheia</emphasis>
Thouars. Under Art.
</taxonomicName>
21.3 of the ICN (
<bibRefCitation id="EFAF4B64FF8EFFF3FE64FEE0A8D69CC1" author="McNeill, J. &amp; F. R. Barrie &amp; W. R. Buck &amp; V. Demoulin &amp; W. Greuter &amp; D. L. Hawksworth &amp; P. S. Herendeen &amp; S. Knapp &amp; K. Marhold &amp; J. Prado &amp; W. F. Prud'homme van Reine &amp; G. F. Smith &amp; J. H. Wiersema &amp; N. J. Turland" box="[450,682,381,406]" journalOrPublisher="Regnum Veg." pageId="2" pageNumber="35" pagination="154" refId="ref29076" refString="McNeill, J., F. R. Barrie, W. R. Buck, V. Demoulin, W. Greuter, D. L. Hawksworth, P. S. Herendeen, S. Knapp, K. Marhold, J. Prado, W. F. Prud'homme van Reine, G. F. Smith, J. H. Wiersema &amp; N. J. Turland (2012). International Code of Nomenclature for algae, fungi, and plants (Melbourne Code): Adopted by the Eighteenth International Botanical Congress Melbourne, Australia, July 2011. Regnum Veg. 154." title="International Code of Nomenclature for algae, fungi, and plants (Melbourne Code): Adopted by the Eighteenth International Botanical Congress Melbourne, Australia, July 2011" type="journal article" year="2012">McNeill et al., 2012</bibRefCitation>
),
<emphasis id="B94AEA87FF8EFFF3FD1DFEE3AAA79CE2" italics="true" pageId="2" pageNumber="35">Eumyriantheia</emphasis>
could not have been validly published as a section within
<taxonomicName id="4C3E4D16FF8EFFF3FF7FFE20AB279C82" box="[217,347,445,469]" class="Magnoliopsida" family="Salicaceae" genus="Myriantheia" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FF7FFE20AB279C82" box="[217,347,445,469]" italics="true" pageId="2" pageNumber="35">Myriantheia</emphasis>
</taxonomicName>
if that genus had been recognized, but it is legal within
<taxonomicName id="4C3E4D16FF8EFFF3FE8DFE40ABE69CA2" box="[299,410,477,501]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FE8DFE40ABE69CA2" box="[299,410,477,501]" italics="true" pageId="2" pageNumber="35">Homalium</emphasis>
</taxonomicName>
.
<taxonomicName id="4C3E4D16FF8EFFF3FE00FE40A8559CA2" box="[422,553,477,501]" class="Magnoliopsida" family="Salicaceae" genus="Myriantheia" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FE00FE40A8559CA2" box="[422,553,477,501]" italics="true" pageId="2" pageNumber="35">Myriantheia</emphasis>
</taxonomicName>
does not enjoy priority at the sectional level, and
<emphasis id="B94AEA87FF8EFFF3FE63FE60A81D9F42" box="[453,609,509,533]" italics="true" pageId="2" pageNumber="35">Eumyriantheia</emphasis>
could have been published either as a replacement name for it or as the name of a new taxon. However, since Sleumer considered the two to be nomenclatural synonyms, his designation of a
<typeStatus id="54858837FF8EFFF3FD33FDC3A8BF9F22" box="[661,707,606,629]" pageId="2" pageNumber="35">type</typeStatus>
for one of them can be considered applicable to both.
</paragraph>
<paragraph id="8B813695FF8EFFF3FF17FD03A8189922" blockId="2.[139,790,285,1941]" pageId="2" pageNumber="35">
The traditional circumscription of sect.
<emphasis id="B94AEA87FF8EFFF3FDC6FD00A8829FE2" box="[608,766,669,693]" italics="true" pageId="2" pageNumber="35">Eumyriantheia</emphasis>
is excessively broad. It was distinguished from other fasciculatestamened sections largely by its lack of the apomorphic characters that were said to characterize those sections; thus it has been defined by plesiomorphic characters, and encompassed a suspicious degree of variation. South Asian, Malesian and Pacific species (see Sleumer, 1954;
<bibRefCitation id="EFAF4B64FF8EFFF3FDA0FCC3A8EB9E21" author="Lescot, M." box="[518,663,862,886]" editor="AubrEville, A. &amp; J. - F. Leroy" pageId="2" pageNumber="35" pagination="3 - 98" part="11" refId="ref28961" refString="Lescot, M. (1970). Flacourtiaceae. In: AubrEville, A. &amp; J. - F. Leroy (ed.), Fl. Cambodge Laos Vietnam 11: 3 - 98." title="Flacourtiaceae" type="journal article" volumeTitle="Fl. Cambodge Laos Vietnam" year="1970">Lescot, 1970</bibRefCitation>
,
<bibRefCitation id="EFAF4B64FF8EFFF3FD04FCC3A8A49E21" author="Lescot, M." box="[674,728,862,886]" editor="AubrEville, A. &amp; J. - F. Leroy" pageId="2" pageNumber="35" pagination="3 - 134" part="9" refId="ref29000" refString="Lescot, M. (1980). Flacourtiaceae. In: AubrEville, A. &amp; J. - F. Leroy (ed.), Fl. Nouvelle-Caledonie et Dependances 9: 3 - 134." title="Flacourtiaceae" type="book chapter" volumeTitle="Fl. Nouvelle-Caledonie et Dependances" year="1980">1980</bibRefCitation>
) have 7 to 12 (rarely 6) narrowly oblong to narrowly elliptical sepals and petals, which are similar in shape, or the sepals shorter and narrower, and long-ciliate or long-pubescent throughout; the calyx cup is usually subcylindrical and prominently ridged and the anthers very small in proportion to the long slender filaments. Flowers of most of these species closely resemble typical flowers of the widespread sect.
<taxonomicName id="4C3E4D16FF8EFFF3FD86FBA0A8E99902" box="[544,661,1085,1109]" class="Magnoliopsida" family="Salicaceae" genus="Blackwellia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FD86FBA0A8E99902" box="[544,661,1085,1109]" italics="true" pageId="2" pageNumber="35">Blackwellia</emphasis>
</taxonomicName>
, except that sect.
<taxonomicName id="4C3E4D16FF8EFFF3FF19FBC0AB499922" box="[191,309,1117,1141]" class="Magnoliopsida" family="Salicaceae" genus="Blackwellia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FF19FBC0AB499922" box="[191,309,1117,1141]" italics="true" pageId="2" pageNumber="35">Blackwellia</emphasis>
</taxonomicName>
has only 1 stamen per petal.
</paragraph>
<paragraph id="8B813695FF8EFFF3FF17FBE3AB609BC2" blockId="2.[139,790,285,1941]" pageId="2" pageNumber="35">
By contrast, the Malagasy species of sect.
<emphasis id="B94AEA87FF8EFFF3FDDFFBE0A96A99C2" box="[633,790,1149,1173]" italics="true" pageId="2" pageNumber="35">Eumyriantheia</emphasis>
(including the
<typeStatus id="54858837FF8EFFF3FE80FB03AB2F99E2" box="[294,339,1182,1205]" pageId="2" pageNumber="35">type</typeStatus>
of the section) have 4 to 5 (rarely 6) sepals and petals, which are relatively broad and variable in shape (the petals often being obovate) and lack long cilia; the calyx cup is usually funnelform or cup-shaped and the anthers different in shape and usually larger. In overall appearance, these species bear much less resemblance to the fasciculate-stamened Asian and Pacific species than to some solitary-stamened Malagasy species. The distribution of morphological variation in
<taxonomicName id="4C3E4D16FF8EFFF3FDD5FAE0A8A398C2" box="[627,735,1405,1429]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FDD5FAE0A8A398C2" box="[627,735,1405,1429]" italics="true" pageId="2" pageNumber="35">Homalium</emphasis>
</taxonomicName>
(and allied genera such as
<taxonomicName id="4C3E4D16FF8EFFF3FECAFA00A82498E2" authority="Jaub. ex Tul." authorityName="Jaub. ex Tul." box="[364,600,1437,1461]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FECAFA00ABB298E2" box="[364,462,1437,1461]" italics="true" pageId="2" pageNumber="35">Calantica</emphasis>
Jaub. ex Tul.
</taxonomicName>
) makes it evident that some character(s) must have evolved homoplasiously. It is this authors opinion that changes in stamen number are clearly homoplasious, and that neither subg.
<taxonomicName id="4C3E4D16FF8EFFF3FDA2FA60A8129B42" box="[516,622,1533,1557]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FDA2FA60A8129B42" box="[516,622,1533,1557]" italics="true" pageId="2" pageNumber="35">Homalium</emphasis>
</taxonomicName>
nor subg.
<taxonomicName id="4C3E4D16FF8EFFF3FD73FA60AAB49B62" class="Magnoliopsida" family="Salicaceae" genus="Blackwellia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA87FF8EFFF3FD73FA60AAB49B62" italics="true" pageId="2" pageNumber="35">Blackwellia</emphasis>
</taxonomicName>
as traditionally defined are natural groups. The exclusion of the Asian and Pacific species from sect.
<emphasis id="B94AEA87FF8EFFF3FD98F9A0A8A89B02" box="[574,724,1597,1621]" italics="true" pageId="2" pageNumber="35">Eumyriantheia</emphasis>
into a restored sect.
<emphasis id="B94AEA87FF8EFFF3FEB5F9C0ABFA9B22" box="[275,390,1629,1653]" italics="true" pageId="2" pageNumber="35">Polyanthera</emphasis>
will therefore be proposed (Applequist, unpubl. data).
</paragraph>
<paragraph id="8B813695FF8EFFF3FF17F900ABF79AC2" blockId="2.[139,790,285,1941]" pageId="2" pageNumber="35">
Almost no fruiting material is available of sect.
<emphasis id="B94AEA87FF8EFFF3FD1CF903AAA69B82" italics="true" pageId="2" pageNumber="35">Eumyriantheia</emphasis>
, as recircumscribed to comprise only the Malagasy species; fruits with large (to
<quantity id="4CC69B70FF8EFFF3FE7EF943A8609BA1" box="[472,540,1758,1782]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="35" unit="mm" value="2.0">2 mm</quantity>
) developing seeds are very rare even in well-collected species. It appears that fruits dehisce promptly as seeds mature. Most flowering parts, except for anthers, usually persist until the maturing fruit is lost as a unit. Species circumscriptions and keys therefore rely solely on flowering characters.
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF8EFFF3FC9DFE92AE2C9C1E" pageId="2" pageNumber="35" type="multiple">
<paragraph id="8B813695FF8EFFF3FC9DFE92AE2C9C1E" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<heading id="D0C981F9FF8EFFF3FC9DFE92AFF99C7E" bold="true" box="[827,1413,271,297]" fontSize="11" level="1" pageId="2" pageNumber="35" reason="0">
<emphasis id="B94AEA87FF8EFFF3FC9DFE92AE8D9C7E" bold="true" box="[827,1265,271,297]" pageId="2" pageNumber="35">Key to the Malagasy species of</emphasis>
<taxonomicName id="4C3E4D16FF8EFFF3FB5FFE92AFF99C7E" box="[1273,1413,271,297]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="section" section="Eumyriantheia">
<emphasis id="B94AEA87FF8EFFF3FB5FFE92AFF99C7E" bold="true" box="[1273,1413,271,297]" pageId="2" pageNumber="35">Homalium</emphasis>
</taxonomicName>
</heading>
sect. Eumyriantheia
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF8EFFF2FC9DFEC3AFD29B3D" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35" type="key">
<keyStep id="30CA2E30FF8EFFF3FC9DFEC3AFB99C81" pageId="2" pageNumber="35">
<paragraph id="8B813695FF8EFFF3FC9DFEC3AFB99CC2" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DFEC3AFB99CC2" pageId="2" pageNumber="35">
1. Stamens in groups of 5; dry forest,
<collectingRegion id="49FAF877FF8EFFF3FB4EFEC0AF1C9C22" box="[1256,1376,349,373]" country="Madagascar" name="Mahajanga" pageId="2" pageNumber="35">Mahajanga</collectingRegion>
.......................................
<emphasis id="B94AEA87FF8EFFF3FA89FEE0AFB99CC2" bold="true" box="[1327,1477,381,405]" pageId="2" pageNumber="35">
1.
<taxonomicName id="4C3E4D16FF8EFFF3FAEDFEE3AFB99CC2" ID-CoL="3MCHF" box="[1355,1477,381,405]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="boinense">
<emphasis id="B94AEA87FF8EFFF3FAEDFEE3AFB99CC2" bold="true" box="[1355,1477,381,405]" italics="true" pageId="2" pageNumber="35">H. boinense</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B813695FF8EFFF3FC9DFE03AFB99C81" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DFE03AFB99C81" pageId="2" pageNumber="35">1a. Stamens in groups of 3 (aberrantly 4); humid and littoral (very rarely to dry) forests, widespread............. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8EFFF3FC9DFE75AFB99FB7" pageId="2" pageNumber="35">
<paragraph id="8B813695FF8EFFF3FC9DFE75AFB99F37" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DFE75AFB99F37" pageId="2" pageNumber="35">
2. Calyx cup cup-shaped, usually elongated, or cylindrical with a broad rounded base; pedicel 0.5-1 (rarely to 3) mm or absent; petals &lt;
<quantity id="4CC69B70FF8EFFF3FBCDFDB5AEC39F17" box="[1131,1215,552,576]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="35" unit="mm" value="3.5">3.5 mm</quantity>
with acute apices....................................
<emphasis id="B94AEA87FF8EFFF3FAB1FDD5AFB99F37" bold="true" box="[1303,1477,584,608]" pageId="2" pageNumber="35">
15.
<taxonomicName id="4C3E4D16FF8EFFF3FA98FDD5AFB99F37" ID-CoL="3MCT6" box="[1342,1477,584,608]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="trigynum">
<emphasis id="B94AEA87FF8EFFF3FA98FDD5AFB99F37" bold="true" box="[1342,1477,584,608]" italics="true" pageId="2" pageNumber="35">H. trigynum</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B813695FF8EFFF3FC9DFDF5AFB99FB7" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DFDF5AFB99FB7" pageId="2" pageNumber="35">
2a. Calyx cup usually funnelform at anthesis (to cup-shaped in fruit or rarely cylindrical); pedicel usually&gt;
<quantity id="4CC69B70FF8EFFF3FADCFD15AFC09FF7" box="[1402,1468,648,672]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="35" unit="mm" value="1.0">1 mm</quantity>
; petals variable, usually&gt;
<quantity id="4CC69B70FF8EFFF3FBDDFD35AEB39F97" box="[1147,1231,680,704]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="35" unit="mm" value="3.5">3.5 mm</quantity>
(rarely &lt;
<quantity id="4CC69B70FF8EFFF3FA9EFD35AFF09F97" box="[1336,1420,680,704]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="35" unit="mm" value="3.5">3.5 mm</quantity>
then apices rounded).............................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8EFFF3FC9DFD6EAFB99EBD" pageId="2" pageNumber="35">
<paragraph id="8B813695FF8EFFF3FC9DFD6EAFB99E1C" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DFD6EAFB99E1C" pageId="2" pageNumber="35">3. Abaxial surface of both sepals and petals densely (to moderately) pubescent over most of surface, often less dense at apical margins........................ 4</keyLead>
</paragraph>
<paragraph id="8B813695FF8EFFF3FC9DFCCEAFB99EBD" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DFCCEAFB99EBD" pageId="2" pageNumber="35">3a. Abaxial surface of sepals and petals sparsely or minutely pubescent to glabrous, or portions of the petals densely pubescent but the sepals glabrous to sparsely (moderately) pubescent (in one species moderately pubescent, densest at base and along margins)...................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8EFFF3FC9DFC63AFB99902" pageId="2" pageNumber="35">
<paragraph id="8B813695FF8EFFF3FC9DFC63AFB99962" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DFC63AFB99962" pageId="2" pageNumber="35">
4. Upper surface of sepal glands at least partly pubescent..............................
<emphasis id="B94AEA87FF8EFFF3FB54FB80AFB99962" bold="true" box="[1266,1477,1053,1077]" pageId="2" pageNumber="35">
9.
<taxonomicName id="4C3E4D16FF8EFFF3FAABFB83AFB99962" ID-CoL="3MCPT" box="[1293,1477,1053,1077]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="oppositifolium">
<emphasis id="B94AEA87FF8EFFF3FAABFB83AFB99962" bold="true" box="[1293,1477,1053,1077]" italics="true" pageId="2" pageNumber="35">H. oppositifolium</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B813695FF8EFFF3FC9DFBA3AFB99902" blockId="2.[827,1478,271,1963]" box="[827,1477,1085,1110]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DFBA3AFB99902" box="[827,1477,1085,1110]" pageId="2" pageNumber="35">4a. Upper surface of sepal glands glabrous............ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8EFFF3FC9DFBF5AFB99B57" pageId="2" pageNumber="35">
<paragraph id="8B813695FF8EFFF3FC9DFBF5AFB99817" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DFBF5AFB99817" pageId="2" pageNumber="35">
5. Sepals 4 (rarely 5), 3-5.5(-6.5) mm, broadly oblong to broadbased obovate or oblong (rarely broadly elliptical) with rounded apex; petals
<quantity id="4CC69B70FF8EFFF3FBF2FB35AEA89997" box="[1108,1236,1192,1216]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="6.2" metricValueMin="3.6" pageId="2" pageNumber="35" unit="mm" value="4.9" valueMax="6.2" valueMin="3.6">3.6-6.2 mm</quantity>
, normally significantly longer than sepals (aberrantly equal in length); neither petals nor sepals at all keeled; petals very densely pubescent at least near base, with fairly short trichomes...........................................
<emphasis id="B94AEA87FF8EFFF3FABCFAB5AFB99817" bold="true" box="[1306,1477,1320,1344]" pageId="2" pageNumber="35">
6.
<taxonomicName id="4C3E4D16FF8EFFF3FA93FAB5AFB99817" ID-CoL="3MCMV" box="[1333,1477,1320,1344]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="laxiflorum">
<emphasis id="B94AEA87FF8EFFF3FA93FAB5AFB99817" bold="true" box="[1333,1477,1320,1344]" italics="true" pageId="2" pageNumber="35">H. laxiflorum</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B813695FF8EFFF3FC9DFAD5AFB99B57" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DFAD5AFB99B57" pageId="2" pageNumber="35">
5a. Sepals 5 (seldom 4 or
<quantity id="4CC69B70FF8EFFF3FBC3FAD5AEED9837" box="[1125,1169,1352,1376]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="35" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="4C3E4D16FF8EFFF3FB3EFAD4AFFD9837" ID-CoL="3MCHL" box="[1176,1409,1352,1376]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="brevipedunculatum">
<emphasis id="B94AEA87FF8EFFF3FB3EFAD4AFFD9837" box="[1176,1409,1352,1376]" italics="true" pageId="2" pageNumber="35">H. brevipedunculatum</emphasis>
</taxonomicName>
),
<quantity id="4CC69B70FF8EFFF3FA3EFAD5A9B898D7" metricMagnitude="-3" metricUnit="m" metricValue="6.45" metricValueMax="8.4" metricValueMin="4.5" pageId="2" pageNumber="35" unit="mm" value="6.45" valueMax="8.4" valueMin="4.5">4.5- 8.4 mm</quantity>
, oblong-lanceolate or elliptical (to obovate) with acute (to obtuse or rounded) apex; petals
<quantity id="4CC69B70FF8EFFF3FAE8FA15AFBD98F7" box="[1358,1473,1416,1440]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="9.0" metricValueMin="4.5" pageId="2" pageNumber="35" unit="mm" value="6.75" valueMax="9.0" valueMin="4.5">4.5-9 mm</quantity>
, slightly longer than (about equal to) sepals; petals and/ or sepals usually slightly keeled towards apex; petals usually in part densely sericeous with long trichomes.... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8EFFF2FC9DF98EA8829CE2" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
<paragraph id="8B813695FF8EFFF3FC9DF98EAFB99A7D" blockId="2.[827,1478,271,1963]" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF3FC9DF98EAFB99A7D" pageId="2" pageNumber="35">
6. Leaves subopposite to alternate (especially proximally), obovate (narrowly or broadly obovate, elliptical), (3.7-) 5.1-11 × 2.1-5.8(-6.5) cm with petiole 3-12(-17) mm; inflorescences 1.5-4(-5.3) cm; abaxial surface of sepals and petals densely pubescent to sericeous (usually only on petals); sepals often shallowly keeled only at apex, petals usually shallowly keeled; petals oblanceolate to narrowly obovate, 4.5-8.5 ×
<quantity id="4CC69B70FF8EFFF3FB3DF96EAF7B9A5D" box="[1179,1287,1779,1803]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="3.0" metricValueMin="1.8" pageId="2" pageNumber="35" unit="mm" value="2.4" valueMax="3.0" valueMin="1.8">1.8-3 mm</quantity>
......................................
<emphasis id="B94AEA87FF8EFFF3FB1AF88EAFB99A7D" bold="true" box="[1212,1477,1810,1835]" pageId="2" pageNumber="35">
2.
<taxonomicName id="4C3E4D16FF8EFFF3FB70F88EAFB99A7D" ID-CoL="3MCHL" box="[1238,1477,1810,1834]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="brevipedunculatum">
<emphasis id="B94AEA87FF8EFFF3FB70F88EAFB99A7D" bold="true" box="[1238,1477,1810,1834]" italics="true" pageId="2" pageNumber="35">H. brevipedunculatum</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B813695FF8EFFF2FC9DF8AEA8829CE2" blockId="2.[827,1478,271,1963]" lastBlockId="3.[116,767,285,1909]" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
<keyLead id="30CF95A0FF8EFFF2FC9DF8AEA8829CE2" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
6a. Leaves usually alternate (seldom subopposite), elliptical to obovate (broadly obovate), (5.5-)6.5-18 × (3-)
<quantity id="4CC69B70FF8EFFF3FAE2F8CEAFF39A3D" box="[1348,1423,1875,1899]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="2" pageNumber="35" unit="cm" value="6.5" valueMax="9.0" valueMin="4.0">4-9 cm</quantity>
with petiole
<quantity id="4CC69B70FF8EFFF3FC18F8EEAE5E9ADD" box="[958,1058,1907,1931]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.3" metricValueMin="0.7" pageId="2" pageNumber="35" unit="mm" value="15.0" valueMax="23.0" valueMin="7.0">7-23 mm</quantity>
; inflorescences (1.5-)2.3-10(-13.5) cm; abaxial surface of petals and sometimes sepals densely sericeous; sepals usually shallowly keeled for half or more of length, petals usually with a thick streak of very dense indument along central axis but not noticeably keeled; petals obovate to broadly obovate with a narrowed base, 5.5-9 ×
<quantity id="4CC69B70FF8FFFF2FF5EFE00AB239CE2" box="[248,351,413,437]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="5.0" metricValueMin="3.4" pageId="3" pageNumber="36" unit="mm" value="4.2" valueMax="5.0" valueMin="3.4">3.4-5 mm</quantity>
.......................
<emphasis id="B94AEA87FF8FFFF2FD2CFE00A8829CE2" bold="true" box="[650,766,413,437]" pageId="3" pageNumber="36">
8.
<taxonomicName id="4C3E4D16FF8FFFF2FD02FE03A8829CE2" ID-CoL="3MCPJ" box="[676,766,413,437]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="nobile">
<emphasis id="B94AEA87FF8FFFF2FD02FE03A8829CE2" bold="true" box="[676,766,413,437]" italics="true" pageId="3" pageNumber="36">H. nobile</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8FFFF2FFD2FE54A8829FF7" pageId="3" pageNumber="36">
<paragraph id="8B813695FF8FFFF2FFD2FE54A8829F17" blockId="3.[116,767,285,1909]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2FE54A8829F17" pageId="3" pageNumber="36">
7. Sepals
<quantity id="4CC69B70FF8FFFF2FF55FE55AB0F9CB7" box="[243,371,456,480]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.6" metricValueMin="5.5" pageId="3" pageNumber="36" unit="mm" value="6.05" valueMax="6.6" valueMin="5.5">5.5-6.6 mm</quantity>
, petals
<quantity id="4CC69B70FF8FFFF2FE67FE55A8459CB7" box="[449,569,456,480]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="10.5" metricValueMin="9.0" pageId="3" pageNumber="36" unit="mm" value="9.75" valueMax="10.5" valueMin="9.0">9-10.5 mm</quantity>
; abaxial surface of sepals usually with a conspicuous broad pale margin more densely pubescent than remainder of surface....................................
<emphasis id="B94AEA87FF8FFFF2FDB3FDB5A8829F17" bold="true" box="[533,766,552,576]" pageId="3" pageNumber="36">
10.
<taxonomicName id="4C3E4D16FF8FFFF2FD98FDB5A8829F17" ID-CoL="3MCQT" box="[574,766,552,576]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="pseudoboinense">
<emphasis id="B94AEA87FF8FFFF2FD98FDB5A8829F17" bold="true" box="[574,766,552,576]" italics="true" pageId="3" pageNumber="36">H. pseudoboinense</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B813695FF8FFFF2FFD2FDD4A8829FF7" blockId="3.[116,767,285,1909]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2FDD4A8829FF7" pageId="3" pageNumber="36">
7a. Sepals to
<quantity id="4CC69B70FF8FFFF2FEB3FDD5AB179F37" box="[277,363,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="36" unit="mm" value="4.6">4.6 mm</quantity>
, petals to
<quantity id="4CC69B70FF8FFFF2FE7AFDD5A84D9F37" box="[476,561,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="36" unit="mm" value="6.5">6.5 mm</quantity>
; abaxial surface of sepals glabrous to sparsely (moderately, minutely) pubescent throughout, the indument not denser near margins..... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8FFFF2FFD2FD2EA8839E7C" pageId="3" pageNumber="36">
<paragraph id="8B813695FF8FFFF2FFD2FD2EA8829FBC" blockId="3.[116,767,285,1909]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2FD2EA8829FBC" pageId="3" pageNumber="36">8. Upper surface of sepal glands densely (to moderately) pubescent...................................... 9</keyLead>
</paragraph>
<paragraph id="8B813695FF8FFFF2FFD2FD6EA8839E7C" blockId="3.[116,767,285,1909]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2FD6EA8839E7C" pageId="3" pageNumber="36">8a. Upper surface of sepal glands glabrous to glabrate, or sparsely or partially pubescent.................. 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8FFFF2FFD2FCA3A8839961" pageId="3" pageNumber="36">
<paragraph id="8B813695FF8FFFF2FFD2FCA3A8839EE2" blockId="3.[116,767,285,1909]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2FCA3A8839EE2" pageId="3" pageNumber="36">
9. Leaves narrowly obovate to narrowly elliptic or oblongelliptic or obovate; sepals
<quantity id="4CC69B70FF8FFFF2FE1FFCC0A84B9E22" box="[441,567,861,885]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.6" metricValueMin="3.7" pageId="3" pageNumber="36" unit="mm" value="4.15" valueMax="4.6" valueMin="3.7">3.7-4.6 mm</quantity>
; petals moderately longer than sepals; littoral or low-elevation forest.......................................
<emphasis id="B94AEA87FF8FFFF2FD22FC00A8839EE2" bold="true" box="[644,767,925,949]" pageId="3" pageNumber="36">
4.
<taxonomicName id="4C3E4D16FF8FFFF2FD06FC03A8839EE2" ID-CoL="3MCKM" box="[672,767,925,949]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="dorrii">
<emphasis id="B94AEA87FF8FFFF2FD06FC03A8839EE2" bold="true" box="[672,767,925,949]" italics="true" pageId="3" pageNumber="36">H. dorrii</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B813695FF8FFFF2FFD2FC23A8839961" blockId="3.[116,767,285,1909]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2FC23A8839961" pageId="3" pageNumber="36">
9a. Leaves elliptical (to narrowly or broadly elliptical, obovate) or broadly obovate (to obovate, broadly elliptical); sepals &lt;
<quantity id="4CC69B70FF8FFFF2FEF7FC60ABD49942" box="[337,424,1021,1045]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="36" unit="mm" value="3.5">3.5 mm</quantity>
; petals almost twice as long as sepals; mid-elevation humid forest.............. 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8FFFF2FFD2FBD5A8829837" pageId="3" pageNumber="36">
<paragraph id="8B813695FF8FFFF2FFD2FBD5A8829997" blockId="3.[116,767,285,1909]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2FBD5A8829997" pageId="3" pageNumber="36">
10. Leaves elliptical (to narrowly or broadly elliptical, obovate); leaf margins toothed or wavy with multiple glands in marginal projections; inflorescences racemose..............................
<emphasis id="B94AEA87FF8FFFF2FDEAFB35A8829997" bold="true" box="[588,766,1192,1216]" pageId="3" pageNumber="36">
7.
<taxonomicName id="4C3E4D16FF8FFFF2FDC1FB35A8829997" ID-CoL="3MCNH" box="[615,766,1192,1216]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="maringitra">
<emphasis id="B94AEA87FF8FFFF2FDC1FB35A8829997" bold="true" box="[615,766,1192,1216]" italics="true" pageId="3" pageNumber="36">H. maringitra</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B813695FF8FFFF2FFD2FB55A8829837" blockId="3.[116,767,285,1909]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2FB55A8829837" pageId="3" pageNumber="36">
10a. Leaves broadly obovate (to obovate, broadly elliptical); leaf margins revolute, often strongly, to subentire with 0 to 2 basal glands sunk in margins; inflorescences paniculate with well-developed branches......................................
<emphasis id="B94AEA87FF8FFFF2FDABFAD5A8829837" bold="true" box="[525,766,1352,1376]" pageId="3" pageNumber="36">
13.
<taxonomicName id="4C3E4D16FF8FFFF2FD90FAD5A8829837" ID-CoL="3MCR3" box="[566,766,1352,1376]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="ranomafanicum">
<emphasis id="B94AEA87FF8FFFF2FD90FAD5A8829837" bold="true" box="[566,766,1352,1376]" italics="true" pageId="3" pageNumber="36">H. ranomafanicum</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8FFFF2FFD2FAEEA8839B7C" pageId="3" pageNumber="36">
<paragraph id="8B813695FF8FFFF2FFD2FAEEA88398BC" blockId="3.[116,767,285,1909]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2FAEEA88398BC" pageId="3" pageNumber="36">11. Inflorescences all or almost completely racemose, with flowers borne singly or in groups of 1-3 on a ridged rachis (at most, clustered flowers rarely forming a very short branch).................................... 12</keyLead>
</paragraph>
<paragraph id="8B813695FF8FFFF2FFD2FA6EA8839B7C" blockId="3.[116,767,285,1909]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2FA6EA8839B7C" pageId="3" pageNumber="36">11a. Inflorescences paniculate, sometimes spiciform with most flowers borne in clusters on short side branches.... 13</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B813695FF8FFFF2FFD2F9A3AFD39F42" blockId="3.[116,767,285,1909]" lastBlockId="3.[804,1455,285,1643]" pageId="3" pageNumber="36">
<keyStep id="30CA2E30FF8FFFF2FFD2F9A3AFD39F42" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FFD2F9A3AFD39F42" pageId="3" pageNumber="36">
12. Leaves oblanceolate to elliptical or narrowly elliptical, 4-9(-13.7) × 1.5-3.6(-4.4) cm; base cuneate to convex or attenuate; apex cuspidate to rounded (acute); leaf margins with 2-4 small basal glands per side; rachis and pedicels usually glabrous (sparsely pubescent); sepal glands 0.6-1 ×
<quantity id="4CC69B70FF8FFFF2FEF0F940ABA19BA2" box="[342,477,1757,1781]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="3" pageNumber="36" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4-0.5 mm</quantity>
; abaxial surface of petals pubescent with short appressed hairs most densely on central basal portion, less so towards margins, usually with a dark line down the center but not a broad glabrate stripe;
<collectingRegion id="49FAF877FF8FFFF2FF52F8C0AB439A22" box="[244,319,1885,1909]" country="Madagascar" name="Toliara" pageId="3" pageNumber="36">Toliara</collectingRegion>
....................
<emphasis id="B94AEA87FF8FFFF2FDE9F8C0A8829A22" bold="true" box="[591,766,1885,1909]" pageId="3" pageNumber="36">
11.
<taxonomicName id="4C3E4D16FF8FFFF2FDDEF8C3A8829A22" ID-CoL="3MCQV" box="[632,766,1885,1909]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="pulchrum">
<emphasis id="B94AEA87FF8FFFF2FDDEF8C3A8829A22" bold="true" box="[632,766,1885,1909]" italics="true" pageId="3" pageNumber="36">H. pulchrum</emphasis>
</taxonomicName>
</emphasis>
12a. Leaves elliptical to obovate, 6-11.5 ×
<quantity id="4CC69B70FF8FFFF2FB50FE80AF129C62" box="[1270,1390,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="6.2" metricValueMin="2.8" pageId="3" pageNumber="36" unit="cm" value="4.5" valueMax="6.2" valueMin="2.8">2.8-6.2 cm</quantity>
; base convex; apex rounded to rounded-cuspidate, roundedobtuse or emarginate; leaf margins with 1-2 small basal glands per side; distal part of rachis and pedicels minutely pubescent; sepal glands 0.9-1.6 ×
<quantity id="4CC69B70FF8FFFF2FB78FE03AF329CE2" box="[1246,1358,414,438]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="3" pageNumber="36" unit="mm" value="0.85" valueMax="1.0" valueMin="0.7">0.7-1 mm</quantity>
; abaxial surface of petals moderately to densely short appressedpubescent with a conspicuous glabrate stripe along the central axis;
<collectingRegion id="49FAF877FF8FFFF2FC79FE60AE2F9F42" box="[991,1107,509,533]" country="Madagascar" name="Toamasina" pageId="3" pageNumber="36">Toamasina</collectingRegion>
.............
<emphasis id="B94AEA87FF8FFFF2FAB2FE60AFD39F42" bold="true" box="[1300,1455,509,533]" pageId="3" pageNumber="36">
14.
<taxonomicName id="4C3E4D16FF8FFFF2FA9BFE63AFD39F42" ID-CoL="3MCRP" box="[1341,1455,509,533]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="schatzii">
<emphasis id="B94AEA87FF8FFFF2FA9BFE63AFD39F42" bold="true" box="[1341,1455,509,533]" italics="true" pageId="3" pageNumber="36">H. schatzii</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30CA2E30FF8FFFF2FC82FDB5AFD39977" pageId="3" pageNumber="36">
<paragraph id="8B813695FF8FFFF2FC82FDB5AFD29E57" blockId="3.[804,1455,285,1643]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FC82FDB5AFD29E57" pageId="3" pageNumber="36">
13. Leaf margins crenate-serrulate for most of length; panicles borne mostly in pairs along length of older twigs, with an open slender appearance, many having one or more well-developed lower branches sometimes subtended by foliose bracts; petals usually only slightly longer than sepals; sepal glands entirely glabrous; lowelevation forest,
<collectingRegion id="49FAF877FF8FFFF2FBA1FD75AEF69E57" box="[1031,1162,744,768]" country="Madagascar" name="Antsiranana" pageId="3" pageNumber="36">Antsiranana</collectingRegion>
......
<emphasis id="B94AEA87FF8FFFF2FB4EFD75AFD29E57" bold="true" box="[1256,1454,744,768]" pageId="3" pageNumber="36">
5.
<taxonomicName id="4C3E4D16FF8FFFF2FAA5FD75AFD29E57" ID-CoL="3MCLK" box="[1283,1454,744,768]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="graciliflorum">
<emphasis id="B94AEA87FF8FFFF2FAA5FD75AFD29E57" bold="true" box="[1283,1454,744,768]" italics="true" pageId="3" pageNumber="36">H. graciliflorum</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B813695FF8FFFF2FC82FC95AFD39977" blockId="3.[804,1455,285,1643]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FC82FC95AFD39977" pageId="3" pageNumber="36">13a. Leaf margins subentire or revolute (occasionally with few glandular teeth); panicles borne singly except when clustered at twig apices, usually confined to younger distal portions of twigs, usually mostly spiciform with very short branches (in part with few well-developed branches, these then not subtended by foliose bracts); petals normally significantly longer than sepals; sepal glands sometimes in part sparsely pubescent in the depressed central portion.................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA2E30FF8FFFF2FC82FBAEAFD29B3D" pageId="3" pageNumber="36">
<paragraph id="8B813695FF8FFFF2FC82FBAEAFD2983C" blockId="3.[804,1455,285,1643]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FC82FBAEAFD2983C" pageId="3" pageNumber="36">
14. Leaves obovate to elliptical, often broadly (rarely to suborbicular, narrowly elliptical, or oblanceolate), (2.5-) 4-7.5(-10.2) × (1.5-)2.1-4.3(-5.1) cm; base cuneate to convex (slightly attenuate); apex rounded (roundedobtuse, shallowly emarginate or rounded-cuspidate, rarely acute or short-acuminate with a rounded tip); margins revolute to subentire or bearing few shallow teeth, with small glands sometimes present and variably placed; panicles normally all spiciform; sepals and petals 4; highto mid-elevation forest;
<collectingRegion id="49FAF877FF8FFFF2FBF6FACEAEB1983C" box="[1104,1229,1363,1387]" country="Madagascar" name="Antsiranana" pageId="3" pageNumber="36">Antsiranana</collectingRegion>
...
<emphasis id="B94AEA87FF8FFFF2FAAEFACEAFD2983C" bold="true" box="[1288,1454,1363,1387]" pageId="3" pageNumber="36">
3.
<taxonomicName id="4C3E4D16FF8FFFF2FA8FFACEAFD2983C" ID-CoL="3MCHX" box="[1321,1454,1363,1387]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="capuronii">
<emphasis id="B94AEA87FF8FFFF2FA8FFACEAFD2983C" bold="true" box="[1321,1454,1363,1387]" italics="true" pageId="3" pageNumber="36">H. capuronii</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B813695FF8FFFF2FC82FAEEAFD29B3D" blockId="3.[804,1455,285,1643]" pageId="3" pageNumber="36">
<keyLead id="30CF95A0FF8FFFF2FC82FAEEAFD29B3D" pageId="3" pageNumber="36">
14a. Leaves elliptical to obovate or broadly (narrowly) elliptical, 6-12.5 ×
<quantity id="4CC69B70FF8FFFF2FC14FA0EAE6298FD" box="[946,1054,1427,1451]" metricMagnitude="-2" metricUnit="m" metricValue="4.699999999999999" metricValueMax="6.6" metricValueMin="2.8" pageId="3" pageNumber="36" unit="cm" value="4.699999999999999" valueMax="6.6" valueMin="2.8">2.8-6.6 cm</quantity>
; base convex to rounded, at extreme base short-attenuate; apex rounded-acute to cuspidate, rounded, or shallowly emarginate; margins minutely revolute with 0 to 2 basal glands just inside margin; panicles sometimes with short but well-developed lower branches; sepals and petals 4(-5); low-elevation forest;
<collectingRegion id="49FAF877FF8FFFF2FBC8F9AEAEC99B1C" box="[1134,1205,1587,1611]" country="Madagascar" name="Toliara" pageId="3" pageNumber="36">Toliara</collectingRegion>
(Marovony)....................................
<emphasis id="B94AEA87FF8FFFF2FB6BF9CEAFD29B3D" bold="true" box="[1229,1454,1618,1643]" italics="true" pageId="3" pageNumber="36">
12.
<taxonomicName id="4C3E4D16FF8FFFF2FB52F9CEAFD29B3D" ID-CoL="3MCR2" box="[1268,1454,1618,1642]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="randrianasoloi">H. randrianasoloi</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>