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<document id="D21E90DB2D09370EC30E6E4E38C6418F" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E51FFDD94FD4BFDE818E9CE" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) indicus subsp. orientalis Kataev 2014" docType="treatment" docVersion="7" lastPageNumber="29" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="29" updateTime="1698562432667" updateUser="plazi">
<mods:mods id="B00D96DE3B0A08649BA59087DE4CAF3D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E70FEEA1C36EC5C1126142FB947CB3CC">
<mods:title id="8C4BCE26E77C6A32ECC5840C7FE86B62">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="2E8D6E0EC7F731CB20657A79345E7766" type="personal">
<mods:role id="FDADD27C886EF10BBD1D0A31135E4996">
<mods:roleTerm id="BF621CCBE73FA8AFA50F3EF5AFF47712">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EF8F5DCC8178C34ADA0D3F8192B60DB3">Kataev, Boris M.</mods:namePart>
</mods:name>
<mods:name id="71B9EE11196305D936B482AE263C1618" type="personal">
<mods:role id="848CDBC493AC70C928C3EE61859DA4F1">
<mods:roleTerm id="1B9690042459CB19697BFE3CBFEADE55">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5EAD67DD521C558FC00E0E58E4991644">Liang, Hongbin</mods:namePart>
</mods:name>
<mods:typeOfResource id="D19D9F470BA53037916C6C89D664C997">text</mods:typeOfResource>
<mods:relatedItem id="921096C207FC508C5FECE0F4731282FA" type="host">
<mods:titleInfo id="2CD42E7B101E831364B1F4EB1818DFC1">
<mods:title id="C6F9AC1B9D4BD74E1BBE5D5EED2ED0A3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BA51629B6CCA013AF30E9462A71FE2B3">
<mods:date id="50631DC3471BAFC94C3FDB87A685D532">2015</mods:date>
<mods:detail id="F9DE25403DC9940FA78C97046F3A00F1" type="volume">
<mods:number id="3BFE855A1066D9121E29300176A11E75">3920</mods:number>
</mods:detail>
<mods:detail id="90972DCA3E57186C495272793020C815" type="issue">
<mods:number id="A09F613C887D6C305285A978105DADBA">1</mods:number>
</mods:detail>
<mods:extent id="9473E2684AE51ACE30BE92B4F924CFFE" unit="page">
<mods:start id="B02C45E424067DDD05C0A0D1210A40CC">1</mods:start>
<mods:end id="15C8D20AD43F310025A9E9BA1326D7CB">39</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="51F460A51D8F4A5E92698CEBD4770128">journal article</mods:classification>
<mods:identifier id="4BBEC830B0CB5A519BDE539895E24D93" type="DOI">10.11646/zootaxa.3920.1.1</mods:identifier>
<mods:identifier id="4A806CD9E2B83453E4A2CB059AC2B27A" type="GBIF-Dataset">7289c86d-0ed1-4e93-b7b3-bc596042d854</mods:identifier>
<mods:identifier id="08BCBBA6F8DC0F4542CD1F7E258E79E1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="35D57C5F5AEC5F90D34B8CC94D02B95C" type="Zenodo-Dep">287886</mods:identifier>
<mods:identifier id="5BC1191BDB18E9CF1D6C4EF895CDC7FE" type="ZooBank">025F9918-F9F6-4049-B7CD-5AC632094F5E</mods:identifier>
</mods:mods>
<treatment id="039787812E51FFDD94FD4BFDE818E9CE" ID-DOI="http://doi.org/10.5281/zenodo.6114292" ID-GBIF-Taxon="127671651" ID-Zenodo-Dep="6114292" LSID="urn:lsid:plazi:treatment:039787812E51FFDD94FD4BFDE818E9CE" httpUri="http://treatment.plazi.org/id/039787812E51FFDD94FD4BFDE818E9CE" lastPageNumber="29" pageId="28" pageNumber="29">
<subSubSection id="C324651C2E51FFDD94FD4BFDE800ED08" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8B8136972E51FFDD94FD4BFDEA42EEEA" blockId="28.[151,869,470,530]" box="[151,869,470,496]" pageId="28" pageNumber="29">
<heading id="D0C981FB2E51FFDD94FD4BFDEA42EEEA" bold="true" box="[151,869,470,496]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<taxonomicName id="4C3E4D142E51FFDD94FD4BFDEA42EEEA" ID-CoL="8MCRH" authority="Kataev, 2014" authorityName="Kataev" authorityYear="2014" box="[151,869,470,496]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="indicus" subGenus="Pseudoophonus" subSpecies="orientalis">
<emphasis id="B94AEA852E51FFDD94FD4BFDEA42EEEA" bold="true" box="[151,869,470,496]" pageId="28" pageNumber="29">
<emphasis id="B94AEA852E51FFDD94FD4BFDE82EEEEA" bold="true" box="[151,265,470,496]" italics="true" pageId="28" pageNumber="29">Harpalus</emphasis>
(
<emphasis id="B94AEA852E51FFDD95704BFDE8FDEEEA" bold="true" box="[282,474,470,496]" italics="true" pageId="28" pageNumber="29">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E51FFDD95814BFDEB9BEEEA" bold="true" box="[491,700,470,496]" italics="true" pageId="28" pageNumber="29">indicus orientalis</emphasis>
<bibRefCitation id="EFAF4B662E51FFDD96A94BFDEA42EEEA" author="Kataev" box="[707,869,470,496]" pageId="28" pageNumber="29" refString="Kataev, B. M. (2014) Systematic and nomenclatorial notes on some taxa of Zabrini and Harpalini from the Palaearctic, Oriental and Australian regions (Coleoptera: Carabidae). Proceedings of the Zoological Institute of the Russian Academy of Sciences, 318 (3), 252 - 267." type="journal article" year="2014">Kataev, 2014</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E51FFDD94FD4BD2E800ED08" blockId="28.[151,869,470,530]" box="[151,295,505,530]" pageId="28" pageNumber="29">
(
<figureCitation id="13052A122E51FFDD94F54BD2E838ED08" box="[159,287,505,530]" captionStart="FIGURES 60 – 67" captionStartId="27.[151,263,1952,1975]" captionTargetBox="[214,1376,206,1918]" captionTargetId="figure@27.[196,1390,193,1932]" captionTargetPageId="27" captionText="FIGURES 60 – 67. Harpalus indicus, median lobe of aedeagus. 60 – 63, H. indicus indicus (Nepal); 64 – 67, H. indicus orientalis (64, Vietnam; 65 – 67, Taiwan). 60, 66, view from the left side; 61, 67, dorsal view; 62 – 65, view from the right side). a, b, folded formations of internal sac. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287898/files/figure.png" pageId="28" pageNumber="29">Figs 64–67</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E51FFDD94FD486AEA03ED4D" box="[151,804,577,599]" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="8B8136972E51FFDD94FD486AEA03ED4D" blockId="28.[151,804,577,599]" box="[151,804,577,599]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="AB2E11B92E51FFDD94FD486AEA03ED4D" box="[151,804,577,599]" pageId="28" pageNumber="29">
<taxonomicName id="4C3E4D142E51FFDD94FD486AEA03ED4D" ID-CoL="8MCRH" authority="Kataev, 2014: 261" authorityName="Kataev" authorityPageNumber="261" authorityYear="2014" box="[151,804,577,599]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="indicus" subGenus="Pseudoophonus" subSpecies="orientalis">
<emphasis id="B94AEA852E51FFDD94FD486AE9D0ED4C" box="[151,247,577,598]" italics="true" pageId="28" pageNumber="29">Harpalus</emphasis>
(
<emphasis id="B94AEA852E51FFDD956F486AE882ED4C" box="[261,421,577,598]" italics="true" pageId="28" pageNumber="29">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E51FFDD95DE486AEB42ED4C" box="[436,613,577,598]" italics="true" pageId="28" pageNumber="29">indicus orientalis</emphasis>
<treatmentCitation id="0A9F10862E51FFDD96014869EA03ED4D" author="Kataev" box="[619,804,577,599]" page="261" pageId="28" pageNumber="29" year="2014">
<bibRefCitation id="EFAF4B662E51FFDD96014869EBD3ED4D" author="Kataev" box="[619,756,577,599]" pageId="28" pageNumber="29" refString="Kataev, B. M. (2014) Systematic and nomenclatorial notes on some taxa of Zabrini and Harpalini from the Palaearctic, Oriental and Australian regions (Coleoptera: Carabidae). Proceedings of the Zoological Institute of the Russian Academy of Sciences, 318 (3), 252 - 267." type="journal article" year="2014">Kataev, 2014</bibRefCitation>
: 261
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E51FFDD94FD48AFEDAAEBA2" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8B8136972E51FFDD94FD48AFEC66EDFE" blockId="28.[151,1437,644,1748]" pageId="28" pageNumber="29">
<emphasis id="B94AEA852E51FFDD94FD48AFE819ED87" bold="true" box="[151,318,644,669]" pageId="28" pageNumber="29">
<typeStatus id="548588352E51FFDD94FD48AFE9F5ED87" box="[151,210,644,669]" pageId="28" pageNumber="29">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548588352E51FFDD952748AFE89DED87" box="[333,442,644,669]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
: ♂, labeled &quot;
<collectingCountry id="F32976072E51FFDD962348AFEB89ED87" box="[585,686,644,669]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
, Lao Cai Prov., env. Sa Pa, Hoang Lien Son Nature Reserve,
<quantity id="4CC69B722E51FFDD910E48AFE988EDA5" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="28" pageNumber="29" unit="m" value="1250.0">1250 m</quantity>
,
<date id="FF8010572E51FFDD94D7488CE87CEDDA" box="[189,347,679,704]" pageId="28" pageNumber="29" value="1998-06-02" valueMax="1998-06-15" valueMin="1998-06-02">2–15.VI.1998</date>
, A. Napolov leg.&quot; (
<collectionCode id="ED2FAE522E51FFDD96554883EB54EDDA" box="[575,627,680,704]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="28" pageNumber="29">ZIN</collectionCode>
).
<typeStatus id="548588352E51FFDD96E34883EBDCEDDA" box="[649,763,680,704]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D38FD1E2E51FFDD976D4883EA97EDDA" box="[775,944,679,704]" pageId="28" pageNumber="29" type="generic">272 specimens</specimenCount>
(males and females) from
<collectingCountry id="F32976072E51FFDD9080488CEC0AEDDA" box="[1258,1325,679,704]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
(Yunnan,
<collectingCountry id="F32976072E51FFDD94FD48E7E9CAEDFF" box="[151,237,716,741]" name="Taiwan" pageId="28" pageNumber="29">Taiwan</collectingCountry>
),
<collectingCountry id="F32976072E51FFDD956A48E7E845EDFF" box="[256,354,716,741]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
,
<collectingCountry id="F32976072E51FFDD950748E6E8FBEDFF" box="[365,476,717,741]" name="Myanmar" pageId="28" pageNumber="29">Myanmar</collectingCountry>
, and
<collectingCountry id="F32976072E51FFDD967C48E6EB6BEDFE" box="[534,588,717,740]" name="Laos" pageId="28" pageNumber="29">Laos</collectingCountry>
(
<collectionCode id="ED2FAE522E51FFDD963648E7EBA8EDFE" box="[604,655,716,740]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="28" pageNumber="29">ZIN</collectionCode>
,
<collectionCode id="ED2FAE522E51FFDD96F348E7EBECEDFE" box="[665,715,716,740]" pageId="28" pageNumber="29">IOZ</collectionCode>
, cWR,
<collectionCode id="ED2FAE522E51FFDD977448E6EA78EDFE" box="[798,863,717,740]" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="28" pageNumber="29">NME</collectionCode>
,
<collectionCode id="ED2FAE522E51FFDD970048E6EA8DEDFE" box="[874,938,717,740]" pageId="28" pageNumber="29">MPU</collectionCode>
,
<collectionCode id="ED2FAE522E51FFDD97DF48E6ED2BEDFE" box="[949,1036,717,740]" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="28" pageNumber="29">NHMB</collectionCode>
,
<collectionCode id="ED2FAE522E51FFDD907D48E6ED71EDFE" box="[1047,1110,717,740]" pageId="28" pageNumber="29">TMB</collectionCode>
,
<collectionCode id="ED2FAE522E51FFDD900B48E6ED83EDFE" box="[1121,1188,717,740]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="28" pageNumber="29">NRM</collectionCode>
,
<collectionCode id="ED2FAE522E51FFDD90C548E7EDCAEDFE" box="[1199,1261,716,740]" pageId="28" pageNumber="29">SIEE</collectionCode>
, cSCI).
</paragraph>
<paragraph id="8B8136972E51FFDD94AD48C4EDAAEBA2" blockId="28.[151,1437,644,1748]" pageId="28" pageNumber="29">
<emphasis id="B94AEA852E51FFDD94AD48C4EB0AEC12" bold="true" box="[199,557,751,776]" pageId="28" pageNumber="29">Additional material examined</emphasis>
.
<collectingCountry id="F32976072E51FFDD965648C4EBA2EC12" box="[572,645,751,776]" name="China" pageId="28" pageNumber="29">
<emphasis id="B94AEA852E51FFDD965648C4EBA2EC12" bold="true" box="[572,645,751,776]" pageId="28" pageNumber="29">China</emphasis>
</collectingCountry>
. XIZANG:
<specimenCount id="9D38FD1E2E51FFDD976448DBEA1DEC1D" box="[782,826,752,776]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, Bomi,
<geoCoordinate id="EE0A50502E51FFDD97F948C4EAC8EC12" box="[915,1007,751,776]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="29.883333">29°53'N</geoCoordinate>
<geoCoordinate id="EE0A50502E51FFDD979D48C4ED74EC12" box="[1015,1107,751,776]" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="95.75">95°45'E</geoCoordinate>
,
<date id="FF8010572E51FFDD903548DBEDD3EC12" box="[1119,1268,752,776]" pageId="28" pageNumber="29" value="2003-08-30">30.VIII.2003</date>
, Ren Guodong leg.(
<collectionCode id="ED2FAE522E51FFDD94A2493FE9DAEC37" box="[200,253,788,813]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD9578493FE818EC36" box="[274,319,788,812]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, 3 ♀, Bomi,
<quantity id="4CC69B722E51FFDD95BC493FEB08EC36" box="[470,559,788,812]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="28" pageNumber="29" unit="m" value="2300.0">2300 m</quantity>
,
<date id="FF8010572E51FFDD9657493FEBD5EC37" box="[573,754,788,813]" pageId="28" pageNumber="29" value="1983-07-25" valueMax="1983-07-26" valueMin="1983-07-25">25–26.VII.1983</date>
, Han Yinheng leg. (
<collectionCode id="ED2FAE522E51FFDD97B2493FED2BEC36" box="[984,1036,788,812]" pageId="28" pageNumber="29">IOZ</collectionCode>
); 1 ♀, same data, but
<date id="FF8010572E51FFDD9160493FECBBEC36" box="[1290,1436,788,812]" pageId="28" pageNumber="29" value="1983-08-14">14.VIII.1983</date>
(
<collectionCode id="ED2FAE522E51FFDD94F54913E9F4EC4A" box="[159,211,824,848]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD948D4913E833EC55" box="[231,276,824,848]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, Medog,
<quantity id="4CC69B722E51FFDD95154913E8F1EC55" box="[383,470,824,848]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="m" value="2000.0">2000 m</quantity>
,
<date id="FF8010572E51FFDD95894913EB69EC4A" box="[483,590,824,848]" pageId="28" pageNumber="29" value="1982-10-07">7.X.1982</date>
, Han Yinheng leg. (
<collectionCode id="ED2FAE522E51FFDD975A4913EA43EC4A" box="[816,868,824,848]" pageId="28" pageNumber="29">IOZ</collectionCode>
); 1 ♀, same data, but
<quantity id="4CC69B722E51FFDD9031491CED94EC55" box="[1115,1203,823,848]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="28" pageNumber="29" unit="m" value="1600.0">1600 m</quantity>
,
<date id="FF8010572E51FFDD90AA4913EC11EC4A" box="[1216,1334,824,848]" pageId="28" pageNumber="29" value="1982-10-28">28.X.1982</date>
(
<collectionCode id="ED2FAE522E51FFDD912C4913EC5DEC4A" box="[1350,1402,824,848]" pageId="28" pageNumber="29">IOZ</collectionCode>
); 1 ♀, same data, but
<quantity id="4CC69B722E51FFDD95054977E89BEC6E" box="[367,444,860,885]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="28" pageNumber="29" unit="m" value="850.0">850 m</quantity>
,
<date id="FF8010572E51FFDD95A14977EB6CEC6E" box="[459,587,860,884]" pageId="28" pageNumber="29" value="1983-03-11">11.III.1983</date>
, (
<collectionCode id="ED2FAE522E51FFDD96084977EBB1EC6E" box="[610,662,860,884]" pageId="28" pageNumber="29">IOZ</collectionCode>
); 1 ♀, same data but
<quantity id="4CC69B722E51FFDD97FD4977ED3BEC6E" box="[919,1052,860,885]" metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="28" pageNumber="29" unit="m" value="875.0" valueMax="950.0" valueMin="800.0">800–950 m</quantity>
,
<date id="FF8010572E51FFDD90414977ED8EEC6E" box="[1067,1193,860,884]" pageId="28" pageNumber="29" value="1983-03-20">20.III.1983</date>
(
<collectionCode id="ED2FAE522E51FFDD90D14977EDC8EC6E" box="[1211,1263,860,884]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD916D4977EC11EC6E" box="[1287,1334,860,884]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, Medog,
<quantity id="4CC69B722E51FFDD94FD4954E812EC8D" box="[151,309,895,920]" metricMagnitude="3" metricUnit="m" metricValue="1.985" metricValueMax="2.05" metricValueMin="1.92" pageId="28" pageNumber="29" unit="m" value="1985.0" valueMax="2050.0" valueMin="1920.0">1920–2050 m</quantity>
,
<date id="FF8010572E51FFDD95284954E88AEC82" box="[322,429,895,920]" pageId="28" pageNumber="29" value="1982-10-06">6.X.1982</date>
, Lin Zai leg. (
<collectionCode id="ED2FAE522E51FFDD962549ABEBA4EC82" box="[591,643,896,920]" pageId="28" pageNumber="29">IOZ</collectionCode>
). GUANGXI:
<specimenCount id="9D38FD1E2E51FFDD974049ABEA70EC8D" box="[810,855,896,920]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, Jinxiu,
<quantity id="4CC69B722E51FFDD97DD4954ED26EC8D" box="[951,1025,895,920]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="28" pageNumber="29" unit="m" value="550.0">550 m</quantity>
,
<date id="FF8010572E51FFDD906449ABEDA3EC82" box="[1038,1156,896,920]" pageId="28" pageNumber="29" value="1999-05-12">12.V.1999</date>
, Liu Dajun leg. (
<collectionCode id="ED2FAE522E51FFDD912F49ABEC5EEC82" box="[1349,1401,896,920]" pageId="28" pageNumber="29">IOZ</collectionCode>
); 1 ♀, Jinxiu,
<quantity id="4CC69B722E51FFDD9579498FE847ECA6" box="[275,352,932,957]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="28" pageNumber="29" unit="m" value="450.0">450 m</quantity>
,
<date id="FF8010572E51FFDD9505498FE8D6ECA6" box="[367,497,932,956]" pageId="28" pageNumber="29" value="2000-06-30">30.VI.2000</date>
, Yao Jian leg. (
<collectionCode id="ED2FAE522E51FFDD96C0498FEBF9ECA6" box="[682,734,932,956]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD969C498FEA02ECA6" box="[758,805,932,956]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, same locality,
<quantity id="4CC69B722E51FFDD97B6498FED0EECA6" box="[988,1065,932,956]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="29" unit="m" value="400.0">400 m</quantity>
,
<date id="FF8010572E51FFDD9052498FED89ECA7" box="[1080,1198,932,957]" pageId="28" pageNumber="29" value="2000-05-16">16.V.2000</date>
, Gao Mingyuan leg. (
<collectionCode id="ED2FAE522E51FFDD94F549E3E9F4ECFA" box="[159,211,968,992]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD948249E3E831ECC5" box="[232,278,968,992]" pageId="28" pageNumber="29" type="generic">2 ♂</specimenCount>
, 1 ♀, same locality,
<quantity id="4CC69B722E51FFDD966B49E3EB7EECC5" box="[513,601,968,992]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="m" value="1000.0">1000 m</quantity>
,
<date id="FF8010572E51FFDD960D49E3EBC3ECFA" box="[615,740,968,992]" pageId="28" pageNumber="29" value="2000-07-02">2.VII.2000</date>
, Chen Yun leg. (
<collectionCode id="ED2FAE522E51FFDD97CE49E3EAFFECFA" box="[932,984,968,992]" pageId="28" pageNumber="29">IOZ</collectionCode>
); 2 ♀, same locality,
<quantity id="4CC69B722E51FFDD90A149E3EC32ECC5" box="[1227,1301,968,992]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="28" pageNumber="29" unit="m" value="900.0">900 m</quantity>
,
<date id="FF8010572E51FFDD914949E3ECB0ECFA" box="[1315,1431,968,992]" pageId="28" pageNumber="29" value="1999-05-17">17.V.1999</date>
, Zhang Guoqing leg. (
<collectionCode id="ED2FAE522E51FFDD95FB49C7E8E2EB1E" box="[401,453,1004,1028]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD95B749C7EB2BEB1E" box="[477,524,1004,1028]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, same locality,
<date id="FF8010572E51FFDD96A949C7EA12EB1E" box="[707,821,1004,1028]" pageId="28" pageNumber="29" value="1984-08">VIII.1984</date>
, Chen Youwei leg. (
<collectionCode id="ED2FAE522E51FFDD904C49C7ED7DEB1E" box="[1062,1114,1004,1028]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD901B49C7ED87EB1E" box="[1137,1184,1004,1028]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, 2 ♀, Napo,
<quantity id="4CC69B722E51FFDD915749C7ECBFEB1E" box="[1341,1432,1004,1029]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="28" pageNumber="29" unit="m" value="1350.0">1350 m</quantity>
,
<date id="FF8010572E51FFDD94FD4E3BE82BEB32" box="[151,268,1040,1064]" pageId="28" pageNumber="29" value="2000-05-18">18.V.2000</date>
, Zhu Chaodong &amp; Li Wenzhu leg. (
<collectionCode id="ED2FAE522E51FFDD96C84E3BEBF1EB32" box="[674,726,1040,1064]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD96804E3BEA30EB3D" box="[746,791,1040,1064]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, 5 ♀, same locality,
<quantity id="4CC69B722E51FFDD906A4E24ED7FEB3D" box="[1024,1112,1039,1064]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="28" pageNumber="29" unit="m" value="1350.0">1350 m</quantity>
,
<date id="FF8010572E51FFDD900E4E3BEDFDEB32" box="[1124,1242,1040,1064]" pageId="28" pageNumber="29" value="2000-05-19">19.V.2000</date>
, Li Wenzhu, Yao Jian, Zhu Chaodong &amp; Chen Jun leg. (
<collectionCode id="ED2FAE522E51FFDD96204E1FEB59EB56" box="[586,638,1076,1100]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD96F84E1FEB98EB56" box="[658,703,1076,1100]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, Fangcheng,
<quantity id="4CC69B722E51FFDD973F4E1FEAB8EB56" box="[853,927,1076,1101]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="28" pageNumber="29" unit="m" value="550.0">550 m</quantity>
,
<date id="FF8010572E51FFDD97C14E1FED07EB57" box="[939,1056,1076,1101]" pageId="28" pageNumber="29" value="1999-05-26">26.V.1999</date>
, Li Wenzhu leg. (
<collectionCode id="ED2FAE522E51FFDD90834E1FEC3AEB56" box="[1257,1309,1076,1100]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD915B4E1FEC79EB56" box="[1329,1374,1076,1100]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, 1 ♀, same data but
<date id="FF8010572E51FFDD95524E7CE88BEB6A" box="[312,428,1111,1136]" pageId="28" pageNumber="29" value="2000-06-05">5.VI.2000</date>
, (
<collectionCode id="ED2FAE522E51FFDD95AA4E73E8D3EB6A" box="[448,500,1112,1136]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD966D4E73EB13EB75" box="[519,564,1112,1136]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, 1 ♀, same data but
<quantity id="4CC69B722E51FFDD97734E7CEA45EB75" box="[793,866,1111,1136]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="29" unit="m" value="250.0">250 m</quantity>
,
<date id="FF8010572E51FFDD97044E73EAC6EB6A" box="[878,993,1112,1136]" pageId="28" pageNumber="29" value="2000-06-03">3.VI.2000</date>
, (
<collectionCode id="ED2FAE522E51FFDD979C4E73ED0DEB6A" box="[1014,1066,1112,1136]" pageId="28" pageNumber="29">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E51FFDD90574E73ED4DEB75" box="[1085,1130,1112,1136]" pageId="28" pageNumber="29" type="generic">1 ♂</specimenCount>
, 1 ♀, same locality,
<quantity id="4CC69B722E51FFDD91244E73ECB0EB75" box="[1358,1431,1112,1136]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="m" value="200.0">200 m</quantity>
,
<date id="FF8010572E51FFDD94FD4E57E82BEB8E" box="[151,268,1148,1172]" pageId="28" pageNumber="29" value="1999-05-23">23.V.1999</date>
, Zhang Yanzhou &amp; Li Wenzhu leg. (
<collectionCode id="ED2FAE522E51FFDD96DB4E57EBC2EB8E" box="[689,741,1148,1172]" pageId="28" pageNumber="29">IOZ</collectionCode>
); 1 ♀, Longzhou,
<quantity id="4CC69B722E51FFDD97DC4E57ED26EB8E" box="[950,1025,1148,1172]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="28" pageNumber="29" unit="m" value="330.0">330 m</quantity>
,
<date id="FF8010572E51FFDD90644E57EDB7EB8F" box="[1038,1168,1148,1173]" pageId="28" pageNumber="29" value="2000-06-15">15.VI.2000</date>
, Li Wenzhu leg. (
<collectionCode id="ED2FAE522E51FFDD91364E57ECB7EB8E" box="[1372,1424,1148,1172]" pageId="28" pageNumber="29">IOZ</collectionCode>
). GUANGDONG: 1 ♀, Dinghushan,
<date id="FF8010572E51FFDD96434E8BEB9DEBA2" box="[553,698,1184,1208]" pageId="28" pageNumber="29" value="1980-05-09" valueMax="1980-05-14" valueMin="1980-05-09">9–14.V.1980</date>
, Wu Xingfang &amp; Du Shaokun leg. (
<collectionCode id="ED2FAE522E51FFDD90264E8BEDA7EBA2" box="[1100,1152,1184,1208]" pageId="28" pageNumber="29">IOZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E51FFDD94AD4EEFED95EA52" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8B8136972E51FFDD94AD4EEFED95EA52" blockId="28.[151,1437,644,1748]" pageId="28" pageNumber="29">
<emphasis id="B94AEA852E51FFDD94AD4EEFE81DEBC7" bold="true" box="[199,314,1220,1245]" pageId="28" pageNumber="29">Diagnosis</emphasis>
. Body size, on average, is greater (
<quantity id="4CC69B722E51FFDD96BE4EEFEA49EBC6" box="[724,878,1220,1245]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3599999999999999" metricValueMin="0.94" pageId="28" pageNumber="29" unit="mm" value="11.5" valueMax="13.6" valueMin="9.4">9.4–13.6 mm</quantity>
) than in the nominotypical subspecies (
<quantity id="4CC69B722E51FFDD91534EEFE9E0EA1A" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.26" metricValueMin="0.89" pageId="28" pageNumber="29" unit="mm" value="10.75" valueMax="12.6" valueMin="8.9">8.9–12.6 mm</quantity>
), elytra are more parallel-sided, metepisterna longer, mesotarsomere 1 of male with adhesive scales in apical part, and medial lobe with a dorsomedial spiny patch in internal sac (
<figureCitation id="13052A122E51FFDD906D4F27EDB4EA3F" box="[1031,1171,1292,1317]" captionStart="FIGURES 60 – 67" captionStartId="27.[151,263,1952,1975]" captionTargetBox="[214,1376,206,1918]" captionTargetId="figure@27.[196,1390,193,1932]" captionTargetPageId="27" captionText="FIGURES 60 – 67. Harpalus indicus, median lobe of aedeagus. 60 – 63, H. indicus indicus (Nepal); 64 – 67, H. indicus orientalis (64, Vietnam; 65 – 67, Taiwan). 60, 66, view from the left side; 61, 67, dorsal view; 62 – 65, view from the right side). a, b, folded formations of internal sac. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287898/files/figure.png" pageId="28" pageNumber="29">Figs 64–67</figureCitation>
); median lobe of the nominotypical subspecies is without dorsomedial spiny patch in the internal sac (
<figureCitation id="13052A122E51FFDD90494F04ED81EA52" box="[1059,1190,1327,1352]" captionStart="FIGURES 60 – 67" captionStartId="27.[151,263,1952,1975]" captionTargetBox="[214,1376,206,1918]" captionTargetId="figure@27.[196,1390,193,1932]" captionTargetPageId="27" captionText="FIGURES 60 – 67. Harpalus indicus, median lobe of aedeagus. 60 – 63, H. indicus indicus (Nepal); 64 – 67, H. indicus orientalis (64, Vietnam; 65 – 67, Taiwan). 60, 66, view from the left side; 61, 67, dorsal view; 62 – 65, view from the right side). a, b, folded formations of internal sac. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287898/files/figure.png" pageId="28" pageNumber="29">Figs 60–63</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E51FFDD94AD4F7FEDF2EAAE" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8B8136972E51FFDD94AD4F7FED6CEA8A" blockId="28.[151,1437,644,1748]" pageId="28" pageNumber="29">
<emphasis id="B94AEA852E51FFDD94AD4F7FE87EEA77" bold="true" box="[199,345,1364,1389]" pageId="28" pageNumber="29">Distribution</emphasis>
. The southern part of
<collectingCountry id="F32976072E51FFDD96EB4F7FEBE3EA77" box="[641,708,1364,1389]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
(Yunnan, Xizang, Guangxi, Guangdong, and
<collectingCountry id="F32976072E51FFDD90944F7FEC71EA77" box="[1278,1366,1364,1389]" name="Taiwan" pageId="28" pageNumber="29">Taiwan</collectingCountry>
). The subspecies is recorded here from Xizang, Guangxi and Guangdong for the first time.
</paragraph>
<paragraph id="8B8136972E51FFDD94AD4FB7EDF2EAAE" blockId="28.[151,1437,644,1748]" box="[199,1237,1436,1461]" pageId="28" pageNumber="29">
<taxonomicName id="4C3E4D142E51FFDD94AD4FB7E8DCEAAE" box="[199,507,1436,1460]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="indicus" subSpecies="orientalis">
<emphasis id="B94AEA852E51FFDD94AD4FB7E8DCEAAE" box="[199,507,1436,1460]" italics="true" pageId="28" pageNumber="29">Harpalus indicus orientalis</emphasis>
</taxonomicName>
is also known from
<collectingCountry id="F32976072E51FFDD968A4FB6EA68EAAF" box="[736,847,1437,1461]" name="Myanmar" pageId="28" pageNumber="29">Myanmar</collectingCountry>
,
<collectingCountry id="F32976072E51FFDD97304FB7EA9EEAAF" box="[858,953,1436,1461]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
and
<collectingCountry id="F32976072E51FFDD97854FB6ED02EAAE" box="[1007,1061,1437,1460]" name="Laos" pageId="28" pageNumber="29">Laos</collectingCountry>
(
<bibRefCitation id="EFAF4B662E51FFDD905E4FB7EDEEEAAE" author="Kataev" box="[1076,1225,1436,1460]" pageId="28" pageNumber="29" refString="Kataev, B. M. (2014) Systematic and nomenclatorial notes on some taxa of Zabrini and Harpalini from the Palaearctic, Oriental and Australian regions (Coleoptera: Carabidae). Proceedings of the Zoological Institute of the Russian Academy of Sciences, 318 (3), 252 - 267." type="journal article" year="2014">Kataev 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E51FFDD94AD4F94E818E9CE" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8B8136972E51FFDD94AD4F94E818E9CE" blockId="28.[151,1437,644,1748]" pageId="28" pageNumber="29">
<emphasis id="B94AEA852E51FFDD94AD4F94E813EAC2" bold="true" box="[199,308,1471,1496]" pageId="28" pageNumber="29">Remarks</emphasis>
.
<taxonomicName id="4C3E4D142E51FFDD95234FEBEB35EAC2" box="[329,530,1472,1496]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94AEA852E51FFDD95234FEBEB35EAC2" box="[329,530,1472,1496]" italics="true" pageId="28" pageNumber="29">Harpalus indicus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C3E4D142E51FFDD976A4FEAED50EAC2" authority="Kataev, 2001" authorityName="Kataev" authorityYear="2001" box="[768,1143,1472,1496]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="meghalayensis">
<emphasis id="B94AEA852E51FFDD976A4FEAEAE9EAC2" box="[768,974,1472,1496]" italics="true" pageId="28" pageNumber="29">H. meghalayensis</emphasis>
<bibRefCitation id="EFAF4B662E51FFDD97B64FEAED50EAC2" author="Kataev" box="[988,1143,1472,1496]" pageId="28" pageNumber="29" refString="Kataev, B. M. (2001) To the knowledge of species of the subgenus Pseudoophonus (genus Harpalus) from the Himalayan region and Southern China (Coleoptera: Carabidae). Zoosystematica Rossica, 9 (2), 2000, 391 - 407." type="journal article" year="2001">Kataev, 2001</bibRefCitation>
</taxonomicName>
from northeastern
<collectingCountry id="F32976072E51FFDD91084F94ECBBEAC2" box="[1378,1436,1471,1496]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
(Meghalaya) in many characteristics including the transverse elytral microsculpture, the hind femora with two or three setigerous pores along hind margin, the nonpubescent elytral intervals and the obtuse, not denticulate, basal angles of pronotum; in addition, like the males of the subspecies
<taxonomicName id="4C3E4D142E51FFDD97A14C06EDE6E95E" box="[971,1217,1580,1604]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="indicus" subSpecies="orientalis">
<emphasis id="B94AEA852E51FFDD97A14C06EDE6E95E" box="[971,1217,1580,1604]" italics="true" pageId="28" pageNumber="29">H. indicus orientalis</emphasis>
</taxonomicName>
, the males of
<taxonomicName id="4C3E4D142E51FFDD91EB4C06E81BE972" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="meghalayensis">
<emphasis id="B94AEA852E51FFDD91EB4C06E81BE972" italics="true" pageId="28" pageNumber="29">H. meghalayensis</emphasis>
</taxonomicName>
possess adhesive scales at apex of the mesotarsomere 1. As compared with adults of
<taxonomicName id="4C3E4D142E51FFDD91754C7AECB2E972" box="[1311,1429,1616,1640]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94AEA852E51FFDD91754C7AECB2E972" box="[1311,1429,1616,1640]" italics="true" pageId="28" pageNumber="29">H. indicus</emphasis>
</taxonomicName>
, those of
<taxonomicName id="4C3E4D142E51FFDD94974C5EE8E0E996" box="[253,455,1652,1676]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="meghalayensis">
<emphasis id="B94AEA852E51FFDD94974C5EE8E0E996" box="[253,455,1652,1676]" italics="true" pageId="28" pageNumber="29">H. meghalayensis</emphasis>
</taxonomicName>
are stouter, with pronotum broader and more strongly depressed at basal angles; the internal sac of aedeagus (
<figureCitation id="13052A122E51FFDD95A84CBCEB6BE9AA" box="[450,588,1687,1712]" captionStart="FIGURES 68 – 71" captionStartId="29.[151,263,1099,1122]" captionTargetBox="[210,1376,201,1070]" captionTargetId="figure@29.[203,1384,195,1078]" captionTargetPageId="29" captionText="FIGURES 68 – 71. Harpalus meghalayensis, median lobe of aedeagus (Meghalaya). 68, view from the left side; 69, dorsal view; 70, 71, view from the right side. a, folded formation of internal sac. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287899/files/figure.png" pageId="28" pageNumber="29">Figs. 68–71</figureCitation>
) has one or two medial spiny patches and comparatively short field &quot;
<emphasis id="B94AEA852E51FFDD91E94CB2ECB6E9AA" box="[1411,1425,1689,1712]" italics="true" pageId="28" pageNumber="29">a</emphasis>
&quot; (
<bibRefCitation id="EFAF4B662E51FFDD94F44C97E815E9CE" author="Kataev" box="[158,306,1724,1748]" pageId="28" pageNumber="29" refString="Kataev, B. M. (2001) To the knowledge of species of the subgenus Pseudoophonus (genus Harpalus) from the Himalayan region and Southern China (Coleoptera: Carabidae). Zoosystematica Rossica, 9 (2), 2000, 391 - 407." type="journal article" year="2001">Kataev 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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