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<document id="A4EBC397701B6028DB4837D76AE1F510" ID-DOI="10.1016/j.sajb.2017.06.011" ID-ISSN="1727-9321" ID-Zenodo-Dep="10496466" IM.bibliography_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1705062396179" checkinUser="felipe" docAuthor="Elliott, T. L. &amp; Muasya, A. M." docDate="2017" docId="0396E93B3413051CFC9314BBFB5ED0EB" docLanguage="en" docName="SAfrJBot.112.354-360.pdf" docOrigin="South African Journal of Botany 112" docSource="http://dx.doi.org/10.1016/j.sajb.2017.06.011" docStyle="DocumentStyle:AD72C92759D651943977BAEA6E2218B4.1:SAfrJBot.2014-.journal_article" docStyleId="AD72C92759D651943977BAEA6E2218B4" docStyleName="SAfrJBot.2014-.journal_article" docStyleVersion="1" docTitle="Schoenus nigricans L." docType="treatment" docVersion="3" lastPageNumber="356" masterDocId="FFAF91433411051EFFB6113AFFC5D72F" masterDocTitle="Taxonomic realignment in the southern African Tetraria (Cyperaceae, tribe Schoeneae; Schoenus clade) *" masterLastPageNumber="360" masterPageNumber="354" pageNumber="356" updateTime="1705514124583" updateUser="felipe">
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<mods:title id="4AE559825156DD233D00AF7B1EEA9FDC">Taxonomic realignment in the southern African Tetraria (Cyperaceae, tribe Schoeneae; Schoenus clade) *</mods:title>
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<mods:namePart id="A32D6B81A67D3EE9060894122D354F2C">Elliott, T. L.</mods:namePart>
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<mods:namePart id="D705122984D30627A2659B3822A16B63">Muasya, A. M.</mods:namePart>
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<mods:title id="4A7DEF5D011E0B374A65BCD7332C4075">South African Journal of Botany</mods:title>
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<mods:date id="0856CCCF6263DDA1FA238CB198F88DD8">2017</mods:date>
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<mods:number id="AD437EE0E01FC03E8F4E5C7E581D4754">2017-09-30</mods:number>
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<treatment id="0396E93B3413051CFC9314BBFB5ED0EB" LSID="urn:lsid:plazi:treatment:0396E93B3413051CFC9314BBFB5ED0EB" httpUri="http://treatment.plazi.org/id/0396E93B3413051CFC9314BBFB5ED0EB" lastPageNumber="356" pageId="2" pageNumber="356">
<subSubSection id="C3250BA63413051CFC9314BBFBCED2BA" box="[805,1035,1409,1429]" pageId="2" pageNumber="356" type="nomenclature">
<paragraph id="8B80582D3413051CFC9314BBFBCED2BA" blockId="2.[805,1035,1409,1429]" box="[805,1035,1409,1429]" pageId="2" pageNumber="356">
<heading id="D0C8EF413413051CFC9314BBFBCED2BA" box="[805,1035,1409,1429]" fontSize="36" level="2" pageId="2" pageNumber="356" reason="3">
<emphasis id="B94B843F3413051CFC9314BBFBCED2BA" box="[805,1035,1409,1429]" italics="true" pageId="2" pageNumber="356">
6.1.
<taxonomicName id="4C3F23AE3413051CFCFC14BBFC32D2BA" ID-CoL="4VF9R" authorityName="L." box="[842,1015,1409,1429]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="356" phylum="Tracheophyta" rank="species" species="nigricans">Schoenus nigricans</taxonomicName>
L.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3250BA63413051CFCF21483FCA7D033" pageId="2" pageNumber="356" type="discussion">
<paragraph id="8B80582D3413051CFCF21483FCA7D033" blockId="2.[805,1475,1465,1988]" pageId="2" pageNumber="356">
<taxonomicName id="4C3F23AE3413051CFCF21483FC35D2E2" ID-CoL="4VF9R" box="[836,1008,1465,1485]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="356" phylum="Tracheophyta" rank="species" species="nigricans">
<emphasis id="B94B843F3413051CFCF21483FC35D2E2" box="[836,1008,1465,1485]" italics="true" pageId="2" pageNumber="356">Schoenus nigricans</emphasis>
</taxonomicName>
(
<figureCitation id="130444A83413051CFC481480FB85D2E2" box="[1022,1088,1466,1485]" captionStart="Fig" captionStartId="3.[113,138,760,774]" captionTargetBox="[239,1372,144,723]" captionTargetId="figure-180@3.[237,1374,142,724]" captionTargetPageId="3" captionText="Fig. 2. Examples of plants from each of the four clades of Schoenus found in southern Africa, showing the corresponding spikelets beneath each plant: A and F—S. nigricans, B and G—T.compar, C and H—Epischoenus sp. and D and I—T.cuspidata.A reticulate-sheathed Tetraria of the Tricostularia clade is shown in E, with its spikelet shown in J.The black scale bar beneath the spikelets in FJ represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496470" httpUri="https://zenodo.org/record/10496470/files/figure.png" pageId="2" pageNumber="356">Fig. 2A</figureCitation>
) is the type species of one of the first 100 genera described by Linnaeus (
<bibRefCitation id="EFAE25DC3413051CFBFC14ECFB1ED2C6" author="Linnaeus, C." box="[1098,1243,1494,1513]" pageId="2" pageNumber="356" pagination="42 - 44" refId="ref8327" refString="Linnaeus, C., 1753. Schoenus. Species Plantarum: exhibentes Plantas Rite Cognitas, Ad Genera Relatas, Cum Differentiis Specificis, Nominibus Trivialibus, Synonymis Selectis, Locis Natalibus, Secundum Systema Sexuale Digestas. Sweden, pp. 42 - 44." type="book chapter" year="1753">Linnaeus, 1753</bibRefCitation>
;
<bibRefCitation id="EFAE25DC3413051CFB5214ECFABED2C6" author="Hitchcock, A. S." box="[1252,1403,1494,1513]" pageId="2" pageNumber="356" pagination="510 - 514" refId="ref7723" refString="Hitchcock, A. S., 1923. Type species of the first 100 genera of Linnaeus' Species Plantarum. American Journal of Botany 10, 510 - 514." type="journal article" year="1923">Hitchcock, 1923</bibRefCitation>
), and a formal
<typeStatus id="5484E68F3413051CFCDE14C8FC05D12A" box="[872,960,1522,1541]" pageId="2" pageNumber="356" type="lectotype">lectotype</typeStatus>
of this species was later designated by Simpson based on a Linnean specimen (LINN 68.6) collected from
<collectingRegion id="49FB96CF3413051CFAAD1734FAAFD10E" box="[1307,1386,1550,1569]" country="Sweden" name="Gotland" pageId="2" pageNumber="356">Gotland</collectingRegion>
,
<collectingCountry id="F32818BD3413051CFAC31734FA07D10E" box="[1397,1474,1550,1569]" name="Sweden" pageId="2" pageNumber="356">Sweden</collectingCountry>
(
<bibRefCitation id="EFAE25DC3413051CFC981713FBD1D112" author="Asberg, M. &amp; Stearn, W. T." box="[814,1044,1577,1597]" pageId="2" pageNumber="356" pagination="1 - 107" refId="ref7307" refString="Asberg, M., Stearn, W. T., 1973. Linnaeus's Oland and Gotland journey 1741. Biological Journal of the Linnean Society 5, 1 - 107." type="journal article" year="1973">Åsberg and Stearn, 1973</bibRefCitation>
;
<bibRefCitation id="EFAE25DC3413051CFBAD1710FB42D112" author="Jarvis, C. E." box="[1051,1159,1578,1597]" pageId="2" pageNumber="356" refId="ref7786" refString="Jarvis, C. E., 1993. A list of Linnaean generic names and their types. Regnum Veg. Balogh Scientific Books, KOnigstein, Germany." type="book" year="1993">Jarvis, 1993</bibRefCitation>
). Key characters of
<taxonomicName id="4C3F23AE3413051CFA8B1713FA66D112" ID-CoL="4VF9R" box="[1341,1443,1577,1597]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="356" phylum="Tracheophyta" rank="species" species="nigricans">
<emphasis id="B94B843F3413051CFA8B1713FA66D112" box="[1341,1443,1577,1597]" italics="true" pageId="2" pageNumber="356">S.nigricans</emphasis>
</taxonomicName>
include black basal sheaths that are notably wider than the leaf blades and loosely ovoid inflorescences with one or two bracts (
<bibRefCitation id="EFAE25DC3413051CFA961758FA5ED15A" author="Tucker, G. C." box="[1312,1435,1634,1653]" pageId="2" pageNumber="356" refId="ref8760" refString="Tucker, G. C., 1993. Schoenus nigricans Linnaeus. In: Flora of North America Editorial Committee (Ed.), Flora of North America North of Mexico (New York and Oxford)." type="book" year="1993">Tucker, 1993</bibRefCitation>
). In addition,
<taxonomicName id="4C3F23AE3413051CFC341747FC28D1BE" ID-CoL="4VF9R" box="[898,1005,1661,1681]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="356" phylum="Tracheophyta" rank="species" species="nigricans">
<emphasis id="B94B843F3413051CFC341747FC28D1BE" box="[898,1005,1661,1681]" italics="true" pageId="2" pageNumber="356">S. nigricans</emphasis>
</taxonomicName>
generally has 1025 flattened, oblong-ellipsoid spikelets with two to three sterile basal scales (
<figureCitation id="130444A83413051CFABE17A3FA89D182" box="[1288,1356,1689,1709]" captionStart="Fig" captionStartId="3.[113,138,760,774]" captionTargetBox="[239,1372,144,723]" captionTargetId="figure-180@3.[237,1374,142,724]" captionTargetPageId="3" captionText="Fig. 2. Examples of plants from each of the four clades of Schoenus found in southern Africa, showing the corresponding spikelets beneath each plant: A and F—S. nigricans, B and G—T.compar, C and H—Epischoenus sp. and D and I—T.cuspidata.A reticulate-sheathed Tetraria of the Tricostularia clade is shown in E, with its spikelet shown in J.The black scale bar beneath the spikelets in FJ represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496470" httpUri="https://zenodo.org/record/10496470/files/figure.png" pageId="2" pageNumber="356">Fig. 2F</figureCitation>
), as well as three to eight medially 1-veined floral scales (i.e. glumes) with acute apices (
<bibRefCitation id="EFAE25DC3413051CFCC717EBFC2AD1CB" author="Tucker, G. C." box="[881,1007,1745,1764]" pageId="2" pageNumber="356" refId="ref8760" refString="Tucker, G. C., 1993. Schoenus nigricans Linnaeus. In: Flora of North America Editorial Committee (Ed.), Flora of North America North of Mexico (New York and Oxford)." type="book" year="1993">Tucker, 1993</bibRefCitation>
). Furthermore,
<taxonomicName id="4C3F23AE3413051CFB3D17EAFB33D1CB" ID-CoL="4VF9R" box="[1163,1270,1744,1764]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="356" phylum="Tracheophyta" rank="species" species="nigricans">
<emphasis id="B94B843F3413051CFB3D17EAFB33D1CB" box="[1163,1270,1744,1764]" italics="true" pageId="2" pageNumber="356">S. nigricans</emphasis>
</taxonomicName>
plants have whitish, ovoid to ellipsoid achenes that are glossy with obtuse apices (
<bibRefCitation id="EFAE25DC3413051CFACA17D7FC91D033" author="Tucker, G. C." pageId="2" pageNumber="356" refId="ref8760" refString="Tucker, G. C., 1993. Schoenus nigricans Linnaeus. In: Flora of North America Editorial Committee (Ed.), Flora of North America North of Mexico (New York and Oxford)." type="book" year="1993">Tucker, 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3250BA63413051CFCF2161EFB5ED0EB" pageId="2" pageNumber="356" type="distribution">
<paragraph id="8B80582D3413051CFCF2161EFB5ED0EB" blockId="2.[805,1475,1465,1988]" pageId="2" pageNumber="356">
<taxonomicName id="4C3F23AE3413051CFCF2161EFC3CD017" ID-CoL="4VF9R" box="[836,1017,1828,1848]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="356" phylum="Tracheophyta" rank="species" species="nigricans">
<emphasis id="B94B843F3413051CFCF2161EFC3CD017" box="[836,1017,1828,1848]" italics="true" pageId="2" pageNumber="356">Schoenus nigricans</emphasis>
</taxonomicName>
is a sub-cosmopolitan species found in North America, Eurasia, Africa and
<collectingCountry id="F32818BD3413051CFB8B167BFB53D07B" box="[1085,1174,1857,1876]" name="Australia" pageId="2" pageNumber="356">Australia</collectingCountry>
, often found growing in damp grasslands (
<figureCitation id="130444A83413051CFC211667FC12D05F" box="[919,983,1885,1904]" captionStart="Fig" captionStartId="3.[113,138,1926,1940]" captionTargetBox="[315,1297,1164,1891]" captionTargetId="figure-265@3.[312,1299,1162,1891]" captionTargetPageId="3" captionText="Fig. 3. Documented locations of Schoenus nigricans and accepted southern African Tetraria and Epischoenus species in the Schoenus clade of Cyperaceae, based on data downloaded from GBIF (Copenhagen: Global Biodiversity Information Facility, 2014). The following groups of species are included in each map: A, S.nigricans; B, T. compar and allies; C,Epischoenus and allies; and D,T.cuspidata and allies. Schoenus nigricans has a cosmopolitan distribution (A), whereas the distribution of the other three groups is limited to southern Africa (B, C, and D)." figureDoi="http://doi.org/10.5281/zenodo.10496472" httpUri="https://zenodo.org/record/10496472/files/figure.png" pageId="2" pageNumber="356">Fig. 3a</figureCitation>
:
<bibRefCitation id="EFAE25DC3413051CFC561667FB9BD05F" author="Tucker, G. C." box="[992,1118,1885,1904]" pageId="2" pageNumber="356" refId="ref8760" refString="Tucker, G. C., 1993. Schoenus nigricans Linnaeus. In: Flora of North America Editorial Committee (Ed.), Flora of North America North of Mexico (New York and Oxford)." type="book" year="1993">Tucker, 1993</bibRefCitation>
;
<bibRefCitation id="EFAE25DC3413051CFBDE1667FAD8D05F" author="Goetghebeur, P." box="[1128,1309,1885,1904]" pageId="2" pageNumber="356" pagination="141 - 190" refId="ref7676" refString="Goetghebeur, P., 1998. Cyperaceae. The Families and Genera of Vascular Plants. Flowering Plants, Monocotyledons: Alismatanae and Commelinanae (except Gramineae). Springer, New York, New York, United States, pp. 141 - 190." type="book chapter" year="1998">Goetghebeur, 1998</bibRefCitation>
). In
<collectingCountry id="F32818BD3413051CFAFF1667FA7AD05F" box="[1353,1471,1885,1904]" name="South Africa" pageId="2" pageNumber="356">South Africa</collectingCountry>
,
<taxonomicName id="4C3F23AE3413051CFC931642FC54D0A3" ID-CoL="4VF9R" box="[805,913,1912,1932]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="356" phylum="Tracheophyta" rank="species" species="nigricans">
<emphasis id="B94B843F3413051CFC931642FC54D0A3" box="[805,913,1912,1932]" italics="true" pageId="2" pageNumber="356">S. nigricans</emphasis>
</taxonomicName>
specimens have been recorded and databased from the Western and
<collectingRegion id="49FB96CF3413051CFC7F16AFFB92D087" box="[969,1111,1941,1960]" country="South Africa" name="Eastern Cape" pageId="2" pageNumber="356">Eastern Cape</collectingRegion>
provinces (Copenhagen: Global Biodiversity Information Facility, 2014).
</paragraph>
</subSubSection>
</treatment>
</document>