209 lines
33 KiB
XML
209 lines
33 KiB
XML
<document id="4316A3CD0509C91A97551113F5F695AD" ID-DOI="10.2478/if-2019-0013" ID-ISSN="2533-4069" ID-Zenodo-Dep="5386203" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630303095841" checkinUser="diego" docAuthor="Friis, Else Marie, Crane, Peter R. & Pedersen, Kaj Raunsgaard" docDate="2019" docId="0396DC10BF24C22FCEDEB599E2B21E64" docLanguage="en" docName="2_Friis.pdf" docOrigin="Fossil Imprint 75 (2)" docSource="http://dx.doi.org/10.2478/if-2019-0013" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Goczania inaequalis E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN 2019, sp. nov." docType="treatment" docVersion="5" lastPageNumber="197" masterDocId="FFAFA468BF0FC203CD58B635E7781C51" masterDocTitle="The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community" masterLastPageNumber="257" masterPageNumber="153" pageNumber="196" updateTime="1699221227565" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="9AE7AB59B3B566C56DA0A4CCD34D3DBA">The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community</mods:title>
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<mods:namePart id="167217B286F1E4C50B061F5C256D667A">Friis, Else Marie</mods:namePart>
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<mods:namePart id="0C0133CA74AA81DF9011421C18C9D848">Crane, Peter R.</mods:namePart>
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<mods:namePart id="FD80492A883631F55A79444F4C86F074">Pedersen, Kaj Raunsgaard</mods:namePart>
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<mods:typeOfResource id="774092F736EA2845F82DFB881329E4EE">text</mods:typeOfResource>
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<mods:title id="AE23F231DC73F4A305E01E3259F7AB0E">Fossil Imprint</mods:title>
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<mods:date id="4E3D58ECDDCAE0518720939CD1C8F31E">2019</mods:date>
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<mods:number id="1C9EC0C6819528488FAAA8D90D08E408">2019-11-25</mods:number>
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<mods:url id="7471C4DE371BF0FFBC2A2E12AF75A983">http://dx.doi.org/10.2478/if-2019-0013</mods:url>
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<subSubSection id="C3253E8DBF24C228CEDEB599E3BA1854" pageId="43" pageNumber="196" type="nomenclature">
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<heading id="D0C8DA6ABF24C228CEDEB599E3861FB4" centered="true" fontSize="8" level="2" pageId="43" pageNumber="196" reason="2">
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<emphasis id="B94BB114BF24C228CEDEB599E3861FB4" bold="true" pageId="43" pageNumber="196">
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<taxonomicName id="4C3F1685BF24C228CEDEB599E3DA1FB2" authority="E.M.FRIIS, P.R.CRANE et K.R.PEDERSEN" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="43" pageNumber="196" phylum="Tracheophyta" rank="species" species="inaequalis" status="sp. nov.">
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<emphasis id="B94BB114BF24C228CEDEB599E31F1F95" bold="true" box="[902,1127,940,964]" italics="true" pageId="43" pageNumber="196">Goczania inaequalis</emphasis>
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E.M.FRIIS, P.R.CRANE et K.R.PEDERSEN
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</taxonomicName>
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<taxonomicNameLabel id="A2780C6FBF24C228C9F1B5F8E3861FB4" box="[1193,1278,973,997]" pageId="43" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B806D06BF24C228C97DB5D8E3BA1854" blockId="43.[1061,1218,1005,1029]" box="[1061,1218,1005,1029]" pageId="43" pageNumber="196">
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<figureCitation id="13047183BF24C228C97DB5D8E3BA1854" box="[1061,1218,1005,1029]" captionStart="Text-fig" captionStartId="45.[125,206,1918,1939]" captionTargetBox="[128,1440,151,1898]" captionTargetId="figure-25@45.[414,884,990,1487]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania inaequalis sp. nov.; Torres Vedras locality, Portugal. a) Holotype; stamen fragment that yielded the pollen in (b–f, k); b) Group of pollen grains showing distal and proximal surfaces and inner surface of anther wall with densely spaced orbicules; c) Inner surface of anther wall showing small, spherical orbicules with microechinate sculpturing; d, e, j–l) Pollen grains in distal and equatorial view (k) showing the long colpus with an irregular margin flanked by narrow bands of poorly differentiated microechinate; note that colpus is aligned perpendicular to the longest axes in elliptical grains and that the pollen wall is almost psilate around the equator and in the distal" figureDoi="http://doi.org/10.5281/zenodo.5386295" httpUri="https://zenodo.org/record/5386295/files/figure.png" pageId="43" pageNumber="196">Text-fig. 28a–l</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3253E8DBF24C22FCE02B228E2B21E64" lastPageId="44" lastPageNumber="197" pageId="43" pageNumber="196" type="description">
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<paragraph id="8B806D06BF24C228CE02B228E3D71805" blockId="43.[820,1459,1052,1108]" pageId="43" pageNumber="196">
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H o l o t y p e. Designated here. S137919 (Torres Vedras sample 44; figured
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<figureCitation id="13047183BF24C228C959B209E3DC1805" box="[1025,1188,1084,1108]" captionStart="Text-fig" captionStartId="45.[125,206,1918,1939]" captionTargetBox="[128,1440,151,1898]" captionTargetId="figure-25@45.[414,884,990,1487]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania inaequalis sp. nov.; Torres Vedras locality, Portugal. a) Holotype; stamen fragment that yielded the pollen in (b–f, k); b) Group of pollen grains showing distal and proximal surfaces and inner surface of anther wall with densely spaced orbicules; c) Inner surface of anther wall showing small, spherical orbicules with microechinate sculpturing; d, e, j–l) Pollen grains in distal and equatorial view (k) showing the long colpus with an irregular margin flanked by narrow bands of poorly differentiated microechinate; note that colpus is aligned perpendicular to the longest axes in elliptical grains and that the pollen wall is almost psilate around the equator and in the distal" figureDoi="http://doi.org/10.5281/zenodo.5386295" httpUri="https://zenodo.org/record/5386295/files/figure.png" pageId="43" pageNumber="196">Text-fig. 28a–f</figureCitation>
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).
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</paragraph>
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<paragraph id="8B806D06BF24C228CE02B259E2C818D2" blockId="43.[820,1456,1132,1188]" box="[858,1456,1132,1156]" pageId="43" pageNumber="196">P l a n t F o s s i l N a m e s R e g i s t r y N u m b e r.</paragraph>
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<paragraph id="8B806D06BF24C228CE6CB2B9E30318F5" blockId="43.[820,1456,1132,1188]" box="[820,1147,1164,1188]" pageId="43" pageNumber="196">PFN000463 (for new species).</paragraph>
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<paragraph id="8B806D06BF24C228CE02B289E2CB1882" blockId="43.[820,1459,1211,1268]" box="[858,1459,1211,1236]" pageId="43" pageNumber="196">P a r a t y p e s. Designated here. S136768, S148011,</paragraph>
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<paragraph id="8B806D06BF24C228CE6CB2EEE3C118A5" blockId="43.[820,1459,1211,1268]" box="[820,1209,1243,1268]" pageId="43" pageNumber="196">S174557 (Torres Vedras sample 44).</paragraph>
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<paragraph id="8B806D06BF24C228CE02B33EE3691932" blockId="43.[820,1460,1291,1379]" pageId="43" pageNumber="196">
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<materialsCitation id="3B57675BBF24C228CE02B33EE3751932" collectionCode="R" country="Sweden" county="Swedish Museum of Natural History" location="Palaeobotanical Collections" municipality="Department of Palaeobiology" pageId="43" pageNumber="196" specimenCount="1" stateProvince="Stockholm">
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<collectionCode id="ED2EF5C3BF24C228CE02B33EE4141973" box="[858,876,1291,1314]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="43" pageNumber="196">R</collectionCode>
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e p o s i t o r y.
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<location id="8EE03BDDBF24C228C954B33EE2561972" LSID="urn:lsid:plazi:treatment:0396DC10BF24C22FCEDEB599E2B21E64:8EE03BDDBF24C228C954B33EE2561972" box="[1036,1326,1291,1315]" country="Sweden" county="Swedish Museum of Natural History" municipality="Department of Palaeobiology" name="Palaeobotanical Collections" pageId="43" pageNumber="196" stateProvince="Stockholm">Palaeobotanical Collections</location>
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,
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<collectingMunicipality id="6BE4F77CBF24C228C861B33EE4941912" pageId="43" pageNumber="196">Department of Palaeobiology</collectingMunicipality>
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, the
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<collectingCounty id="62E1158ABF24C228C978B31EE2D71912" box="[1056,1455,1323,1347]" pageId="43" pageNumber="196">Swedish Museum of Natural History</collectingCounty>
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,
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<collectingRegion id="49FBA3E4BF24C228CE6CB37EE4D21932" box="[820,938,1355,1379]" country="Sweden" name="Stockholm" pageId="43" pageNumber="196">Stockholm</collectingRegion>
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,
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<collectingCountry id="F3282D96BF24C228CEEDB37EE3751932" box="[949,1037,1355,1379]" name="Sweden" pageId="43" pageNumber="196">Sweden</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B806D06BF24C228CE02B34EE39D19E2" blockId="43.[820,1459,1403,1459]" pageId="43" pageNumber="196">E t y m o l o g y. From Latin: inaequalis referring to the uneven distribution of the microechinae.</paragraph>
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<paragraph id="8B806D06BF24C228CE02B3FEE2CB19B3" blockId="43.[820,1459,1482,1539]" box="[858,1459,1482,1507]" pageId="43" pageNumber="196">T y p e l o c a l i t y. Torres Vedras (NE of Forte de</paragraph>
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<paragraph id="8B806D06BF24C228CE6CB3DEE3D81A52" blockId="43.[820,1459,1482,1539]" box="[820,1184,1513,1539]" pageId="43" pageNumber="196">
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Forca;
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<geoCoordinate id="EE0B0BC1BF24C228CE27B3DCE37F1A53" box="[895,1031,1513,1538]" degrees="39" direction="north" minutes="06" orientation="latitude" pageId="43" pageNumber="196" precision="15" seconds="13" value="39.10361">39°06′13″ N</geoCoordinate>
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,
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<geoCoordinate id="EE0B0BC1BF24C228C94AB3DCE3EB1A52" box="[1042,1171,1513,1539]" degrees="9" direction="west" minutes="14" orientation="longitude" pageId="43" pageNumber="196" precision="15" seconds="47" value="-9.246389">9°14′47″ W</geoCoordinate>
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).
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</paragraph>
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<paragraph id="8B806D06BF24C228CE02B02FE4B01A23" blockId="43.[820,1459,1562,1650]" pageId="43" pageNumber="196">T y p e s t r a t u m a n d a g e. Lower member of the Almargem Formation; Early Cretaceous (late Barremianearly Aptian).</paragraph>
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<paragraph id="8B806D06BF24C228CE02B0BFE4B41B32" blockId="43.[820,1459,1674,1891]" pageId="43" pageNumber="196">D i a g n o s i s. As for the genus with the following additions: Equatorial outline of pollen circular to transversely elliptical. Tectum microperforate and microechinate, not rugulate. Perforations minute and scattered. Microechinae confined to the proximal surface and bordering the colpus on the distal surface; otherwise the distal face and equatorial region psilate.</paragraph>
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<paragraph id="8B806D06BF24C228CE02B14EE3361462" blockId="43.[820,1459,1915,2100]" pageId="43" pageNumber="196">
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D i s t i n g u i s h i n g f e a t u r e s. The species is distinguished from
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<taxonomicName id="4C3F1685BF24C228C956B1AEE3B21BE2" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" box="[1038,1226,1947,1971]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="43" pageNumber="196" phylum="Tracheophyta" rank="species" species="rugosa">
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<emphasis id="B94BB114BF24C228C956B1AEE3B21BE2" box="[1038,1226,1947,1971]" italics="true" pageId="43" pageNumber="196">Goczania rugosa</emphasis>
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</taxonomicName>
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by the psilate, nonrugulate, surface of the pollen and the uneven distribution of the supratectal ornamentation.
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<taxonomicName id="4C3F1685BF24C228C9EAB1E9E2EC1BA2" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" box="[1202,1428,2011,2035]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="43" pageNumber="196" phylum="Tracheophyta" rank="species" species="inaequalis">
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<emphasis id="B94BB114BF24C228C9EAB1E9E2EC1BA2" box="[1202,1428,2011,2035]" italics="true" pageId="43" pageNumber="196">Goczania inaequalis</emphasis>
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</taxonomicName>
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is distinguished from
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<taxonomicName id="4C3F1685BF24C228C948B1C9E3E71442" box="[1040,1183,2044,2067]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="43" pageNumber="196" phylum="Tracheophyta" rank="species" species="punctatus">
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<emphasis id="B94BB114BF24C228C948B1C9E3E71442" box="[1040,1183,2044,2067]" italics="true" pageId="43" pageNumber="196">G. punctatus</emphasis>
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</taxonomicName>
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by the less pronounced perforations in the tectum.
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</paragraph>
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<paragraph id="8B806D06BF23C22FCDFEB6ADE4781CFE" blockId="44.[129,768,151,208]" box="[166,768,151,175]" pageId="44" pageNumber="197">D i m e n s i o n s. Equatorial diameter of pollen grains:</paragraph>
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<paragraph id="8B806D06BF23C22FCDD9B68DE78C1C81" blockId="44.[129,768,151,208]" box="[129,244,184,208]" pageId="44" pageNumber="197">11–12 µm.</paragraph>
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<paragraph id="8B806D06BF23C22FCDFEB6DDE6591E53" blockId="44.[128,768,231,968]" pageId="44" pageNumber="197">
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D e s c r i p t i o n a n d r e m a r k s.
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<taxonomicName id="4C3F1685BF23C22FCF7AB6DDE5871CAE" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" box="[546,767,231,255]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="inaequalis">
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<emphasis id="B94BB114BF23C22FCF7AB6DDE5871CAE" box="[546,767,231,255]" italics="true" pageId="44" pageNumber="197">Goczania inaequalis</emphasis>
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</taxonomicName>
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is based on four anther fragments with in situ pollen grains (
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<figureCitation id="13047183BF23C22FCDD0B71DE6481D11" box="[136,304,296,320]" captionStart="Text-fig" captionStartId="45.[125,206,1918,1939]" captionTargetBox="[128,1440,151,1898]" captionTargetId="figure-25@45.[414,884,990,1487]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania inaequalis sp. nov.; Torres Vedras locality, Portugal. a) Holotype; stamen fragment that yielded the pollen in (b–f, k); b) Group of pollen grains showing distal and proximal surfaces and inner surface of anther wall with densely spaced orbicules; c) Inner surface of anther wall showing small, spherical orbicules with microechinate sculpturing; d, e, j–l) Pollen grains in distal and equatorial view (k) showing the long colpus with an irregular margin flanked by narrow bands of poorly differentiated microechinate; note that colpus is aligned perpendicular to the longest axes in elliptical grains and that the pollen wall is almost psilate around the equator and in the distal" figureDoi="http://doi.org/10.5281/zenodo.5386295" httpUri="https://zenodo.org/record/5386295/files/figure.png" pageId="44" pageNumber="197">Text-fig. 28a, g</figureCitation>
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) as well as isolated pollen grains found in coprolites (
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<figureCitation id="13047183BF23C22FCDA0B77DE6E71D30" box="[248,415,328,353]" captionStart="Text-fig" captionStartId="75.[125,206,1931,1952]" captionTargetBox="[128,1440,1066,1910]" captionTargetId="figure-511@75.[406,961,1063,1912]" captionTargetPageId="75" captionText="Text-fig. 48. Scanning electron microscope (SEM) images of monocolpate pollen from a coprolite; Torres Vedras locality, Portugal. a) Coprolite that yielded the pollen in this Text-figure as well as several other kinds of pollen; b) Reticulum of pollen grain showing the tall, sharp muri and obconical columellae; c–e) Grains of Goczania inaequalis sp. nov. intermixed with pollen grains that have a coarse, loosely attached reticulum with tall, sharp muri and sparse columellae; note the weak transverse striations on the muri and that the reticulum is much larger than the main body of the grain that it encloses. Specimen, TV44-S148023. Scale bars 300 Μm (a), 6 Μm (c–e), 3 Μm (b)." figureDoi="http://doi.org/10.5281/zenodo.5386363" httpUri="https://zenodo.org/record/5386363/files/figure.png" pageId="44" pageNumber="197">Text-fig. 48c, e</figureCitation>
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). The inner surface of the anther wall is covered by densely spaced orbicules (
|
||
<figureCitation id="13047183BF23C22FCF0FB75CE7F41DF0" captionStart="Text-fig" captionStartId="45.[125,206,1918,1939]" captionTargetBox="[128,1440,151,1898]" captionTargetId="figure-25@45.[414,884,990,1487]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania inaequalis sp. nov.; Torres Vedras locality, Portugal. a) Holotype; stamen fragment that yielded the pollen in (b–f, k); b) Group of pollen grains showing distal and proximal surfaces and inner surface of anther wall with densely spaced orbicules; c) Inner surface of anther wall showing small, spherical orbicules with microechinate sculpturing; d, e, j–l) Pollen grains in distal and equatorial view (k) showing the long colpus with an irregular margin flanked by narrow bands of poorly differentiated microechinate; note that colpus is aligned perpendicular to the longest axes in elliptical grains and that the pollen wall is almost psilate around the equator and in the distal" figureDoi="http://doi.org/10.5281/zenodo.5386295" httpUri="https://zenodo.org/record/5386295/files/figure.png" pageId="44" pageNumber="197">Text-fig. 28b, c, f</figureCitation>
|
||
). Orbicules are also scattered over the surface of the pollen grains (
|
||
<figureCitation id="13047183BF23C22FCD97B79FE6211D93" box="[207,345,426,450]" captionStart="Text-fig" captionStartId="45.[125,206,1918,1939]" captionTargetBox="[128,1440,151,1898]" captionTargetId="figure-25@45.[414,884,990,1487]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania inaequalis sp. nov.; Torres Vedras locality, Portugal. a) Holotype; stamen fragment that yielded the pollen in (b–f, k); b) Group of pollen grains showing distal and proximal surfaces and inner surface of anther wall with densely spaced orbicules; c) Inner surface of anther wall showing small, spherical orbicules with microechinate sculpturing; d, e, j–l) Pollen grains in distal and equatorial view (k) showing the long colpus with an irregular margin flanked by narrow bands of poorly differentiated microechinate; note that colpus is aligned perpendicular to the longest axes in elliptical grains and that the pollen wall is almost psilate around the equator and in the distal" figureDoi="http://doi.org/10.5281/zenodo.5386295" httpUri="https://zenodo.org/record/5386295/files/figure.png" pageId="44" pageNumber="197">Text-fig. 28l</figureCitation>
|
||
). The orbicules are small, spherical and with a surface ornamentation of microechinate sculpturing (
|
||
<figureCitation id="13047183BF23C22FCDD0B7DFE66D1E53" box="[136,277,490,514]" captionStart="Text-fig" captionStartId="45.[125,206,1918,1939]" captionTargetBox="[128,1440,151,1898]" captionTargetId="figure-25@45.[414,884,990,1487]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania inaequalis sp. nov.; Torres Vedras locality, Portugal. a) Holotype; stamen fragment that yielded the pollen in (b–f, k); b) Group of pollen grains showing distal and proximal surfaces and inner surface of anther wall with densely spaced orbicules; c) Inner surface of anther wall showing small, spherical orbicules with microechinate sculpturing; d, e, j–l) Pollen grains in distal and equatorial view (k) showing the long colpus with an irregular margin flanked by narrow bands of poorly differentiated microechinate; note that colpus is aligned perpendicular to the longest axes in elliptical grains and that the pollen wall is almost psilate around the equator and in the distal" figureDoi="http://doi.org/10.5281/zenodo.5386295" httpUri="https://zenodo.org/record/5386295/files/figure.png" pageId="44" pageNumber="197">Text-fig. 28c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B806D06BF23C22FCDFEB43EE7A41F99" blockId="44.[128,768,231,968]" pageId="44" pageNumber="197">
|
||
The pollen grains are small, circular to transversely elliptical in equatorial outline, and about 11–12 µm in equatorial diameter (
|
||
<figureCitation id="13047183BF23C22FCC33B47EE5121E32" box="[363,618,587,611]" captionStart="Text-fig" captionStartId="45.[125,206,1918,1939]" captionTargetBox="[128,1440,151,1898]" captionTargetId="figure-25@45.[414,884,990,1487]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania inaequalis sp. nov.; Torres Vedras locality, Portugal. a) Holotype; stamen fragment that yielded the pollen in (b–f, k); b) Group of pollen grains showing distal and proximal surfaces and inner surface of anther wall with densely spaced orbicules; c) Inner surface of anther wall showing small, spherical orbicules with microechinate sculpturing; d, e, j–l) Pollen grains in distal and equatorial view (k) showing the long colpus with an irregular margin flanked by narrow bands of poorly differentiated microechinate; note that colpus is aligned perpendicular to the longest axes in elliptical grains and that the pollen wall is almost psilate around the equator and in the distal" figureDoi="http://doi.org/10.5281/zenodo.5386295" httpUri="https://zenodo.org/record/5386295/files/figure.png" pageId="44" pageNumber="197">Text-fig. 28d–f, h, j–l</figureCitation>
|
||
). The colpus is long, reaching almost to the equator. In grains that are transversely elliptical, the colpus is oriented perpendicular to the longest axis of the grain (
|
||
<figureCitation id="13047183BF23C22FCCAAB498E5C51E94" box="[498,701,685,709]" captionStart="Text-fig" captionStartId="45.[125,206,1918,1939]" captionTargetBox="[128,1440,151,1898]" captionTargetId="figure-25@45.[414,884,990,1487]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania inaequalis sp. nov.; Torres Vedras locality, Portugal. a) Holotype; stamen fragment that yielded the pollen in (b–f, k); b) Group of pollen grains showing distal and proximal surfaces and inner surface of anther wall with densely spaced orbicules; c) Inner surface of anther wall showing small, spherical orbicules with microechinate sculpturing; d, e, j–l) Pollen grains in distal and equatorial view (k) showing the long colpus with an irregular margin flanked by narrow bands of poorly differentiated microechinate; note that colpus is aligned perpendicular to the longest axes in elliptical grains and that the pollen wall is almost psilate around the equator and in the distal" figureDoi="http://doi.org/10.5281/zenodo.5386295" httpUri="https://zenodo.org/record/5386295/files/figure.png" pageId="44" pageNumber="197">Text-fig. 28d, k, l</figureCitation>
|
||
). The colpus margin is distinct with a slightly irregular outline. The pollen wall is tectate with a supratectal ornamentation of solitary microechinae on the proximal surface and also bordering the colpus. Microechinae are absent from the distal face between the colpus margin and the equator where the tectum is psilate (
|
||
<figureCitation id="13047183BF23C22FCC65B55AE5531FD6" box="[317,555,879,903]" captionStart="Text-fig" captionStartId="45.[125,206,1918,1939]" captionTargetBox="[128,1440,151,1898]" captionTargetId="figure-25@45.[414,884,990,1487]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania inaequalis sp. nov.; Torres Vedras locality, Portugal. a) Holotype; stamen fragment that yielded the pollen in (b–f, k); b) Group of pollen grains showing distal and proximal surfaces and inner surface of anther wall with densely spaced orbicules; c) Inner surface of anther wall showing small, spherical orbicules with microechinate sculpturing; d, e, j–l) Pollen grains in distal and equatorial view (k) showing the long colpus with an irregular margin flanked by narrow bands of poorly differentiated microechinate; note that colpus is aligned perpendicular to the longest axes in elliptical grains and that the pollen wall is almost psilate around the equator and in the distal" figureDoi="http://doi.org/10.5281/zenodo.5386295" httpUri="https://zenodo.org/record/5386295/files/figure.png" pageId="44" pageNumber="197">Text-fig. 28d–e, h, j–l</figureCitation>
|
||
). As seen in several broken grains the infratectal layer is thin and granular (Textfig. 28i).
|
||
</paragraph>
|
||
<paragraph id="8B806D06BF23C22FCE29B6ADE34D1D60" blockId="44.[843,1482,152,565]" pageId="44" pageNumber="197">
|
||
A f f i n i t y a n d o t h e r o c c u r r e n c e s.
|
||
<taxonomicName id="4C3F1685BF23C22FC83BB6ADE4C21C81" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="inaequalis">
|
||
<emphasis id="B94BB114BF23C22FC83BB6ADE4C21C81" italics="true" pageId="44" pageNumber="197">Goczania inaequalis</emphasis>
|
||
</taxonomicName>
|
||
, like
|
||
<taxonomicName id="4C3F1685BF23C22FC943B68DE26C1C81" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[1051,1300,184,208]" class="Magnoliopsida" genus="Appomattoxia" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="minuta">
|
||
<emphasis id="B94BB114BF23C22FC943B68DE26C1C81" box="[1051,1300,184,208]" italics="true" pageId="44" pageNumber="197">Appomattoxia minuta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F1685BF23C22FC86CB68DE4991CA0" class="Magnoliopsida" genus="Appomattoxia" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="ancistrophora">
|
||
<emphasis id="B94BB114BF23C22FC86CB68DE4991CA0" italics="true" pageId="44" pageNumber="197">Appomattoxia ancistrophora</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F1685BF23C22FCEA0B6ECE3B31CA0" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[1016,1227,217,241]" class="Magnoliopsida" family="Chloranthaceae" genus="Appofructus" kingdom="Plantae" order="Chloranthales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="nudus">
|
||
<emphasis id="B94BB114BF23C22FCEA0B6ECE3B31CA0" box="[1016,1227,217,241]" italics="true" pageId="44" pageNumber="197">Appofructus nudus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F1685BF23C22FC84AB6ECE2BB1CA1" box="[1298,1475,217,241]" class="Magnoliopsida" family="Chloranthaceae" genus="Appofructus" kingdom="Plantae" order="Chloranthales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B94BB114BF23C22FC84AB6ECE2EC1CA0" box="[1298,1428,217,241]" italics="true" pageId="44" pageNumber="197">Appofructus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, as well as
|
||
<taxonomicName id="4C3F1685BF23C22FCE9DB6CCE3071D40" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" box="[965,1151,249,273]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="rugosa">
|
||
<emphasis id="B94BB114BF23C22FCE9DB6CCE3071D40" box="[965,1151,249,273]" italics="true" pageId="44" pageNumber="197">Goczania rugosa</emphasis>
|
||
</taxonomicName>
|
||
, is probably related to extant
|
||
<taxonomicName id="4C3F1685BF23C22FCE13B72CE4D51D60" box="[843,941,281,305]" class="Magnoliopsida" kingdom="Plantae" order="Piperales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="order">Piperales</taxonomicName>
|
||
(see above).
|
||
</paragraph>
|
||
<paragraph id="8B806D06BF23C22FCE29B70FE2B21E64" blockId="44.[843,1482,152,565]" pageId="44" pageNumber="197">
|
||
<taxonomicName id="4C3F1685BF23C22FCE29B70FE32F1D03" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" box="[881,1111,314,338]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="inaequalis">
|
||
<emphasis id="B94BB114BF23C22FCE29B70FE32F1D03" box="[881,1111,314,338]" italics="true" pageId="44" pageNumber="197">Goczania inaequalis</emphasis>
|
||
</taxonomicName>
|
||
is currently known only from the Torres Vedras mesofossil flora. Pollen grains reported as
|
||
<emphasis id="B94BB114BF23C22FCE29B74EE4831DC2" box="[881,1019,379,403]" italics="true" pageId="44" pageNumber="197">Tucanopollis</emphasis>
|
||
cf.
|
||
<emphasis id="B94BB114BF23C22FC96EB74EE3981DC2" box="[1078,1248,379,403]" italics="true" pageId="44" pageNumber="197">crispopollenites</emphasis>
|
||
from the early-late Aptian of Algarve,
|
||
<collectingCountry id="F3282D96BF23C22FC97CB7AEE3061DE2" box="[1060,1150,411,435]" name="Portugal" pageId="44" pageNumber="197">Portugal</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFAE10F7BF23C22FC9C8B7AEE2F21DE5" author="Heimhofer, U. & Hochuli, P. A. & Burla, S. & Weissert, H." box="[1168,1418,411,436]" pageId="44" pageNumber="197" pagination="39 - 76" refId="ref78283" refString="Heimhofer, U., Hochuli, P. A., Burla, S., Weissert, H. (2007): New records of Early Cretaceous angiosperm pollen from Portuguese coastal deposits: Implications for the timing of the early angiosperm radiation. - Review of Palaeobotany and Palynology, 144: 39 - 76. https: // doi. org / 10.1016 / j. revpalbo. 2005.09.006" type="journal article" year="2007">Heimhofer et al. 2007</bibRefCitation>
|
||
) may be conspecific with
|
||
<taxonomicName id="4C3F1685BF23C22FC97EB789E3861D85" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" box="[1062,1278,444,468]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="inaequalis">
|
||
<emphasis id="B94BB114BF23C22FC97EB789E3861D85" box="[1062,1278,444,468]" italics="true" pageId="44" pageNumber="197">Goczania inaequalis</emphasis>
|
||
</taxonomicName>
|
||
, but details of the tectum ornamentation/perforation are not clear from the LM illustrations and at this resolution it may be difficult to distinguish between
|
||
<taxonomicName id="4C3F1685BF23C22FC942B42BE3DA1E64" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" box="[1050,1186,541,565]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="inaequalis">
|
||
<emphasis id="B94BB114BF23C22FC942B42BE3DA1E64" box="[1050,1186,541,565]" italics="true" pageId="44" pageNumber="197">G. inaequalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F1685BF23C22FC989B42BE2311E64" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[1233,1353,542,565]" class="Magnoliopsida" family="Schisandraceae" genus="Samylinaea" kingdom="Plantae" order="Austrobaileyales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="punctata">
|
||
<emphasis id="B94BB114BF23C22FC989B42BE2311E64" box="[1233,1353,542,565]" italics="true" pageId="44" pageNumber="197">G. punctata</emphasis>
|
||
</taxonomicName>
|
||
(see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |