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<document id="1335473496EE207932759DDA78C39EDE" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E5EFFD594FD48A6E84EED7A" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) tridens Morawitz 1862" docType="treatment" docVersion="7" lastPageNumber="21" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="20" updateTime="1698562432667" updateUser="plazi">
<mods:mods id="BEC2C189A223F69FB7EA22C6F2547654" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="12409ED5CD86959C4909B18EC374E347">
<mods:title id="1640599C890791851310D0A9D7DF3FED">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="7546EFF4434C9B8FDF827CB521493A46" type="personal">
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</mods:role>
<mods:namePart id="087920EDC57201EE8439945ADB248DD7">Kataev, Boris M.</mods:namePart>
</mods:name>
<mods:name id="0758F3A6C3C24B158DDD62EC0DE2B8C2" type="personal">
<mods:role id="9884B55C1EF7358531873D16395DE512">
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</mods:role>
<mods:namePart id="BD54F85F476CA4DC69189D75BA1C19C4">Liang, Hongbin</mods:namePart>
</mods:name>
<mods:typeOfResource id="25B81E9C3C631BBFF0BF6F488945FE7B">text</mods:typeOfResource>
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<mods:title id="A5EC41A8EE55E3CEA0CFACE5A2FCA566">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DBA7FB9F1E1D61DA150A086AFEAA450B">
<mods:date id="542AE8112ACE6E458CD5961E86AA5C73">2015</mods:date>
<mods:detail id="AB1125D8F099EC206DBAB9D04E41DA2B" type="volume">
<mods:number id="AC8A69E29B873DC02BDB9E9899D79E50">3920</mods:number>
</mods:detail>
<mods:detail id="09A4F3B95872CA4223B9005ECEA481D0" type="issue">
<mods:number id="D1B02BBAF48A4B059CFF18FC8F164038">1</mods:number>
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<mods:classification id="C17A6519352CCACDE524744F3C61C691">journal article</mods:classification>
<mods:identifier id="B9626A4CEE64F73B148BA723C42B6E0D" type="DOI">10.11646/zootaxa.3920.1.1</mods:identifier>
<mods:identifier id="02E172CB96769F1ED07F6694F73685AC" type="GBIF-Dataset">7289c86d-0ed1-4e93-b7b3-bc596042d854</mods:identifier>
<mods:identifier id="E5935C89FB9F5EDD57E70A903AFE0B91" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="039787812E5EFFD594FD48A6E84EED7A" ID-DOI="http://doi.org/10.5281/zenodo.6114268" ID-GBIF-Taxon="127671639" ID-Zenodo-Dep="6114268" LSID="urn:lsid:plazi:treatment:039787812E5EFFD594FD48A6E84EED7A" httpUri="http://treatment.plazi.org/id/039787812E5EFFD594FD48A6E84EED7A" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="C324651C2E5EFFD294FD48A6E800EDD2" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8B8136972E5EFFD294FD48A6EA2EEDBD" blockId="19.[151,777,653,712]" box="[151,777,653,679]" pageId="19" pageNumber="20">
<heading id="D0C981FB2E5EFFD294FD48A6EA2EEDBD" bold="true" box="[151,777,653,679]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="4C3E4D142E5EFFD294FD48A6EA2EEDBD" ID-CoL="6LDV8" authority="Morawitz, 1862" authorityName="Morawitz" authorityYear="1862" box="[151,777,653,679]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tridens" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E5EFFD294FD48A6EA2EEDBD" bold="true" box="[151,777,653,679]" pageId="19" pageNumber="20">
<emphasis id="B94AEA852E5EFFD294FD48A6E82EEDBD" bold="true" box="[151,265,653,679]" italics="true" pageId="19" pageNumber="20">Harpalus</emphasis>
(
<emphasis id="B94AEA852E5EFFD2957048A6E8FDEDBD" bold="true" box="[282,474,653,679]" italics="true" pageId="19" pageNumber="20">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E5EFFD2958148A6EB19EDBD" bold="true" box="[491,574,653,679]" italics="true" pageId="19" pageNumber="20">tridens</emphasis>
Morawitz, 1862
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E5EFFD294FD4884E800EDD2" blockId="19.[151,777,653,712]" box="[151,295,687,712]" pageId="19" pageNumber="20">
(
<figureCitation id="13052A122E5EFFD294F54884E838EDD2" box="[159,287,687,712]" captionStart="FIGURES 40 – 43" captionStartId="20.[151,263,1670,1693]" captionTargetBox="[319,1267,654,1642]" captionTargetId="figure@20.[309,1278,645,1649]" captionTargetPageId="20" captionText="FIGURES 40 – 43. Harpalus, median lobe of aedeagus. 40 – 41, H. tridens (Shaanxi); 42 – 43, H. suensoni (Zhejiang). 40, 42, dorsal view; 41, 43, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287894/files/figure.png" pageId="19" pageNumber="20">Figs 40–41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5EFFD294FD48D3ECB6EC92" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="8B8136972E5EFFD294FD48D3EAA7EC14" blockId="19.[151,1425,760,904]" box="[151,896,760,782]" pageId="19" pageNumber="20">
<taxonomicName id="4C3E4D142E5EFFD294FD48D3EB06EC14" ID-CoL="6LDV8" authority="Morawitz, 1862: 245" authorityName="Morawitz" authorityPageNumber="245" authorityYear="1862" box="[151,545,760,782]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E5EFFD294FD48D3E864EC17" box="[151,323,760,781]" italics="true" pageId="19" pageNumber="20">Harpalus tridens</emphasis>
Morawitz, 1862: 245
</taxonomicName>
.
<typeStatus id="548588352E5EFFD2964148D2EB7AEC14" box="[555,605,761,782]" pageId="19" pageNumber="20">Type</typeStatus>
locality: &quot;Hakodate&quot;,
<collectingCountry id="F32976072E5EFFD2975548D2EA5BEC14" box="[831,892,761,782]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
.
</paragraph>
<paragraph id="8B8136972E5EFFD294FD493CED9EEC51" blockId="19.[151,1425,760,904]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="AB2E11B92E5EFFD294FD493CED9EEC51" pageId="19" pageNumber="20">
<taxonomicName id="4C3E4D142E5EFFD294FD493CEBAAEC36" authority="Pecirkai Jedlicka, 1928: 93" authorityName="Pecirkai Jedlicka" authorityPageNumber="93" authorityYear="1928" box="[151,653,790,812]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subGenus" subGenus="Pardileus">
<emphasis id="B94AEA852E5EFFD294FD493CE9D0EC36" box="[151,247,791,812]" italics="true" pageId="19" pageNumber="20">Harpalus</emphasis>
(
<emphasis id="B94AEA852E5EFFD2956F493CE840EC36" box="[261,359,791,812]" italics="true" pageId="19" pageNumber="20">Pardileus</emphasis>
)
<emphasis id="B94AEA852E5EFFD2951F493CE8ECEC36" box="[373,459,791,812]" italics="true" pageId="19" pageNumber="20">Pecirkai</emphasis>
Jedlička, 1928: 93
</taxonomicName>
.
<typeStatus id="548588352E5EFFD296FD493CEBEEEC36" box="[663,713,791,812]" pageId="19" pageNumber="20">Type</typeStatus>
locality: &quot;Sumpanting&quot; [=? Songpan], Sichuan,
<collectingCountry id="F32976072E5EFFD290D3493DEDDEEC36" box="[1209,1273,790,812]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
.
<taxonomicName id="4C3E4D142E5EFFD294FD491EEA2EEC51" ID-CoL="3JTJY" authority="Schauberger, 1929: 190" authorityName="Schauberger" authorityPageNumber="190" authorityYear="1929" box="[151,777,821,843]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="magnodentatus" subGenus="Pardileus">
<emphasis id="B94AEA852E5EFFD294FD491EE9D0EC50" box="[151,247,821,842]" italics="true" pageId="19" pageNumber="20">Harpalus</emphasis>
(
<emphasis id="B94AEA852E5EFFD2956F491EE840EC50" box="[261,359,821,842]" italics="true" pageId="19" pageNumber="20">Pardileus</emphasis>
)
<emphasis id="B94AEA852E5EFFD2951F491EEB37EC50" box="[373,528,821,842]" italics="true" pageId="19" pageNumber="20">magnodentatus</emphasis>
<treatmentCitation id="0A9F10862E5EFFD2967C491EEA2EEC51" author="Schauberger" box="[534,777,821,843]" page="190" pageId="19" pageNumber="20" year="1929">
<bibRefCitation id="EFAF4B662E5EFFD2967C491EEBF1EC51" author="Schauberger" box="[534,726,821,843]" pageId="19" pageNumber="20" refString="Schauberger, E. (1929) Beitrag zur Kenntnis der palaarktischen Harpalinen (V. Beitrag). Coleopterologisches Centralblatt, 3 (5 - 6), 179 - 196." type="journal article" year="1929">Schauberger, 1929</bibRefCitation>
: 190
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E5EFFD29779491DEA62EC51" box="[787,837,822,843]" pageId="19" pageNumber="20">Type</typeStatus>
locality: &quot;Tsinchow&quot;, Gansu,
<collectingCountry id="F32976072E5EFFD2901F491EED92EC51" box="[1141,1205,821,843]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136972E5EFFD294FD497FECB6EC92" blockId="19.[151,1425,760,904]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="AB2E11B92E5EFFD294FD497FECB6EC92" infered="true" pageId="19" pageNumber="20">
<taxonomicName id="4C3E4D142E5EFFD294FD497FEB6EEC70" ID-CoL="8LG22" authority="Huang, 1993: 451" authorityName="Huang" authorityPageNumber="451" authorityYear="1993" box="[151,585,852,874]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hypogeomysis">
<emphasis id="B94AEA852E5EFFD294FD497FE8ACEC73" box="[151,395,852,873]" italics="true" pageId="19" pageNumber="20">Harpalus hypogeomysis</emphasis>
<treatmentCitation id="0A9F10862E5EFFD295FB497EEB6EEC70" author="Huang" box="[401,585,852,874]" page="451" pageId="19" pageNumber="20" year="1993">
<bibRefCitation id="EFAF4B662E5EFFD295FB497EEB30EC70" author="Huang" box="[401,535,852,874]" pageId="19" pageNumber="20" refString="Huang, T. (1993) Three new species of the genus Harpalus from China (Coleoptera: Carabidae). Acta Zootaxonomica Sinica, 18 (4), 451 - 455." type="journal article" year="1993">Huang, 1993</bibRefCitation>
: 451
</treatmentCitation>
</taxonomicName>
, 454,
<emphasis id="B94AEA852E5EFFD296EC497FEBEBEC73" bold="true" box="[646,716,852,873]" pageId="19" pageNumber="20">syn. n.</emphasis>
<typeStatus id="548588352E5EFFD296B8497EEA23EC70" box="[722,772,853,874]" pageId="19" pageNumber="20">Type</typeStatus>
locality: Beibei,
<geoCoordinate id="EE0A50502E5EFFD297D9497FED21EC70" box="[947,1030,852,874]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="29.85">29°51'N</geoCoordinate>
<geoCoordinate id="EE0A50502E5EFFD29067497FED4BEC70" box="[1037,1132,852,874]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="106.45">106°27'E</geoCoordinate>
, Chongqing,
<collectingCountry id="F32976072E5EFFD2909E497FEC13EC70" box="[1268,1332,852,874]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
.
<taxonomicName id="4C3E4D142E5EFFD294FD4958EB61EC92" ID-CoL="8LG29" authority="Huang" authorityName="Huang" box="[151,582,882,904]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pilosus" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E5EFFD294FD4958E9D0EC92" box="[151,247,883,904]" italics="true" pageId="19" pageNumber="20">Harpalus</emphasis>
(
<emphasis id="B94AEA852E5EFFD2956F4958E882EC92" box="[261,421,883,904]" italics="true" pageId="19" pageNumber="20">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E5EFFD295DE4958E8DBEC92" box="[436,508,883,904]" italics="true" pageId="19" pageNumber="20">pilosus</emphasis>
Huang
</taxonomicName>
in
<bibRefCitation id="EFAF4B662E5EFFD2960C4958EA6AEC92" author="Huang" box="[614,845,882,904]" pageId="19" pageNumber="20" refString="Huang, T. &amp; Zhang, J. (1995) Three new species of the genus Harpalus Latreille (Coleoptera: Carabidae). Entomotaxonomia, 17 (2), 113 - 117." type="journal article" year="1995">Huang &amp; Zhang, 1995</bibRefCitation>
: 113,116,
<emphasis id="B94AEA852E5EFFD297DE4958EADDEC92" bold="true" box="[948,1018,883,904]" pageId="19" pageNumber="20">syn. n.</emphasis>
<typeStatus id="548588352E5EFFD2906A4958ED15EC92" box="[1024,1074,883,904]" pageId="19" pageNumber="20">Type</typeStatus>
locality: Wuyi Mt., Fujian,
<collectingCountry id="F32976072E5EFFD291274959ECAAEC92" box="[1357,1421,882,904]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5EFFD294FD499EEB77EAB8" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8B8136972E5EFFD294FD499EEBBFEB0C" blockId="19.[151,1437,949,2018]" pageId="19" pageNumber="20">
<emphasis id="B94AEA852E5EFFD294FD499EE81AECD4" bold="true" box="[151,317,949,974]" pageId="19" pageNumber="20">
<typeStatus id="548588352E5EFFD294FD499EE9F5ECD4" box="[151,210,949,974]" pageId="19" pageNumber="20">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548588352E5EFFD29521499DE89CECD4" box="[331,443,950,974]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C3E4D142E5EFFD2958A499DEB77ECD7" box="[480,592,949,973]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E5EFFD2958A499DEB77ECD7" box="[480,592,949,973]" italics="true" pageId="19" pageNumber="20">H. tridens</emphasis>
</taxonomicName>
[designated by
<bibRefCitation id="EFAF4B662E5EFFD29768499DEAC6ECD4" author="Kataev" box="[770,993,950,974]" pageId="19" pageNumber="20" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev (1997: 129)</bibRefCitation>
]: ♂, with labels &quot;Hakodate, 132–3&quot; and &quot;132,
<taxonomicName id="4C3E4D142E5EFFD294B649F2E848ECE8" authority="Mor." authorityName="Mor." box="[220,367,985,1010]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E5EFFD294B649F2E80DECEB" box="[220,298,985,1009]" italics="true" pageId="19" pageNumber="20">tridens</emphasis>
Mor.
</taxonomicName>
&quot;; and 4
<typeStatus id="548588352E5EFFD295A449F2EB57ECE8" box="[462,624,985,1010]" pageId="19" pageNumber="20" type="paralectotype">paralectotypes</typeStatus>
(
<specimenCount id="9D38FD1E2E5EFFD296E849F1EB96ECEB" box="[642,689,986,1010]" pageId="19" pageNumber="20" type="generic">2 ♂</specimenCount>
, 2 ♀) with geographical labels as in
<typeStatus id="548588352E5EFFD2903749F2EDE0ECE8" box="[1117,1223,985,1010]" pageId="19" pageNumber="20" type="lectotype">lectotype</typeStatus>
, but with numbers 132–1, 132–2, 132–4, and 132–5 (all in
<collectionCode id="ED2FAE522E5EFFD2963249D5EBACEB0C" box="[600,651,1022,1046]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="20">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E5EFFD294AD4E09EA4BEB98" blockId="19.[151,1437,949,2018]" pageId="19" pageNumber="20">
<typeStatus id="548588352E5EFFD294AD4E09E810EB20" box="[199,311,1058,1082]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C3E4D142E5EFFD2950A4E09E8C7EB23" box="[352,480,1057,1081]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pecirkai">
<emphasis id="B94AEA852E5EFFD2950A4E09E8C7EB23" box="[352,480,1057,1081]" italics="true" pageId="19" pageNumber="20">H. pecirkai</emphasis>
</taxonomicName>
[designated by
<bibRefCitation id="EFAF4B662E5EFFD296FD4E09EA5CEB20" author="Kataev" box="[663,891,1058,1082]" pageId="19" pageNumber="20" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev (1997: 129)</bibRefCitation>
]: ♂, with labels &quot;Szetschwan, Sunpanting, Exp. Stotzner&quot;, &quot;
<typeStatus id="548588352E5EFFD2957C4E6DE872EB44" box="[278,341,1094,1118]" pageId="19" pageNumber="20">Type</typeStatus>
&quot;, and &quot;
<taxonomicName id="4C3E4D142E5EFFD295C84E6EE8D9EB47" box="[418,510,1093,1117]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pecirkai">
<emphasis id="B94AEA852E5EFFD295C84E6EE8D9EB47" box="[418,510,1093,1117]" italics="true" pageId="19" pageNumber="20">pecirkai</emphasis>
</taxonomicName>
mihi, sp. n., det. Ing. Jedlička&quot;, &quot;Mus. Nat. Pragae, Inv. 21337&quot;; and
<typeStatus id="548588352E5EFFD290944E6EECBFEB44" box="[1278,1432,1093,1118]" pageId="19" pageNumber="20" type="paralectotype">paralectotype</typeStatus>
, ♂, with labels as in
<typeStatus id="548588352E5EFFD2951C4E42E8F8EB98" box="[374,479,1129,1154]" pageId="19" pageNumber="20" type="lectotype">lectotype</typeStatus>
, but number 21338 (both in
<collectionCode id="ED2FAE522E5EFFD297744E41EA47EB98" box="[798,864,1130,1154]" httpUri="http://grbio.org/cool/fx6k-enrh" name="Natal Museum" pageId="19" pageNumber="20">NMP</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E5EFFD294AD4EA5ED86EBD0" blockId="19.[151,1437,949,2018]" pageId="19" pageNumber="20">
<typeStatus id="548588352E5EFFD294AD4EA5E813EBBC" box="[199,308,1166,1190]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
of
<taxonomicName id="4C3E4D142E5EFFD295344EA5E8F5EBBF" box="[350,466,1165,1189]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B94AEA852E5EFFD295344EA5E8F5EBBF" box="[350,466,1165,1189]" italics="true" pageId="19" pageNumber="20">H. pilosus</emphasis>
</taxonomicName>
:
<specimenCount id="9D38FD1E2E5EFFD295894EA5EB35EBBF" box="[483,530,1166,1190]" pageId="19" pageNumber="20" type="generic">1 ♂</specimenCount>
, 1 ♀, &quot;Fujian, Wuyishan, Sangang,
<quantity id="4CC69B722E5EFFD297D94EA5EADBEBBF" box="[947,1020,1166,1190]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="19" pageNumber="20" unit="m" value="740.0">740 m</quantity>
/ light trap, 1982.
<date id="FF8010572E5EFFD290BB4EA5EC35EBBC" box="[1233,1298,1166,1190]" pageId="19" pageNumber="20" value="1917-09">IX.17</date>
, Liao Subai collector&quot; [in
<taxonomicName id="4C3E4D142E5EFFD2955B4E9AE8B6EBD0" box="[305,401,1201,1226]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="19" pageNumber="37" rank="genus">Chinese</taxonomicName>
], and &quot;
<taxonomicName id="4C3E4D142E5EFFD295B74E9AEBBAEBD3" box="[477,669,1201,1225]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pilosus" status="sp. nov.">
<emphasis id="B94AEA852E5EFFD295B74E9AEBBAEBD3" box="[477,669,1201,1225]" italics="true" pageId="19" pageNumber="20">Harpalus pilosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27957FE2E5EFFD296CF4E99EBC6EBD3" box="[677,737,1202,1225]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
1993.6.23, det Huang Tongling&quot; (
<collectionCode id="ED2FAE522E5EFFD2900A4E99EDB3EBD0" box="[1120,1172,1202,1226]" pageId="19" pageNumber="20">IOZ</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E5EFFD294AD4EFEEB77EAB8" blockId="19.[151,1437,949,2018]" pageId="19" pageNumber="20">
<emphasis id="B94AEA852E5EFFD294AD4EFEEB1FEBF4" bold="true" box="[199,568,1237,1262]" pageId="19" pageNumber="20">Additional material examined</emphasis>
. More than 1800 specimens (1686 from
<collectingCountry id="F32976072E5EFFD290404EFEED56EBF4" box="[1066,1137,1237,1262]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
), including the following specimens from Hunan and Guizhou.
<collectingCountry id="F32976072E5EFFD2962F4ED2EBA9EA08" box="[581,654,1273,1298]" name="China" pageId="19" pageNumber="20">
<emphasis id="B94AEA852E5EFFD2962F4ED2EBA9EA08" bold="true" box="[581,654,1273,1298]" pageId="19" pageNumber="20">China</emphasis>
</collectingCountry>
.
<collectionCode id="ED2FAE522E5EFFD296F74ED1EA25EA08" box="[669,770,1274,1298]" pageId="19" pageNumber="20">HUNAN</collectionCode>
:
<specimenCount id="9D38FD1E2E5EFFD297784ED1EA18EA0B" box="[786,831,1274,1298]" pageId="19" pageNumber="20" type="generic">1 ♂</specimenCount>
, no additional locality,
<date id="FF8010572E5EFFD290204ED2ED9CEA08" box="[1098,1211,1273,1298]" pageId="19" pageNumber="20" value="1987-09-06">6.IX.1987</date>
(
<collectionCode id="ED2FAE522E5EFFD290A64ED1EC27EA08" box="[1228,1280,1274,1298]" pageId="19" pageNumber="20">IOZ</collectionCode>
); GUIZHOU:
<specimenCount id="9D38FD1E2E5EFFD294FD4F35E808EA2C" box="[151,303,1309,1334]" pageId="19" pageNumber="20" type="generic">42 specimens</specimenCount>
(♂ and ♀), &quot;Kouy-Tcheou [= Guizhou], Reg. de Pin-Fa, Pere Cavalerie, 1908&quot; (
<collectionCode id="ED2FAE522E5EFFD290C54F35EC2CEA2C" LSID="urn:lsid:biocol.org:col:34988" box="[1199,1291,1310,1334]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="20">MNHN</collectionCode>
); 1 ♀,
<quantity id="4CC69B722E5EFFD2913C4F35ECBBEA2C" box="[1366,1436,1309,1334]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="km" value="20.0">20 km</quantity>
NW of Jiangkou, Fanjing Shan—Kuaichang,
<date id="FF8010572E5EFFD296F84F6AEA68EA40" box="[658,847,1345,1370]" pageId="19" pageNumber="20" value="1995-05-27" valueMax="1995-06-03" valueMin="1995-05-27">
27.V.–3.
<bibRefCitation id="EFAF4B662E5EFFD296854F6AEA68EA40" author="VI" box="[751,847,1345,1370]" pageId="19" pageNumber="20" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V., Shilenkov, V. G. (1995) A checklist of the groundbeetles of Russia and adjacent lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia - Moscow, 272 pp." type="book" year="1995">VI.1995</bibRefCitation>
</date>
, E. Jendek &amp; O. Sauša leg. (cITO);
<specimenCount id="9D38FD1E2E5EFFD290B74F69EC2FEA43" box="[1245,1288,1346,1370]" pageId="19" pageNumber="20" type="generic">1 ♂</specimenCount>
, Fanjingshan Mts,
<date id="FF8010572E5EFFD294B24F4EE870EA64" box="[216,343,1381,1406]" pageId="19" pageNumber="20" value="1986-06-25">25.VI.1986</date>
(
<collectionCode id="ED2FAE522E5EFFD295044F4DE885EA64" box="[366,418,1382,1406]" pageId="19" pageNumber="20">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E5EFFD295D64F4DE8D7EA67" box="[444,496,1382,1406]" pageId="19" pageNumber="20" type="generic">1 ♂</specimenCount>
, &quot;Guizhou, Guiyang / 1981.X&quot; [in
<taxonomicName id="4C3E4D142E5EFFD297C94F4EED24EA64" box="[931,1027,1381,1406]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="19" pageNumber="37" rank="genus">Chinese</taxonomicName>
], &quot;
<taxonomicName id="4C3E4D142E5EFFD2904D4F4EECBFEA64" authority="Huang" authorityName="Huang" box="[1063,1432,1381,1406]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hypogeomysis">
<emphasis id="B94AEA852E5EFFD2904D4F4EEC1BEA67" box="[1063,1340,1381,1405]" italics="true" pageId="19" pageNumber="20">Harpalus hypogeomysis</emphasis>
Huang
</taxonomicName>
, 1993.6.23, det Huang Tongling&quot; (
<collectionCode id="ED2FAE522E5EFFD296654FA1EB64EAB8" box="[527,579,1418,1442]" pageId="19" pageNumber="20">IOZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5EFFD294AD4F86E8A7E94F" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8B8136972E5EFFD294AD4F86E8A7E94F" blockId="19.[151,1437,949,2018]" pageId="19" pageNumber="20">
<emphasis id="B94AEA852E5EFFD294AD4F86E81DEADC" bold="true" box="[199,314,1453,1478]" pageId="19" pageNumber="20">Diagnosis</emphasis>
. Well recognizable by the combination of the following characters: pronotum with basal angles sharp at apex; apical spur of protibia markedly dentate at margins (as in Figs 4–6); elytral striae smooth, impunctate; elytral disc in baso-medial portion with distinct transverse microsculpture; all elytral intervals punctate apically; and internal sac of median lobe of aedeagus (
<figureCitation id="13052A122E5EFFD296934C32EA5EE928" box="[761,889,1561,1586]" captionStart="FIGURES 40 – 43" captionStartId="20.[151,263,1670,1693]" captionTargetBox="[319,1267,654,1642]" captionTargetId="figure@20.[309,1278,645,1649]" captionTargetPageId="20" captionText="FIGURES 40 – 43. Harpalus, median lobe of aedeagus. 40 – 41, H. tridens (Shaanxi); 42 – 43, H. suensoni (Zhejiang). 40, 42, dorsal view; 41, 43, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287894/files/figure.png" pageId="19" pageNumber="20">Figs 40–41</figureCitation>
) without any distinct sclerotized elements. Body length 8.8–14.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5EFFD294AD4C4AED0DE86C" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8B8136972E5EFFD294AD4C4AE9D8E810" blockId="19.[151,1437,949,2018]" pageId="19" pageNumber="20">
<emphasis id="B94AEA852E5EFFD294AD4C4AE87EE960" bold="true" box="[199,345,1633,1658]" pageId="19" pageNumber="20">Distribution</emphasis>
.
<collectingCountry id="F32976072E5EFFD295064C4AE894E960" box="[364,435,1633,1658]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
: the north-eastern, eastern and southern parts. The species is known from Liaoning, Hebei, Hubei, Shanxi, Shaanxi, Gansu, Sichuan, Yunnan, Jiangsu, Shanghai, Jiangxi, Zhejiang, Fujian and Guangdong (
<bibRefCitation id="EFAF4B662E5EFFD295424C81E8DBE9D8" author="Kataev" box="[296,508,1705,1730]" pageId="19" pageNumber="20" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E5EFFD295EA4C81E89FE9DB" box="[384,440,1705,1729]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2003
</bibRefCitation>
).
<bibRefCitation id="EFAF4B662E5EFFD296644C82EBCCE9D8" author="Schauberger" box="[526,747,1705,1730]" pageId="19" pageNumber="20" refString="Schauberger, E. (1930) Zur Kenntnis der palaarktischen Harpalinen (VII. Beitrag). Coleopterologisches Centralblatt, 4, 169 - 218." type="journal article" year="1930">Schauberger (1930)</bibRefCitation>
also reported this species from
<collectingCountry id="F32976072E5EFFD290254C82EDEEE9D8" box="[1103,1225,1705,1730]" name="Hong Kong" pageId="19" pageNumber="20">Hongkong</collectingCountry>
. According to
<bibRefCitation id="EFAF4B662E5EFFD291074C81E9C3E9FC" author="Hua" pageId="19" pageNumber="20" refString="Hua, L. (2002) List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, 612 pp." type="book" year="2002">Hua (2002)</bibRefCitation>
, it also occurs in Henan, Anhui and Jiangsu. We provided here the additional material from Hunan and Guizhou.
</paragraph>
<paragraph id="8B8136972E5EFFD294AD4D3EED0DE86C" blockId="19.[151,1437,949,2018]" pageId="19" pageNumber="20">
The species is also distributed in the Russian Far East (the south of Maritime Territory), North and
<collectingCountry id="F32976072E5EFFD291314D3EE9FBE84B" name="South Korea" pageId="19" pageNumber="20">South Korea</collectingCountry>
including Jejudo, and
<collectingCountry id="F32976072E5EFFD295894D11EB04E84B" box="[483,547,1850,1873]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
(Hokkaido, Honshu, Shikoku, and Kyushu Islands) (
<bibRefCitation id="EFAF4B662E5EFFD290EC4D12EC2FE848" author="Habu" box="[1158,1288,1849,1874]" pageId="19" pageNumber="20" refString="Habu, A. (1973) Carabidae: Harpalini (Insecta, Coleoptera). Fauna Japonica. Keigaku Publishing Co., Tokio, xiii + 430 pp." type="book" year="1973">Habu 1973</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5EFFD2917C4D12ECB0E848" author="Lafer" box="[1302,1431,1849,1874]" pageId="19" pageNumber="20" refString="Lafer, G. Sh. (1989) Fam. Carabidae-The ground-beetles. In: Lehr, P. A. (Ed.), Opredelitel' nasekomykh Dal'nego Vostoka SSSR v shesti tomakh. Tom III. Zhestkokrylye, ili zhuki. Chast' 1. Nauka, Leningrad, pp. 70 - 222. [in Russian]" type="journal article" year="1989">Lafer 1989</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5EFFD294FD4D76E89AE86C" author="Kryzhanovskij" box="[151,445,1885,1910]" pageId="19" pageNumber="20" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V., Shilenkov, V. G. (1995) A checklist of the groundbeetles of Russia and adjacent lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia - Moscow, 272 pp." type="book" year="1995">
Kryzhanovskij
<emphasis id="B94AEA852E5EFFD295294D75E853E86F" box="[323,372,1885,1909]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. 1995
</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5EFFD295A24D75EB7DE86C" author="Kataev" box="[456,602,1886,1910]" pageId="19" pageNumber="20" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5EFFD2960C4D75EA11E86C" author="Kataev" box="[614,822,1885,1910]" pageId="19" pageNumber="20" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E5EFFD296D64D75EBCBE86F" box="[700,748,1885,1909]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. 2003
</bibRefCitation>
; Moon &amp;
<bibRefCitation id="EFAF4B662E5EFFD297C24D76ED39E86C" author="Paik" box="[936,1054,1885,1910]" pageId="19" pageNumber="20" refString="Moon, Ch. - S. &amp; Paik, J. - Ch. (2006) Notes on Harpalini (Coleoptera, Carabidae) from Korea (1 - 3). Korean Journal of Soil Zoology, 11 (1 - 2), 30 - 84." type="journal article" year="2006">Paik 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5EFFD294AD4DAAECBAE8F8" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8B8136972E5EFFD294AD4DAAECBAE8F8" blockId="19.[151,1437,949,2018]" pageId="19" pageNumber="20">
<emphasis id="B94AEA852E5EFFD294AD4DAAE813E880" bold="true" box="[199,308,1921,1946]" pageId="19" pageNumber="20">Remarks</emphasis>
. The geographical and individual variation of
<taxonomicName id="4C3E4D142E5EFFD2973E4DA9EAEFE883" box="[852,968,1921,1945]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E5EFFD2973E4DA9EAEFE883" box="[852,968,1921,1945]" italics="true" pageId="19" pageNumber="20">H. tridens</emphasis>
</taxonomicName>
was discussed by
<bibRefCitation id="EFAF4B662E5EFFD290CF4DA9EC6EE880" author="Kataev" box="[1189,1353,1922,1946]" pageId="19" pageNumber="20" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev (1997)</bibRefCitation>
on the basis of the material available at that time. The species demonstrates variation in shape of the pronotal basal angles, elytral punctation and pubescence of abdominal sternites, with most distinctive specimens occurring in Fujian.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E59FFD594FD4AB3E84EED7A" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="8B8136972E59FFD594FD4AB3E84EED7A" blockId="20.[151,1436,151,608]" pageId="20" pageNumber="21">
<taxonomicName id="4C3E4D142E59FFD594FD4AB3E882EFAA" ID-CoL="8LG22" box="[151,421,152,176]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hypogeomysis">
<emphasis id="B94AEA852E59FFD594FD4AB3E882EFAA" box="[151,421,152,176]" italics="true" pageId="20" pageNumber="21">Harpalus hypogeomysis</emphasis>
</taxonomicName>
was described from two specimens (the male
<typeStatus id="548588352E59FFD597C54ABCED36EFAA" box="[943,1041,151,176]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
and the female
<typeStatus id="548588352E59FFD590AC4AB3EC0FEFAA" box="[1222,1320,152,176]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
) collected together in Beibei, Chongqing as similar to
<taxonomicName id="4C3E4D142E59FFD596174A96EBCBEFCE" ID-CoL="6LDV8" box="[637,748,188,212]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E59FFD596174A96EBCBEFCE" box="[637,748,188,212]" italics="true" pageId="20" pageNumber="21">H. tridens</emphasis>
</taxonomicName>
but differing in the presence of sparse punctation on intervals of elytra. Based on the original description and the male from Guizhou determined by Huang Tongling as
<taxonomicName id="4C3E4D142E59FFD591EB4ACAE813EE06" ID-CoL="8LG22" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hypogeomysis">
<emphasis id="B94AEA852E59FFD591EB4ACAE813EE06" italics="true" pageId="20" pageNumber="21">H. hypogeomysis</emphasis>
</taxonomicName>
(see material above), we consider this taxon as conspecific with
<taxonomicName id="4C3E4D142E59FFD590774B2EEDA8EE06" ID-CoL="6LDV8" box="[1053,1167,260,284]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E59FFD590774B2EEDA8EE06" box="[1053,1167,260,284]" italics="true" pageId="20" pageNumber="21">H. tridens</emphasis>
</taxonomicName>
. Although the elytra of this male from Guizhou are impunctate on inner intervals, the sparse micropunctation is occasionally present in specimens of
<taxonomicName id="4C3E4D142E59FFD595544B66E893EE7E" ID-CoL="6LDV8" box="[318,436,332,356]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E59FFD595544B66E893EE7E" box="[318,436,332,356]" italics="true" pageId="20" pageNumber="21">H. tridens</emphasis>
</taxonomicName>
from various localities.
<taxonomicName id="4C3E4D142E59FFD596B54B67EA81EE7E" ID-CoL="8LG29" box="[735,934,332,356]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B94AEA852E59FFD596B54B67EA81EE7E" box="[735,934,332,356]" italics="true" pageId="20" pageNumber="21">Harpalus pilosus</emphasis>
</taxonomicName>
was described from seven specimens, all collected in Wuyi Mountain (Fujian), as similar to
<taxonomicName id="4C3E4D142E59FFD596B64B5AED42EE92" ID-CoL="6LDV8" authority="Huang &amp; Zhang 1995" authorityName="Huang &amp; Zhang" authorityYear="1995" box="[732,1125,367,392]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E59FFD596B64B5AEA6AEE92" box="[732,845,368,392]" italics="true" pageId="20" pageNumber="21">H. tridens</emphasis>
(
<bibRefCitation id="EFAF4B662E59FFD597374B5BED7AEE92" author="Huang" box="[861,1117,367,392]" pageId="20" pageNumber="21" refString="Huang, T. &amp; Zhang, J. (1995) Three new species of the genus Harpalus Latreille (Coleoptera: Carabidae). Entomotaxonomia, 17 (2), 113 - 117." type="journal article" year="1995">Huang &amp; Zhang 1995</bibRefCitation>
)
</taxonomicName>
. The synonymization of
<taxonomicName id="4C3E4D142E59FFD591EB4B5AE9C1EEB6" ID-CoL="8LG29" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B94AEA852E59FFD591EB4B5AE9C1EEB6" italics="true" pageId="20" pageNumber="21">H. pilosus</emphasis>
</taxonomicName>
with
<taxonomicName id="4C3E4D142E59FFD595464BBEE887EEB6" ID-CoL="6LDV8" box="[300,416,404,428]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E59FFD595464BBEE887EEB6" box="[300,416,404,428]" italics="true" pageId="20" pageNumber="21">H. tridens</emphasis>
</taxonomicName>
is based on the examination of the original description and two
<typeStatus id="548588352E59FFD590FE4BBEEDD8EEB7" box="[1172,1279,405,429]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
(see material above). In our opinion, all distinctive characters mentioned in the original description for
<taxonomicName id="4C3E4D142E59FFD590E24B92EDDEEECA" ID-CoL="8LG29" box="[1160,1273,440,464]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B94AEA852E59FFD590E24B92EDDEEECA" box="[1160,1273,440,464]" italics="true" pageId="20" pageNumber="21">H. pilosus</emphasis>
</taxonomicName>
are within the variation of
<taxonomicName id="4C3E4D142E59FFD5954A4BF6E8B7EEEE" ID-CoL="6LDV8" box="[288,400,476,500]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E59FFD5954A4BF6E8B7EEEE" box="[288,400,476,500]" italics="true" pageId="20" pageNumber="21">H. tridens</emphasis>
</taxonomicName>
. Both
<typeStatus id="548588352E59FFD595B14BF6EB60EEEF" box="[475,583,477,501]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
have elytral intervals densely punctate and pubescent apically, and protibial apical spur dentate, though, according to the original description, apical spur of
<taxonomicName id="4C3E4D142E59FFD5904C482AEDBEED02" ID-CoL="8LG29" box="[1062,1177,512,536]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B94AEA852E59FFD5904C482AEDBEED02" box="[1062,1177,512,536]" italics="true" pageId="20" pageNumber="21">H. pilosus</emphasis>
</taxonomicName>
is &quot;obtusely angulated laterally, not dentated&quot;. The examination of the
<typeStatus id="548588352E59FFD596DC480FEA3FED27" box="[694,792,548,573]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3E4D142E59FFD5972A480EEA94ED26" ID-CoL="8LG29" box="[832,947,548,572]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B94AEA852E59FFD5972A480EEA94ED26" box="[832,947,548,572]" italics="true" pageId="20" pageNumber="21">H. pilosus</emphasis>
</taxonomicName>
will be very useful for confirmation of the present synonymy.
</paragraph>
</subSubSection>
</treatment>
</document>
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