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<document id="99CC27795AF6E17D12B37A663688F391" ID-DOI="10.5281/zenodo.178539" ID-GBIF-Dataset="dda53384-9974-4b83-b780-a9cd51fffacf" ID-ISSN="1175-5326" ID-Zenodo-Dep="178539" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459899930198" checkinUser="plazi" docAuthor="Freitag, Hendrik &amp; Jäch, Manfred A." docDate="2007" docId="03939707FFBF2545FF2CFCDFE5BDFC9E" docLanguage="en" docName="zt01590p059.pdf" docOrigin="Zootaxa 1590" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ancyronyx" docType="key" docVersion="9" lastPageNumber="41" masterDocId="FFAAEF7FFFBC2541FFBBFFBDE027FF97" masterDocTitle="The genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) in Palawan and Busuanga, (Philippines) with descriptions of six new species" masterLastPageNumber="59" masterPageNumber="37" pageNumber="40" updateTime="1698228715028" updateUser="plazi">
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<mods:title id="E18BB2550A1723FB1D3DF0114DA24114">The genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) in Palawan and Busuanga, (Philippines) with descriptions of six new species</mods:title>
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<mods:namePart id="67C335299CDAD640A78B61924411C407">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="46C532764A7B23C056B638BF666C0D6C">Jäch, Manfred A.</mods:namePart>
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<mods:title id="6630D5B3C5F03ABBF98C22B881F64CD0">Zootaxa</mods:title>
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<mods:date id="F1FE2C9F2CCE0A79DC3729BDC06EAF1D">2007</mods:date>
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<mods:number id="C0DF87CC29371C9FA4E694B214753FF1">1590</mods:number>
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<mods:classification id="BBD7278033599661E32AD1FD74522C40">journal article</mods:classification>
<mods:identifier id="BC1030A6F95D859F77011178323AFEFD" type="DOI">10.5281/zenodo.178539</mods:identifier>
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<mods:identifier id="0602304774AB4861A1EF9D096BDA6C15" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03939707FFBF2545FF2CFCDFE5BDFC9E" ID-DOI="http://doi.org/10.5281/zenodo.6243447" ID-GBIF-Taxon="119372902" ID-Zenodo-Dep="6243447" LSID="urn:lsid:plazi:treatment:03939707FFBF2545FF2CFCDFE5BDFC9E" httpUri="http://treatment.plazi.org/id/03939707FFBF2545FF2CFCDFE5BDFC9E" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40">
<subSubSection id="C320759AFFBF2542FF2CFCDFE28FFCEB" box="[151,680,866,892]" pageId="3" pageNumber="40" type="nomenclature">
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<emphasis id="B94EFA03FFBF2542FF2CFCDFE28FFCEB" bold="true" box="[151,680,866,892]" pageId="3" pageNumber="40">
Key to the
<collectingCountry id="F32D6681FFBF2542FEA4FCDFE187FCEB" box="[287,416,866,892]" name="Philippines" pageId="3" pageNumber="40">Philippine</collectingCountry>
species of
<taxonomicName id="4C3A5D92FFBF2542FD9FFCDFE28FFCEB" ID-CoL="VNF" box="[548,680,866,892]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B94EFA03FFBF2542FD9FFCDFE28FFCEB" bold="true" box="[548,680,866,892]" italics="true" pageId="3" pageNumber="40">Ancyronyx</emphasis>
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<paragraph id="8B852611FFBF2542FF2CFC10E5BDFC78" blockId="3.[151,1437,941,2007]" pageId="3" pageNumber="40">
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1. CL&gt;
<quantity id="4CC28BF4FFBF2542FEA9FC10E155FC50" box="[274,370,941,967]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="40" unit="mm" value="1.9">1.9 mm</quantity>
; legs very long (length of elytra: metafemur&gt; 0.5); ovipositor (coxite) short and stout (comp.
<figureCitation id="13013A94FFBF2542FE9BFC68E154FC78" box="[288,371,981,1007]" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="3" pageNumber="40">Fig. 16</figureCitation>
d) ......................................................................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8B852611FFBF2542FF2CFC40E5BDFBA8" blockId="3.[151,1437,941,2007]" pageId="3" pageNumber="40">
<keyLead id="30CB8524FFBF2542FF2CFC40E5BDFBA8" pageId="3" pageNumber="40">
- CL &lt;
<quantity id="4CC28BF4FFBF2542FEB7FC40E14DFB80" box="[268,362,1021,1047]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="40" unit="mm" value="1.8">1.8 mm</quantity>
; legs long to moderately long (length of elytra: metafemur &lt;0.5); ovipositor (coxite) long and slender (
<figureCitation id="13013A94FFBF2542FEE6FB98E19BFBA8" box="[349,444,1061,1087]" captionStart="FIGURE 11" captionStartId="7.[151,255,1901,1925]" captionTargetBox="[188,1386,183,1846]" captionTargetId="figure@7.[188,1386,183,1846]" captionTargetPageId="7" captionText="FIGURE 11. Ancyronyx patrolus sp. n., paratypes from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) ovipositor in ventral view (right half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) female sternite VIII (strioles omitted); h) male sternite VIII (strioles omitted); i) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178544/files/figure.png" pageId="3" pageNumber="40">Figs. 11</figureCitation>
d; 13d; 14d) ............................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8B852611FFBF2542FF2CFBF0E5BEFB48" blockId="3.[151,1437,941,2007]" pageId="3" pageNumber="40">
<keyLead id="30CB8524FFBF2542FF2CFBF0E5BEFB48" pageId="3" pageNumber="40">
2. Pronotum predominantly dark, anteriorly yellowish; elytra with yellowish “X”-shaped, colour pattern. Aedeagal parameres moderately long (ca. 0.5 of entire aedeagus), clearly not reaching opening of ventral sac, with numerous long apical setae. Ovipositor medially without, but apically with few trichoid spines.
<emphasis id="B94EFA03FFBF2542FF77FB78E46EFB49" box="[204,1097,1221,1246]" italics="true" pageId="3" pageNumber="40">..........................................................................................................................</emphasis>
<taxonomicName id="4C3A5D92FFBF2542FBF6FB78E5BEFB48" ID-CoL="DPT5" authority="Jach, 1994" authorityName="Jach" authorityYear="1994" box="[1101,1433,1221,1247]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B94EFA03FFBF2542FBF6FB78E532FB49" box="[1101,1301,1221,1246]" italics="true" pageId="3" pageNumber="40">A. schillhammeri</emphasis>
<bibRefCitation id="EFAB5BE0FFBF2542FAA7FB78E5BEFB48" author="Jach" box="[1308,1433,1221,1247]" pageId="3" pageNumber="59" refString="Jach, M. A. (1994) A taxonomic review of the Oriental species of the genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae). Revue suisse de Zoologie, 101, 601 - 622." type="journal article" year="1994">Jäch, 1994</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B852611FFBF2542FF2CFB50E5BDFA30" blockId="3.[151,1437,941,2007]" pageId="3" pageNumber="40">
<keyLead id="30CB8524FFBF2542FF2CFB50E5BDFA30" pageId="3" pageNumber="40">
- Pronotum predominantly yellowish, anteriorly brown; elytra with large yellowish median marking, sometimes extended into bands at the edges (
<figureCitation id="13013A94FFBF2542FD27FAA8E2D3FAB8" box="[668,756,1301,1327]" captionStart="FIGURES 7 – 10" captionStartId="6.[151,269,1956,1980]" captionTargetBox="[156,1436,194,1932]" captionTargetId="figure@6.[151,1436,194,1932]" captionTargetPageId="6" captionText="FIGURES 7 – 10. Habitus of 7) Ancyronyx minerva sp. n., paratype Ψ; 8) A. minerva sp. n., paratype Ψ, winged, partly completed; 9) A. minutulus sp. n., holotype ♂, partly completed; 10) A. helgeschneideri sp. n., paratype Ψ." httpUri="https://zenodo.org/record/178543/files/figure.png" pageId="3" pageNumber="40">Fig. 10</figureCitation>
). Aedeagal parameres very long (ca. 0.65 of entire aedeagus), surpassing opening of ventral sac, almost reaching apex of penis, without conspicuous setae (
<figureCitation id="13013A94FFBF2542FAD6FA80E0C2FAE8" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="3" pageNumber="40">Fig. 16</figureCitation>
a, b). Ovipositor without apical spines, but with one long median spine (at apical margin of valvifer) (
<figureCitation id="13013A94FFBF2542FF74FA30E104FA30" box="[207,291,1421,1447]" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="3" pageNumber="40">Fig. 16</figureCitation>
d)
<emphasis id="B94EFA03FFBF2542FEF2FA30E4BAFA31" box="[329,1181,1421,1446]" italics="true" pageId="3" pageNumber="40">
......................................................................................
<taxonomicName id="4C3A5D92FFBF2542FC78FA30E4BAFA31" ID-CoL="DPSH" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[963,1181,1421,1446]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="helgeschneideri" status="sp. nov.">A. helgeschneideri</taxonomicName>
</emphasis>
Freitag &amp; Jäch
<emphasis id="B94EFA03FFBF2542FAE3FA30E5BDFA30" bold="true" box="[1368,1434,1421,1447]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A27D4778FFBF2542FAE3FA30E5BDFA30" box="[1368,1434,1421,1447]" pageId="3" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE3EB4FFBF2542FF2CFA08E5BDF9D0" pageId="3" pageNumber="40">
<paragraph id="8B852611FFBF2542FF2CFA08E5BDF988" blockId="3.[151,1437,941,2007]" pageId="3" pageNumber="40">
<keyLead id="30CB8524FFBF2542FF2CFA08E5BDF988" pageId="3" pageNumber="40">
3. Elytra with four pairs of elongately oval, yellow patches; total length ca.
<quantity id="4CC28BF4FFBF2542FBFFFA08E482FA58" box="[1092,1189,1461,1487]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="40" unit="mm" value="1.3">1.3 mm</quantity>
. Aedeagus conically tapered; parameres moderately long (ca. 0.6 of entire aedeagus), with few very short apical setae ............ ............................................................................................................................
<taxonomicName id="4C3A5D92FFBF2542FBDFF9B8E5BDF988" ID-CoL="DPT8" authority="Jach, 2004" authorityName="Jach" authorityYear="2004" box="[1124,1434,1541,1567]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="sophiemarie">
<emphasis id="B94EFA03FFBF2542FBDFF9B8E532F989" box="[1124,1301,1541,1566]" italics="true" pageId="3" pageNumber="40">A. sophiemarie</emphasis>
<bibRefCitation id="EFAB5BE0FFBF2542FAA7F9B8E5BDF988" author="Jach" box="[1308,1434,1541,1567]" pageId="3" pageNumber="59" refString="Jach, M. A. (2004) Descriptions of two new species of Ancyronyx Erichson (Insecta: Coleoptera: Elmidae). Annalen des Naturhistorischen Museums in Wien, 105 B, 389 - 395." type="journal article" year="2004">Jäch, 2004</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8B852611FFBF2542FF2CF990E5BDF9D0" blockId="3.[151,1437,941,2007]" box="[151,1434,1581,1607]" pageId="3" pageNumber="40">
<keyLead id="30CB8524FFBF2542FF2CF990E5BDF9D0" box="[151,1434,1581,1607]" pageId="3" pageNumber="40">
- Elytra with two pairs of yellowish patches or entirely dark (
<figureCitation id="13013A94FFBF2542FC3BF990E3D4F9D0" box="[896,1011,1581,1607]" captionStart-0="FIGURES 3 – 6" captionStart-1="FIGURES 7 – 10" captionStartId-0="5.[151,269,1843,1867]" captionStartId-1="6.[151,269,1956,1980]" captionTargetBox-0="[153,1436,208,1819]" captionTargetBox-1="[156,1436,194,1932]" captionTargetId-0="figure@5.[151,1436,194,1819]" captionTargetId-1="figure@6.[151,1436,194,1932]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed." captionText-1="FIGURES 7 – 10. Habitus of 7) Ancyronyx minerva sp. n., paratype Ψ; 8) A. minerva sp. n., paratype Ψ, winged, partly completed; 9) A. minutulus sp. n., holotype ♂, partly completed; 10) A. helgeschneideri sp. n., paratype Ψ." httpUri-0="https://zenodo.org/record/178542/files/figure.png" httpUri-1="https://zenodo.org/record/178543/files/figure.png" pageId="3" pageNumber="40">Figs. 3–9</figureCitation>
). Aedeagus different ..................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE3EB4FFBF2542FF2CF9E8E5BDF8C8" pageId="3" pageNumber="40">
<paragraph id="8B852611FFBF2542FF2CF9E8E5BDF928" blockId="3.[151,1437,941,2007]" pageId="3" pageNumber="40">
<keyLead id="30CB8524FFBF2542FF2CF9E8E5BDF928" pageId="3" pageNumber="40">
4. Elytra with two pairs of large longitudinal elongate yellowish patches (
<figureCitation id="13013A94FFBF2542FBB6F9E8E472F9F8" box="[1037,1109,1621,1647]" captionStart="FIGURES 7 – 10" captionStartId="6.[151,269,1956,1980]" captionTargetBox="[156,1436,194,1932]" captionTargetId="figure@6.[151,1436,194,1932]" captionTargetPageId="6" captionText="FIGURES 7 – 10. Habitus of 7) Ancyronyx minerva sp. n., paratype Ψ; 8) A. minerva sp. n., paratype Ψ, winged, partly completed; 9) A. minutulus sp. n., holotype ♂, partly completed; 10) A. helgeschneideri sp. n., paratype Ψ." httpUri="https://zenodo.org/record/178543/files/figure.png" pageId="3" pageNumber="40">Fig. 9</figureCitation>
); CL ca.
<quantity id="4CC28BF4FFBF2542FB7FF9E8E505F9F8" box="[1220,1314,1621,1647]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="40" unit="mm" value="1.1">1.1 mm</quantity>
. Aedeagal parameres moderately long (ca. 0.5 of entire aedeagus), with two very long apical setae (&gt; 0.5 paramere length) (
<figureCitation id="13013A94FFBF2542FE92F918E159F928" box="[297,382,1701,1727]" captionStart="FIGURE 15" captionStartId="17.[151,255,1743,1767]" captionTargetBox="[151,1436,183,1683]" captionTargetId="figure@17.[151,1436,183,1683]" captionTargetPageId="17" captionText="FIGURE 15. Ancyronyx minutulus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male sternite VIII (strioles omitted); e) male tergite VIII (setae and structures of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178548/files/figure.png" pageId="3" pageNumber="40">Fig. 15</figureCitation>
a, b) ................................................................................
<taxonomicName id="4C3A5D92FFBF2542FBB3F918E577F928" ID-CoL="DPSS" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1032,1360,1701,1727]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="minutulus" status="sp. nov.">
<emphasis id="B94EFA03FFBF2542FBB3F918E4BAF929" box="[1032,1181,1701,1726]" italics="true" pageId="3" pageNumber="40">A. minutulus</emphasis>
Freitag &amp; Jäch
</taxonomicName>
<emphasis id="B94EFA03FFBF2542FAE3F918E5BDF928" bold="true" box="[1368,1434,1701,1727]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A27D4778FFBF2542FAE3F918E5BDF928" box="[1368,1434,1701,1727]" pageId="3" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8B852611FFBF2542FF2CF970E5BDF8C8" blockId="3.[151,1437,941,2007]" pageId="3" pageNumber="40">
<keyLead id="30CB8524FFBF2542FF2CF970E5BDF8C8" pageId="3" pageNumber="40">
- Elytra with one subapical pair of longitudinal elongate yellowish patch and one pair of round or transversally elongate, yellowish patches or entirely dark (
<figureCitation id="13013A94FFBF2542FCA7F948E3A9F898" box="[796,910,1781,1807]" captionStart-0="FIGURES 3 – 6" captionStart-1="FIGURES 7 – 10" captionStartId-0="5.[151,269,1843,1867]" captionStartId-1="6.[151,269,1956,1980]" captionTargetBox-0="[153,1436,208,1819]" captionTargetBox-1="[156,1436,194,1932]" captionTargetId-0="figure@5.[151,1436,194,1819]" captionTargetId-1="figure@6.[151,1436,194,1932]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed." captionText-1="FIGURES 7 – 10. Habitus of 7) Ancyronyx minerva sp. n., paratype Ψ; 8) A. minerva sp. n., paratype Ψ, winged, partly completed; 9) A. minutulus sp. n., holotype ♂, partly completed; 10) A. helgeschneideri sp. n., paratype Ψ." httpUri-0="https://zenodo.org/record/178542/files/figure.png" httpUri-1="https://zenodo.org/record/178543/files/figure.png" pageId="3" pageNumber="40">Figs. 3–8</figureCitation>
); CL usually&gt;
<quantity id="4CC28BF4FFBF2542FBF2F948E48FF898" box="[1097,1192,1781,1807]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="40" unit="mm" value="1.1">1.1 mm</quantity>
. Aedeagus different; parameres short (&lt;0.5 of entire aedeagus length), with&gt; 10 long setae or without long setae (
<figureCitation id="13013A94FFBF2542FA9DF8A0E5A1F8A0" box="[1318,1414,1821,1847]" captionStart="FIGURE 11" captionStartId="7.[151,255,1901,1925]" captionTargetBox="[188,1386,183,1846]" captionTargetId="figure@7.[188,1386,183,1846]" captionTargetPageId="7" captionText="FIGURE 11. Ancyronyx patrolus sp. n., paratypes from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) ovipositor in ventral view (right half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) female sternite VIII (strioles omitted); h) male sternite VIII (strioles omitted); i) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178544/files/figure.png" pageId="3" pageNumber="40">Figs. 11</figureCitation>
b, 12b, 13b, 14b) ............................................................................................................................................. 5
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B852611FFBF2542FF2CF8D0E5BBF838" blockId="3.[151,1437,941,2007]" pageId="3" pageNumber="40">
<keyLead id="30CB8524FFBF2542FF2CF8D0E5BBF838" pageId="3" pageNumber="40">
<keyStep id="30CE3EB4FFBF2542FF2CF8D0E5BBF838" pageId="3" pageNumber="40">
5. Pronotum anteriorly with transverse white to yellowish patch or band (
<figureCitation id="13013A94FFBF2542FBB8F8D0E454F810" box="[1027,1139,1901,1927]" captionStart="FIGURES 7 – 10" captionStartId="6.[151,269,1956,1980]" captionTargetBox="[156,1436,194,1932]" captionTargetId="figure@6.[151,1436,194,1932]" captionTargetPageId="6" captionText="FIGURES 7 – 10. Habitus of 7) Ancyronyx minerva sp. n., paratype Ψ; 8) A. minerva sp. n., paratype Ψ, winged, partly completed; 9) A. minutulus sp. n., holotype ♂, partly completed; 10) A. helgeschneideri sp. n., paratype Ψ." httpUri="https://zenodo.org/record/178543/files/figure.png" pageId="3" pageNumber="40">Figs. 7, 8</figureCitation>
); CL &lt;
<quantity id="4CC28BF4FFBF2542FB6BF8D0E51BF810" box="[1232,1340,1901,1927]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="3" pageNumber="40" unit="mm" value="1.35">1.35 mm</quantity>
. Aedeagus conically tapered; parameres with&gt; 10 long apical setae (
<figureCitation id="13013A94FFBF2542FC22F828E3C9F838" box="[921,1006,1941,1967]" captionStart="FIGURE 14" captionStartId="14.[151,255,1867,1891]" captionTargetBox="[172,1415,178,1812]" captionTargetId="figure@14.[172,1416,178,1812]" captionTargetPageId="14" captionText="FIGURE 14. Ancyronyx minerva sp. n., ♂ holotype, Ψ paratype from same locality: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) ovipositor in ventral view (left half); e) male ventrite 5 (setae and strioles of left half omitted); f) female ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) male tergite VIII (setae and structures of left half omitted); i) female tergite VIII (setae and strioles of left half omitted); j) female sternite VIII (strioles omitted)." httpUri="https://zenodo.org/record/178547/files/figure.png" pageId="3" pageNumber="40">Fig. 14</figureCitation>
a, b)
<taxonomicName id="4C3A5D92FFBF2542FB99F828E573F838" ID-CoL="DPSR" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1058,1364,1941,1967]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="minerva" status="sp. nov.">
<emphasis id="B94EFA03FFBF2542FB99F828E485F839" box="[1058,1186,1941,1966]" italics="true" pageId="3" pageNumber="40">A. minerva</emphasis>
Freitag &amp; Jäch
</taxonomicName>
<emphasis id="B94EFA03FFBF2542FAE0F828E5BBF838" bold="true" box="[1371,1436,1941,1967]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A27D4778FFBF2542FAE0F828E5BBF838" box="[1371,1436,1941,1967]" pageId="3" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B852611FFBF2542FF2CF800E5BBF840" blockId="3.[151,1437,941,2007]" box="[151,1436,1981,2007]" pageId="3" pageNumber="40">
<keyLead id="30CB8524FFBF2542FF2CF800E5BBF840" box="[151,1436,1981,2007]" pageId="3" pageNumber="40">
- Pronotum entirely dark, without transverse patch or band (
<figureCitation id="13013A94FFBF2542FCC5F800E3D6F840" box="[894,1009,1981,2007]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,269,1843,1867]" captionTargetBox="[153,1436,208,1819]" captionTargetId="figure@5.[151,1436,194,1819]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed." httpUri="https://zenodo.org/record/178542/files/figure.png" pageId="3" pageNumber="40">Figs. 3–6</figureCitation>
); CL&gt;
<quantity id="4CC28BF4FFBF2542FBF4F800E499F840" box="[1103,1214,1981,2007]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="3" pageNumber="40" unit="mm" value="1.35">1.35 mm</quantity>
. Aedeagus subapi-
</keyLead>
</paragraph>
<paragraph id="8B852611FFB82545FF7CFF25E5BDFEEE" blockId="4.[151,1437,152,778]" pageId="4" pageNumber="41">
cally more or less widened, not conically tapered; parameres with &lt;10 short to moderately long apical setae (
<figureCitation id="13013A94FFB82545FEAAFF7DE157FF4D" box="[273,368,192,218]" captionStart="FIGURE 11" captionStartId="7.[151,255,1901,1925]" captionTargetBox="[188,1386,183,1846]" captionTargetId="figure@7.[188,1386,183,1846]" captionTargetPageId="7" captionText="FIGURE 11. Ancyronyx patrolus sp. n., paratypes from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) ovipositor in ventral view (right half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) female sternite VIII (strioles omitted); h) male sternite VIII (strioles omitted); i) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178544/files/figure.png" pageId="4" pageNumber="41">Figs. 11</figureCitation>
a, 12a, 13a)........................................................................................................................... 6
<keyLead id="30CB8524FFB82545FF2CFF55E5BDFEEE" pageId="4" pageNumber="41">
<keyStep id="30CE3EB4FFB82545FF2CFF55E5BDFEEE" pageId="4" pageNumber="41">
6. [Determination in dark specimens unambiguously possible by sexual characters only]: Subbasal elytral patches more or less equally round, not reaching posterior or lateral elytral margin; legs entirely dark (
<figureCitation id="13013A94FFB82545FAD6FEADE0FFFEC5" captionStart="FIGURES 3 – 6" captionStartId="5.[151,269,1843,1867]" captionTargetBox="[153,1436,208,1819]" captionTargetId="figure@5.[151,1436,194,1819]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed." httpUri="https://zenodo.org/record/178542/files/figure.png" pageId="4" pageNumber="41">Fig. 6</figureCitation>
). Aedeagal median lobe long and moderately slender, subapically slightly widened (ca. 0.3 times as wide as long); ventral sac almost reaching apex (
<figureCitation id="13013A94FFB82545FCB9FEDDE370FEED" box="[770,855,352,378]" captionStart="FIGURE 13" captionStartId="11.[151,255,1841,1865]" captionTargetBox="[242,1346,200,1776]" captionTargetId="figure@11.[242,1347,200,1777]" captionTargetPageId="11" captionText="FIGURE 13. Ancyronyx punkti sp. n., paratypes from site “ Zettel (59) ”: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) ovipositor in ventral view (left half); e) female tergite VIII (setae and strioles of left half omitted); f) female ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female sternite VIII (strioles omitted); i) male tergite VIII (setae and structures of left half omitted); j) male ventrite 5 (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178546/files/figure.png" pageId="4" pageNumber="41">Fig. 13</figureCitation>
a, b)
<emphasis id="B94EFA03FFB82545FC28FEDDE4BBFEEE" box="[915,1180,352,377]" italics="true" pageId="4" pageNumber="41">
.....................
<taxonomicName id="4C3A5D92FFB82545FB88FEDDE4BBFEEE" ID-CoL="DPSZ" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1075,1180,352,377]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="punkti" status="sp. nov.">A. punkti</taxonomicName>
</emphasis>
Freitag &amp; Jäch
<emphasis id="B94EFA03FFB82545FAE3FEE2E5BDFEEE" bold="true" box="[1368,1434,351,377]" pageId="4" pageNumber="41">
<taxonomicNameLabel id="A27D4778FFB82545FAE3FEE2E5BDFEEE" box="[1368,1434,351,377]" pageId="4" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B852611FFB82545FF2CFE35E5BDFE65" blockId="4.[151,1437,152,778]" pageId="4" pageNumber="41">
<keyLead id="30CB8524FFB82545FF2CFE35E5BDFE65" pageId="4" pageNumber="41">
- Subbasal elytral patches transversely elongate (cross-stripe-like), at least reaching elytral margin at shoul- der; legs partly yellowish to pale brown (
<figureCitation id="13013A94FFB82545FD00FE0DE316FE5D" box="[699,817,432,458]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,269,1843,1867]" captionTargetBox="[153,1436,208,1819]" captionTargetId="figure@5.[151,1436,194,1819]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed." httpUri="https://zenodo.org/record/178542/files/figure.png" pageId="4" pageNumber="41">Figs. 3, 5</figureCitation>
). Aedeagal median lobe subapically distinctly widened; ventral sac not reaching apical 0.1 of aedeagus (
<figureCitation id="13013A94FFB82545FCFBFE65E3B8FE65" box="[832,927,472,498]" captionStart="FIGURE 11" captionStartId="7.[151,255,1901,1925]" captionTargetBox="[188,1386,183,1846]" captionTargetId="figure@7.[188,1386,183,1846]" captionTargetPageId="7" captionText="FIGURE 11. Ancyronyx patrolus sp. n., paratypes from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) ovipositor in ventral view (right half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) female sternite VIII (strioles omitted); h) male sternite VIII (strioles omitted); i) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178544/files/figure.png" pageId="4" pageNumber="41">Figs. 11</figureCitation>
a, b; 12a, b) .............................................. 7
</keyLead>
</paragraph>
<keyStep id="30CE3EB4FFB82545FF2CFDBDE5BDFC9E" pageId="4" pageNumber="41">
<paragraph id="8B852611FFB82545FF2CFDBDE5BDFD06" blockId="4.[151,1437,152,778]" pageId="4" pageNumber="41">
<keyLead id="30CB8524FFB82545FF2CFDBDE5BDFD06" pageId="4" pageNumber="41">
7. Femora predominantly yellowish to pale brown (except for articulations), ventral (inner) side without conspicuous setae; basal elytral patches &lt;0.65 times as long as broad (
<figureCitation id="13013A94FFB82545FC44FD95E462FDD5" box="[1023,1093,552,578]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,269,1843,1867]" captionTargetBox="[153,1436,208,1819]" captionTargetId="figure@5.[151,1436,194,1819]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed." httpUri="https://zenodo.org/record/178542/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
), in dark specimens medially interrupted, reduced or lacking (
<figureCitation id="13013A94FFB82545FDF9FDEDE2ABFDFD" box="[578,652,592,618]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,269,1843,1867]" captionTargetBox="[153,1436,208,1819]" captionTargetId="figure@5.[151,1436,194,1819]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed." httpUri="https://zenodo.org/record/178542/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
). Aedeagal median lobe long and slender (ca. 0.2 times as wide as long;
<figureCitation id="13013A94FFB82545FEB1FDC5E17BFD05" box="[266,348,632,658]" captionStart="FIGURE 11" captionStartId="7.[151,255,1901,1925]" captionTargetBox="[188,1386,183,1846]" captionTargetId="figure@7.[188,1386,183,1846]" captionTargetPageId="7" captionText="FIGURE 11. Ancyronyx patrolus sp. n., paratypes from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) ovipositor in ventral view (right half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) female sternite VIII (strioles omitted); h) male sternite VIII (strioles omitted); i) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178544/files/figure.png" pageId="4" pageNumber="41">Fig. 11</figureCitation>
a)
<emphasis id="B94EFA03FFB82545FECEFDC5E4BBFD06" box="[373,1180,632,657]" italics="true" pageId="4" pageNumber="41">
............................................................................................
<taxonomicName id="4C3A5D92FFB82545FBA7FDC5E4BBFD06" ID-CoL="DPSV" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1052,1180,632,657]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="patrolus" status="sp. nov.">A. patrolus</taxonomicName>
</emphasis>
Freitag &amp; Jäch
<emphasis id="B94EFA03FFB82545FAE3FDCAE5BDFD06" bold="true" box="[1368,1434,631,657]" pageId="4" pageNumber="41">
<taxonomicNameLabel id="A27D4778FFB82545FAE3FDCAE5BDFD06" box="[1368,1434,631,657]" pageId="4" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B852611FFB82545FF2CFD1DE5BDFC9E" blockId="4.[151,1437,152,778]" pageId="4" pageNumber="41">
<keyLead id="30CB8524FFB82545FF2CFD1DE5BDFC9E" pageId="4" pageNumber="41">
- Dorsal side of femora entirely dark brown; ventral side with conspicuous setae; basal elytral patches large, ca. 0.8 times as long as broad (
<figureCitation id="13013A94FFB82545FDFBFD75E2ACFD75" box="[576,651,712,738]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,269,1843,1867]" captionTargetBox="[153,1436,208,1819]" captionTargetId="figure@5.[151,1436,194,1819]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed." httpUri="https://zenodo.org/record/178542/files/figure.png" pageId="4" pageNumber="41">Fig. 5</figureCitation>
). Aedeagal median lobe short and stout (ca. 0.4 times as wide as long;
<figureCitation id="13013A94FFB82545FEB1FD4DE179FC9D" box="[266,350,752,778]" captionStart="FIGURE 12" captionStartId="10.[151,255,1319,1343]" captionTargetBox="[162,1427,157,1263]" captionTargetId="figure@10.[162,1427,157,1265]" captionTargetPageId="10" captionText="FIGURE 12. Ancyronyx pseudopatrolus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male ventrite 5 (left half)." httpUri="https://zenodo.org/record/178545/files/figure.png" pageId="4" pageNumber="41">Fig. 12</figureCitation>
a)
<emphasis id="B94EFA03FFB82545FEC7FD4DE4BAFC9E" box="[380,1181,752,777]" italics="true" pageId="4" pageNumber="41">
................................................................................
<taxonomicName id="4C3A5D92FFB82545FC72FD4DE4BAFC9E" ID-CoL="DPSX" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[969,1181,752,777]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="pseudopatrolus" status="sp. nov.">A. pseudopatrolus</taxonomicName>
</emphasis>
Freitag &amp; Jäch
<emphasis id="B94EFA03FFB82545FAE3FD52E5BDFC9E" bold="true" box="[1368,1434,751,777]" pageId="4" pageNumber="41">
<taxonomicNameLabel id="A27D4778FFB82545FAE3FD52E5BDFC9E" box="[1368,1434,751,777]" pageId="4" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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</subSubSection>
</treatment>
</document>
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