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<document id="F2DA1939A21E9751A5BEF9A978545337" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E58FFD794FD48FEE818E831" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) sinicus Hope 1845" docType="treatment" docVersion="7" lastPageNumber="23" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="22" updateTime="1698562432667" updateUser="plazi">
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<mods:title id="4F9A123D6835CECB99B60CD2B9732A5D">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
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<treatment id="039787812E58FFD794FD48FEE818E831" ID-DOI="http://doi.org/10.5281/zenodo.6114274" ID-GBIF-Taxon="127671645" ID-Zenodo-Dep="6114274" LSID="urn:lsid:plazi:treatment:039787812E58FFD794FD48FEE818E831" httpUri="http://treatment.plazi.org/id/039787812E58FFD794FD48FEE818E831" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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<heading id="D0C981FB2E58FFD494FD48FEEBF4EDF5" bold="true" box="[151,723,725,751]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="4C3E4D142E58FFD494FD48FEEBF4EDF5" ID-CoL="6LDVG" authority="Hope, 1845" authorityName="Hope" authorityYear="1845" box="[151,723,725,751]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sinicus" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E58FFD494FD48FEEBF4EDF5" bold="true" box="[151,723,725,751]" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD494FD48FEE82EEDF5" bold="true" box="[151,265,725,751]" italics="true" pageId="21" pageNumber="22">Harpalus</emphasis>
(
<emphasis id="B94AEA852E58FFD4957048FEE8FDEDF5" bold="true" box="[282,474,725,751]" italics="true" pageId="21" pageNumber="22">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E58FFD4958148FEEB67EDF5" bold="true" box="[491,576,725,751]" italics="true" pageId="21" pageNumber="22">sinicus</emphasis>
<bibRefCitation id="EFAF4B662E58FFD4962D48FEEBF4EDF5" author="Hope" box="[583,723,725,751]" pageId="21" pageNumber="22" refString="Hope, F. W. (1845) On the entomology of China, with descriptions of the species sent by Dr. Cantor from Chusan et Canton. The Transactions of the Royal Entomological Society of London, 4, 4 - 17. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1845. tb 01326. x" type="journal article" year="1845">Hope, 1845</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8B8136972E58FFD494FD48DCE878EC0A" blockId="21.[151,723,725,784]" box="[151,351,759,784]" pageId="21" pageNumber="22">(Figs 4–5, 44–45)</paragraph>
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<treatmentCitationGroup id="AB2E11B92E58FFD494FD496BEA11EC4C" box="[151,822,832,854]" pageId="21" pageNumber="22">
<taxonomicName id="4C3E4D142E58FFD494FD496BE8CBEC4C" ID-CoL="8KWCG" authority="Sinicus Hope, 1845: 14" authorityName="Sinicus Hope" authorityPageNumber="14" authorityYear="1845" box="[151,492,832,854]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA852E58FFD494FD496BE861EC4F" box="[151,326,832,853]" italics="true" pageId="21" pageNumber="22">Harpalus Sinicus</emphasis>
<treatmentCitation id="0A9F10862E58FFD49526496AE8CBEC4C" author="Hope" box="[332,492,832,854]" page="14" pageId="21" pageNumber="22" year="1845">
<bibRefCitation id="EFAF4B662E58FFD49526496AE8E1EC4C" author="Hope" box="[332,454,832,854]" pageId="21" pageNumber="22" refString="Hope, F. W. (1845) On the entomology of China, with descriptions of the species sent by Dr. Cantor from Chusan et Canton. The Transactions of the Royal Entomological Society of London, 4, 4 - 17. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1845. tb 01326. x" type="journal article" year="1845">Hope, 1845</bibRefCitation>
: 14
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E58FFD4959C496AEB0FEC4C" box="[502,552,833,854]" pageId="21" pageNumber="22">Type</typeStatus>
locality: &quot;Canton&quot;,
<collectingCountry id="F32976072E58FFD49698496BEA15EC4C" box="[754,818,832,854]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
.
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</paragraph>
<paragraph id="8B8136972E58FFD494FD4974EA51EC6E" blockId="21.[151,1437,832,1068]" box="[151,886,862,884]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB2E11B92E58FFD494FD4974EA51EC6E" box="[151,886,862,884]" pageId="21" pageNumber="22">
<taxonomicName id="4C3E4D142E58FFD494FD4974EBBEEC6E" ID-CoL="4JNPJ" authority="Motschulsky, 1860: 5" authorityName="Motschulsky" authorityPageNumber="5" authorityYear="1860" box="[151,665,862,884]" class="Insecta" family="Carabidae" genus="Platus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="rugicollis" subGenus="Harpalus">
<emphasis id="B94AEA852E58FFD494FD4974E9FFEC6E" box="[151,216,863,884]" italics="true" pageId="21" pageNumber="22">Platus</emphasis>
(
<emphasis id="B94AEA852E58FFD4948C4974E861EC6E" box="[230,326,863,884]" italics="true" pageId="21" pageNumber="22">Harpalus</emphasis>
)
<emphasis id="B94AEA852E58FFD4953E4974E892EC6E" box="[340,437,863,884]" italics="true" pageId="21" pageNumber="22">rugicollis</emphasis>
<treatmentCitation id="0A9F10862E58FFD495D64975EBBEEC6E" author="Motschulsky" box="[444,665,862,884]" page="5" pageId="21" pageNumber="22" year="1860">
<bibRefCitation id="EFAF4B662E58FFD495D64975EBA7EC6E" author="Motschulsky" box="[444,640,862,884]" pageId="21" pageNumber="22" refString="Motschulsky, V. (1860) Insectes du Japon. Etudes Entomologiques, 9, 4 - 39." type="journal article" year="1860">Motschulsky, 1860</bibRefCitation>
: 5
</treatmentCitation>
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.
<typeStatus id="548588352E58FFD496C94974EBF2EC6E" box="[675,725,863,884]" pageId="21" pageNumber="22">Type</typeStatus>
locality:
<collectingCountry id="F32976072E58FFD4975F4974EA55EC6E" box="[821,882,863,884]" name="Japan" pageId="21" pageNumber="22">Japan</collectingCountry>
.
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</paragraph>
<paragraph id="8B8136972E58FFD494FD4956EA8EEC89" blockId="21.[151,1437,832,1068]" box="[151,937,893,915]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB2E11B92E58FFD494FD4956EA8EEC89" box="[151,937,893,915]" pageId="21" pageNumber="22">
<taxonomicName id="4C3E4D142E58FFD494FD4956EB6DEC89" ID-CoL="3JTFT" authority="Morawitz, 1862b: 327" authorityName="Morawitz" authorityPageNumber="327" authorityYear="1862" box="[151,586,893,915]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B94AEA852E58FFD494FD4956E847EC89" box="[151,352,893,915]" italics="true" pageId="21" pageNumber="22">Harpalus japonicus</emphasis>
<treatmentCitation id="0A9F10862E58FFD4950F4956EB6DEC89" author="Morawitz" box="[357,586,893,915]" page="327" pageId="21" pageNumber="22" year="1862">
<bibRefCitation id="EFAF4B662E58FFD4950F4956EB30EC89" author="Morawitz" box="[357,535,893,915]" pageId="21" pageNumber="22" refString="Morawitz, A. (1862 b) Vorlaufige Diagnosen neuer Carabiciden aus Hakodate. Bulletin de l'Academie Imperiale des Sciences de St. - Petersbourg, 5, 321 - 328." type="journal article" year="1862" yearSuffix="b">Morawitz, 1862b</bibRefCitation>
: 327
</treatmentCitation>
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.
<typeStatus id="548588352E58FFD4963E4955EBA1EC89" box="[596,646,894,915]" pageId="21" pageNumber="22">Type</typeStatus>
locality: &quot;Hakodate&quot;,
<collectingCountry id="F32976072E58FFD497024955EA82EC89" box="[872,933,894,915]" name="Japan" pageId="21" pageNumber="22">Japan</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136972E58FFD494FD49B7E8E4ECCA" blockId="21.[151,1437,832,1068]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB2E11B92E58FFD494FD49B7E8E4ECCA" pageId="21" pageNumber="22">
<taxonomicName id="4C3E4D142E58FFD494FD49B7E8E5ECAB" ID-CoL="6LDVG" box="[151,450,924,946]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sinicus" subGenus="Pardileus">
<emphasis id="B94AEA852E58FFD494FD49B7E9D0ECAB" box="[151,247,924,945]" italics="true" pageId="21" pageNumber="22">Harpalus</emphasis>
(
<emphasis id="B94AEA852E58FFD4956349B7E84CECAB" box="[265,363,924,945]" italics="true" pageId="21" pageNumber="22">Pardileus</emphasis>
)
<emphasis id="B94AEA852E58FFD4951649B7E8E5ECAB" box="[380,450,924,945]" italics="true" pageId="21" pageNumber="22">sinicus</emphasis>
</taxonomicName>
<taxonomicName id="4C3E4D142E58FFD495A749B6EAB8ECA8" authority="Schauberger, 1930: 171" authorityName="Schauberger" authorityPageNumber="171" authorityYear="1930" box="[461,927,924,946]" pageId="21" pageNumber="22" rank="variety" variety="kiautschouensis">
var.
<emphasis id="B94AEA852E58FFD4959649B7EBBBECAB" box="[508,668,924,945]" italics="true" pageId="21" pageNumber="22">kiautschouensis</emphasis>
<treatmentCitation id="0A9F10862E58FFD496CF49B7EAB8ECA8" author="Schauberger" box="[677,927,924,946]" page="171" pageId="21" pageNumber="22" year="1930">
<bibRefCitation id="EFAF4B662E58FFD496CF49B7EA4EECA8" author="Schauberger" box="[677,873,924,946]" pageId="21" pageNumber="22" refString="Schauberger, E. (1930) Zur Kenntnis der palaarktischen Harpalinen (VII. Beitrag). Coleopterologisches Centralblatt, 4, 169 - 218." type="journal article" year="1930">Schauberger, 1930</bibRefCitation>
: 171
</treatmentCitation>
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.
<typeStatus id="548588352E58FFD497C649B6EAF9ECA8" box="[940,990,925,946]" pageId="21" pageNumber="22">Type</typeStatus>
locality: &quot;Kiautschou, Schantung&quot; [= Jiao Xian, Shandong],
<collectingCountry id="F32976072E58FFD495154991E898ECCA" box="[383,447,954,976]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
.
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</paragraph>
<paragraph id="8B8136972E58FFD494FD49F2EC5EECF5" blockId="21.[151,1437,832,1068]" box="[151,1401,985,1007]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB2E11B92E58FFD494FD49F2EC5EECF5" box="[151,1401,985,1007]" pageId="21" pageNumber="22">
<taxonomicName id="4C3E4D142E58FFD494FD49F2EA3BECF5" ID-CoL="6LDCS" authority="Schauberger, 1930: 172" authorityName="Schauberger" authorityPageNumber="172" authorityYear="1930" box="[151,796,985,1007]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="sinicus" subGenus="Pardileus" subSpecies="yunnanus">
<emphasis id="B94AEA852E58FFD494FD49F2E9D0ECF4" box="[151,247,985,1006]" italics="true" pageId="21" pageNumber="22">Harpalus</emphasis>
(
<emphasis id="B94AEA852E58FFD4956F49F2E840ECF4" box="[261,359,985,1006]" italics="true" pageId="21" pageNumber="22">Pardileus</emphasis>
)
<emphasis id="B94AEA852E58FFD4951F49F1EB04ECF5" box="[373,547,986,1007]" italics="true" pageId="21" pageNumber="22">sinicus yunnanus</emphasis>
<treatmentCitation id="0A9F10862E58FFD4964349F2EA3BECF5" author="Schauberger" box="[553,796,985,1007]" page="172" pageId="21" pageNumber="22" year="1930">
<bibRefCitation id="EFAF4B662E58FFD4964349F2EBCEECF5" author="Schauberger" box="[553,745,985,1007]" pageId="21" pageNumber="22" refString="Schauberger, E. (1930) Zur Kenntnis der palaarktischen Harpalinen (VII. Beitrag). Coleopterologisches Centralblatt, 4, 169 - 218." type="journal article" year="1930">Schauberger, 1930</bibRefCitation>
: 172
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E58FFD4974F49F1EA70ECF5" box="[805,855,986,1007]" pageId="21" pageNumber="22">Type</typeStatus>
locality: &quot;Yunnan-sen und Mengtze&quot;, Yunnan,
<collectingCountry id="F32976072E58FFD4915F49F2EC52ECF5" box="[1333,1397,985,1007]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136972E58FFD494FD49D3EC44EB14" blockId="21.[151,1437,832,1068]" box="[151,1379,1016,1038]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB2E11B92E58FFD494FD49D3EC44EB14" box="[151,1379,1016,1038]" pageId="21" pageNumber="22">
<taxonomicName id="4C3E4D142E58FFD494FD49D3EBCBEB14" ID-CoL="8LG28" authority="Huang, 1992: 59" authorityName="Huang" authorityPageNumber="59" authorityYear="1992" box="[151,748,1016,1038]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="periglabellus" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E58FFD494FD49D3E9D0EB17" box="[151,247,1016,1037]" italics="true" pageId="21" pageNumber="22">Harpalus</emphasis>
(
<emphasis id="B94AEA852E58FFD4956F49D3E882EB17" box="[261,421,1016,1037]" italics="true" pageId="21" pageNumber="22">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E58FFD495DE49D3EB1DEB17" box="[436,570,1016,1037]" italics="true" pageId="21" pageNumber="22">periglabellus</emphasis>
<treatmentCitation id="0A9F10862E58FFD4962A49D2EBCBEB14" author="Huang" box="[576,748,1016,1038]" page="59" pageId="21" pageNumber="22" year="1992">
<bibRefCitation id="EFAF4B662E58FFD4962A49D2EBE1EB14" author="Huang" box="[576,710,1016,1038]" pageId="21" pageNumber="22" refString="Huang, T. (1992) Four new species of the genus Harpalus Latreille (Coleoptera: Carabidae). Entomotaxonomia, 14 (1), 59 - 65." type="journal article" year="1992">Huang, 1992</bibRefCitation>
: 59
</treatmentCitation>
</taxonomicName>
, 63,
<emphasis id="B94AEA852E58FFD4977649D3EA45EB17" bold="true" box="[796,866,1016,1037]" pageId="21" pageNumber="22">syn. n.</emphasis>
<typeStatus id="548588352E58FFD4970249D2EABDEB14" box="[872,922,1017,1038]" pageId="21" pageNumber="22">Type</typeStatus>
locality: Jinyun Mt.,
<quantity id="4CC69B722E58FFD4901E49D3ED90EB14" box="[1140,1207,1016,1038]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="22" unit="m" value="900.0">900 m</quantity>
, Sichuan,
<collectingCountry id="F32976072E58FFD4917549D3EC78EB14" box="[1311,1375,1016,1038]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
.
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</paragraph>
<paragraph id="8B8136972E58FFD494FD4E3CECBAEB36" blockId="21.[151,1437,832,1068]" box="[151,1437,1046,1068]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB2E11B92E58FFD494FD4E3CECBAEB36" box="[151,1437,1046,1068]" pageId="21" pageNumber="22">
<taxonomicName id="4C3E4D142E58FFD494FD4E3CEA39EB36" ID-CoL="8LG25" authority="Lei &amp; Huang, 1997: 111" authorityName="Lei &amp; Huang" authorityPageNumber="111" authorityYear="1997" box="[151,798,1046,1068]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="longihornus" subGenus="Pseudophonus">
<emphasis id="B94AEA852E58FFD494FD4E3CE9D0EB36" box="[151,247,1047,1068]" italics="true" pageId="21" pageNumber="22">Harpalus</emphasis>
(
<emphasis id="B94AEA852E58FFD4956F4E3CE8BEEB36" box="[261,409,1047,1068]" italics="true" pageId="21" pageNumber="22">Pseudophonus</emphasis>
)
<emphasis id="B94AEA852E58FFD495CD4E3CEB06EB36" box="[423,545,1047,1068]" italics="true" pageId="21" pageNumber="22">longihornus</emphasis>
<treatmentCitation id="0A9F10862E58FFD4964D4E3DEA39EB36" author="Lei" box="[551,798,1046,1068]" page="111" pageId="21" pageNumber="22" year="1997">
<bibRefCitation id="EFAF4B662E58FFD4964D4E3DEBC8EB36" author="Lei" box="[551,751,1046,1068]" pageId="21" pageNumber="22" refString="Lei, H. &amp; Huang, T. (1997) A new species of the genus Harpalus (Coleoptera: Carabidae) from China. Entomotaxonomia, 19 (2), 111 - 113." type="journal article" year="1997">Lei &amp; Huang, 1997</bibRefCitation>
: 111
</treatmentCitation>
</taxonomicName>
, 113,
<emphasis id="B94AEA852E58FFD497304E3CEAB8EB36" bold="true" box="[858,927,1047,1068]" pageId="21" pageNumber="22">syn. n.</emphasis>
<typeStatus id="548588352E58FFD497CF4E3CEAF0EB36" box="[933,983,1047,1068]" pageId="21" pageNumber="22">Type</typeStatus>
locality: Chuxiong,
<quantity id="4CC69B722E58FFD490CD4E3DEDD2EB36" box="[1191,1269,1046,1068]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="21" pageNumber="22" unit="m" value="2400.0">2400 m</quantity>
, Yunnan,
<collectingCountry id="F32976072E58FFD491334E3DECBEEB36" box="[1369,1433,1046,1068]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E58FFD494FD4E72EA32E994" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8B8136972E58FFD494FD4E72E807EB8C" blockId="21.[151,1436,1113,2002]" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD494FD4E72E818EB68" bold="true" box="[151,319,1113,1138]" pageId="21" pageNumber="22">
<typeStatus id="548588352E58FFD494FD4E72E9F5EB68" box="[151,210,1113,1138]" pageId="21" pageNumber="22">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548588352E58FFD495244E71E899EB68" box="[334,446,1114,1138]" pageId="21" pageNumber="22" type="lectotype">Lectotype</typeStatus>
(present designation) of
<taxonomicName id="4C3E4D142E58FFD496B14E71EA4CEB6B" box="[731,875,1114,1137]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B94AEA852E58FFD496B14E71EA4CEB6B" box="[731,875,1114,1137]" italics="true" pageId="21" pageNumber="22">H. japonicus</emphasis>
</taxonomicName>
: ♀, labeled &quot;Hakodate, 133–1.&quot;, &quot;133.
<taxonomicName id="4C3E4D142E58FFD491404E71ECBBEB6B" box="[1322,1436,1114,1137]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B94AEA852E58FFD491404E71ECBBEB6B" box="[1322,1436,1114,1137]" italics="true" pageId="21" pageNumber="22">Japonicus</emphasis>
</taxonomicName>
Mor.&quot; (
<collectionCode id="ED2FAE522E58FFD4948D4E55E83EEB8C" box="[231,281,1150,1174]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="21" pageNumber="22">ZIN</collectionCode>
)
</paragraph>
<paragraph id="8B8136972E58FFD494AD4E8AE847EA3C" blockId="21.[151,1436,1113,2002]" pageId="21" pageNumber="22">
<typeStatus id="548588352E58FFD494AD4E8AE808EBA0" box="[199,303,1185,1210]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C3E4D142E58FFD495324E89EB24EBA3" box="[344,515,1185,1209]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="longihornus">
<emphasis id="B94AEA852E58FFD495324E89EB24EBA3" box="[344,515,1185,1209]" italics="true" pageId="21" pageNumber="22">H. longihornus</emphasis>
</taxonomicName>
: ♂ (teneral), labeled &quot;Yunnan, Chuxiong,
<quantity id="4CC69B722E58FFD497804E89ED64EBA3" box="[1002,1091,1186,1210]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="21" pageNumber="22" unit="m" value="2400.0">2400 m</quantity>
, 1982.
<date id="FF8010572E58FFD490F84E89ED9FEBA0" box="[1170,1208,1186,1210]" pageId="21" pageNumber="22">V.8</date>
, Yu Peiyu leg. &quot; [in
<taxonomicName id="4C3E4D142E58FFD494FD4EEEE9D0EBC4" box="[151,247,1221,1246]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="21" pageNumber="37" rank="genus">Chinese</taxonomicName>
], &quot;
<taxonomicName id="4C3E4D142E58FFD495754EEEEBDFEBC7" box="[287,760,1221,1246]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="longihornus" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E58FFD495754EEEE8AEEBC7" box="[287,393,1221,1245]" italics="true" pageId="21" pageNumber="22">Harpalus</emphasis>
(
<emphasis id="B94AEA852E58FFD495CE4EEEEB72EBC7" box="[420,597,1221,1245]" italics="true" pageId="21" pageNumber="22">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E58FFD4961B4EEEEBDFEBC7" box="[625,760,1221,1245]" italics="true" pageId="21" pageNumber="22">longihornus</emphasis>
</taxonomicName>
&quot; [Huang's handwriting] (
<collectionCode id="ED2FAE522E58FFD4905D4EEDED4CEBC4" box="[1079,1131,1222,1246]" pageId="21" pageNumber="22">IOZ</collectionCode>
).
<typeStatus id="548588352E58FFD490E34EEDEDD7EBC4" box="[1161,1264,1222,1246]" pageId="21" pageNumber="22" type="paratype">Paratype</typeStatus>
: ♂, &quot;Yunnan, Chuxiong,
<quantity id="4CC69B722E58FFD4957F4EC1E84CEA1B" box="[277,363,1258,1282]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="21" pageNumber="22" unit="m" value="2400.0">2400 m</quantity>
/ 1982.
<date id="FF8010572E58FFD495AE4EC1E8CCEA18" box="[452,491,1258,1282]" pageId="21" pageNumber="22">V.8</date>
, Liao Subai leg.&quot; [in
<taxonomicName id="4C3E4D142E58FFD496B54EC2EA18EA18" box="[735,831,1257,1282]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="21" pageNumber="37" rank="genus">Chinese</taxonomicName>
], &quot;
<taxonomicName id="4C3E4D142E58FFD497374EC2EC32EA1B" box="[861,1301,1257,1282]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="longihornus" subGenus="Pseudophonus">
<emphasis id="B94AEA852E58FFD497374EC2EAE0EA1B" box="[861,967,1257,1281]" italics="true" pageId="21" pageNumber="22">Harpalus</emphasis>
(
<emphasis id="B94AEA852E58FFD497B34EC2ED5AEA1B" box="[985,1149,1257,1281]" italics="true" pageId="21" pageNumber="22">Pseudophonus</emphasis>
)
<emphasis id="B94AEA852E58FFD490E44EC2EC32EA1B" box="[1166,1301,1257,1281]" italics="true" pageId="21" pageNumber="22">longihornus</emphasis>
</taxonomicName>
[det Huang Tongling]&quot; (
<collectionCode id="ED2FAE522E58FFD4954A4F25E873EA3C" box="[288,340,1294,1318]" pageId="21" pageNumber="22">IOZ</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E58FFD494AD4F1AEA32E994" blockId="21.[151,1436,1113,2002]" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD494AD4F1AEB10EA50" bold="true" box="[199,567,1329,1354]" pageId="21" pageNumber="22">Additional material examined</emphasis>
. More than 1750 specimens (more than 1610 from
<collectingCountry id="F32976072E58FFD490CF4F1AEDCBEA50" box="[1189,1260,1329,1354]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
), including the following specimens from
<collectingCountry id="F32976072E58FFD495A34F7EEB2BEA74" box="[457,524,1365,1390]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
and
<collectingCountry id="F32976072E58FFD4962C4F7DEBA6EA77" box="[582,641,1366,1389]" name="Laos" pageId="21" pageNumber="22">Laos</collectingCountry>
:
<collectingCountry id="F32976072E58FFD496E74F7EEBF1EA74" box="[653,726,1365,1390]" name="China" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD496E74F7EEBF1EA74" bold="true" box="[653,726,1365,1390]" pageId="21" pageNumber="22">China</emphasis>
</collectingCountry>
. BEIJING:
<specimenCount id="9D38FD1E2E58FFD4970B4F7DEAA8EA77" box="[865,911,1366,1390]" pageId="21" pageNumber="22" type="generic">1 ♂</specimenCount>
, Pinggu Co.,
<date id="FF8010572E58FFD490474F7EED9FEA74" box="[1069,1208,1365,1390]" pageId="21" pageNumber="22" value="1975-07-10">10.VII.1975</date>
, no collector (IZB). YUNNAN: 1 ♀, &quot;Xishuangbanna, Mengzhe,
<quantity id="4CC69B722E58FFD496C24F51EA24EA8B" box="[680,771,1402,1426]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="22" unit="m" value="1200.0">1200 m</quantity>
, 1958.
<date id="FF8010572E58FFD4973F4F52EAB1EA88" box="[853,918,1401,1426]" pageId="21" pageNumber="22" value="1915-06">VI.15</date>
, Pu Fuji Collector&quot; [in
<taxonomicName id="4C3E4D142E58FFD490DB4F52EC36EA88" box="[1201,1297,1401,1426]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="21" pageNumber="37" rank="genus">Chinese</taxonomicName>
], &quot;
<taxonomicName id="4C3E4D142E58FFD491584F52EB76EAAC" authority="Huang" authorityName="Huang" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="periglabellus" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E58FFD491584F52ECBBEA8B" box="[1330,1436,1401,1425]" italics="true" pageId="21" pageNumber="22">Harpalus</emphasis>
(
<emphasis id="B94AEA852E58FFD494CA4FB6E876EAAF" box="[160,337,1437,1461]" italics="true" pageId="21" pageNumber="22">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E58FFD4950F4FB6E8DFEAAF" box="[357,504,1437,1461]" italics="true" pageId="21" pageNumber="22">periglabellus</emphasis>
Huang
</taxonomicName>
, 1993.6.23, det Huang Tongling&quot; (
<collectionCode id="ED2FAE522E58FFD497834FB5ED3AEAAC" box="[1001,1053,1438,1462]" pageId="21" pageNumber="22">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E58FFD4905E4FB5ED43EAAF" box="[1076,1124,1438,1462]" pageId="21" pageNumber="22" type="generic">1 ♂</specimenCount>
, &quot;Xishuangbanna, Mengla,
<quantity id="4CC69B722E58FFD494FD4FEAE811EAC3" box="[151,310,1473,1498]" metricMagnitude="3" metricUnit="m" metricValue="1.065" metricValueMax="1.08" metricValueMin="1.05" pageId="21" pageNumber="22" unit="m" value="1065.0" valueMax="1080.0" valueMin="1050.0">1050–1080 m</quantity>
, 1958.
<date id="FF8010572E58FFD495E94FEAE89BEAC0" box="[387,444,1473,1498]" pageId="21" pageNumber="22" value="1923-10">X.23</date>
, Chen Zhizi collector&quot; [in
<taxonomicName id="4C3E4D142E58FFD496854FEAEA68EAC0" box="[751,847,1473,1498]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="21" pageNumber="37" rank="genus">Chinese</taxonomicName>
], &quot;
<taxonomicName id="4C3E4D142E58FFD497044FEAECB0EAC0" authority="Huang" authorityName="Huang" box="[878,1431,1473,1498]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="periglabellus" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E58FFD497044FEAEAFFEAC3" box="[878,984,1473,1497]" italics="true" pageId="21" pageNumber="22">Harpalus</emphasis>
(
<emphasis id="B94AEA852E58FFD497804FEAEDBCEAC3" box="[1002,1179,1473,1497]" italics="true" pageId="21" pageNumber="22">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E58FFD490C74FEAEC67EAC3" box="[1197,1344,1473,1497]" italics="true" pageId="21" pageNumber="22">periglabellus</emphasis>
Huang
</taxonomicName>
, 1993.6.23, det Huang Tongling&quot; (
<collectionCode id="ED2FAE522E58FFD496654FCDEB64EAE4" box="[527,579,1510,1534]" pageId="21" pageNumber="22">IOZ</collectionCode>
). GUIZHOU:
<specimenCount id="9D38FD1E2E58FFD4968E4FCDEA5BEAE4" box="[740,892,1509,1534]" pageId="21" pageNumber="22" type="generic">14 specimens</specimenCount>
(♂ and ♀), &quot;Kouy-Tcheou [= Guizhou], Reg. de Pin-Fa, Pere Cavalerie, 1908&quot; (
<collectionCode id="ED2FAE522E58FFD496694C21EB78E938" LSID="urn:lsid:biocol.org:col:34988" box="[515,607,1546,1570]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="22">MNHN</collectionCode>
); 1 ♀, Guiyang, Huaxi,
<date id="FF8010572E58FFD497144C22EADBE938" box="[894,1020,1545,1570]" pageId="21" pageNumber="22" value="2007-04-15">15.IV.2007</date>
, Y. Liu leg. (
<collectionCode id="ED2FAE522E58FFD490F64C21EDF7E938" box="[1180,1232,1546,1570]" pageId="21" pageNumber="22">IOZ</collectionCode>
). HAINAN: 1 ♀, Yinggeling,
<quantity id="4CC69B722E58FFD495494C05E84AE95F" box="[291,365,1582,1606]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="21" pageNumber="22" unit="m" value="410.0">410 m</quantity>
,
<date id="FF8010572E58FFD495114C05E8D4E95C" box="[379,499,1582,1606]" pageId="21" pageNumber="22" value="2007-10-18">18.X.2007</date>
, Wang Fang let. (
<collectionCode id="ED2FAE522E58FFD496D44C05EBD5E95C" box="[702,754,1582,1606]" pageId="21" pageNumber="22">IOZ</collectionCode>
).
<collectingCountry id="F32976072E58FFD4976C4C06EA67E95C" box="[774,832,1581,1606]" name="Laos" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD4976C4C06EA67E95C" bold="true" box="[774,832,1581,1606]" pageId="21" pageNumber="22">Laos</emphasis>
</collectingCountry>
.
<specimenCount id="9D38FD1E2E58FFD497254C05EA5BE95F" box="[847,892,1582,1606]" pageId="21" pageNumber="22" type="generic">3 ♂</specimenCount>
, 1 ♀, NE
<collectingCountry id="F32976072E58FFD497994C05ED0BE95F" box="[1011,1068,1582,1605]" name="Laos" pageId="21" pageNumber="22">Laos</collectingCountry>
, Pr. Hua Phan Ban Saleui, Phou Pan (Mt.), ca
<geoCoordinate id="EE0A50502E58FFD4955C4C79E8B5E970" box="[310,402,1618,1642]" direction="north" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="20.2">20°12'N</geoCoordinate>
<geoCoordinate id="EE0A50502E58FFD495F14C79EB22E970" box="[411,517,1618,1642]" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="104.01667">104°01'E</geoCoordinate>
,
<quantity id="4CC69B722E58FFD496794C79EB95E973" box="[531,690,1618,1642]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="21" pageNumber="22" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">1300–1900 m</quantity>
,
<date id="FF8010572E58FFD496D54C7AEAA2E970" box="[703,901,1617,1642]" pageId="21" pageNumber="22" value="2012-04-11" valueMax="2012-05-15" valueMin="2012-04-11">11.IV.–15.V.2012</date>
, C. Holzschuh leg. (
<collectionCode id="ED2FAE522E58FFD4901B4C79ED92E970" box="[1137,1205,1618,1642]" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="21" pageNumber="22">NME</collectionCode>
);
<specimenCount id="9D38FD1E2E58FFD490A04C79EDDFE973" box="[1226,1272,1618,1642]" pageId="21" pageNumber="22" type="generic">4 ♂</specimenCount>
, 9 ♀, same as preceding, but
<date id="FF8010572E58FFD495544C5DE8E9E994" box="[318,462,1654,1678]" pageId="21" pageNumber="22" value="2011-05-01" valueMax="2011-05-31" valueMin="2011-05-01">1–31.V.2011</date>
, local collector (
<collectionCode id="ED2FAE522E58FFD496E24C5DEBEDE994" box="[648,714,1654,1678]" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="21" pageNumber="22">NME</collectionCode>
,
<collectionCode id="ED2FAE522E58FFD496BC4C5DEA2EE994" box="[726,777,1654,1678]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="21" pageNumber="22">ZIN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E58FFD494AD4CB2EC1EE807" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8B8136972E58FFD494AD4CB2EC1EE807" blockId="21.[151,1436,1113,2002]" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD494AD4CB2E81DE9A8" bold="true" box="[199,314,1689,1714]" pageId="21" pageNumber="22">Diagnosis</emphasis>
. Well recognizable by combination of pronotum with basal angles narrowly rounded or blunt at apex, never denticulate, and apical spur of protibia markedly dentate at margins (as in Figs 4–6). Elytral microsculpture is present throughout, consisting of isodiametric meshes. The internal sac of aedeagus (
<figureCitation id="13052A122E58FFD4917E4CCAECB2E9E0" box="[1300,1429,1761,1786]" captionStart="FIGURES 44 – 47" captionStartId="22.[151,263,1178,1201]" captionTargetBox="[229,1355,184,1149]" captionTargetId="figure@22.[222,1365,177,1157]" captionTargetPageId="22" captionText="FIGURES 44 – 47. Harpalus, median lobe of aedeagus. 44 – 45, H. sinicus (Hanoi )); 46 – 47, H. meridianus (India). 44, 46, view from the left side; 45, 47, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287895/files/figure.png" pageId="21" pageNumber="22">Figs 44–45</figureCitation>
) is armed with two or three spiny patches in middle portion of the median lobe. Body length
<quantity id="4CC69B722E58FFD490F74D2EEC12E807" box="[1181,1333,1797,1822]" metricMagnitude="-2" metricUnit="m" metricValue="1.255" metricValueMax="1.56" metricValueMin="0.95" pageId="21" pageNumber="22" unit="mm" value="12.55" valueMax="15.6" valueMin="9.5">9.5–15.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E58FFD794AD4D02EA4EEA61" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8B8136972E58FFD494AD4D02E8CFE8C8" blockId="21.[151,1436,1113,2002]" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD494AD4D02E87EE858" bold="true" box="[199,345,1833,1858]" pageId="21" pageNumber="22">Distribution</emphasis>
.
<collectingCountry id="F32976072E58FFD495024D02E888E858" box="[360,431,1833,1858]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
: widely distributed in the north-eastern, eastern and south-eastern parts. The species was known from Liaoning, Hebei, Shandong, Henan, Shaanxi, Gansu, Sichuan, Hubei, Anhui, Hunan, Jiangsu, Shanghai, Jiangxi, Zhejiang, Fujian, Guangdong,
<collectingCountry id="F32976072E58FFD496BF4D5AEA69E890" box="[725,846,1905,1930]" name="Hong Kong" pageId="21" pageNumber="22">Hongkong</collectingCountry>
, Guangxi, Yunnan and
<collectingCountry id="F32976072E58FFD4900B4D5AED94E890" box="[1121,1203,1905,1930]" name="Taiwan" pageId="21" pageNumber="22">Taiwan</collectingCountry>
(
<bibRefCitation id="EFAF4B662E58FFD490AF4D5AECBFE890" author="Schauberger" box="[1221,1432,1905,1930]" pageId="21" pageNumber="22" refString="Schauberger, E. (1930) Zur Kenntnis der palaarktischen Harpalinen (VII. Beitrag). Coleopterologisches Centralblatt, 4, 169 - 218." type="journal article" year="1930">Schauberger 1930</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E58FFD494FD4DBDE84EE8B4" author="Kataev" box="[151,361,1941,1966]" pageId="21" pageNumber="22" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E58FFD494874DBDE802E8B7" box="[237,293,1941,1965]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2003
</bibRefCitation>
). We examined additionally specimens from Beijing, Guizhou and Hainan.
<bibRefCitation id="EFAF4B662E58FFD490D24DBDEC1FE8B4" author="Hua" box="[1208,1336,1942,1966]" pageId="21" pageNumber="22" refString="Hua, L. (2002) List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, 612 pp." type="book" year="2002">Hua (2002)</bibRefCitation>
reported this species also from Xizang.
</paragraph>
<caption id="DF41661F2E5BFFD794FD4EB1EBE3EBD5" httpUri="https://zenodo.org/record/287895/files/figure.png" pageId="22" pageNumber="23" targetBox="[229,1355,184,1149]" targetPageId="22">
<paragraph id="8B8136972E5BFFD794FD4EB1EBE3EBD5" blockId="22.[151,1436,1178,1231]" pageId="22" pageNumber="23">
<emphasis id="B94AEA852E5BFFD794FD4EB1E874EBAA" bold="true" box="[151,339,1178,1201]" pageId="22" pageNumber="23">FIGURES 44–47.</emphasis>
<taxonomicName id="4C3E4D142E5BFFD795334EB0E89EEBAA" box="[345,441,1179,1200]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA852E5BFFD795334EB0E89EEBAA" box="[345,441,1179,1200]" italics="true" pageId="22" pageNumber="23">Harpalus</emphasis>
</taxonomicName>
, median lobe of aedeagus. 44–45,
<taxonomicName id="4C3E4D142E5BFFD797734EB7EA59EBAA" box="[793,894,1179,1201]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="B94AEA852E5BFFD797734EB7EA59EBAA" box="[793,894,1179,1201]" italics="true" pageId="22" pageNumber="23">H. sinicus</emphasis>
</taxonomicName>
(Hanoi)); 46–47,
<taxonomicName id="4C3E4D142E5BFFD7905C4EB7EC33EBAB" authority="(India)" baseAuthorityName="India" box="[1078,1300,1179,1201]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="meridianus">
<emphasis id="B94AEA852E5BFFD7905C4EB7EDE0EBAA" box="[1078,1223,1179,1201]" italics="true" pageId="22" pageNumber="23">H. meridianus</emphasis>
(India)
</taxonomicName>
. 44, 46, view from the left side; 45, 47, dorsal view. Scale = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B8136972E5BFFD794AD4EDDEA4EEA61" blockId="22.[151,1437,1270,1835]" pageId="22" pageNumber="23">
The species also occurs in the Russian Far East (the south of Maritime Territory), North and
<collectingCountry id="F32976072E5BFFD791664EDDECBCEA15" box="[1292,1435,1270,1295]" name="South Korea" pageId="22" pageNumber="23">South Korea</collectingCountry>
including Jejudo,
<collectingCountry id="F32976072E5BFFD7950E4F30E883EA28" box="[356,420,1307,1330]" name="Japan" pageId="22" pageNumber="23">Japan</collectingCountry>
(Hokkaido, Honshu, Shikoku, Kyushu, and Ryukyu Islands),
<collectingCountry id="F32976072E5BFFD7901B4F30EDC7EA29" box="[1137,1248,1307,1331]" name="Myanmar" pageId="22" pageNumber="23">Myanmar</collectingCountry>
,
<collectingCountry id="F32976072E5BFFD790844F31EC6AEA29" box="[1262,1357,1306,1331]" name="Vietnam" pageId="22" pageNumber="23">Vietnam</collectingCountry>
(
<bibRefCitation id="EFAF4B662E5BFFD791344F31E9F5EA4D" author="Habu" pageId="22" pageNumber="23" refString="Habu, A. (1973) Carabidae: Harpalini (Insecta, Coleoptera). Fauna Japonica. Keigaku Publishing Co., Tokio, xiii + 430 pp." type="book" year="1973">Habu 1973</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5BFFD794B44F15E87AEA4D" author="Lafer" box="[222,349,1342,1367]" pageId="22" pageNumber="23" refString="Lafer, G. Sh. (1989) Fam. Carabidae-The ground-beetles. In: Lehr, P. A. (Ed.), Opredelitel' nasekomykh Dal'nego Vostoka SSSR v shesti tomakh. Tom III. Zhestkokrylye, ili zhuki. Chast' 1. Nauka, Leningrad, pp. 70 - 222. [in Russian]" type="journal article" year="1989">Lafer 1989</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5BFFD795024F15EBA9EA4D" author="Kryzhanovskij" box="[360,654,1342,1367]" pageId="22" pageNumber="23" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V., Shilenkov, V. G. (1995) A checklist of the groundbeetles of Russia and adjacent lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia - Moscow, 272 pp." type="book" year="1995">
Kryzhanovskij
<emphasis id="B94AEA852E5BFFD7967E4F6BEB62EA4D" box="[532,581,1343,1367]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. 1995
</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5BFFD796F04F6BEA0BEA4D" author="Kataev" box="[666,812,1343,1367]" pageId="22" pageNumber="23" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5BFFD7975D4F6BED2EEA4D" author="Kataev" box="[823,1033,1343,1367]" pageId="22" pageNumber="23" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E5BFFD797E44F6BEA98EA4D" box="[910,959,1343,1367]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. 2003
</bibRefCitation>
; Moon &amp;
<bibRefCitation id="EFAF4B662E5BFFD790114F15EDC9EA4D" author="Paik" box="[1147,1262,1342,1367]" pageId="22" pageNumber="23" refString="Moon, Ch. - S. &amp; Paik, J. - Ch. (2006) Notes on Harpalini (Coleoptera, Carabidae) from Korea (1 - 3). Korean Journal of Soil Zoology, 11 (1 - 2), 30 - 84." type="journal article" year="2006">Paik 2006</bibRefCitation>
) and
<collectingCountry id="F32976072E5BFFD791464F6BEC41EA4D" box="[1324,1382,1344,1367]" name="Laos" pageId="22" pageNumber="23">Laos</collectingCountry>
. The species is recorded here from Hainan and
<collectingCountry id="F32976072E5BFFD796074F48EB84EA60" box="[621,675,1379,1402]" name="Laos" pageId="22" pageNumber="23">Laos</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5BFFD794AD4FADE818E831" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8B8136972E5BFFD794AD4FADEAEBE980" blockId="22.[151,1437,1270,1835]" pageId="22" pageNumber="23">
<emphasis id="B94AEA852E5BFFD794AD4FADE813EA85" bold="true" box="[199,308,1414,1439]" pageId="22" pageNumber="23">Remarks</emphasis>
.
<taxonomicName id="4C3E4D142E5BFFD795284FACEB61EA85" box="[322,582,1415,1439]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="periglabellus">
<emphasis id="B94AEA852E5BFFD795284FACEB61EA85" box="[322,582,1415,1439]" italics="true" pageId="22" pageNumber="23">Harpalus periglabellus</emphasis>
</taxonomicName>
was described on the basis of four specimens from Sichuan (Jinyun Mt. and Jianyang) as similar to
<taxonomicName id="4C3E4D142E5BFFD795F24F80EB2FEAD8" box="[408,520,1451,1474]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="B94AEA852E5BFFD795F24F80EB2FEAD8" box="[408,520,1451,1474]" italics="true" pageId="22" pageNumber="23">H. sinicus</emphasis>
</taxonomicName>
but differing in &quot;its hind femur with four setae near hind margin, invisible cilia on sternites 5&quot;.
<taxonomicName id="4C3E4D142E5BFFD7954D4FE4EB06EAFD" box="[295,545,1487,1511]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="longihornus">
<emphasis id="B94AEA852E5BFFD7954D4FE4EB06EAFD" box="[295,545,1487,1511]" italics="true" pageId="22" pageNumber="23">Harpalus longihornus</emphasis>
</taxonomicName>
was described from two males collected in Chuxiong, Yunnan as close to
<taxonomicName id="4C3E4D142E5BFFD791EB4FFBE87BE911" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="disaogashimensis">
<emphasis id="B94AEA852E5BFFD791EB4FFBE87BE911" italics="true" pageId="22" pageNumber="23">H. disaogashimensis</emphasis>
</taxonomicName>
[=
<taxonomicName id="4C3E4D142E5BFFD795EC4FD8EB14E911" box="[390,563,1523,1547]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="hauserianus">
<emphasis id="B94AEA852E5BFFD795EC4FD8EB14E911" box="[390,563,1523,1547]" italics="true" pageId="22" pageNumber="23">H. hauserianus</emphasis>
</taxonomicName>
] but differing in &quot;apical spur with lateral dentate, first tarsal segment of mid leg of male with adhesive hairs, median lobe curved to right side, parameres emarginated laterally near base&quot;. Examination of the
<typeStatus id="548588352E5BFFD7951C4C10E881E949" box="[374,422,1595,1619]" pageId="22" pageNumber="23">type</typeStatus>
series of the latter species and the original description of the former species together with the specimens from Xishuangbanna (Yunnan) determined by Huang Tongling as
<taxonomicName id="4C3E4D142E5BFFD7902E4C4BEDD9E96D" box="[1092,1278,1631,1655]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="periglabellus">
<emphasis id="B94AEA852E5BFFD7902E4C4BEDD9E96D" box="[1092,1278,1631,1655]" italics="true" pageId="22" pageNumber="23">H. periglabellus</emphasis>
</taxonomicName>
revealed that both
<taxonomicName id="4C3E4D142E5BFFD794A54CA8E8A2E981" box="[207,389,1667,1691]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="periglabellus">
<emphasis id="B94AEA852E5BFFD794A54CA8E8A2E981" box="[207,389,1667,1691]" italics="true" pageId="22" pageNumber="23">H. periglabellus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3E4D142E5BFFD795D64CA8EB42E981" box="[444,613,1667,1691]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="longihornus">
<emphasis id="B94AEA852E5BFFD795D64CA8EB42E981" box="[444,613,1667,1691]" italics="true" pageId="22" pageNumber="23">H. longihornus</emphasis>
</taxonomicName>
are conspecific with
<taxonomicName id="4C3E4D142E5BFFD7973F4CA8EAE2E980" box="[853,965,1667,1690]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="B94AEA852E5BFFD7973F4CA8EAE2E980" box="[853,965,1667,1690]" italics="true" pageId="22" pageNumber="23">H. sinicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8136972E5BFFD794AD4C8CE818E831" blockId="22.[151,1437,1270,1835]" pageId="22" pageNumber="23">
<taxonomicName id="4C3E4D142E5BFFD794AD4C8CE8A0E9A5" box="[199,391,1703,1727]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="B94AEA852E5BFFD794AD4C8CE8A0E9A5" box="[199,391,1703,1727]" italics="true" pageId="22" pageNumber="23">Harpalus sinicus</emphasis>
</taxonomicName>
is very similar in external morphology and male genitalia to
<taxonomicName id="4C3E4D142E5BFFD790284C83EC7AE9A5" authority="Andrewes" authorityName="Andrewes" box="[1090,1373,1702,1727]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="meridianus">
<emphasis id="B94AEA852E5BFFD790284C83EDC4E9A5" box="[1090,1251,1703,1727]" italics="true" pageId="22" pageNumber="23">H. meridianus</emphasis>
Andrewes
</taxonomicName>
from
<collectingCountry id="F32976072E5BFFD794FD4CE1E9F6E9F9" box="[151,209,1738,1763]" name="India" pageId="22" pageNumber="23">India</collectingCountry>
and
<collectingCountry id="F32976072E5BFFD795664CE1E849E9F9" box="[268,366,1738,1763]" name="Pakistan" pageId="22" pageNumber="23">Pakistan</collectingCountry>
, which differs from the former species mainly in having nondentate apical spur of protibiae, more elytral intervals punctate and male genitalia (
<figureCitation id="13052A122E5BFFD796B94CC5EA73E81D" box="[723,852,1774,1799]" captionStart="FIGURES 44 – 47" captionStartId="22.[151,263,1178,1201]" captionTargetBox="[229,1355,184,1149]" captionTargetId="figure@22.[222,1365,177,1157]" captionTargetPageId="22" captionText="FIGURES 44 – 47. Harpalus, median lobe of aedeagus. 44 – 45, H. sinicus (Hanoi )); 46 – 47, H. meridianus (India). 44, 46, view from the left side; 45, 47, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287895/files/figure.png" pageId="22" pageNumber="23">Figs 46–47</figureCitation>
) with differently-organized armature of internal sac (
<bibRefCitation id="EFAF4B662E5BFFD794F44D38E815E831" author="Kataev" box="[158,306,1811,1835]" pageId="22" pageNumber="23" refString="Kataev, B. M. (2001) To the knowledge of species of the subgenus Pseudoophonus (genus Harpalus) from the Himalayan region and Southern China (Coleoptera: Carabidae). Zoosystematica Rossica, 9 (2), 2000, 391 - 407." type="journal article" year="2001">Kataev 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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