162 lines
20 KiB
XML
162 lines
20 KiB
XML
<document id="BA9E05F5B5665A8CC72A108C3B31626F" ID-DOI="10.11646/zootaxa.3272.1.1" ID-GBIF-Dataset="02e53dfe-221c-4c20-aa11-82fc7e6e5318" ID-ISSN="1175-5326" ID-Zenodo-Dep="215455" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1460472935642" checkinUser="plazi" docAuthor="Rooijen, Johan Van & Vogel, Gernot" docDate="2012" docId="039579531E0E8F542E86FA3ED8FDFE1E" docLanguage="en" docName="zt03272p025.pdf" docOrigin="Zootaxa 3272" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Dendrelaphis philippinensis Gunther 1879" docType="treatment" docVersion="10" lastPageNumber="19" masterDocId="FFAC012B1E1E8F462E11FFD9DB07FFD1" masterDocTitle="A revision of the taxonomy of Dendrelaphis caudolineatus (Gray, 1834) (Serpentes: Colubridae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="17" updateTime="1698306253900" updateUser="plazi">
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<mods:mods id="E248C8F28DA982EF3727C74F36D93369" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E72E02F0BD34B450D99DB28562E10CE2">
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<mods:title id="05C2F453D67ADB6E50CF866B4473AEBA">A revision of the taxonomy of Dendrelaphis caudolineatus (Gray, 1834) (Serpentes: Colubridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="715640CD2C26D4B1E5D959460047500F" type="personal">
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<mods:roleTerm id="011193C5F09D33E601E5F2DF7E2956B2">Author</mods:roleTerm>
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<mods:namePart id="A22B259668195DA567C405B55D5E5A60">Rooijen, Johan Van</mods:namePart>
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<mods:affiliation id="E12D4EF09B0BF0D57C1468A94BC3BA7A">Netherlands Centre for Biodiversity Naturalis, P. O. Box 9517, 2300 RA Leiden, The Netherlands & Tulpentuin 313, 2272 EH, Voorburg, The Netherlands</mods:affiliation>
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<mods:nameIdentifier id="4722844B0A2327F03ADE6A2AB8761F14" type="email">j1.van.rooijen@hetnet.nl</mods:nameIdentifier>
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<mods:name id="12BE0C6F95AEFC3E466E2C310D4E991A" type="personal">
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<mods:roleTerm id="555A2BEE0F4BADD450301AAB3AB64D78">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="37B2343AFB2C0933E9878137253D5A44">Vogel, Gernot</mods:namePart>
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<mods:affiliation id="0132205D3DD1042E7A5B5FE5218E1E0B">Society for Southeast Asian Herpetology, Im Sand 3, D- 69115 Heidelberg, Germany</mods:affiliation>
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<mods:nameIdentifier id="6871E66F9961354D858C59A846C4BB86" type="email">Gernot.Vogel@t-online.de</mods:nameIdentifier>
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<mods:title id="6BE89D632F9665988563764BE9161884">Zootaxa</mods:title>
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<mods:part id="09DB6986188CF17D4A0E1DE371378807">
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<mods:date id="2A831B8E75158722193F78B1D374A865">2012</mods:date>
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<mods:number id="E596639B5B8265FB03618BC04FBE9FC7">2012-04-19</mods:number>
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<mods:detail id="676B9E7C7E1601E76A91345E6CF7F3A8" type="volume">
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<mods:number id="1A1E8F6D01135472FAC13E8723E3F2E0">3272</mods:number>
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<mods:start id="9816E1A20321978A8D26772D4DE4E590">1</mods:start>
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<mods:classification id="52A49EA68BDA7C7995327926BB2C9A24">journal article</mods:classification>
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<mods:identifier id="F14F268F4021EBBEA849EB30676983A3" type="DOI">10.11646/zootaxa.3272.1.1</mods:identifier>
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<mods:identifier id="E93D87C63EB86A48FDE8376B55AFF66B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="FA4125E6BA7C5BAFFD3353B6132EA370" type="Zenodo-Dep">215455</mods:identifier>
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<treatment id="039579531E0E8F542E86FA3ED8FDFE1E" ID-DOI="http://doi.org/10.5281/zenodo.6318694" ID-GBIF-Taxon="119515811" ID-Zenodo-Dep="6318694" LSID="urn:lsid:plazi:treatment:039579531E0E8F542E86FA3ED8FDFE1E" httpUri="http://treatment.plazi.org/id/039579531E0E8F542E86FA3ED8FDFE1E" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
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<subSubSection id="C3269BCE1E0E8F562E86FA3ED9A4F9D0" box="[151,675,1511,1537]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="8B83C8451E0E8F562E86FA3ED9A4F9D0" blockId="16.[151,675,1511,1570]" box="[151,675,1511,1537]" pageId="16" pageNumber="17">
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<heading id="D0CB7F291E0E8F562E86FA3ED9A4F9D0" bold="true" box="[151,675,1511,1537]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<taxonomicName id="4C3CB3C61E0E8F562E86FA3ED9A4F9D0" authority="Gunther, 1879" authorityName="Gunther" authorityYear="1879" box="[151,675,1511,1537]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="philippinensis">
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<emphasis id="B94814571E0E8F562E86FA3ED9A4F9D0" bold="true" box="[151,675,1511,1537]" pageId="16" pageNumber="17">
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<emphasis id="B94814571E0E8F562E86FA3EDAEDF9D0" bold="true" box="[151,490,1511,1537]" italics="true" pageId="16" pageNumber="17">Dendrelaphis philippinensis</emphasis>
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<bibRefCitation id="EFADB5B41E0E8F562FE3FA3ED9A4F9D0" author="Gunther" box="[498,675,1511,1537]" firstAuthor="Gunther" pageId="16" pageNumber="17" pagination="74 - 79" refId="ref13686" refString="Gunther, A. (1879) List of the Mammals, Reptiles, and Batrachians sent by Mr. Everett from the Philippine Islands. Proceedings of the Zoological Society, London, January 14: 74 - 79. Plate IV - Dendrophis philippinensis Gunther, 1879 - Norte de Mindanao. Del. R. Mintern." type="journal article" year="1879">Günther, 1879</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3269BCE1E0E8F562E86F9D3DF34F8CF" pageId="16" pageNumber="17" type="description">
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<paragraph id="8B83C8451E0E8F562E86F9D3DA25F9F3" blockId="16.[151,675,1511,1570]" box="[151,290,1546,1570]" pageId="16" pageNumber="17">
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<figureCitation id="1307D4C01E0E8F562E86F9D3DBF4F9F3" box="[151,243,1546,1570]" captionStart="FIGURE 12" captionStartId="17.[151,249,1323,1346]" captionTargetBox="[151,1435,445,1300]" captionTargetId="figure@17.[151,1436,445,1302]" captionTargetPageId="17" captionText="FIGURE 12. Dendrelaphis philippinensis Günther, 1879, adult from Mindanao, holotype (BMNH 1946.1.6.69; TL 106 cm.)." httpUri="https://zenodo.org/record/215467/files/figure.png" pageId="16" pageNumber="17">Figs. 12</figureCitation>
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,
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<figureCitation id="1307D4C01E0E8F562EEEF9D3DA19F9F3" box="[255,286,1546,1570]" captionStart="FIGURE 13" captionStartId="17.[151,249,1980,2003]" captionTargetBox="[151,1435,1395,1958]" captionTargetId="figure@17.[151,1436,1395,1959]" captionTargetPageId="17" captionText="FIGURE 13. Dendrelaphis philippinensis Günther, 1879, adult from Catanduanes (USNM 319029; TL 103.5)." httpUri="https://zenodo.org/record/215468/files/figure.png" pageId="16" pageNumber="17">13</figureCitation>
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.
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</paragraph>
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<paragraph id="8B83C8451E0E8F562E86F98BDF34F8CF" blockId="16.[151,1436,1618,2002]" pageId="16" pageNumber="17">
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<emphasis id="B94814571E0E8F562E86F98BDA2EF9BA" bold="true" box="[151,297,1618,1643]" pageId="16" pageNumber="17">Description.</emphasis>
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A species of the
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<taxonomicName id="4C3CB3C61E0E8F562FF8F98BD823F9BB" box="[489,804,1618,1642]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="caudolineatus">
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<emphasis id="B94814571E0E8F562FF8F98BD823F9BB" box="[489,804,1618,1642]" italics="true" pageId="16" pageNumber="17">Dendrelaphis caudolineatus</emphasis>
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</taxonomicName>
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group, characterized by the combination of 1) a reduced pattern of black longitudinal stripes, 2–6 black longitudinal stripes at midbody; 2) a pale ventrolateral line is absent or faintly present; 3) a postocular stripe is present and extends onto the neck; 4) 162–170 ventrals (males), 161–172 [179] ventrals (females); 5) 94–108 subcaudals; 6) relative tail-length 0.268–0.298 (males), 0.252–0.290 (females); 7) 6–7 temporal scales on each side; 8) 9–11 infralabials on each side; 9) 2–5 scales touching the posterior borders of the parietal scales; 10) maximum total length males 89.0 cm, females
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<quantity id="4CC465A01E0E8F562DD5F8DFDF28F8CF" box="[964,1071,1798,1822]" metricMagnitude="0" metricUnit="m" metricValue="1.1949999999999998" pageId="16" pageNumber="17" unit="cm" value="119.5">119.5 cm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3269BCE1E0E8F572ED6F8F3D822FEEE" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="8B83C8451E0E8F572ED6F8F3D822FEEE" blockId="16.[151,1436,1618,2002]" lastBlockId="17.[151,1436,151,391]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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<emphasis id="B94814571E0E8F562ED6F8F3DAEEF892" bold="true" box="[199,489,1834,1859]" pageId="16" pageNumber="17">
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Description of
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<typeStatus id="548776E71E0E8F562F93F8F2DAEEF892" box="[386,489,1835,1859]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
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</emphasis>
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.
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<materialsCitation id="3B54C2181E0E8F562FECF8F2D919F8B7" ID-GBIF-Occurrence="3753898302" collectionCode="BMNH" collectorName="A. Everett" location="North Mindanao" pageId="16" pageNumber="17" specimenCode="BMNH 1946.1.6.69" specimenCount="1">
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<specimenCode id="DB9A603E1E0E8F562FECF8F2D9E4F893" box="[509,739,1834,1858]" pageId="16" pageNumber="17">BMNH 1946.1.6.69</specimenCode>
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(formerly
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<collectionCode id="ED2D50801E0E8F562D79F8F2D8BBF893" box="[872,956,1835,1858]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="16" pageNumber="17">BMNH</collectionCode>
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1877.10.9.62),
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<specimenCount id="9D3A03CC1E0E8F562A68F8F3DE0DF893" box="[1145,1290,1834,1858]" pageId="16" pageNumber="17" type="adult">adult female</specimenCount>
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from
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<location id="8EE39E9E1E0E8F562B4AF8F3DA0EF8B7" LSID="urn:lsid:plazi:treatment:039579531E0E8F542E86FA3ED8FDFE1E:8EE39E9E1E0E8F562B4AF8F3DA0EF8B7" name="North Mindanao" pageId="16" pageNumber="17">North Mindanao</location>
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collected by
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<collectorName id="26C9AD931E0E8F562FB7F896D919F8B7" box="[422,542,1871,1895]" pageId="16" pageNumber="17">A. Everett</collectorName>
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</materialsCitation>
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; SVL 76.0 cm; TAIL 30.0 cm; 164 ventral scales; 100 subcaudal scales; anal shield divided; dorsal scales in
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<date id="FF82EE851E0E8F562FE5F8ABD95EF85B" box="[500,601,1906,1930]" pageId="16" pageNumber="17" value="1913-11-13">13-13-11</date>
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rows; 9 supralabials, supralabials 4,5,6 (L) 5,6 (R) touch the eye; 10 (L), 9 (R) infralalabials, infralabials 6 and 7 touch first sublabial; infralabials 1–5 touch first chinshield, infralabials 5,6 touch second chinshield; 2 postoculars; temporal formula 2:2:2/2:2:2; vertebral scales not enlarged; eye-diameter
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<quantity id="4CC465A01E0F8F572E86FF4EDBF5FF7E" box="[151,242,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="17" pageNumber="18" unit="mm" value="5.4">5.4 mm</quantity>
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; eye-nostril distance
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<quantity id="4CC465A01E0F8F572FF5FF4ED938FF7E" box="[484,575,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="17" pageNumber="18" unit="mm" value="5.8">5.8 mm</quantity>
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; snout-width
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<quantity id="4CC465A01E0F8F572CCBFF4ED832FF7E" box="[730,821,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="17" pageNumber="18" unit="mm" value="6.2">6.2 mm</quantity>
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; posterior edge of parietal scales touched by 3 scales; head-length
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<quantity id="4CC465A01E0F8F572F33FF65DA8DFF05" box="[290,394,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" pageId="17" pageNumber="18" unit="mm" value="29.5">29.5 mm</quantity>
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; ground-colour olive-brown; supralabials, chin, throat yellow; venter yellow; black temporal stripe covers more than half of temporal region, extends onto neck where it fades away; 4 black stripes anteriorly (including extensions of temporal stripes), disappear towards middle of body, reappear posteriorly; faint ventrolateral line present; black stripe on underside of tail.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3269BCE1E0F8F572ED6FE92DAD3FE56" pageId="17" pageNumber="18" type="description">
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<paragraph id="8B83C8451E0F8F572ED6FE92DAD3FE56" blockId="17.[151,1436,151,391]" pageId="17" pageNumber="18">
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<emphasis id="B94814571E0F8F572ED6FE92DAA8FEB5" bold="true" box="[199,431,331,356]" pageId="17" pageNumber="18">Sexual dimorphism</emphasis>
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. Females grow larger than males (see description). None of the other examined characters exhibits sexual dimorphism.
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</paragraph>
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</subSubSection>
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<caption id="DF4398CD1E0F8F572E86FAF2DE88FA90" box="[151,1423,1323,1346]" httpUri="https://zenodo.org/record/215467/files/figure.png" pageId="17" pageNumber="18" targetBox="[151,1435,445,1300]" targetPageId="17">
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<paragraph id="8B83C8451E0F8F572E86FAF2DE88FA90" blockId="17.[151,1423,1323,1346]" box="[151,1423,1323,1346]" pageId="17" pageNumber="18">
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<emphasis id="B94814571E0F8F572E86FAF2DA18FA91" bold="true" box="[151,287,1323,1346]" pageId="17" pageNumber="18">FIGURE 12.</emphasis>
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<taxonomicName id="4C3CB3C61E0F8F572F34FAF2D9DBFA90" authority="Gunther, 1879" authorityName="Gunther" authorityYear="1879" box="[293,732,1323,1345]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="philippinensis">
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<emphasis id="B94814571E0F8F572F34FAF2D945FA90" box="[293,578,1323,1345]" italics="true" pageId="17" pageNumber="18">Dendrelaphis philippinensis</emphasis>
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<bibRefCitation id="EFADB5B41E0F8F572C58FAF2D9DBFA90" author="Gunther" box="[585,732,1323,1345]" firstAuthor="Gunther" pageId="17" pageNumber="18" pagination="74 - 79" refId="ref13686" refString="Gunther, A. (1879) List of the Mammals, Reptiles, and Batrachians sent by Mr. Everett from the Philippine Islands. Proceedings of the Zoological Society, London, January 14: 74 - 79. Plate IV - Dendrophis philippinensis Gunther, 1879 - Norte de Mindanao. Del. R. Mintern." type="journal article" year="1879">Günther, 1879</bibRefCitation>
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</taxonomicName>
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, adult from Mindanao, holotype (BMNH 1946.1.6.69; TL 106 cm.).
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</paragraph>
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</caption>
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<caption id="DF4398CD1E0F8F572E86F865DFFAF803" box="[151,1277,1980,2003]" httpUri="https://zenodo.org/record/215468/files/figure.png" pageId="17" pageNumber="18" targetBox="[151,1435,1395,1958]" targetPageId="17">
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<paragraph id="8B83C8451E0F8F572E86F865DFFAF803" blockId="17.[151,1277,1980,2003]" box="[151,1277,1980,2003]" pageId="17" pageNumber="18">
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<emphasis id="B94814571E0F8F572E86F865DA18F803" bold="true" box="[151,287,1980,2003]" pageId="17" pageNumber="18">FIGURE 13.</emphasis>
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<taxonomicName id="4C3CB3C61E0F8F572F34F864D9DBF803" authority="Gunther, 1879" authorityName="Gunther" authorityYear="1879" box="[293,732,1980,2003]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="philippinensis">
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<emphasis id="B94814571E0F8F572F34F864D945F802" box="[293,578,1981,2003]" italics="true" pageId="17" pageNumber="18">Dendrelaphis philippinensis</emphasis>
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<bibRefCitation id="EFADB5B41E0F8F572C58F865D9DBF803" author="Gunther" box="[585,732,1980,2002]" firstAuthor="Gunther" pageId="17" pageNumber="18" pagination="74 - 79" refId="ref13686" refString="Gunther, A. (1879) List of the Mammals, Reptiles, and Batrachians sent by Mr. Everett from the Philippine Islands. Proceedings of the Zoological Society, London, January 14: 74 - 79. Plate IV - Dendrophis philippinensis Gunther, 1879 - Norte de Mindanao. Del. R. Mintern." type="journal article" year="1879">Günther, 1879</bibRefCitation>
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</taxonomicName>
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, adult from Catanduanes (USNM 319029; TL 103.5).
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</paragraph>
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</caption>
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<subSubSection id="C3269BCE1E0C8F542ED6FF4ED88AFE56" pageId="18" pageNumber="19" type="biology_ecology">
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<paragraph id="8B83C8451E0C8F542ED6FF4ED88AFE56" blockId="18.[151,1436,151,463]" pageId="18" pageNumber="19">
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<emphasis id="B94814571E0C8F542ED6FF4EDACFFF61" bold="true" box="[199,456,151,176]" pageId="18" pageNumber="19">Geographic variation</emphasis>
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. In spite of the large number of geographically isolated populations,
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<taxonomicName id="4C3CB3C61E0C8F542ACAFF40DE9CFF61" box="[1243,1435,152,176]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="philippinensis">
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<emphasis id="B94814571E0C8F542ACAFF40DE9CFF61" box="[1243,1435,152,176]" italics="true" pageId="18" pageNumber="19">D. philippinensis</emphasis>
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</taxonomicName>
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appears to exhibit little inter-population variation in its morphology. However, populations do differ in their coloration. For instance, in the population from Mindanao (
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<figureCitation id="1307D4C01E0C8F542D5BFF06D8A7FF26" box="[842,928,223,247]" captionStart="FIGURE 12" captionStartId="17.[151,249,1323,1346]" captionTargetBox="[151,1435,445,1300]" captionTargetId="figure@17.[151,1436,445,1302]" captionTargetPageId="17" captionText="FIGURE 12. Dendrelaphis philippinensis Günther, 1879, adult from Mindanao, holotype (BMNH 1946.1.6.69; TL 106 cm.)." httpUri="https://zenodo.org/record/215467/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
|
||
), the pattern of longitudinal black stripes is reduced to two narrow stripes that occupy little more than the outer edges of the ventral scales whereas the populations from South Luzon and Catanduanes (
|
||
<figureCitation id="1307D4C01E0C8F542CDDFEFED818FEEE" box="[716,799,295,319]" captionStart="FIGURE 13" captionStartId="17.[151,249,1980,2003]" captionTargetBox="[151,1435,1395,1958]" captionTargetId="figure@17.[151,1436,1395,1959]" captionTargetPageId="17" captionText="FIGURE 13. Dendrelaphis philippinensis Günther, 1879, adult from Catanduanes (USNM 319029; TL 103.5)." httpUri="https://zenodo.org/record/215468/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
|
||
) exhibit comparatively pronounced longitudinal stripes. This may be taxonomically significant. On the other hand, differences in the degree to which the pattern of longitudinal stripes is manifest, were also noted within populations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3269BCE1E0C8F542ED6FE4AD8FDFE1E" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="8B83C8451E0C8F542ED6FE4AD8FDFE1E" blockId="18.[151,1436,151,463]" pageId="18" pageNumber="19">
|
||
<emphasis id="B94814571E0C8F542ED6FE4ADA5EFE7D" bold="true" box="[199,345,403,428]" pageId="18" pageNumber="19">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="4C3CB3C61E0C8F542F7BFE4DD9AEFE7D" box="[362,681,404,428]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="philippinensis">
|
||
<emphasis id="B94814571E0C8F542F7BFE4DD9AEFE7D" box="[362,681,404,428]" italics="true" pageId="18" pageNumber="19">Dendrelaphis philippinensis</emphasis>
|
||
</taxonomicName>
|
||
inhabits the
|
||
<collectingCountry id="F32B88D51E0C8F542D50FE4DDF0BFE7D" box="[833,1036,404,428]" name="Philippines" pageId="18" pageNumber="19">Philippine islands</collectingCountry>
|
||
Basilan, Mindanao, Samar, Leyte, Bohol, Cebu, Polillo, Kalotkot, Catanduanes as well as South Luzon (Albay).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |