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<mods:title id="F8995E855F6FEE010885522CC9084C45">Synonymic notes in the Southeast Asian genus Lutera Westwood, 1875 (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
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<mods:namePart id="2DA464364C9984EF8F8E69618912B0F7">Xu, Hao</mods:namePart>
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<mods:namePart id="CEC5F69A5376A8961EB916A2F0DFA6EE">Zhao, Ming-Zhi</mods:namePart>
<mods:affiliation id="C4866A158B8CDA80824C8B68E66B5795">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, P. R. China.</mods:affiliation>
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<paragraph id="8B82814FC744A30DFF4DF8DEFE5C325E" blockId="7.[151,596,1794,1857]" box="[151,484,1794,1821]" pageId="7" pageNumber="538">
<heading id="D0CA3623C744A30DFF4DF8DEFE5C325E" bold="true" box="[151,484,1794,1821]" fontSize="11" level="1" pageId="7" pageNumber="538" reason="1">
<taxonomicName id="4C3DFACCC744A30DFF4DF8DEFE5C325E" ID-CoL="3WG3V" authority="Ohaus, 1930" authorityName="Ohaus" authorityYear="1930" box="[151,484,1794,1821]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B9495D5DC744A30DFF4DF8DEFE5C325E" bold="true" box="[151,484,1794,1821]" pageId="7" pageNumber="538">
<emphasis id="B9495D5DC744A30DFF4DF8DEFEFA325E" bold="true" box="[151,322,1794,1821]" italics="true" pageId="7" pageNumber="538">Lutera nigrita</emphasis>
<bibRefCitation id="EFACFCBEC744A30DFE93F8DFFE5C325E" author="Ohaus, F." box="[329,484,1794,1820]" pageId="7" pageNumber="538" pagination="138 - 158" refId="ref7844" refString="Ohaus, F. (1930) XXVI. Beitrag zur Kenntnis der Ruteliden (Col. Lamell.). Deutsche Entomologische Zeitschrift, 1930, 138 - 158. https: // doi. org / 10.1002 / mmnd. 48019300205" type="journal article" year="1930">Ohaus, 1930</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C327D2C4C744A30DFF4DF8FBFDEC3202" box="[151,596,1830,1857]" pageId="7" pageNumber="538" type="description">
<paragraph id="8B82814FC744A30DFF4DF8FBFDEC3202" blockId="7.[151,596,1794,1857]" box="[151,596,1830,1857]" pageId="7" pageNumber="538">
(
<figureCitation id="13069DCAC744A30DFF45F8FBFE903203" box="[159,296,1830,1857]" captionStart-0="FIGURES 1724" captionStart-1="FIGURES 2530" captionStartId-0="5.[151,264,1568,1592]" captionStartId-1="7.[151,264,1660,1684]" captionTargetBox-0="[232,1377,193,1522]" captionTargetBox-1="[190,1392,182,1629]" captionTargetId-0="figure-18@5.[202,1384,181,1544]" captionTargetId-1="figure-18@7.[183,1404,178,1638]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionText-0="FIGURES 1724. Type specimens of Lutera spp. 1718, female holotype of L. nigromaculata Ohaus, 1900 and labels (©MNHN); 1920, inauthentic type of L. nigromaculata in MFNB (photograph by Bernd Jäger); 2124, female holotype of L. nigrita Ohaus, 1930 (in dorsal and lateral view), head, and labels (photograph by Bernd Jäger, MFNB)." captionText-1="FIGURES 2530. Lutera nigrita Ohaus, 1930 with different color patterns.25, 27, Dingalan; 26, Don Salvador; 28, Compostela; 29, Kapatagan; 30, Wao." figureDoi-0="http://doi.org/10.5281/zenodo.13949866" figureDoi-1="http://doi.org/10.5281/zenodo.13949868" httpUri-0="https://zenodo.org/record/13949866/files/figure.png" httpUri-1="https://zenodo.org/record/13949868/files/figure.png" pageId="7" pageNumber="538">Figs. 1930</figureCitation>
,
<figureCitation id="13069DCAC744A30DFEE9F8FBFD9B3203" box="[307,547,1830,1857]" captionStart="FIGURES 3145" captionStartId="8.[151,264,1924,1948]" captionTargetBox="[189,1398,275,1888]" captionTargetId="figure-77@8.[176,1411,260,1900]" captionTargetPageId="8" captionText="FIGURES 3145. Male genitalia of Lutera species (3135, dorsal view; 3640, lateral view; 4145, ventral view;). 31, 36, 41, L. nigromaculata Ohaus, 1900 from Thailand; 3235, 3740, 4235, L. nigrita Ohaus, 1930: 32, 37, 42, from Luzon Island; 33, 38, 43, from Mindoro Island; 34, 39, 44, from Negros Island; 35, 40, 45, from Mindanao Island" figureDoi="http://doi.org/10.5281/zenodo.13949870" httpUri="https://zenodo.org/record/13949870/files/figure.png" pageId="7" pageNumber="538">3235, 3740, 4245</figureCitation>
,
<figureCitation id="13069DCAC744A30DFDF5F8FBFDF43202" box="[559,588,1830,1856]" captionStart="FIGURE 46" captionStartId="9.[151,250,1970,1994]" captionTargetBox="[164,1423,1173,1947]" captionTargetId="figure-488@9.[164,1423,1173,1947]" captionTargetPageId="9" captionText="FIGURE 46. Known distribution of Lutera nigromaculata Ohaus, 1900 (black dots) and L. nigrita Ohaus, 1930 (black triangles). Type localities are underlined, and the open dot in Nepal is a record from Wada (1998) without precise location." figureDoi="http://doi.org/10.5281/zenodo.13949872" httpUri="https://zenodo.org/record/13949872/files/figure.png" pageId="7" pageNumber="538">46</figureCitation>
)
</paragraph>
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<paragraph id="8B82814FC744A30DFF4DF8B5FD6732FE" blockId="7.[151,1437,1895,2042]" pageId="7" pageNumber="538">
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<taxonomicName id="4C3DFACCC744A30DFF4DF8B5FECD323D" ID-CoL="3WG3W" authorityName="Ohaus" authorityYear="1900" box="[151,373,1895,1919]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC744A30DFF4DF8B5FECD323D" box="[151,373,1895,1919]" italics="true" pageId="7" pageNumber="538">Lutera nigromaculata</emphasis>
</taxonomicName>
(nec
<bibRefCitation id="EFACFCBEC744A30DFE74F8BAFD8B323D" author="Ohaus, F." box="[430,563,1895,1919]" pageId="7" pageNumber="538" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus, 1900</bibRefCitation>
):
<treatmentCitation id="0A9CA75EC744A30DFD9FF8BAFD5E323D" author="Ohaus, F." box="[581,742,1895,1919]" page="97" pageId="7" pageNumber="538" year="1905">
<bibRefCitation id="EFACFCBEC744A30DFD9FF8BAFD5E323D" author="Ohaus, F." box="[581,742,1895,1919]" pageId="7" pageNumber="538" pagination="81 - 99" refId="ref7756" refString="Ohaus, F. (1905) Beitrage zur Kenntnis der Ruteliden. Deutsche Entomologische Zeitschrift, 1905, 81 - 99. https: // doi. org / 10.1002 / mmnd. 48119050214" type="journal article" year="1905">Ohaus 1905: 97</bibRefCitation>
</treatmentCitation>
(a male from Camarines,
<collectingCountry id="F32AC1DFC744A30DFC2AF8BAFBDF323D" box="[1008,1127,1895,1919]" name="Philippines" pageId="7" pageNumber="538">Philippines</collectingCountry>
);
<treatmentCitation id="0A9CA75EC744A30DFBA2F8BAFAA1323D" author="Ohaus, F." box="[1144,1305,1895,1919]" page="39" pageId="7" pageNumber="538" year="1918">
<bibRefCitation id="EFACFCBEC744A30DFBA2F8BAFAA1323D" author="Ohaus, F." box="[1144,1305,1895,1919]" pageId="7" pageNumber="538" pagination="1 - 241" refId="ref7791" refString="Ohaus, F. (1918) Scarabaeidae: Phaenomerinae, Euchirinae, Rutelinae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum catalogus. Vo. XX. Pars 66. W. Junk, Berlin, pp. 1 - 241." type="book chapter" year="1918">Ohaus 1918: 39</bibRefCitation>
</treatmentCitation>
(catalogued, partim);
<treatmentCitation id="0A9CA75EC744A30DFEF9F85BFE6E32DC" author="Ohaus, F." box="[291,470,1926,1950]" page="112" pageId="7" pageNumber="538" year="1934">
<bibRefCitation id="EFACFCBEC744A30DFEF9F85BFE6E32DC" author="Ohaus, F." box="[291,470,1926,1950]" pageId="7" pageNumber="538" pagination="1 - 172" refId="ref7920" refString="Ohaus, F. (1934) Coleoptera Lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. Erster Teil. In: Wytsman, P. (Ed.), Genera Insectorum. Coleoptera. Fascicule 199 A. Louis Desmet-Verteneuil, Bruxelles, pp. 1 - 172." type="book chapter" year="1934">Ohaus 1934: 112</bibRefCitation>
</treatmentCitation>
(catalogued, partim);
<treatmentCitation id="0A9CA75EC744A30DFD1BF85BFCCD32DC" author="Kuijten, P. J." box="[705,885,1926,1950]" page="87" pageId="7" pageNumber="538" year="1988">
<bibRefCitation id="EFACFCBEC744A30DFD1BF85BFCCD32DC" author="Kuijten, P. J." box="[705,885,1926,1950]" pageId="7" pageNumber="538" pagination="75 - 90" refId="ref7400" refString="Kuijten, P. J. (1988) Rutelarcha, Lutera and Cyphelytra: notes on taxonomy and nomenclature, diagnoses, and keys (Coleoptera: Rutelinae, Rutelini). Zoologische Mededelingen, 62 (5 - 9), 75 - 90." type="journal article" year="1988">Kuijten 1988: 87</bibRefCitation>
</treatmentCitation>
, figs. 2024 [redescription and illustration based on specimens from the
<collectingCountry id="F32AC1DFC744A30DFE4EF879FDB432FE" box="[404,524,1956,1980]" name="Philippines" pageId="7" pageNumber="538">Philippines</collectingCountry>
]. (misidentification).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B82814FC744A30DFF4DF819FEF532B8" blockId="7.[151,1437,1895,2042]" pageId="7" pageNumber="538">
<treatmentCitationGroup id="AB2DA661C744A30DFF4DF819FEF532B8" pageId="7" pageNumber="538">
<treatmentCitation id="0A9CA75EC744A30DFF4DF819FD0F3299" author="Ohaus, F." box="[151,695,1987,2011]" page="140" pageId="7" pageNumber="538" year="1930">
<taxonomicName id="4C3DFACCC744A30DFF4DF819FD0F3299" ID-CoL="3WG3V" authority="Ohaus, 1930: 140" authorityName="Ohaus" authorityPageNumber="140" authorityYear="1930" box="[151,695,1987,2011]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="variety" species="nigromaculata" variety="nigrita">
<emphasis id="B9495D5DC744A30DFF4DF819FEC03299" box="[151,376,1987,2011]" italics="true" pageId="7" pageNumber="538">Lutera nigromaculata</emphasis>
var.
<emphasis id="B9495D5DC744A30DFE6AF819FE4D3299" box="[432,501,1988,2011]" italics="true" pageId="7" pageNumber="538">nigrita</emphasis>
<bibRefCitation id="EFACFCBEC744A30DFE24F81EFD0F3299" author="Ohaus, F." box="[510,695,1987,2011]" pageId="7" pageNumber="538" pagination="138 - 158" refId="ref7844" refString="Ohaus, F. (1930) XXVI. Beitrag zur Kenntnis der Ruteliden (Col. Lamell.). Deutsche Entomologische Zeitschrift, 1930, 138 - 158. https: // doi. org / 10.1002 / mmnd. 48019300205" type="journal article" year="1930">Ohaus, 1930: 140</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54863FEDC744A30DFD1DF81EFD4C3299" box="[711,756,1987,2011]" pageId="7" pageNumber="538">type</typeStatus>
locality: Mt. Banahao, Luzon Is.,
<collectingCountry id="F32AC1DFC744A30DFBBAF81EFB6F3299" box="[1120,1239,1987,2011]" name="Philippines" pageId="7" pageNumber="538">Philippines</collectingCountry>
);
<treatmentCitation id="0A9CA75EC744A30DFB31F81EFA243299" author="Ohaus, F." box="[1259,1436,1987,2011]" page="112" pageId="7" pageNumber="538" year="1934">
<bibRefCitation id="EFACFCBEC744A30DFB31F81EFA243299" author="Ohaus, F." box="[1259,1436,1987,2011]" pageId="7" pageNumber="538" pagination="1 - 172" refId="ref7920" refString="Ohaus, F. (1934) Coleoptera Lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. Erster Teil. In: Wytsman, P. (Ed.), Genera Insectorum. Coleoptera. Fascicule 199 A. Louis Desmet-Verteneuil, Bruxelles, pp. 1 - 172." type="book chapter" year="1934">Ohaus 1934: 112</bibRefCitation>
</treatmentCitation>
(catalogued).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B82814FC74BA302FF4DFF4AFBA235ED" blockId="8.[151,1436,150,236]" box="[151,1050,150,175]" pageId="8" pageNumber="539">
<treatmentCitationGroup id="AB2DA661C74BA302FF4DFF4AFBA235ED" box="[151,1050,150,175]" pageId="8" pageNumber="539">
<taxonomicName id="4C3DFACCC74BA302FF4DFF4AFE1435ED" authority="(Ohaus)" baseAuthorityName="Ohaus" box="[151,428,151,175]" class="Insecta" family="Scarabaeidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B9495D5DC74BA302FF4DFF4AFEE835EC" box="[151,336,151,174]" italics="true" pageId="8" pageNumber="539">Parastasia nigrita</emphasis>
(Ohaus)
</taxonomicName>
:
<treatmentCitation id="0A9CA75EC74BA302FE6CFF4AFD2335EC" author="Machatschke, J. W." box="[438,667,150,175]" page="45" pageId="8" pageNumber="539" year="1972">
<bibRefCitation id="EFACFCBEC74BA302FE6CFF4AFD2335EC" author="Machatschke, J. W." box="[438,667,150,175]" pageId="8" pageNumber="539" pagination="1 - 361" refId="ref7615" refString="Machatschke, J. W. (1972) Scarabaeoidea: Melolonthidae Rutelinae. In: Wilcox, J. A. (Ed.), Coleopterorum Catalogus, Supplementa. Pars 66. Fascicle I. Editio Secunda. W. Junk, The Hague, pp. 1 - 361." type="book chapter" year="1972">Machatschke 1972: 45</bibRefCitation>
</treatmentCitation>
(catalogued, raised to species status).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B82814FC74BA302FF4DFF6BFDC435AE" blockId="8.[151,1436,150,236]" pageId="8" pageNumber="539">
<treatmentCitationGroup id="AB2DA661C74BA302FF4DFF6BFDC435AE" pageId="8" pageNumber="539">
<taxonomicName id="4C3DFACCC74BA302FF4DFF6BFED6358F" ID-CoL="3WG3V" authority="Ohaus" authorityName="Ohaus" authorityYear="1930" box="[151,366,181,205]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B9495D5DC74BA302FF4DFF6BFE9C358F" box="[151,292,182,205]" italics="true" pageId="8" pageNumber="539">Lutera nigrita</emphasis>
Ohaus
</taxonomicName>
:
<treatmentCitation id="0A9CA75EC74BA302FEAFFF68FDAF358F" author="Wada, K." box="[373,535,181,205]" page="360" pageId="8" pageNumber="539" year="1998">
<bibRefCitation id="EFACFCBEC74BA302FEAFFF68FDAF358F" author="Wada, K." box="[373,535,181,205]" pageId="8" pageNumber="539" pagination="359 - 367" refId="ref8045" refString="Wada, K. (1998) Two new species of the genus Lutera (Coleoptera, Scarabaeidae, Rutelinae) from Borneo and Sumatra. Elytra, 26 (2), 359 - 367." type="journal article" year="1998">Wada 1998: 360</bibRefCitation>
</treatmentCitation>
(in key), figs. 59 habitus;
<treatmentCitation id="0A9CA75EC74BA302FCFFFF68FC75358F" author="Krajcik, M." box="[805,973,181,205]" page="79" pageId="8" pageNumber="539" year="2007">
<bibRefCitation id="EFACFCBEC74BA302FCFFFF68FC75358F" author="Krajcik, M." box="[805,973,181,205]" pageId="8" pageNumber="539" pagination="1 - 139" refId="ref7338" refString="Krajcik, M. (2007) Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. x Supplementu m, 4, 1 - 139." type="journal article" year="2007">Krajčík 2007: 79</bibRefCitation>
</treatmentCitation>
(catalogued);
<treatmentCitation id="0A9CA75EC74BA302FB87FF68FAAA358F" author="Krajcik, M." box="[1117,1298,181,205]" page="150" pageId="8" pageNumber="539" year="2012">
<bibRefCitation id="EFACFCBEC74BA302FB87FF68FAAA358F" author="Krajcik, M." box="[1117,1298,181,205]" pageId="8" pageNumber="539" pagination="1 - 139" refId="ref7376" refString="Krajcik, M. (2012) Checklist of the world Scarabaeoidea. Animma. x Supplementum, 5, 1 - 139." type="journal article" year="2012">Krajčík 2012: 150</bibRefCitation>
</treatmentCitation>
(catalogued);
<treatmentCitation id="0A9CA75EC74BA302FF1DFF09FEEC35AE" author="Wada, K." box="[199,340,212,236]" page="9" pageId="8" pageNumber="539" year="2015">
<bibRefCitation id="EFACFCBEC74BA302FF1DFF09FEEC35AE" author="Wada, K." box="[199,340,212,236]" pageId="8" pageNumber="539" pagination="9 - 11" refId="ref8082" refString="Wada, K. (2015) Records of Lutera boettcheri for the first time in 99 years after its discovery. Saikaku-Tsushin, 31, 9 - 11." type="journal article" year="2015">Wada 2015: 9</bibRefCitation>
</treatmentCitation>
(figs. 812 color variations).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF42D1C7C74BA302FF4DF859FBE632A6" ID-DOI="http://doi.org/10.5281/zenodo.13949870" ID-Zenodo-Dep="13949870" httpUri="https://zenodo.org/record/13949870/files/figure.png" pageId="8" pageNumber="539" startId="8.[151,264,1924,1948]" targetBox="[189,1398,275,1888]" targetPageId="8" targetType="figure">
<paragraph id="8B82814FC74BA302FF4DF859FBE632A6" blockId="8.[151,1436,1924,2021]" pageId="8" pageNumber="539">
<emphasis id="B9495D5DC74BA302FF4DF859FEEC32DE" bold="true" box="[151,340,1924,1948]" pageId="8" pageNumber="539">FIGURES 3145.</emphasis>
Male genitalia of
<taxonomicName id="4C3DFACCC74BA302FDD6F858FD2732DE" box="[524,671,1924,1948]" pageId="8" pageNumber="539">
<emphasis id="B9495D5DC74BA302FDD6F858FDE832DE" box="[524,592,1925,1948]" italics="true" pageId="8" pageNumber="539">Lutera</emphasis>
species
</taxonomicName>
(3135, dorsal view; 3640, lateral view; 4145, ventral view;).
<emphasis id="B9495D5DC74BA302FAEBF859FA2432DE" bold="true" box="[1329,1436,1924,1948]" pageId="8" pageNumber="539">31, 36, 41,</emphasis>
<taxonomicName id="4C3DFACCC74BA302FF4DF874FE743282" authority="Ohaus, 1900" authorityName="Ohaus" authorityYear="1900" box="[151,460,1960,1984]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC74BA302FF4DF874FEFE3282" box="[151,326,1960,1984]" italics="true" pageId="8" pageNumber="539">L. nigromaculata</emphasis>
<bibRefCitation id="EFACFCBEC74BA302FE96F875FE743282" author="Ohaus, F." box="[332,460,1960,1984]" pageId="8" pageNumber="539" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus, 1900</bibRefCitation>
</taxonomicName>
from Thailand;
<emphasis id="B9495D5DC74BA302FDB5F875FCF43282" bold="true" box="[623,844,1960,1984]" pageId="8" pageNumber="539">3235, 3740, 4235,</emphasis>
<taxonomicName id="4C3DFACCC74BA302FC88F874FBE63282" authority="Ohaus, 1930: 32" authorityName="Ohaus" authorityPageNumber="32" authorityYear="1930" box="[850,1118,1960,1984]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B9495D5DC74BA302FC88F874FC093282" box="[850,945,1961,1984]" italics="true" pageId="8" pageNumber="539">L. nigrita</emphasis>
<bibRefCitation id="EFACFCBEC74BA302FC6DF875FBE63282" author="Ohaus, F." box="[951,1118,1960,1984]" pageId="8" pageNumber="539" pagination="138 - 158" refId="ref7844" refString="Ohaus, F. (1930) XXVI. Beitrag zur Kenntnis der Ruteliden (Col. Lamell.). Deutsche Entomologische Zeitschrift, 1930, 138 - 158. https: // doi. org / 10.1002 / mmnd. 48019300205" type="journal article" year="1930">
Ohaus, 1930:
<emphasis id="B9495D5DC74BA302FB99F875FBE63282" bold="true" box="[1091,1118,1960,1984]" pageId="8" pageNumber="539">32</emphasis>
</bibRefCitation>
</taxonomicName>
<emphasis id="B9495D5DC74BA302FB84F875FB163282" bold="true" box="[1118,1198,1960,1984]" pageId="8" pageNumber="539">, 37, 42,</emphasis>
from Luzon Island;
<emphasis id="B9495D5DC74BA302FAA7F875FF6532A6" bold="true" pageId="8" pageNumber="539">33, 38, 43,</emphasis>
from Mindoro Island;
<emphasis id="B9495D5DC74BA302FE1FF811FD8932A6" bold="true" box="[453,561,1996,2020]" pageId="8" pageNumber="539">34, 39, 44,</emphasis>
from Negros Island;
<emphasis id="B9495D5DC74BA302FCD3F811FCCD32A6" bold="true" box="[777,885,1996,2020]" pageId="8" pageNumber="539">35, 40, 45,</emphasis>
from Mindanao Island
</paragraph>
</caption>
<subSubSection id="C327D2C4C74AA303FF4DFF4AFDA7313F" pageId="9" pageNumber="540" type="materials_examined">
<paragraph id="8B82814FC74AA303FF4DFF4AFE6735BB" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="540">
<emphasis id="B9495D5DC74AA303FF4DFF4AFE7E35F2" bold="true" box="[151,454,150,177]" pageId="9" pageNumber="540">Type material examined.</emphasis>
<taxonomicName id="4C3DFACCC74AA303FE0AFF4AFC4835F3" authority="Ohaus, 1930" authorityName="Ohaus" authorityYear="1930" box="[464,1008,151,177]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="variety" species="nigromaculata" variety="nigrita">
<emphasis id="B9495D5DC74AA303FE0AFF4AFD7035F3" box="[464,712,151,177]" italics="true" pageId="9" pageNumber="540">Lutera nigromaculata</emphasis>
<materialsCitation id="3B558B12C74AA303FD08FF4AFC4835F3" box="[722,1008,151,177]" collectingDate="1914-06-16" collectionCode="MFNB" collectorName="G. Boettcher" location="Mt. Banahao" municipality="Luzon" pageId="9" pageNumber="540" specimenCount="2" specimenCount-female="2" stateProvince="Berlin" typeStatus="holotype">
var.
<emphasis id="B9495D5DC74AA303FCDCFF4AFCEA35F2" box="[774,850,151,176]" italics="true" pageId="9" pageNumber="540">nigrita</emphasis>
<bibRefCitation id="EFACFCBEC74AA303FC86FF4AFC4835F3" author="Ohaus, F." box="[860,1008,151,177]" pageId="9" pageNumber="540" pagination="138 - 158" refId="ref7844" refString="Ohaus, F. (1930) XXVI. Beitrag zur Kenntnis der Ruteliden (Col. Lamell.). Deutsche Entomologische Zeitschrift, 1930, 138 - 158. https: // doi. org / 10.1002 / mmnd. 48019300205" type="journal article" year="1930">Ohaus, 1930</bibRefCitation>
</materialsCitation>
</taxonomicName>
,
<emphasis id="B9495D5DC74AA303FC24FF4AFBDD35F3" bold="true" box="[1022,1125,151,177]" pageId="9" pageNumber="540">
<typeStatus id="54863FEDC74AA303FC24FF4AFBDD35F3" box="[1022,1125,151,177]" pageId="9" pageNumber="540" type="holotype">holotype</typeStatus>
</emphasis>
, by monotypy,
<specimenCount id="9D3B4AC6C74AA303FAC7FF4AFA8A35F3" box="[1309,1330,151,177]" count="1" pageId="9" pageNumber="540" type="female"></specimenCount>
(
<collectionCode id="ED2C198AC74AA303FA9EFF4AFA2035F3" box="[1348,1432,151,177]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
,
<figureCitation id="13069DCAC74AA303FF4DFF66FEA13597" box="[151,281,187,213]" captionStart="FIGURES 1724" captionStartId="5.[151,264,1568,1592]" captionTargetBox="[232,1377,193,1522]" captionTargetId="figure-18@5.[202,1384,181,1544]" captionTargetPageId="5" captionText="FIGURES 1724. Type specimens of Lutera spp. 1718, female holotype of L. nigromaculata Ohaus, 1900 and labels (©MNHN); 1920, inauthentic type of L. nigromaculata in MFNB (photograph by Bernd Jäger); 2124, female holotype of L. nigrita Ohaus, 1930 (in dorsal and lateral view), head, and labels (photograph by Bernd Jäger, MFNB)." figureDoi="http://doi.org/10.5281/zenodo.13949866" httpUri="https://zenodo.org/record/13949866/files/figure.png" pageId="9" pageNumber="540">Figs.2124</figureCitation>
): “
<collectingMunicipality id="6BE61B35C74AA303FEECFF66FE3A3597" box="[310,386,187,213]" pageId="9" pageNumber="540">Luzon</collectingMunicipality>
,
<location id="8EE2D794C74AA303FE54FF66FD993597" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FE54FF66FD993597" box="[398,545,187,213]" municipality="Luzon" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Berlin">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FDF1FF66FD263597" box="[555,670,187,213]" pageId="9" pageNumber="540" value="1914-06-16">
<collectingDate id="EFC75E67C74AA303FDF1FF66FD263597" box="[555,670,187,213]" pageId="9" pageNumber="540" value="1914-06-16">16.6.1914</collectingDate>
</date>
, leg.
<collectorName id="26C8E499C74AA303FD03FF66FCD03597" box="[729,872,187,213]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
//
<specimenCount id="9D3B4AC6C74AA303FC5EFF66FC213597" box="[900,921,187,213]" count="1" pageId="9" pageNumber="540" type="female"></specimenCount>
//
<emphasis id="B9495D5DC74AA303FC6CFF66FC763596" box="[950,974,187,212]" italics="true" pageId="9" pageNumber="540">R.</emphasis>
<taxonomicName id="4C3DFACCC74AA303FC0FFF66FA253597" authority="Ohs." authorityName="Ohaus" authorityYear="1930" box="[981,1437,187,213]" class="Insecta" family="Scarabaeidae" genus="Lutera" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="variety" species="nigromaculata" variety="nigrita">
[sic!]
<emphasis id="B9495D5DC74AA303FBCCFF66FB023597" box="[1046,1210,187,213]" italics="true" pageId="9" pageNumber="540">nigromaculata</emphasis>
Ohs. v.
<emphasis id="B9495D5DC74AA303FACFFF66FAD93596" box="[1301,1377,187,212]" italics="true" pageId="9" pageNumber="540">nigrita</emphasis>
Ohs.
</taxonomicName>
// Type // Zool. Mus.
<collectingRegion id="49F94FADC74AA303FE5FFF02FE6935BB" box="[389,465,223,249]" country="Germany" name="Berlin" pageId="9" pageNumber="540">Berlin</collectingRegion>
”.
</paragraph>
<paragraph id="8B82814FC74AA303FF1DFEDFFDA7313F" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="540">
<emphasis id="B9495D5DC74AA303FF1DFEDFFE45345E" bold="true" box="[199,509,258,284]" pageId="9" pageNumber="540">
<typeStatus id="54863FEDC74AA303FF1DFEDFFE45345E" box="[199,509,258,284]" isOtherTypeStatus="true" pageId="9" pageNumber="540">Other material examined</typeStatus>
</emphasis>
(
<specimenCount id="9D3B4AC6C74AA303FDCAFEDEFDE2345F" box="[528,602,259,285]" count="27" pageId="9" pageNumber="540" type="male">27♂♂</specimenCount>
,
<specimenCount id="9D3B4AC6C74AA303FDB0FEDEFD08345F" box="[618,688,259,285]" count="47" pageId="9" pageNumber="540" type="female">47♀♀</specimenCount>
).
<emphasis id="B9495D5DC74AA303FD11FEDFFC2E345F" bold="true" box="[715,918,258,285]" pageId="9" pageNumber="540">Origin unknown</emphasis>
:
<materialsCitation id="3B558B12C74AA303FC73FEDEFD623403" collectionCode="MFNB" collectorName="Ohs" location="Lakhon" municipality="Tonkin" pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D3B4AC6C74AA303FC73FEDEFC74345F" box="[937,972,259,285]" count="1" pageId="9" pageNumber="540" type="female">1♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FC05FEDEFB8B345F" box="[991,1075,259,285]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
,
<figureCitation id="13069DCAC74AA303FB98FEDEFB76345F" box="[1090,1230,259,285]" captionStart="FIGURES 1724" captionStartId="5.[151,264,1568,1592]" captionTargetBox="[232,1377,193,1522]" captionTargetId="figure-18@5.[202,1384,181,1544]" captionTargetPageId="5" captionText="FIGURES 1724. Type specimens of Lutera spp. 1718, female holotype of L. nigromaculata Ohaus, 1900 and labels (©MNHN); 1920, inauthentic type of L. nigromaculata in MFNB (photograph by Bernd Jäger); 2124, female holotype of L. nigrita Ohaus, 1930 (in dorsal and lateral view), head, and labels (photograph by Bernd Jäger, MFNB)." figureDoi="http://doi.org/10.5281/zenodo.13949866" httpUri="https://zenodo.org/record/13949866/files/figure.png" pageId="9" pageNumber="540">Figs. 1920</figureCitation>
),
<collectingMunicipality id="6BE61B35C74AA303FB3CFEDEFA80345F" box="[1254,1336,259,285]" pageId="9" pageNumber="540">Tonkin</collectingMunicipality>
,
<location id="8EE2D794C74AA303FA9DFEDEFA25345F" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FA9DFEDEFA25345F" box="[1351,1437,259,285]" municipality="Tonkin" name="Lakhon" pageId="9" pageNumber="540">Lakhon</location>
[unreliable data] //
<taxonomicName id="4C3DFACCC74AA303FEB0FEFAFDE73403" authorityName="Ohaus" authorityYear="1900" box="[362,607,295,321]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC74AA303FEB0FEFAFDE73403" box="[362,607,295,321]" italics="true" pageId="9" pageNumber="540">Lutera nigromaculata</emphasis>
</taxonomicName>
,
<typeStatus id="54863FEDC74AA303FDB1FEFAFD1B3403" box="[619,675,295,321]" pageId="9" pageNumber="540">Type</typeStatus>
<collectorName id="26C8E499C74AA303FD73FEFAFD623403" box="[681,730,295,321]" pageId="9" pageNumber="540">Ohs</collectorName>
</materialsCitation>
.
<materialsCitation id="3B558B12C74AA303FD39FEFAFEAA3427" location="Zool. Mus. Berlin" pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D3B4AC6C74AA303FD39FEFAFD413403" box="[739,761,295,321]" count="1" pageId="9" pageNumber="540" type="female"></specimenCount>
. //
<location id="8EE2D794C74AA303FCC1FEFAFC5A3403" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FCC1FEFAFC5A3403" box="[795,994,295,321]" name="Zool. Mus. Berlin" pageId="9" pageNumber="540">Zool. Mus. Berlin</location>
//!
<bibRefCitation id="EFACFCBEC74AA303FBD7FEFAFB1D3403" author="Ohaus, F." box="[1037,1189,295,321]" pageId="9" pageNumber="540" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus (1900)</bibRefCitation>
described this species on a single
</materialsCitation>
<materialsCitation id="3B558B12C74AA303FEC0FE96FE2F34CA" collectorName="Kuijten Leiden" country="Philippines" location="PHILIPPINES" pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D3B4AC6C74AA303FEC0FE96FE973427" box="[282,303,331,357]" count="1" pageId="9" pageNumber="540" type="female"></specimenCount>
with 2 transverse series of spots on pronotum, conseq. these 5 are no [sic!]
<typeStatus id="54863FEDC74AA303FB57FE96FB743427" box="[1165,1228,331,357]" pageId="9" pageNumber="540">types</typeStatus>
. P.
<collectorName id="26C8E499C74AA303FB28FE96FA243427" box="[1266,1436,331,357]" pageId="9" pageNumber="540">Kuijten Leiden</collectorName>
1986;
<collectingCountry id="F32AC1DFC74AA303FF3AFEB3FE2F34CA" box="[224,407,366,392]" name="Philippines" pageId="9" pageNumber="540">
<emphasis id="B9495D5DC74AA303FF3AFEB3FE2F34CA" bold="true" box="[224,407,366,392]" pageId="9" pageNumber="540">PHILIPPINES</emphasis>
</collectingCountry>
</materialsCitation>
<emphasis id="B9495D5DC74AA303FE4DFEB3FE2434CA" bold="true" box="[407,412,366,392]" pageId="9" pageNumber="540">:</emphasis>
<materialsCitation id="3B558B12C74AA303FE7EFEB2FCCE34EF" collectionCode="MFNB" collectorName="Maes C. &amp; Kuijten's" country="Philippines" location="Zool. Mus. Berlin" pageId="9" pageNumber="540" specimenCount="1" specimenCount-male="1" stateProvince="Camarines Sur">
<specimenCount id="9D3B4AC6C74AA303FE7EFEB2FE7F34CB" box="[420,455,367,393]" count="1" pageId="9" pageNumber="540" type="male">1♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FE02FEB2FD9534CB" box="[472,557,367,393]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<collectingRegion id="49F94FADC74AA303FD98FEB2FD5134CB" box="[578,745,367,393]" country="Philippines" name="Camarines Sur" pageId="9" pageNumber="540">Camarines Sur</collectingRegion>
<collectorName id="26C8E499C74AA303FD2BFEB2FCE834CB" box="[753,848,367,393]" pageId="9" pageNumber="540">Maes C.</collectorName>
//
<taxonomicName id="4C3DFACCC74AA303FCABFEB2FB4634CB" authority="Ohs. Cotype" authorityName="Ohs. Cotype" box="[881,1278,367,393]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC74AA303FCABFEB2FBD034CB" box="[881,1128,367,393]" italics="true" pageId="9" pageNumber="540">Lutera nigromaculata</emphasis>
Ohs. Cotype
</taxonomicName>
//
<location id="8EE2D794C74AA303FAC5FEB2FF6534EF" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FAC5FEB2FF6534EF" country="Philippines" name="Zool. Mus. Berlin" pageId="9" pageNumber="540" stateProvince="Camarines Sur">Zool. Mus. Berlin</location>
// [
<collectorName id="26C8E499C74AA303FED9FE4EFED534EF" box="[259,365,403,429]" pageId="9" pageNumber="540">Kuijtens</collectorName>
handwriting, indicated genital apparatus lost]
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FC59FE4EFAC134EF" box="[899,1401,403,429]" collectingDate="1914-06" collectionCode="MFNB" country="Philippines" location="Luzon" municipality="Mt. Banahao" pageId="9" pageNumber="540" specimenCount="2" specimenCount-female="2" stateProvince="Camarines Sur">
<specimenCount id="9D3B4AC6C74AA303FC59FE4EFC0334EF" box="[899,955,403,429]" count="2" pageId="9" pageNumber="540" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FC11FE4EFB9834EF" box="[971,1056,403,429]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FBEEFE4EFB7134EF" box="[1076,1225,403,429]" pageId="9" pageNumber="540">Mt. Banahao</collectingMunicipality>
,
<location id="8EE2D794C74AA303FB0FFE4EFAA534EF" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FB0FFE4EFAA534EF" box="[1237,1309,403,429]" country="Philippines" municipality="Mt. Banahao" name="Luzon" pageId="9" pageNumber="540" stateProvince="Camarines Sur">Luzon</location>
<date id="FF83A78FC74AA303FAFEFE4EFAC134EF" box="[1316,1401,403,429]" pageId="9" pageNumber="540" value="1914-06">
<collectingDate id="EFC75E67C74AA303FAFEFE4EFAC134EF" box="[1316,1401,403,429]" pageId="9" pageNumber="540" value="1914-06">vi.1914</collectingDate>
</date>
</materialsCitation>
G. Boettcher //
<taxonomicName id="4C3DFACCC74AA303FEFAFE6AFD1D3493" authority="Ohs. Cotype" authorityName="Ohs. Cotype" box="[288,677,439,465]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC74AA303FEFAFE6AFDAC3493" box="[288,532,439,465]" italics="true" pageId="9" pageNumber="540">Lutera nigromaculata</emphasis>
Ohs. Cotype
</taxonomicName>
// Zool. Mus. Berlin;
<materialsCitation id="3B558B12C74AA303FC4FFE6AFBD934B7" collectionCode="MFNB" collectorName="S. Luzon Mt. Isarog Bottcher &amp; Zool. Mus. Berlin" country="Philippines" location="Camarin." pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1" stateProvince="Camarines Sur">
<specimenCount id="9D3B4AC6C74AA303FC4FFE6AFC003493" box="[917,952,439,465]" count="1" pageId="9" pageNumber="540" type="female">1♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FC1CFE6AFBA33493" box="[966,1051,439,465]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<collectorName id="26C8E499C74AA303FBF7FE6AFAC93493" box="[1069,1393,439,465]" pageId="9" pageNumber="540">S. Luzon Mt. Isarog Böttcher</collectorName>
S. /
<location id="8EE2D794C74AA303FF4DFE06FF4734B7" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FF4DFE06FF4734B7" box="[151,255,475,501]" country="Philippines" name="Camarin." pageId="9" pageNumber="540" stateProvince="Camarines Sur">Camarin.</location>
iv. 16 [reverse side] //
<taxonomicName id="4C3DFACCC74AA303FE2DFE06FCC434B7" authority="Ohs. Cotype" authorityName="Ohs. Cotype" box="[503,892,475,501]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC74AA303FE2DFE06FD5434B7" box="[503,748,475,501]" italics="true" pageId="9" pageNumber="540">Lutera nigromaculata</emphasis>
Ohs. Cotype
</taxonomicName>
//
<collectorName id="26C8E499C74AA303FC4DFE06FBD934B7" box="[919,1121,475,501]" pageId="9" pageNumber="540">Zool. Mus. Berlin</collectorName>
</materialsCitation>
;
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(
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),
<location id="8EE2D794C74AA303FADBFE06FA2134B7" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FADBFE06FA2134B7" box="[1281,1433,475,501]" country="Philippines" name="Mt. Makiling" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Makiling</location>
,
<collectingRegion id="49F94FADC74AA303FF4DFE22FF56375B" box="[151,238,511,537]" country="Philippines" name="Laguna" pageId="9" pageNumber="540">Laguna</collectingRegion>
, P.I., V.1.[19]
<quantity id="4CC52CAAC74AA303FE5AFE22FE4D375B" box="[384,501,511,537]" metricMagnitude="3" metricUnit="m" metricValue="9.87552" pageId="9" pageNumber="540" unit="ft" value="32400.0">
<elevation id="0010667CC74AA303FE5AFE22FE4D375B" box="[384,501,511,537]" metricMagnitude="3" metricUnit="m" metricValue="9.87552" pageId="9" pageNumber="540" unit="ft" value="32400.0">32, 400 ft.</elevation>
</quantity>
elev. //
<collectorName id="26C8E499C74AA303FD90FE22FCE9375B" box="[586,849,511,537]" pageId="9" pageNumber="540">F.C. Hadden Collection</collectorName>
//
<determinerName id="7E5E6D8BC74AA303FCB6FE22FC0C375B" box="[876,948,511,537]" pageId="9" pageNumber="540">Ohaus</determinerName>
determ.,
<taxonomicName id="4C3DFACCC74AA303FBC1FE22FAFD375B" authority="Ohs." authorityName="Ohaus" authorityYear="1900" box="[1051,1349,511,537]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="species" species="nigromaculata">Lutera nigromaculata Ohs.</taxonomicName>
//
<collectorName id="26C8E499C74AA303FABAFE22FE9B377F" pageId="9" pageNumber="540">Zool. Mus. Berlin</collectorName>
</materialsCitation>
;
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<specimenCount id="9D3B4AC6C74AA303FEF5FDFEFEDF377F" box="[303,359,547,573]" count="3" pageId="9" pageNumber="540" type="male">3♂♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FEADFDFEFE74377F" box="[375,460,547,573]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<location id="8EE2D794C74AA303FE3BFDFEFDCF377F" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FE3BFDFEFDCF377F" box="[481,631,547,573]" country="Philippines" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FD59FDFEFCBF377F" box="[643,775,547,573]" pageId="9" pageNumber="540" value="1914-06-11">
<collectingDate id="EFC75E67C74AA303FD59FDFEFCBF377F" box="[643,775,547,573]" pageId="9" pageNumber="540" value="1914-06-11">11 VI 1914</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B558B12C74AA303FCCEFDFEFC55377F" box="[788,1005,547,573]" collectorName="Luzon" country="Philippines" location="Boettcher" pageId="9" pageNumber="540" specimenCount="1" stateProvince="Laguna">
<location id="8EE2D794C74AA303FCCEFDFEFC39377F" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FCCEFDFEFC39377F" box="[788,897,547,573]" country="Philippines" name="Boettcher" pageId="9" pageNumber="540" stateProvince="Laguna">Boettcher</location>
//
<collectorName id="26C8E499C74AA303FC78FDFEFC55377F" box="[930,1005,547,573]" pageId="9" pageNumber="540">Luzon</collectorName>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FC20FDFEFE7F3723" collectingDate="1914-06" collectionCode="MFNB" collectorName="G. Boettcher" country="Philippines" location="Luzon" municipality="Mt. Banahao" pageId="9" pageNumber="540" specimenCount="2" specimenCount-male="2" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FC20FDFEFB8A377F" box="[1018,1074,547,573]" count="2" pageId="9" pageNumber="540" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FB98FDFEFB2F377F" box="[1090,1175,547,573]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FB76FDFEFAF9377F" box="[1196,1345,547,573]" pageId="9" pageNumber="540">Mt. Banahao</collectingMunicipality>
,
<location id="8EE2D794C74AA303FA94FDFEFA20377F" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FA94FDFEFA20377F" box="[1358,1432,547,573]" country="Philippines" municipality="Mt. Banahao" name="Luzon" pageId="9" pageNumber="540" stateProvince="Laguna">Luzon</location>
,
<date id="FF83A78FC74AA303FF4DFD9AFF4F3723" box="[151,247,583,609]" pageId="9" pageNumber="540" value="1914-06">
<collectingDate id="EFC75E67C74AA303FF4DFD9AFF4F3723" box="[151,247,583,609]" pageId="9" pageNumber="540" value="1914-06">VI 1914</collectingDate>
</date>
,
<collectorName id="26C8E499C74AA303FED9FD9AFE2B3723" box="[259,403,583,609]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FE0EFD9AFB763723" box="[468,1230,583,609]" collectingDate="1914-06-11" collectionCode="MFNB" collectorName="G. Boettcher" country="Philippines" location="Mt. Banahao" municipality="Luzon" pageId="9" pageNumber="540" specimenCount="8" specimenCount-female="8" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FE0EFD9AFDB93723" box="[468,513,583,609]" count="8" pageId="9" pageNumber="540" type="female">8♀♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FDD3FD9AFDDF3723" box="[521,615,583,609]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FDA6FD9AFD7E3723" box="[636,710,583,609]" pageId="9" pageNumber="540">Luzon</collectingMunicipality>
,
<location id="8EE2D794C74AA303FD09FD9AFCDF3723" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FD09FD9AFCDF3723" box="[723,871,583,609]" country="Philippines" municipality="Luzon" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FCA9FD9AFC4D3723" box="[883,1013,583,609]" pageId="9" pageNumber="540" value="1914-06-11">
<collectingDate id="EFC75E67C74AA303FCA9FD9AFC4D3723" box="[883,1013,583,609]" pageId="9" pageNumber="540" value="1914-06-11">11 VI 1914</collectingDate>
</date>
, leg.
<collectorName id="26C8E499C74AA303FBE8FD9AFB763723" box="[1074,1230,583,609]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FB03FD9AFDD037C7" collectingDate="1914-06-10" collectionCode="MFNB" collectorName="G. Boettcher" country="Philippines" location="Mt. Banahao" pageId="9" pageNumber="540" specimenCount="1" specimenCount-male="1" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FB03FD9AFB443723" box="[1241,1276,583,609]" count="1" pageId="9" pageNumber="540" type="male">1♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FAD6FD9AFAD93723" box="[1292,1377,583,609]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<location id="8EE2D794C74AA303FAAFFD9AFF4437C7" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FAAFFD9AFF4437C7" country="Philippines" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FED0FDB6FE3137C7" box="[266,393,619,645]" pageId="9" pageNumber="540" value="1914-06-10">
<collectingDate id="EFC75E67C74AA303FED0FDB6FE3137C7" box="[266,393,619,645]" pageId="9" pageNumber="540" value="1914-06-10">10. 6 1914</collectingDate>
</date>
, leg.
<collectorName id="26C8E499C74AA303FE13FDB6FDD037C7" box="[457,616,619,645]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FDAFFDB6FBEA37C7" box="[629,1106,619,645]" collectingDate="1914-06-11" collectionCode="MFNB" country="Philippines" location="Mt. Banahao" pageId="9" pageNumber="540" specimenCount="2" specimenCount-male="2" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FDAFFDB6FD1537C7" box="[629,685,619,645]" count="2" pageId="9" pageNumber="540" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FD64FDB6FCAB37C7" box="[702,787,619,645]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<location id="8EE2D794C74AA303FCF3FDB6FC0737C7" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FCF3FDB6FC0737C7" box="[809,959,619,645]" country="Philippines" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FC16FDB6FBEA37C7" box="[972,1106,619,645]" pageId="9" pageNumber="540" value="1914-06-11">
<collectingDate id="EFC75E67C74AA303FC16FDB6FBEA37C7" box="[972,1106,619,645]" pageId="9" pageNumber="540" value="1914-06-11">11 VI 1914</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B558B12C74AA303FBBAFDB6FE9A37EB" collectorName="Zool. Mus. Berlin" country="Philippines" location="Luzon" municipality="Boettcher" pageId="9" pageNumber="540" specimenCount="1" stateProvince="Laguna">
<collectingMunicipality id="6BE61B35C74AA303FBBAFDB6FB7537C7" box="[1120,1229,619,645]" pageId="9" pageNumber="540">Boettcher</collectingMunicipality>
//
<location id="8EE2D794C74AA303FB35FDB6FA8137C7" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FB35FDB6FA8137C7" box="[1263,1337,619,645]" country="Philippines" municipality="Boettcher" name="Luzon" pageId="9" pageNumber="540" stateProvince="Laguna">Luzon</location>
, //
<collectorName id="26C8E499C74AA303FABAFDB6FE9A37EB" pageId="9" pageNumber="540">Zool. Mus. Berlin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FEF5FD52FCBF37EB" box="[303,775,655,681]" collectingDate="1914-06-10" collectionCode="MFNB" country="Philippines" location="Mt. Banahao" pageId="9" pageNumber="540" specimenCount="2" specimenCount-male="2" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FEF5FD52FEDF37EB" box="[303,359,655,681]" count="2" pageId="9" pageNumber="540" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FEADFD52FE7437EB" box="[375,460,655,681]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<location id="8EE2D794C74AA303FE3AFD52FDCE37EB" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FE3AFD52FDCE37EB" box="[480,630,655,681]" country="Philippines" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FD58FD52FCBF37EB" box="[642,775,655,681]" pageId="9" pageNumber="540" value="1914-06-10">
<collectingDate id="EFC75E67C74AA303FD58FD52FCBF37EB" box="[642,775,655,681]" pageId="9" pageNumber="540" value="1914-06-10">10 VI 1914</collectingDate>
</date>
</materialsCitation>
, G. Boettcher // Luzon;
<materialsCitation id="3B558B12C74AA303FBFEFD52FF6A378F" collectingDate="1914-06-10" collectionCode="MFNB" country="Philippines" location="Mt. Banahao" pageId="9" pageNumber="540" specimenCount="1" specimenCount-male="1" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FBFEFD52FBFF37EB" box="[1060,1095,655,681]" count="1" pageId="9" pageNumber="540" type="male">1♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FB82FD52FB1537EB" box="[1112,1197,655,681]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<location id="8EE2D794C74AA303FB1BFD52FAEE37EB" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FB1BFD52FAEE37EB" box="[1217,1366,655,681]" country="Philippines" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FAB9FD52FF6A378F" pageId="9" pageNumber="540" value="1914-06-10">
<collectingDate id="EFC75E67C74AA303FAB9FD52FF6A378F" pageId="9" pageNumber="540" value="1914-06-10">10. 6 1914</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B558B12C74AA303FF07FD6EFE0C378F" box="[221,436,691,717]" collectorName="Luzon" country="Philippines" location="Boettcher" pageId="9" pageNumber="540" specimenCount="1" stateProvince="Laguna">
<location id="8EE2D794C74AA303FF07FD6EFEF2378F" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FF07FD6EFEF2378F" box="[221,330,691,717]" country="Philippines" name="Boettcher" pageId="9" pageNumber="540" stateProvince="Laguna">Boettcher</location>
//
<collectorName id="26C8E499C74AA303FEBCFD6EFE0C378F" box="[358,436,691,717]" pageId="9" pageNumber="540">Luzon</collectorName>
</materialsCitation>
,;
<materialsCitation id="3B558B12C74AA303FE19FD6EFB2D378F" box="[451,1173,691,717]" collectingDate="1914-06-09" collectionCode="MFNB" collectorName="G. Boettcher" country="Philippines" location="Mt. Banahao" municipality="Luzon" pageId="9" pageNumber="540" specimenCount="1" specimenCount-male="1" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FE19FD6EFE5E378F" box="[451,486,691,717]" count="1" pageId="9" pageNumber="540" type="male">1♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FE2EFD6EFDF1378F" box="[500,585,691,717]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FD86FD6EFD1E378F" box="[604,678,691,717]" pageId="9" pageNumber="540">Luzon</collectingMunicipality>
,
<location id="8EE2D794C74AA303FD6BFD6EFCFC378F" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FD6BFD6EFCFC378F" box="[689,836,691,717]" country="Philippines" municipality="Luzon" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FC95FD6EFC7A378F" box="[847,962,691,717]" pageId="9" pageNumber="540" value="1914-06-09">
<collectingDate id="EFC75E67C74AA303FC95FD6EFC7A378F" box="[847,962,691,717]" pageId="9" pageNumber="540" value="1914-06-09">9 VI 1914</collectingDate>
</date>
, leg.
<collectorName id="26C8E499C74AA303FC26FD6EFB2D378F" box="[1020,1173,691,717]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FB7AFD6EFD3737B3" collectingDate="1914-06-10" collectionCode="MFNB" collectorName="G. Boettcher" country="Philippines" location="Mt. Banahao" municipality="Luzon" pageId="9" pageNumber="540" specimenCount="6" specimenCount-female="6" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FB7AFD6EFB6F378F" box="[1184,1239,691,717]" count="6" pageId="9" pageNumber="540" type="female">6♀♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FB3CFD6EFA83378F" box="[1254,1339,691,717]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FA94FD6EFA20378F" box="[1358,1432,691,717]" pageId="9" pageNumber="540">Luzon</collectingMunicipality>
,
<location id="8EE2D794C74AA303FF4DFD0AFE9337B3" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FF4DFD0AFE9337B3" box="[151,299,727,753]" country="Philippines" municipality="Luzon" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FEECFD0AFE0137B3" box="[310,441,727,753]" pageId="9" pageNumber="540" value="1914-06-10">
<collectingDate id="EFC75E67C74AA303FEECFD0AFE0137B3" box="[310,441,727,753]" pageId="9" pageNumber="540" value="1914-06-10">10 VI 1914</collectingDate>
</date>
, leg.
<collectorName id="26C8E499C74AA303FE2EFD0AFD3737B3" box="[500,655,727,753]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FD40FD0BFBD237B3" box="[666,1130,726,753]" collectingDate="1914-06-11" collectionCode="MFNB" country="Philippines" location="Mt. Banahao" pageId="9" pageNumber="540" specimenCount="5" specimenCount-female="5" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FD40FD0BFD6A37B3" box="[666,722,726,753]" count="5" pageId="9" pageNumber="540" type="female">5♀♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FD3BFD0AFC8E37B3" box="[737,822,727,753]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<location id="8EE2D794C74AA303FC93FD0AFC6537B3" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FC93FD0AFC6537B3" box="[841,989,727,753]" country="Philippines" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
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<collectingDate id="EFC75E67C74AA303FC32FD0AFBD237B3" box="[1000,1130,727,753]" pageId="9" pageNumber="540" value="1914-06-11">11 VI 1914</collectingDate>
</date>
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,
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<location id="8EE2D794C74AA303FBAFFD0AFB5A37B3" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FBAFFD0AFB5A37B3" box="[1141,1250,727,753]" country="Philippines" name="Boettcher" pageId="9" pageNumber="540" stateProvince="Laguna">Boettcher</location>
//
<collectorName id="26C8E499C74AA303FADAFD0AFAF337B3" box="[1280,1355,727,753]" pageId="9" pageNumber="540">Luzon</collectorName>
</materialsCitation>
;
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<specimenCount id="9D3B4AC6C74AA303FA8CFD0AFA2437B3" box="[1366,1436,727,753]" count="12" pageId="9" pageNumber="540" type="female">12♀♀</specimenCount>
(
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),
<collectingMunicipality id="6BE61B35C74AA303FEDDFD26FEE93657" box="[263,337,763,789]" pageId="9" pageNumber="540">Luzon</collectingMunicipality>
,
<location id="8EE2D794C74AA303FE86FD26FE573657" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FE86FD26FE573657" box="[348,495,763,789]" country="Philippines" municipality="Luzon" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FE20FD26FDC23657" box="[506,634,763,789]" pageId="9" pageNumber="540" value="1914-06-10">
<collectingDate id="EFC75E67C74AA303FE20FD26FDC23657" box="[506,634,763,789]" pageId="9" pageNumber="540" value="1914-06-10">10. 6. 1914</collectingDate>
</date>
, leg.
<collectorName id="26C8E499C74AA303FD6EFD26FCF53657" box="[692,845,763,789]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FC82FD26FAB73657" box="[856,1295,763,789]" collectingDate="1914-06-11" collectionCode="MFNB" country="Philippines" location="Mt. Banahao" pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FC82FD26FCC23657" box="[856,890,763,789]" count="1" pageId="9" pageNumber="540" type="female">1♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FC53FD26FC663657" box="[905,990,763,789]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<location id="8EE2D794C74AA303FC2BFD26FB3C3657" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FC2BFD26FB3C3657" box="[1009,1156,763,789]" country="Philippines" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FB54FD26FAB73657" box="[1166,1295,763,789]" pageId="9" pageNumber="540" value="1914-06-11">
<collectingDate id="EFC75E67C74AA303FB54FD26FAB73657" box="[1166,1295,763,789]" pageId="9" pageNumber="540" value="1914-06-11">11 VI 1914</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B558B12C74AA303FAC3FD26FF5D367B" collectorName="Luzon" country="Philippines" location="Boettcher" pageId="9" pageNumber="540" specimenCount="1" stateProvince="Laguna">
<location id="8EE2D794C74AA303FAC3FD26FA3E3657" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FAC3FD26FA3E3657" box="[1305,1414,763,789]" country="Philippines" name="Boettcher" pageId="9" pageNumber="540" stateProvince="Laguna">Boettcher</location>
//
<collectorName id="26C8E499C74AA303FF4DFCC2FF5D367B" box="[151,229,799,825]" pageId="9" pageNumber="540">Luzon</collectorName>
</materialsCitation>
,;
<materialsCitation id="3B558B12C74AA303FF2EFCC2FCEC367B" box="[244,852,799,825]" collectingDate="1914-06" collectionCode="MFNB" collectorName="G. Boettcher" country="Philippines" location="Mt. Banahao" pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FF2EFCC2FEAF367B" box="[244,279,799,825]" count="1" pageId="9" pageNumber="540" type="female">1♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FEFFFCC2FEC2367B" box="[293,378,799,825]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<location id="8EE2D794C74AA303FE57FCC2FD98367B" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FE57FCC2FD98367B" box="[397,544,799,825]" country="Philippines" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FDF3FCC2FD30367B" box="[553,648,799,825]" pageId="9" pageNumber="540" value="1914-06">
<collectingDate id="EFC75E67C74AA303FDF3FCC2FD30367B" box="[553,648,799,825]" pageId="9" pageNumber="540" value="1914-06">VI 1914</collectingDate>
</date>
,
<collectorName id="26C8E499C74AA303FD49FCC2FC99367B" box="[659,801,799,825]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FC85FCC2FE88361F" collectingDate="1914-06-11" collectionCode="MFNB" collectorName="G. Boettcher" country="Philippines" location="Mt. Banahao" municipality="Luzon" pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FC85FCC2FC3A367B" box="[863,898,799,825]" count="1" pageId="9" pageNumber="540" type="female">1♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FC4AFCC2FC5D367B" box="[912,997,799,825]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FC22FCC2FBFA367B" box="[1016,1090,799,825]" pageId="9" pageNumber="540">Luzon</collectingMunicipality>
,
<location id="8EE2D794C74AA303FB97FCC2FB58367B" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FB97FCC2FB58367B" box="[1101,1248,799,825]" country="Philippines" municipality="Luzon" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FB30FCC2FAD1367B" box="[1258,1385,799,825]" pageId="9" pageNumber="540" value="1914-06-11">
<collectingDate id="EFC75E67C74AA303FB30FCC2FAD1367B" box="[1258,1385,799,825]" pageId="9" pageNumber="540" value="1914-06-11">11. 6. 1914</collectingDate>
</date>
, leg.
<collectorName id="26C8E499C74AA303FF4DFC9EFE88361F" box="[151,304,835,861]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FEE1FC9EFBB5361F" box="[315,1037,835,861]" collectingDate="1914-06-09" collectionCode="MFNB" collectorName="G. Boettcher" country="Philippines" location="Mt. Banahao" municipality="Luzon" pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FEE1FC9EFEE5361F" box="[315,349,835,861]" count="1" pageId="9" pageNumber="540" type="female">1♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FEB6FC9EFE79361F" box="[364,449,835,861]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FE0EFC9EFDA6361F" box="[468,542,835,861]" pageId="9" pageNumber="540">Luzon</collectingMunicipality>
,
<location id="8EE2D794C74AA303FDF3FC9EFD04361F" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FDF3FC9EFD04361F" box="[553,700,835,861]" country="Philippines" municipality="Luzon" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FD1CFC9EFC81361F" box="[710,825,835,861]" pageId="9" pageNumber="540" value="1914-06-09">
<collectingDate id="EFC75E67C74AA303FD1CFC9EFC81361F" box="[710,825,835,861]" pageId="9" pageNumber="540" value="1914-06-09">9 VI 1914</collectingDate>
</date>
, leg.
<collectorName id="26C8E499C74AA303FCAEFC9EFBB5361F" box="[884,1037,835,861]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FBCDFC9EFE4536C3" collectingDate="1914-06-11" collectionCode="MFNB" collectorName="G. Boettcher" country="Philippines" location="Mt. Banahao" municipality="Luzon" pageId="9" pageNumber="540" specimenCount="1" specimenCount-male="1" stateProvince="Laguna">
<specimenCount id="9D3B4AC6C74AA303FBCDFC9EFB82361F" box="[1047,1082,835,861]" count="1" pageId="9" pageNumber="540" type="male">1♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FB92FC9EFB25361F" box="[1096,1181,835,861]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="540">MFNB</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FB6AFC9EFB42361F" box="[1200,1274,835,861]" pageId="9" pageNumber="540">Luzon</collectingMunicipality>
,
<location id="8EE2D794C74AA303FADFFC9EFA20361F" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FADFFC9EFA20361F" box="[1285,1432,835,861]" country="Philippines" municipality="Luzon" name="Mt. Banahao" pageId="9" pageNumber="540" stateProvince="Laguna">Mt. Banahao</location>
,
<date id="FF83A78FC74AA303FF4DFCBAFEA536C3" box="[151,285,871,897]" pageId="9" pageNumber="540" value="1914-06-11">
<collectingDate id="EFC75E67C74AA303FF4DFCBAFEA536C3" box="[151,285,871,897]" pageId="9" pageNumber="540" value="1914-06-11">11 VI 1914</collectingDate>
</date>
, leg.
<collectorName id="26C8E499C74AA303FE84FCBAFE4536C3" box="[350,509,871,897]" pageId="9" pageNumber="540">G. Boettcher</collectorName>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FDD0FCBAFB8836C3" box="[522,1072,871,897]" collectingDate="2018-04" collectionCode="ZMPC" country="Philippines" location="Dingalan" municipality="Luzon" pageId="9" pageNumber="540" specimenCount="1" specimenCount-male="1" stateProvince="Aurora">
<specimenCount id="9D3B4AC6C74AA303FDD0FCBAFD9536C3" box="[522,557,871,897]" count="1" pageId="9" pageNumber="540" type="male">1♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FDE5FCBAFD2936C3" box="[575,657,871,897]" pageId="9" pageNumber="540">ZMPC</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FD7CFCBAFD4836C3" box="[678,752,871,897]" pageId="9" pageNumber="540">Luzon</collectingMunicipality>
,
<collectingRegion id="49F94FADC74AA303FD27FCBAFCF736C3" box="[765,847,871,897]" country="Philippines" name="Aurora" pageId="9" pageNumber="540">Aurora</collectingRegion>
,
<location id="8EE2D794C74AA303FC87FCBAFC7F36C3" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FC87FCBAFC7F36C3" box="[861,967,871,897]" country="Philippines" municipality="Luzon" name="Dingalan" pageId="9" pageNumber="540" stateProvince="Aurora">Dingalan</location>
,
<date id="FF83A78FC74AA303FC0EFCBAFB8836C3" box="[980,1072,871,897]" pageId="9" pageNumber="540" value="2018-04">
<collectingDate id="EFC75E67C74AA303FC0EFCBAFB8836C3" box="[980,1072,871,897]" pageId="9" pageNumber="540" value="2018-04">IV.2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FBE4FCBAFD5536E7" collectingDate="2018-07" collectionCode="ZMPC" country="Philippines" county="Luzon" location="Mountains Range" municipality="Dingalan" pageId="9" pageNumber="540" specimenCount="2" specimenCount-female="2" stateProvince="Aurora">
<specimenCount id="9D3B4AC6C74AA303FBE4FCBAFBCE36C3" box="[1086,1142,871,897]" count="2" pageId="9" pageNumber="540" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FB52FCBAFB6236C3" box="[1160,1242,871,897]" pageId="9" pageNumber="540">ZMPC</collectionCode>
),
<collectingCounty id="62E3F9C3C74AA303FB35FCBAFA8136C3" box="[1263,1337,871,897]" pageId="9" pageNumber="540">Luzon</collectingCounty>
,
<collectingRegion id="49F94FADC74AA303FA9CFCBAFA2036C3" box="[1350,1432,871,897]" country="Philippines" name="Aurora" pageId="9" pageNumber="540">Aurora</collectingRegion>
,
<collectingMunicipality id="6BE61B35C74AA303FF4DFC56FEB936E7" box="[151,257,907,933]" pageId="9" pageNumber="540">Dingalan</collectingMunicipality>
, Sierra Madre
<location id="8EE2D794C74AA303FE76FC56FDCF36E7" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FE76FC56FDCF36E7" box="[428,631,907,933]" country="Philippines" county="Luzon" municipality="Dingalan" name="Mountains Range" pageId="9" pageNumber="540" stateProvince="Aurora">Mountains Range</location>
,
<date id="FF83A78FC74AA303FD5EFC56FD5536E7" box="[644,749,907,933]" pageId="9" pageNumber="540" value="2018-07">
<collectingDate id="EFC75E67C74AA303FD5EFC56FD5536E7" box="[644,749,907,933]" pageId="9" pageNumber="540" value="2018-07">VII.2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FD21FC56FAB636E7" box="[763,1294,907,933]" collectionCode="ZMPC" country="Philippines" location="Mt. Halcon" municipality="Mindoro" pageId="9" pageNumber="540" specimenCount="2" specimenCount-male="2" stateProvince="Aurora">
<specimenCount id="9D3B4AC6C74AA303FD21FC56FC8B36E7" box="[763,819,907,933]" count="2" pageId="9" pageNumber="540" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FC9EFC56FC2E36E7" box="[836,918,907,933]" pageId="9" pageNumber="540">ZMPC</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FC71FC56FBA836E7" box="[939,1040,907,933]" pageId="9" pageNumber="540">Mindoro</collectingMunicipality>
,
<location id="8EE2D794C74AA303FBC7FC56FB1A36E7" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FBC7FC56FB1A36E7" box="[1053,1186,907,933]" country="Philippines" municipality="Mindoro" name="Mt. Halcon" pageId="9" pageNumber="540" stateProvince="Aurora">Mt. Halcon</location>
, 2021.VI
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FAC6FC56FBDB368B" collectingDate="2017-04" collectionCode="ZMPC" country="Philippines" location="Don Salvador Benedicto" municipality="Visayas Region" pageId="9" pageNumber="540" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Negros Occidental">
<specimenCount id="9D3B4AC6C74AA303FAC6FC56FAEF36E7" box="[1308,1367,907,933]" count="2" pageId="9" pageNumber="540" type="male">2♂♂</specimenCount>
,
<specimenCount id="9D3B4AC6C74AA303FABEFC56FA2436E7" box="[1380,1436,907,933]" count="2" pageId="9" pageNumber="540" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FF45FC72FF49368B" box="[159,241,943,969]" pageId="9" pageNumber="540">ZMPC</collectionCode>
), W.
<collectingMunicipality id="6BE61B35C74AA303FEEAFC72FE5D368B" box="[304,485,943,969]" pageId="9" pageNumber="540">Visayas Region</collectingMunicipality>
,
<collectingRegion id="49F94FADC74AA303FE2EFC72FD73368B" box="[500,715,943,969]" country="Philippines" name="Negros Occidental" pageId="9" pageNumber="540">Negros Occidental</collectingRegion>
,
<location id="8EE2D794C74AA303FD00FC72FC4F368B" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FD00FC72FC4F368B" box="[730,1015,943,969]" country="Philippines" municipality="Visayas Region" name="Don Salvador Benedicto" pageId="9" pageNumber="540" stateProvince="Negros Occidental">Don Salvador Benedicto</location>
,
<date id="FF83A78FC74AA303FBDCFC72FBDB368B" box="[1030,1123,943,969]" pageId="9" pageNumber="540" value="2017-04">
<collectingDate id="EFC75E67C74AA303FBDCFC72FBDB368B" box="[1030,1123,943,969]" pageId="9" pageNumber="540" value="2017-04">IV.2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FBABFC72FE5236AF" collectingDate="2018-06" collectionCode="ZMPC" country="Philippines" location="Wao" municipality="Mindanao" pageId="9" pageNumber="540" specimenCount="3" specimenCount-male="3" stateProvince="Lanao del Sur">
<specimenCount id="9D3B4AC6C74AA303FBABFC72FB11368B" box="[1137,1193,943,969]" count="3" pageId="9" pageNumber="540" type="male">3♂♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FB61FC72FAB5368B" box="[1211,1293,943,969]" pageId="9" pageNumber="540">ZMPC</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FAFEFC72FA20368B" box="[1316,1432,943,969]" pageId="9" pageNumber="540">Mindanao</collectingMunicipality>
,
<collectingRegion id="49F94FADC74AA303FF4DFC0EFE8336AF" box="[151,315,979,1005]" country="Philippines" name="Lanao del Sur" pageId="9" pageNumber="540">Lanao del Sur</collectingRegion>
,
<location id="8EE2D794C74AA303FE9DFC0EFEC536AF" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FE9DFC0EFEC536AF" box="[327,381,979,1005]" country="Philippines" municipality="Mindanao" name="Wao" pageId="9" pageNumber="540" stateProvince="Lanao del Sur">Wao</location>
,
<date id="FF83A78FC74AA303FE50FC0EFE5236AF" box="[394,490,979,1005]" pageId="9" pageNumber="540" value="2018-06">
<collectingDate id="EFC75E67C74AA303FE50FC0EFE5236AF" box="[394,490,979,1005]" pageId="9" pageNumber="540" value="2018-06">VI.2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FE2DFC0EFBDE36AF" box="[503,1126,979,1005]" collectingDate="2017-03" collectionCode="ZMPC" country="Philippines" location="Wao" municipality="Mindanao" pageId="9" pageNumber="540" specimenCount="1" specimenCount-male="1" stateProvince="Lanao del Sur">
<specimenCount id="9D3B4AC6C74AA303FE2DFC0EFDA236AF" box="[503,538,979,1005]" count="1" pageId="9" pageNumber="540" type="male">1♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FDF1FC0EFDC536AF" box="[555,637,979,1005]" pageId="9" pageNumber="540">ZMPC</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FD49FC0EFCBF36AF" box="[659,775,979,1005]" pageId="9" pageNumber="540">Mindanao</collectingMunicipality>
,
<collectingRegion id="49F94FADC74AA303FCCEFC0EFC0036AF" box="[788,952,979,1005]" country="Philippines" name="Lanao del Sur" pageId="9" pageNumber="540">Lanao del Sur</collectingRegion>
,
<location id="8EE2D794C74AA303FC1EFC0EFC4236AF" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FC1EFC0EFC4236AF" box="[964,1018,979,1005]" country="Philippines" municipality="Mindanao" name="Wao" pageId="9" pageNumber="540" stateProvince="Lanao del Sur">Wao</location>
,
<date id="FF83A78FC74AA303FBDDFC0EFBDE36AF" box="[1031,1126,979,1005]" pageId="9" pageNumber="540" value="2017-03">
<collectingDate id="EFC75E67C74AA303FBDDFC0EFBDE36AF" box="[1031,1126,979,1005]" pageId="9" pageNumber="540" value="2017-03">III.2017</collectingDate>
</date>
</materialsCitation>
;
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<specimenCount id="9D3B4AC6C74AA303FBA9FC0EFB1436AF" box="[1139,1196,979,1005]" count="2" pageId="9" pageNumber="540" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FB66FC0EFAB636AF" box="[1212,1294,979,1005]" pageId="9" pageNumber="540">ZMPC</collectionCode>
),
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,
<collectingRegion id="49F94FADC74AA303FF4DFC2AFE833153" box="[151,315,1015,1041]" country="Philippines" name="Davao del Sur" pageId="9" pageNumber="540">Davao del Sur</collectingRegion>
,
<location id="8EE2D794C74AA303FE9CFC2AFE793153" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FE9CFC2AFE793153" box="[326,449,1015,1041]" country="Philippines" municipality="Mindanao" name="Kapatagan" pageId="9" pageNumber="540" stateProvince="Davao del Sur">Kapatagan</location>
,
<date id="FF83A78FC74AA303FE16FC2AFD8D3153" box="[460,565,1015,1041]" pageId="9" pageNumber="540" value="2018-07">
<collectingDate id="EFC75E67C74AA303FE16FC2AFD8D3153" box="[460,565,1015,1041]" pageId="9" pageNumber="540" value="2018-07">VII.2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FD9BFC2AFAB83153" box="[577,1280,1015,1041]" collectingDate="2021-08" collectionCode="ZMPC" country="Philippines" location="Kapatagan" municipality="Mindanao" pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1" stateProvince="Davao del Sur">
<specimenCount id="9D3B4AC6C74AA303FD9BFC2AFDDC3153" box="[577,612,1015,1041]" count="1" pageId="9" pageNumber="540" type="female">1♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FDA9FC2AFD7D3153" box="[627,709,1015,1041]" pageId="9" pageNumber="540">ZMPC</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FD03FC2AFCF53153" box="[729,845,1015,1041]" pageId="9" pageNumber="540">Mindanao</collectingMunicipality>
,
<collectingRegion id="49F94FADC74AA303FC83FC2AFC443153" box="[857,1020,1015,1041]" country="Philippines" name="Davao del Sur" pageId="9" pageNumber="540">Davao del Sur</collectingRegion>
,
<location id="8EE2D794C74AA303FBDDFC2AFB3A3153" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FBDDFC2AFB3A3153" box="[1031,1154,1015,1041]" country="Philippines" municipality="Mindanao" name="Kapatagan" pageId="9" pageNumber="540" stateProvince="Davao del Sur">Kapatagan</location>
,
<date id="FF83A78FC74AA303FB57FC2AFAB83153" box="[1165,1280,1015,1041]" pageId="9" pageNumber="540" value="2021-08">
<collectingDate id="EFC75E67C74AA303FB57FC2AFAB83153" box="[1165,1280,1015,1041]" pageId="9" pageNumber="540" value="2021-08">VIII.2021</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FAD6FC2AFD253177" collectingDate="2022-02" collectionCode="ZMPC" country="Philippines" location="Kapatagan" municipality="Mindanao" pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1" stateProvince="Davao del Sur">
<specimenCount id="9D3B4AC6C74AA303FAD6FC2AFA973153" box="[1292,1327,1015,1041]" count="1" pageId="9" pageNumber="540" type="female">1♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FAE4FC2AFA283153" box="[1342,1424,1015,1041]" pageId="9" pageNumber="540">ZMPC</collectionCode>
),
<collectingMunicipality id="6BE61B35C74AA303FF4DFBC6FEB33177" box="[151,267,1051,1077]" pageId="9" pageNumber="540">Mindanao</collectingMunicipality>
,
<collectingRegion id="49F94FADC74AA303FECCFBC6FE003177" box="[278,440,1051,1077]" country="Philippines" name="Davao del Sur" pageId="9" pageNumber="540">Davao del Sur</collectingRegion>
,
<location id="8EE2D794C74AA303FE18FBC6FD853177" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FE18FBC6FD853177" box="[450,573,1051,1077]" country="Philippines" municipality="Mindanao" name="Kapatagan" pageId="9" pageNumber="540" stateProvince="Davao del Sur">Kapatagan</location>
,
<date id="FF83A78FC74AA303FD92FBC6FD253177" box="[584,669,1051,1077]" pageId="9" pageNumber="540" value="2022-02">
<collectingDate id="EFC75E67C74AA303FD92FBC6FD253177" box="[584,669,1051,1077]" pageId="9" pageNumber="540" value="2022-02">II.2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FD72FBC6FE8E311B" collectingDate="2018-06" collectionCode="ZMPC" country="Philippines" county="Mindanao" location="Ilomavis" municipality="Kidapawan" pageId="9" pageNumber="540" specimenCount="1" specimenCount-male="1" stateProvince="North Cotabato">
<specimenCount id="9D3B4AC6C74AA303FD72FBC6FD733177" box="[680,715,1051,1077]" count="1" pageId="9" pageNumber="540" type="male">1♂</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FD03FBC6FC933177" box="[729,811,1051,1077]" pageId="9" pageNumber="540">ZMPC</collectionCode>
),
<collectingCounty id="62E3F9C3C74AA303FCE4FBC6FC0A3177" box="[830,946,1051,1077]" pageId="9" pageNumber="540">Mindanao</collectingCounty>
,
<collectingRegion id="49F94FADC74AA303FC67FBC6FBD53177" box="[957,1133,1051,1077]" country="Philippines" name="North Cotabato" pageId="9" pageNumber="540">North Cotabato</collectingRegion>
,
<collectingMunicipality id="6BE61B35C74AA303FBA2FBC6FB423177" box="[1144,1274,1051,1077]" pageId="9" pageNumber="540">Kidapawan</collectingMunicipality>
,
<location id="8EE2D794C74AA303FADFFBC6FAD23177" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FADFFBC6FAD23177" box="[1285,1386,1051,1077]" country="Philippines" county="Mindanao" municipality="Kidapawan" name="Ilomavis" pageId="9" pageNumber="540" stateProvince="North Cotabato">Ilomavis</location>
,
<location id="8EE2D794C74AA303FAAFFBC6FF72311B" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FAAFFBC6FF72311B" country="Philippines" county="Mindanao" municipality="Kidapawan" name="Mt. Apo" pageId="9" pageNumber="540" stateProvince="North Cotabato">Mt. Apo</location>
,
<date id="FF83A78FC74AA303FF0FFBE2FE8E311B" box="[213,310,1087,1113]" pageId="9" pageNumber="540" value="2018-06">
<collectingDate id="EFC75E67C74AA303FF0FFBE2FE8E311B" box="[213,310,1087,1113]" pageId="9" pageNumber="540" value="2018-06">VI.2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B558B12C74AA303FE9BFBE2FAAC311B" box="[321,1300,1087,1113]" collectingDate="2018-06" collectionCode="ZMPC" country="Philippines" location="Mindanao" pageId="9" pageNumber="540" specimenCount="1" specimenCount-female="1" stateProvince="Compostela">
<specimenCount id="9D3B4AC6C74AA303FE9BFBE2FEDC311B" box="[321,356,1087,1113]" count="1" pageId="9" pageNumber="540" type="female">1♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FEA9FBE2FE7D311B" box="[371,453,1087,1113]" pageId="9" pageNumber="540">ZMPC</collectionCode>
),
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,
<collectingRegion id="49F94FADC74AA303FD82FBE2FC95311B" box="[600,813,1087,1113]" country="Philippines" name="Compostela" pageId="9" pageNumber="540">Compostela Valley</collectingRegion>
[current
<collectingRegion id="49F94FADC74AA303FC4EFBE2FC65311B" box="[916,989,1087,1113]" country="Philippines" name="Davao" pageId="9" pageNumber="540">Davao</collectingRegion>
de Oro Province],
<date id="FF83A78FC74AA303FB6EFBE2FAAC311B" box="[1204,1300,1087,1113]" pageId="9" pageNumber="540" value="2018-06">
<collectingDate id="EFC75E67C74AA303FB6EFBE2FAAC311B" box="[1204,1300,1087,1113]" pageId="9" pageNumber="540" value="2018-06">VI.2018</collectingDate>
</date>
</materialsCitation>
;
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<specimenCount id="9D3B4AC6C74AA303FAC5FBE2FAFA311B" box="[1311,1346,1087,1113]" count="1" pageId="9" pageNumber="540" type="female">1♀</specimenCount>
(
<collectionCode id="ED2C198AC74AA303FA8BFBE2FA2C311B" box="[1361,1428,1087,1113]" pageId="9" pageNumber="540">CJPC</collectionCode>
)
<location id="8EE2D794C74AA303FF4DFBBEFEF0313F" LSID="urn:lsid:plazi:treatment:03943059C744A300FF4DF8DEFD25311A:8EE2D794C74AA303FF4DFBBEFEF0313F" box="[151,328,1123,1149]" country="Philippines" name="Mountain Prov." pageId="9" pageNumber="540" stateProvince="Compostela">Mountain Prov.</location>
,
<collectorName id="26C8E499C74AA303FE89FBBEFE1F313F" box="[339,423,1123,1149]" pageId="9" pageNumber="540">Bontoc</collectorName>
, 2022.VII
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF42D1C7C74AA303FF4DF86FFABE32AC" ID-DOI="http://doi.org/10.5281/zenodo.13949872" ID-Zenodo-Dep="13949872" httpUri="https://zenodo.org/record/13949872/files/figure.png" pageId="9" pageNumber="540" startId="9.[151,250,1970,1994]" targetBox="[164,1423,1173,1947]" targetPageId="9" targetType="figure">
<paragraph id="8B82814FC74AA303FF4DF86FFABE32AC" blockId="9.[151,1437,1970,2031]" pageId="9" pageNumber="540">
<emphasis id="B9495D5DC74AA303FF4DF86FFEA63288" bold="true" box="[151,286,1970,1994]" pageId="9" pageNumber="540">FIGURE 46.</emphasis>
Known distribution of
<taxonomicName id="4C3DFACCC74AA303FDDFF86EFCDC3289" authority="Ohaus, 1900" authorityName="Ohaus" authorityYear="1900" box="[517,868,1970,1995]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC74AA303FDDFF86EFD593288" box="[517,737,1970,1994]" italics="true" pageId="9" pageNumber="540">Lutera nigromaculata</emphasis>
<bibRefCitation id="EFACFCBEC74AA303FD3CF86FFCDC3289" author="Ohaus, F." box="[742,868,1970,1995]" pageId="9" pageNumber="540" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus, 1900</bibRefCitation>
</taxonomicName>
(black dots) and
<taxonomicName id="4C3DFACCC74AA303FBD7F86EFB563289" authority="Ohaus, 1930" authorityName="Ohaus" authorityYear="1930" box="[1037,1262,1970,1995]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B9495D5DC74AA303FBD7F86EFBD33288" box="[1037,1131,1971,1994]" italics="true" pageId="9" pageNumber="540">L. nigrita</emphasis>
<bibRefCitation id="EFACFCBEC74AA303FBB5F86FFB563289" author="Ohaus, F." box="[1135,1262,1970,1995]" pageId="9" pageNumber="540" pagination="138 - 158" refId="ref7844" refString="Ohaus, F. (1930) XXVI. Beitrag zur Kenntnis der Ruteliden (Col. Lamell.). Deutsche Entomologische Zeitschrift, 1930, 138 - 158. https: // doi. org / 10.1002 / mmnd. 48019300205" type="journal article" year="1930">Ohaus, 1930</bibRefCitation>
</taxonomicName>
(black triangles). Type localities are underlined, and the open dot in Nepal is a record from
<bibRefCitation id="EFACFCBEC74AA303FC58F80BFBBD32AD" author="Wada, K." box="[898,1029,2006,2031]" pageId="9" pageNumber="540" pagination="359 - 367" refId="ref8045" refString="Wada, K. (1998) Two new species of the genus Lutera (Coleoptera, Scarabaeidae, Rutelinae) from Borneo and Sumatra. Elytra, 26 (2), 359 - 367." type="journal article" year="1998">Wada (1998)</bibRefCitation>
without precise location.
</paragraph>
</caption>
<subSubSection id="C327D2C4C749A300FF1DFF4BFDDC3402" pageId="10" pageNumber="541" type="description">
<paragraph id="8B82814FC749A300FF1DFF4BFC1135F3" blockId="10.[151,1437,150,1112]" box="[199,937,150,177]" pageId="10" pageNumber="541">
<emphasis id="B9495D5DC749A300FF1DFF4BFEC635F2" bold="true" box="[199,382,150,176]" pageId="10" pageNumber="541">Measurements.</emphasis>
Body length
<quantity id="4CC52CAAC749A300FDCFFF4BFD0235F3" box="[533,698,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" metricValueMax="1.83" metricValueMin="1.45" pageId="10" pageNumber="541" unit="mm" value="16.4" valueMax="18.3" valueMin="14.5">14.518.3 mm</quantity>
, width
<quantity id="4CC52CAAC749A300FCD6FF4BFC1C35F3" box="[780,932,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.6" metricValueMin="8.4" pageId="10" pageNumber="541" unit="mm" value="9.5" valueMax="10.6" valueMin="8.4">8.410.6 mm</quantity>
.
</paragraph>
<paragraph id="8B82814FC749A300FF1DFF67FDDC3402" blockId="10.[151,1437,150,1112]" pageId="10" pageNumber="541">
<emphasis id="B9495D5DC749A300FF1DFF67FEF53596" bold="true" box="[199,333,186,212]" pageId="10" pageNumber="541">Variability.</emphasis>
Body of this species usually yellow with black maculae on dorsal surface, but this color pattern is also variable (
<figureCitation id="13069DCAC749A300FEEFFF03FE0635BB" box="[309,446,222,249]" captionStart="FIGURES 2530" captionStartId="7.[151,264,1660,1684]" captionTargetBox="[190,1392,182,1629]" captionTargetId="figure-18@7.[183,1404,178,1638]" captionTargetPageId="7" captionText="FIGURES 2530. Lutera nigrita Ohaus, 1930 with different color patterns.25, 27, Dingalan; 26, Don Salvador; 28, Compostela; 29, Kapatagan; 30, Wao." figureDoi="http://doi.org/10.5281/zenodo.13949868" httpUri="https://zenodo.org/record/13949868/files/figure.png" pageId="10" pageNumber="541">Figs. 2530</figureCitation>
), and
<bibRefCitation id="EFACFCBEC749A300FDD8FF03FD2B35BB" author="Wada, K." box="[514,659,222,249]" pageId="10" pageNumber="541" pagination="9 - 11" refId="ref8082" refString="Wada, K. (2015) Records of Lutera boettcheri for the first time in 99 years after its discovery. Saikaku-Tsushin, 31, 9 - 11." type="journal article" year="2015">Wada (2015)</bibRefCitation>
presented a totally black individual. Male genitalia displayed as
<figureCitation id="13069DCAC749A300FAB6FF03FE34345F" captionStart="FIGURES 3145" captionStartId="8.[151,264,1924,1948]" captionTargetBox="[189,1398,275,1888]" captionTargetId="figure-77@8.[176,1411,260,1900]" captionTargetPageId="8" captionText="FIGURES 3145. Male genitalia of Lutera species (3135, dorsal view; 3640, lateral view; 4145, ventral view;). 31, 36, 41, L. nigromaculata Ohaus, 1900 from Thailand; 3235, 3740, 4235, L. nigrita Ohaus, 1930: 32, 37, 42, from Luzon Island; 33, 38, 43, from Mindoro Island; 34, 39, 44, from Negros Island; 35, 40, 45, from Mindanao Island" figureDoi="http://doi.org/10.5281/zenodo.13949870" httpUri="https://zenodo.org/record/13949870/files/figure.png" pageId="10" pageNumber="541">Figs 3235, 3740, 4245</figureCitation>
. Parameres symmetric, the width of apices and concavity of ventral margin vary slightly in profile across different local populations.
</paragraph>
</subSubSection>
<subSubSection id="C327D2C4C749A300FF1DFE97FEFA34EF" pageId="10" pageNumber="541" type="diagnosis">
<paragraph id="8B82814FC749A300FF1DFE97FEFA34EF" blockId="10.[151,1437,150,1112]" pageId="10" pageNumber="541">
<emphasis id="B9495D5DC749A300FF1DFE97FE773426" bold="true" box="[199,463,330,356]" pageId="10" pageNumber="541">Differential diagnosis.</emphasis>
This species resembles
<taxonomicName id="4C3DFACCC749A300FD3EFE96FBFB3426" authority="Ohaus, 1916" authorityName="Ohaus" authorityYear="1916" box="[740,1091,330,356]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="boettcheri">
<emphasis id="B9495D5DC749A300FD3EFE96FC113426" box="[740,937,330,356]" italics="true" pageId="10" pageNumber="541">Lutera boettcheri</emphasis>
Ohaus, 1916
</taxonomicName>
, but is usually smaller in size. Parameres short and simple, in contrast with long parameres and sinuate dorsal margins in
<taxonomicName id="4C3DFACCC749A300FB5CFEB2FAAD34CA" authorityName="Ohaus" authorityYear="1916" box="[1158,1301,366,392]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="boettcheri">
<emphasis id="B9495D5DC749A300FB5CFEB2FAAD34CA" box="[1158,1301,366,392]" italics="true" pageId="10" pageNumber="541">L. boettcheri</emphasis>
</taxonomicName>
(see figures in
<bibRefCitation id="EFACFCBEC749A300FF69FE4FFE8F34EF" author="Wada, K." box="[179,311,402,429]" pageId="10" pageNumber="541" pagination="9 - 11" refId="ref8082" refString="Wada, K. (2015) Records of Lutera boettcheri for the first time in 99 years after its discovery. Saikaku-Tsushin, 31, 9 - 11." type="journal article" year="2015">Wada 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327D2C4C749A300FF1DFE6BFC143492" box="[199,940,438,465]" pageId="10" pageNumber="541" type="distribution">
<paragraph id="8B82814FC749A300FF1DFE6BFC143492" blockId="10.[151,1437,150,1112]" box="[199,940,438,465]" pageId="10" pageNumber="541">
<emphasis id="B9495D5DC749A300FF1DFE6BFED93492" bold="true" box="[199,353,438,464]" pageId="10" pageNumber="541">Distribution.</emphasis>
The
<collectingCountry id="F32AC1DFC749A300FE43FE6BFDAF3492" box="[409,535,438,464]" name="Philippines" pageId="10" pageNumber="541">Philippines</collectingCountry>
(Luzon Island to Mindanao Island).
</paragraph>
</subSubSection>
<subSubSection id="C327D2C4C749A300FF1DFE07FD25311A" pageId="10" pageNumber="541" type="discussion">
<paragraph id="8B82814FC749A300FF1DFE07FD25311A" blockId="10.[151,1437,150,1112]" pageId="10" pageNumber="541">
<emphasis id="B9495D5DC749A300FF1DFE07FE8334B6" bold="true" box="[199,315,474,500]" pageId="10" pageNumber="541">Remarks.</emphasis>
The Philippine specimen from Camarines, Luzon Island (possibly the male bearing label of “
<collectingRegion id="49F94FADC749A300FAF8FE07FF78375A" country="Philippines" name="Camarines Sur" pageId="10" pageNumber="541">Camarines Sur</collectingRegion>
” in the MFNB) was the first known individual of
<taxonomicName id="4C3DFACCC749A300FD25FE22FC26375A" authorityName="Ohaus" authorityYear="1930" box="[767,926,511,536]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B9495D5DC749A300FD25FE22FC26375A" box="[767,926,511,536]" italics="true" pageId="10" pageNumber="541">Lutera nigrita</emphasis>
</taxonomicName>
, but
<bibRefCitation id="EFACFCBEC749A300FC00FE23FBCD375B" author="Ohaus, F." box="[986,1141,510,537]" pageId="10" pageNumber="541" pagination="81 - 99" refId="ref7756" refString="Ohaus, F. (1905) Beitrage zur Kenntnis der Ruteliden. Deutsche Entomologische Zeitschrift, 1905, 81 - 99. https: // doi. org / 10.1002 / mmnd. 48119050214" type="journal article" year="1905">Ohaus (1905)</bibRefCitation>
misidentified it as a male of the Indo-Chinese species
<taxonomicName id="4C3DFACCC749A300FE03FDFEFD24377E" authorityName="Ohaus" authorityYear="1900" box="[473,668,546,572]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC749A300FE03FDFEFD24377E" box="[473,668,546,572]" italics="true" pageId="10" pageNumber="541">L. nigromaculata</emphasis>
</taxonomicName>
and attributed the difference in color pattern to sexual dimorphism. Interestingly,
<bibRefCitation id="EFACFCBEC749A300FEE8FD9BFE753722" author="Ohaus, F." box="[306,461,582,608]" pageId="10" pageNumber="541" pagination="138 - 158" refId="ref7844" refString="Ohaus, F. (1930) XXVI. Beitrag zur Kenntnis der Ruteliden (Col. Lamell.). Deutsche Entomologische Zeitschrift, 1930, 138 - 158. https: // doi. org / 10.1002 / mmnd. 48019300205" type="journal article" year="1930">Ohaus (1930)</bibRefCitation>
made the same misidentification again when more conspecific beetles were available to him, including some females collected near Mount Banahao. All were treated as
<taxonomicName id="4C3DFACCC749A300FBEFFDB6FB4F37C6" authorityName="Ohaus" authorityYear="1900" box="[1077,1271,618,644]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC749A300FBEFFDB6FB4F37C6" box="[1077,1271,618,644]" italics="true" pageId="10" pageNumber="541">L. nigromaculata</emphasis>
</taxonomicName>
except for one black specimen with yellow spots (
<figureCitation id="13069DCAC749A300FDFBFD53FD1237EA" box="[545,682,654,680]" captionStart="FIGURES 1724" captionStartId="5.[151,264,1568,1592]" captionTargetBox="[232,1377,193,1522]" captionTargetId="figure-18@5.[202,1384,181,1544]" captionTargetPageId="5" captionText="FIGURES 1724. Type specimens of Lutera spp. 1718, female holotype of L. nigromaculata Ohaus, 1900 and labels (©MNHN); 1920, inauthentic type of L. nigromaculata in MFNB (photograph by Bernd Jäger); 2124, female holotype of L. nigrita Ohaus, 1930 (in dorsal and lateral view), head, and labels (photograph by Bernd Jäger, MFNB)." figureDoi="http://doi.org/10.5281/zenodo.13949866" httpUri="https://zenodo.org/record/13949866/files/figure.png" pageId="10" pageNumber="541">Figs. 2124</figureCitation>
), which he named as the variety
<taxonomicName id="4C3DFACCC749A300FBC3FD52FAE637EA" authorityName="Ohaus" authorityYear="1930" box="[1049,1374,654,681]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="variety" species="nigromaculata" variety="nigrita">
<emphasis id="B9495D5DC749A300FBC3FD52FB6237EA" box="[1049,1242,654,680]" italics="true" pageId="10" pageNumber="541">L. nigromaculata</emphasis>
var.
<emphasis id="B9495D5DC749A300FAC8FD52FAE637EA" box="[1298,1374,655,680]" italics="true" pageId="10" pageNumber="541">nigrita</emphasis>
</taxonomicName>
, now considered as a valid species.
<bibRefCitation id="EFACFCBEC749A300FE35FD6FFD2E378F" author="Paulian, R." box="[495,662,690,717]" pageId="10" pageNumber="541" pagination="73 - 105" refId="ref7972" refString="Paulian, R. (1959) Coleopteres Scarabeides de l'Indochine (Rutelines et Cetonines). Annales de la Societe entomologique de France, 127, 73 - 105. https: // doi. org / 10.1080 / 21686351.1958.12279112" type="journal article" year="1959">Paulian (1959)</bibRefCitation>
recognized that the specimens from Indo-China and the
<collectingCountry id="F32AC1DFC749A300FAC4FD6FFA24378E" box="[1310,1436,690,716]" name="Philippines" pageId="10" pageNumber="541">Philippines</collectingCountry>
represented different species and that
<bibRefCitation id="EFACFCBEC749A300FD98FD0BFD6D37B3" author="Ohaus, F." box="[578,725,726,753]" pageId="10" pageNumber="541" pagination="81 - 99" refId="ref7756" refString="Ohaus, F. (1905) Beitrage zur Kenntnis der Ruteliden. Deutsche Entomologische Zeitschrift, 1905, 81 - 99. https: // doi. org / 10.1002 / mmnd. 48119050214" type="journal article" year="1905">Ohaus (1905</bibRefCitation>
,
<bibRefCitation id="EFACFCBEC749A300FD38FD0BFCA337B2" author="Ohaus, F." box="[738,795,726,752]" pageId="10" pageNumber="541" pagination="138 - 158" refId="ref7844" refString="Ohaus, F. (1930) XXVI. Beitrag zur Kenntnis der Ruteliden (Col. Lamell.). Deutsche Entomologische Zeitschrift, 1930, 138 - 158. https: // doi. org / 10.1002 / mmnd. 48019300205" type="journal article" year="1930">1930</bibRefCitation>
) had confused them. Consequently,
<bibRefCitation id="EFACFCBEC749A300FB6CFD0BFA2437B2" author="Machatschke, J. W." box="[1206,1436,726,752]" pageId="10" pageNumber="541" pagination="1 - 361" refId="ref7615" refString="Machatschke, J. W. (1972) Scarabaeoidea: Melolonthidae Rutelinae. In: Wilcox, J. A. (Ed.), Coleopterorum Catalogus, Supplementa. Pars 66. Fascicle I. Editio Secunda. W. Junk, The Hague, pp. 1 - 361." type="book chapter" year="1972">Machatschke (1972)</bibRefCitation>
removed Camarines from the distribution range of
<taxonomicName id="4C3DFACCC749A300FD08FD26FC2C3656" authorityName="Ohaus" authorityYear="1900" box="[722,916,762,788]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC749A300FD08FD26FC2C3656" box="[722,916,762,788]" italics="true" pageId="10" pageNumber="541">L. nigromaculata</emphasis>
</taxonomicName>
and raised the variety to species rank.
<bibRefCitation id="EFACFCBEC749A300FA92FD27FF5A367A" author="Kuijten, P. J." pageId="10" pageNumber="541" pagination="75 - 90" refId="ref7400" refString="Kuijten, P. J. (1988) Rutelarcha, Lutera and Cyphelytra: notes on taxonomy and nomenclature, diagnoses, and keys (Coleoptera: Rutelinae, Rutelini). Zoologische Mededelingen, 62 (5 - 9), 75 - 90." type="journal article" year="1988">Kuijten (1988)</bibRefCitation>
tried to clarify this confusion, but the
<typeStatus id="54863FEDC749A300FD55FCC3FD49367A" box="[655,753,798,824]" pageId="10" pageNumber="541" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3DFACCC749A300FCCFFCC2FC6F367A" authorityName="Ohaus" authorityYear="1900" box="[789,983,798,824]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC749A300FCCFFCC2FC6F367A" box="[789,983,798,824]" italics="true" pageId="10" pageNumber="541">L. nigromaculata</emphasis>
</taxonomicName>
, the sole specimen of this species at that time, had unfortunately disappeared. In contrast,
<specimenCount id="9D3B4AC6C749A300FD66FC9FFCDB361E" box="[700,867,834,860]" count="5" pageId="10" pageNumber="541" type="generic">five specimens</specimenCount>
of
<taxonomicName id="4C3DFACCC749A300FC5DFC9EFC49361E" authorityName="Ohaus" authorityYear="1930" box="[903,1009,835,860]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B9495D5DC749A300FC5DFC9EFC49361E" box="[903,1009,835,860]" italics="true" pageId="10" pageNumber="541">L. nigrita</emphasis>
</taxonomicName>
deposited at MFNB (see
<figureCitation id="13069DCAC749A300FAD5FC9FFA2C361E" box="[1295,1428,834,860]" captionStart="FIGURES 1724" captionStartId="5.[151,264,1568,1592]" captionTargetBox="[232,1377,193,1522]" captionTargetId="figure-18@5.[202,1384,181,1544]" captionTargetPageId="5" captionText="FIGURES 1724. Type specimens of Lutera spp. 1718, female holotype of L. nigromaculata Ohaus, 1900 and labels (©MNHN); 1920, inauthentic type of L. nigromaculata in MFNB (photograph by Bernd Jäger); 2124, female holotype of L. nigrita Ohaus, 1930 (in dorsal and lateral view), head, and labels (photograph by Bernd Jäger, MFNB)." figureDoi="http://doi.org/10.5281/zenodo.13949866" httpUri="https://zenodo.org/record/13949866/files/figure.png" pageId="10" pageNumber="541">Figs. 1920</figureCitation>
) were incorrectly labeled as type or cotype of
<taxonomicName id="4C3DFACCC749A300FD45FCBAFCDA36C2" authorityName="Ohaus" authorityYear="1900" box="[671,866,870,896]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC749A300FD45FCBAFCDA36C2" box="[671,866,870,896]" italics="true" pageId="10" pageNumber="541">L. nigromaculata</emphasis>
</taxonomicName>
. As
<bibRefCitation id="EFACFCBEC749A300FC42FCBBFB8736C2" author="Kuijten, P. J." box="[920,1087,870,896]" pageId="10" pageNumber="541" pagination="75 - 90" refId="ref7400" refString="Kuijten, P. J. (1988) Rutelarcha, Lutera and Cyphelytra: notes on taxonomy and nomenclature, diagnoses, and keys (Coleoptera: Rutelinae, Rutelini). Zoologische Mededelingen, 62 (5 - 9), 75 - 90." type="journal article" year="1988">Kuijten (1988)</bibRefCitation>
pointed out, the type status of these specimens is obviously invalid because
<taxonomicName id="4C3DFACCC749A300FD4BFC56FCE936E6" authorityName="Ohaus" authorityYear="1900" box="[657,849,906,932]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC749A300FD4BFC56FCE936E6" box="[657,849,906,932]" italics="true" pageId="10" pageNumber="541">L. nigromaculata</emphasis>
</taxonomicName>
was originally described from a single specimen and the color patterns of this series clearly differ from the original description of
<taxonomicName id="4C3DFACCC749A300FC3EFC72FB1D368A" authorityName="Ohaus" authorityYear="1900" box="[996,1189,942,968]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC749A300FC3EFC72FB1D368A" box="[996,1189,942,968]" italics="true" pageId="10" pageNumber="541">L. nigromaculata</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFACFCBEC749A300FB16FC73FADF368A" author="Ohaus, F." box="[1228,1383,942,968]" pageId="10" pageNumber="541" pagination="225 - 266" refId="ref7723" refString="Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 1900, 225 - 266. https: // doi. org / 10.1002 / mmnd. 48019000210" type="journal article" year="1900">Ohaus (1900)</bibRefCitation>
. The validity of these two species was later verified by the authentic
<typeStatus id="54863FEDC749A300FC80FC0FFC0436AE" box="[858,956,978,1004]" pageId="10" pageNumber="541" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3DFACCC749A300FC3AFC0EFB1936AE" authorityName="Ohaus" authorityYear="1900" box="[992,1185,978,1004]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC749A300FC3AFC0EFB1936AE" box="[992,1185,978,1004]" italics="true" pageId="10" pageNumber="541">L. nigromaculata</emphasis>
</taxonomicName>
rediscovered by Mary Liz Jameson at MNHN (
<bibRefCitation id="EFACFCBEC749A300FE6BFC2BFD8E3152" author="Wada, K." box="[433,566,1014,1040]" pageId="10" pageNumber="541" pagination="359 - 367" refId="ref8045" refString="Wada, K. (1998) Two new species of the genus Lutera (Coleoptera, Scarabaeidae, Rutelinae) from Borneo and Sumatra. Elytra, 26 (2), 359 - 367." type="journal article" year="1998">Wada 1998</bibRefCitation>
). The two species can be easily distinguished by their black body markings:
<taxonomicName id="4C3DFACCC749A300FF4DFBC6FEE13176" authorityName="Ohaus" authorityYear="1900" box="[151,345,1050,1076]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="B9495D5DC749A300FF4DFBC6FEE13176" box="[151,345,1050,1076]" italics="true" pageId="10" pageNumber="541">L. nigromaculata</emphasis>
</taxonomicName>
has several isolated, rounded spots in generally stable positions, whereas
<taxonomicName id="4C3DFACCC749A300FB41FBC6FABE3176" authorityName="Ohaus" authorityYear="1930" box="[1179,1286,1051,1076]" class="Insecta" family="Scarabaeidae" genus="Lutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B9495D5DC749A300FB41FBC6FABE3176" box="[1179,1286,1051,1076]" italics="true" pageId="10" pageNumber="541">L. nigrita</emphasis>
</taxonomicName>
has typically irregular markings with significant variability.
</paragraph>
</subSubSection>
</treatment>
</document>