treatments-xml/data/03/93/8B/03938B6C6211304663F6E757FEDCFDC0.xml

<document id="CCB2D3412723A123CED99A87BCA9ABA9" ID-DOI="10.5281/zenodo.195786" ID-GBIF-Dataset="c447671e-d700-4c8e-b275-d551bcdb3a1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="195786" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460234666671" checkinUser="plazi" docAuthor="Vernoux, Julien, Garrouste, Romain &amp; Nel, Andre" docDate="2010" docId="03938B6C6211304663F6E757FEDCFDC0" docLanguage="en" docName="zt02499p068.pdf" docOrigin="Zootaxa 2499" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Isometopsallops prokopi Vernoux, Garrouste &amp; Nel, 2010, sp. nov." docType="treatment" docVersion="8" lastPageNumber="68" masterDocId="FFAAF314621130436361E005FFCAFFE7" masterDocTitle="The first psallopinous bug from Lowermost Eocene French amber (Hemiptera: Heteroptera: Miridae)" masterLastPageNumber="68" masterPageNumber="63" pageNumber="63" updateTime="1698239222712" updateUser="plazi">
<mods:mods id="628900993BF8CFD7C99668E539609FAD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4F1A2E242D24A5AEE78A6BBBC03B2198">
<mods:title id="6044C9AB14E69D8E2757B52FFB5D8847">The first psallopinous bug from Lowermost Eocene French amber (Hemiptera: Heteroptera: Miridae)</mods:title>
</mods:titleInfo>
<mods:name id="AFBBA13137119C7F2620FBF6AD305068" type="personal">
<mods:role id="B6FA79E202715424F1C7001068D02EAE">
<mods:roleTerm id="78F29CAE8066A6F916771B4208BF99D1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FCBF680434DD8691C63FEBCC3AEAC30B">Vernoux, Julien</mods:namePart>
</mods:name>
<mods:name id="16C1A20ADC70546E53A6F7D9449CADF1" type="personal">
<mods:role id="0EA69ED289F2806C9D8AC45A281DAE90">
<mods:roleTerm id="2A516E6DDFC5D9CDD7B0BE350504B31E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="86A42EEC84A4CA4950C38105E39664BE">Garrouste, Romain</mods:namePart>
</mods:name>
<mods:name id="F09A4E987ECF7CD673CB7AD31AAA5919" type="personal">
<mods:role id="A23C02FBD3C0C3F6483EA88826E544C8">
<mods:roleTerm id="E0587991A5CF6DB021382AF5195D5CE2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="77054F1D0C8095A9018CC0BABADBDC17">Nel, Andre</mods:namePart>
</mods:name>
<mods:typeOfResource id="FC5FAB150F4B6FDF77ABC56A2061EE33">text</mods:typeOfResource>
<mods:relatedItem id="9205F4957E1EF908599332B0B49F1AF9" type="host">
<mods:titleInfo id="973E76C7DDF9F3A2D64E4457A97DA7A0">
<mods:title id="5EDC72BE43509B1AC7B1CC725BF14C73">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="85320D26BF1980B50C54F90D73B50A57">
<mods:date id="F3C33E49F6BE1814F70449276FA5E232">2010</mods:date>
<mods:detail id="F9746B4D9A4493768E22B2D5DEC6AE84" type="volume">
<mods:number id="1F6697B3AF6ED21A6AB188CEBD5465A2">2499</mods:number>
</mods:detail>
<mods:extent id="8BD52E6A13B04D6D840781CC4C499961" unit="page">
<mods:start id="D5873A0A2A2223D21A536076413B1F2F">63</mods:start>
<mods:end id="23B6840765E79A3B3F8F1EC05B9069DF">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="57807AF044A3AFE5336F2BFC435344A1">journal article</mods:classification>
<mods:identifier id="C548F00B57F2A9E7E469410915394CF9" type="DOI">10.5281/zenodo.195786</mods:identifier>
<mods:identifier id="B58EB4E3300DCE646D038A4D269BBFA1" type="GBIF-Dataset">c447671e-d700-4c8e-b275-d551bcdb3a1b</mods:identifier>
<mods:identifier id="7E2CE18C54FBE19851DA6B755B71EAAC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="85FD14CAFF06590E9A2AED29DF97DD60" type="Zenodo-Dep">195786</mods:identifier>
</mods:mods>
<treatment id="03938B6C6211304663F6E757FEDCFDC0" ID-DOI="http://doi.org/10.5281/zenodo.6204699" ID-GBIF-Taxon="119387793" ID-Zenodo-Dep="6204699" LSID="urn:lsid:plazi:treatment:03938B6C6211304663F6E757FEDCFDC0" httpUri="http://treatment.plazi.org/id/03938B6C6211304663F6E757FEDCFDC0" lastPageId="5" lastPageNumber="68" pageId="0" pageNumber="63">
<subSubSection id="C32069F16211304363F6E757FF36F869" pageId="0" pageNumber="63" type="nomenclature">
<paragraph id="8B853A7A6211304363F6E757FDEBF88C" blockId="0.[151,545,1873,1934]" box="[151,545,1873,1900]" pageId="0" pageNumber="63">
<heading id="D0CD8D166211304363F6E757FDEBF88C" bold="true" box="[151,545,1873,1900]" fontSize="11" level="1" pageId="0" pageNumber="63" reason="1">
<emphasis id="B94EE6686211304363F6E757FDEBF88C" bold="true" box="[151,545,1873,1900]" pageId="0" pageNumber="63">
<taxonomicName id="4C3A41F96211304363F6E757FE77F88B" ID-CoL="6MZ5C" box="[151,445,1874,1900]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="prokopi" status="sp. nov.">
<emphasis id="B94EE6686211304363F6E757FE77F88B" bold="true" box="[151,445,1874,1900]" italics="true" pageId="0" pageNumber="63">Isometopsallops prokopi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D5B136211304362A5E754FDEBF88C" box="[452,545,1873,1899]" pageId="0" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B853A7A6211304363F6E771FF36F869" blockId="0.[151,545,1873,1934]" box="[151,252,1908,1934]" pageId="0" pageNumber="63">
<figureCitation id="130126FF6211304363F6E771FF36F869" box="[151,252,1908,1934]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,255,1403,1427]" captionStartId-1="2.[151,255,1611,1635]" captionStartId-2="3.[151,255,1211,1235]" captionStartId-3="4.[151,255,1076,1100]" captionTargetBox-0="[156,1436,274,1376]" captionTargetBox-1="[151,1434,570,1587]" captionTargetBox-2="[226,1360,607,1186]" captionTargetBox-3="[151,1434,194,1053]" captionTargetId-0="figure@1.[156,1441,274,1376]" captionTargetId-1="figure@2.[151,1436,570,1587]" captionTargetId-2="figure@3.[226,1360,607,1187]" captionTargetId-3="figure@4.[151,1436,194,1055]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Isometopsallops prokopi sp. nov. holotype MNHN A 30851, photograph of dorsal view." captionText-1="FIGURE 2. Isometopsallops prokopi sp. nov. holotype MNHN A 30851, photograph of ventral view." captionText-2="FIGURE 3. Isometopsallops prokopi sp. nov. paratype MNHN A 30854, photograph of dorsal view." captionText-3="FIGURE 4. Isometopsallops prokopi sp. nov. allotype MNHN A 30852, photograph of hind leg." httpUri-0="https://zenodo.org/record/195787/files/figure.png" httpUri-1="https://zenodo.org/record/195788/files/figure.png" httpUri-2="https://zenodo.org/record/195789/files/figure.png" httpUri-3="https://zenodo.org/record/195790/files/figure.png" pageId="0" pageNumber="63">Figs 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32069F16211304263F6E7BAFC64FF30" lastPageId="1" lastPageNumber="64" pageId="0" pageNumber="63" type="materials_examined">
<paragraph id="8B853A7A6211304263F6E7BAFC64FF30" blockId="0.[151,1436,1983,2009]" lastBlockId="1.[151,1437,152,215]" lastPageId="1" lastPageNumber="64" pageId="0" pageNumber="63">
<emphasis id="B94EE6686211304363F6E7BAFECEF83E" bold="true" box="[151,260,1983,2009]" pageId="0" pageNumber="63">Material</emphasis>
. 
<typeStatus id="548184D8621130436273E7BAFEB5F83E" box="[274,383,1983,2009]" pageId="0" pageNumber="63" type="holotype">Holotype</typeStatus>
<collectionCode id="ED2BA2BF6211304362E6E7BAFE2BF83E" LSID="urn:lsid:biocol.org:col:34988" box="[391,481,1983,2009]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="63">MNHN</collectionCode>
A30851 (PA 8543, male), allotype 
<collectionCode id="ED2BA2BF6211304360E0E7BAFC11F83E" LSID="urn:lsid:biocol.org:col:34988" box="[897,987,1983,2009]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="63">MNHN</collectionCode>
A30852 (PA 6524, female), 
<typeStatus id="548184D862113043664DE7BAFA56F83E" box="[1324,1436,1983,2009]" pageId="0" pageNumber="63" type="paratype">paratypes</typeStatus>
<collectionCode id="ED2BA2BF6210304263F6E09DFF39FF55" LSID="urn:lsid:biocol.org:col:34988" box="[151,243,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="64">MNHN</collectionCode>
A30853 (PA 8584, male), and 
<collectionCode id="ED2BA2BF621030426117E09DFD18FF55" LSID="urn:lsid:biocol.org:col:34988" box="[630,722,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="64">MNHN</collectionCode>
A30854 (PA 9493, male), all stored at the Laboratoire de Paléontologie, Muséum national d’Histoire naturelle, Paris, 
<collectingCountry id="F32D7AEA621030426037E0B8FC60FF30" box="[854,938,189,215]" name="France" pageId="1" pageNumber="64">France</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF456AF26210304263F6E57EFB0FFA74" box="[151,1221,1403,1427]" httpUri="https://zenodo.org/record/195787/files/figure.png" pageId="1" pageNumber="64" targetBox="[156,1436,274,1376]" targetPageId="1">
<paragraph id="8B853A7A6210304263F6E57EFB0FFA74" blockId="1.[151,1221,1403,1427]" box="[151,1221,1403,1427]" pageId="1" pageNumber="64">
<emphasis id="B94EE6686210304263F6E57EFED3FA74" bold="true" box="[151,281,1403,1427]" pageId="1" pageNumber="64">FIGURE 1.</emphasis>
<taxonomicName id="4C3A41F9621030426241E579FDECFA74" box="[288,550,1404,1427]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="prokopi" status="sp. nov.">
<emphasis id="B94EE668621030426241E579FDECFA74" box="[288,550,1404,1427]" italics="true" pageId="1" pageNumber="64">Isometopsallops prokopi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D5B1362103042614DE57EFD48FA74" box="[556,642,1403,1427]" pageId="1" pageNumber="64" rank="species">
<emphasis id="B94EE66862103042614DE57EFDB1FA74" bold="true" box="[556,635,1403,1427]" pageId="1" pageNumber="64">sp. nov</emphasis>
.
</taxonomicNameLabel>
holotype MNHN A30851, photograph of dorsal view.
</paragraph>
</caption>
<subSubSection id="C32069F16210304263A4E5BEFAE1FA31" box="[197,1323,1467,1494]" pageId="1" pageNumber="64" type="etymology">
<paragraph id="8B853A7A6210304263A4E5BEFAE1FA31" blockId="1.[151,1437,1467,2016]" box="[197,1323,1467,1494]" pageId="1" pageNumber="64">
<emphasis id="B94EE6686210304263A4E5BEFE80FA32" bold="true" box="[197,330,1467,1493]" pageId="1" pageNumber="64">Etymology</emphasis>
. Named after our friend and colleague Dr Jakub Prokop (Charles University, Praha).
</paragraph>
</subSubSection>
<subSubSection id="C32069F16210304263A4E5E5FD40F951" pageId="1" pageNumber="64" type="diagnosis">
<paragraph id="8B853A7A6210304263A4E5E5FD40F951" blockId="1.[151,1437,1467,2016]" pageId="1" pageNumber="64">
<emphasis id="B94EE6686210304263A4E5E5FEF7FA1D" bold="true" box="[197,317,1504,1530]" pageId="1" pageNumber="64">Diagnosis</emphasis>
. tarsi three-segmented; a subapical tooth on each claw; parempodia setiform; no pulvilli; broad femora; seven metafemoral trichobothria and six mesofemoral trichobothria; large eyes semi-globular; antennae placed basally on inner side of eyes; head nearly vertical; no distinct collar; pronotum trapezoidal, ratio length/width pronotum 0.4; membrane of hemelytra crumpled and with two cells, one small and one large; body two times as long as wide, with dorsal surface smooth, with dense hairs; male right genital style (or paramere) slightly longer than left one.
</paragraph>
</subSubSection>
<subSubSection id="C32069F16210304163A4E6C5FF19F837" lastPageId="2" lastPageNumber="65" pageId="1" pageNumber="64" type="description">
<paragraph id="8B853A7A6210304163A4E6C5FD55FDE5" blockId="1.[151,1437,1467,2016]" lastBlockId="2.[151,1437,152,514]" lastPageId="2" lastPageNumber="65" pageId="1" pageNumber="64">
<emphasis id="B94EE6686210304263A4E6C5FEA9F93D" bold="true" box="[197,355,1728,1754]" pageId="1" pageNumber="64">Description</emphasis>
. General coloration light brown or brownish; macropterous; dorsal surface smooth, impunctate, with moderately short, erect and sub-erect hairs, densely covering dorsal surface of body and posteriorly directed. Head large, nearly spherical in frontal view, nearly vertical, and nearly twice as wide as long; antenna cylindrical, inserted at inner side of eyes about 1/3 of their lower level; eyes very large, almost semi-globular, covering much of dorsal surface of head, bare, emarginated along inner side of genae; interocular space much narrower than width of eye, hind margins of eyes touching anterior margin of pronotum; mandibular plates not enlarged; labium four-segmented, inserted ventrally on head, and reaching at least a third of abdomen, with segment 1 reaching posterior margin of head; trapezoidal pronotum a little longer than head, with ratio length/width = 0.4, collar absent; no calli; lateral margins and posterior margin not carinate nor emarginated; mesoscutum two times shorter than scutellum; claval commissure present; cuneus and costal fracture clearly distinct; a white spot or light-coloured spot around the costal fracture; hemelytral membrane with a large cell and a small cell which branches to a supplementary vein, membrane distinctly crumpled, hairs absent on membrane; femora of hind legs broad and long, tibia nearly as long as femora, seven metafemoral trichobothria and six mesofemoral trichobothria; tibia with delicate spines distally, three-segmented tarsi more or less of equal size, first tarsal segment well separated from second but suture between third and second segments is weaker than that between first and second (probably presence of a pseudo-trisegmentation sensu 
<bibRefCitation id="EFAB478B62133041629BE1C7FD0EFE3B" author="Gorczyca" box="[506,708,450,476]" pageId="2" pageNumber="68" refString="Gorczyca, J. (2000) A systematic study on Cylapinae with a revision of the Afrotropical region (Heteroptera, Miridae). Wydawnictwo Uniwersystetu Slaskiego, Katowice, 176 pp." type="book" year="2000">Gorczyca [2000]</bibRefCitation>
); claws of equal size and almost straight; a subapical tooth on each claw; parempodia setiform; no pulvilli.
</paragraph>
<caption id="DF456AF26213304163F6E64EFB04F985" box="[151,1230,1611,1635]" httpUri="https://zenodo.org/record/195788/files/figure.png" pageId="2" pageNumber="65" targetBox="[151,1434,570,1587]" targetPageId="2">
<paragraph id="8B853A7A6213304163F6E64EFB04F985" blockId="2.[151,1230,1611,1635]" box="[151,1230,1611,1635]" pageId="2" pageNumber="65">
<emphasis id="B94EE6686213304163F6E64EFED3F984" bold="true" box="[151,281,1611,1635]" pageId="2" pageNumber="65">FIGURE 2.</emphasis>
<taxonomicName id="4C3A41F9621330416241E64EFDECF985" box="[288,550,1611,1634]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="prokopi" status="sp. nov.">
<emphasis id="B94EE668621330416241E64EFDECF985" box="[288,550,1611,1634]" italics="true" pageId="2" pageNumber="65">Isometopsallops prokopi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D5B1362133041614DE64EFD48F985" box="[556,642,1611,1635]" pageId="2" pageNumber="65" rank="species">
<emphasis id="B94EE66862133041614DE64EFDB1F984" bold="true" box="[556,635,1611,1635]" pageId="2" pageNumber="65">sp. nov</emphasis>
.
</taxonomicNameLabel>
holotype MNHN A30851, photograph of ventral view.
</paragraph>
</caption>
<paragraph id="8B853A7A6213304163A4E68EFF19F837" blockId="2.[151,1438,1675,2037]" pageId="2" pageNumber="65">
<emphasis id="B94EE6686213304163A4E68EFE86F942" bold="true" box="[197,332,1675,1701]" pageId="2" pageNumber="65">Dimension</emphasis>
. (taken on PA 8543 and PA 9493 for accessibility): Length of body from apex of hemelytra 
<quantity id="4CC2979F6213304163F6E6B4FEC9F92C" box="[151,259,1713,1739]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="2" pageNumber="65" unit="mm" value="2.25">2.25 mm</quantity>
(around 
<quantity id="4CC2979F621330416212E6B4FE1BF92C" box="[371,465,1713,1739]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="65" unit="mm" value="2.5">2.5 mm</quantity>
for the PA 8584 and PA6524); width 
<quantity id="4CC2979F6213304160C3E6B4FBE8F92C" box="[930,1058,1713,1739]" metricMagnitude="-3" metricUnit="m" metricValue="1.025" pageId="2" pageNumber="65" unit="mm" value="1.025">1.025 mm</quantity>
; length of hemelytra 
<quantity id="4CC2979F621330416646E6B4FA5DF92C" box="[1319,1431,1713,1739]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="2" pageNumber="65" unit="mm" value="1.75">1.75 mm</quantity>
; length of head between 0.25 and 
<quantity id="4CC2979F621330416179E6D3FD45F917" box="[536,655,1750,1776]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" pageId="2" pageNumber="65" unit="mm" value="0.375">0.375 mm</quantity>
(for different bending of head); width of head 
<quantity id="4CC2979F6213304167CFE6D3FAE2F917" box="[1198,1320,1750,1776]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" pageId="2" pageNumber="65" unit="mm" value="0.625">0.625 mm</quantity>
; height of head 
<quantity id="4CC2979F6213304163B9E6FEFE9CF8F2" box="[216,342,1787,1813]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" pageId="2" pageNumber="65" unit="mm" value="0.375">0.375 mm</quantity>
; width of eye 
<quantity id="4CC2979F621330416165E6FEFDAEF8F2" box="[516,612,1787,1813]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="65" unit="mm" value="0.3">0.3 mm</quantity>
; distance between eyes 
<quantity id="4CC2979F6213304160E6E6FEFBCFF8F2" box="[903,1029,1787,1813]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="65" unit="mm" value="0.075">0.075 mm</quantity>
; antennal segments IV 
<quantity id="4CC2979F621330416640E6FEFA56F8F2" box="[1313,1436,1787,1813]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" pageId="2" pageNumber="65" unit="mm" value="0.225">0.225 mm</quantity>
long, III 
<quantity id="4CC2979F621330416263E724FEB6F8DC" box="[258,380,1825,1851]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" pageId="2" pageNumber="65" unit="mm" value="0.275">0.275 mm</quantity>
long, II 
<quantity id="4CC2979F621330416287E724FD8CF8DC" box="[486,582,1825,1851]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="65" unit="mm" value="0.4">0.4 mm</quantity>
, I 
<quantity id="4CC2979F621330416104E724FD2AF8DC" box="[613,736,1825,1851]" metricMagnitude="-4" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="65" unit="mm" value="0.115">0.115 mm</quantity>
, antennal segment II the longest; length of pronotum 
<quantity id="4CC2979F621330416609E724FF03F887" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="65" unit="mm" value="0.35">0.35 mm</quantity>
, width 
<quantity id="4CC2979F621330416243E743FE56F887" box="[290,412,1862,1888]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" pageId="2" pageNumber="65" unit="mm" value="0.875">0.875 mm</quantity>
(max.), 
<quantity id="4CC2979F621330416164E743FDB5F887" box="[517,639,1862,1888]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" pageId="2" pageNumber="65" unit="mm" value="0.375">0.375 mm</quantity>
(min.); length of mesoscutum 
<quantity id="4CC2979F621330416099E743FBADF887" box="[1016,1127,1862,1888]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="65" unit="mm" value="0.15">0.15 mm</quantity>
; length of scutellum 
<quantity id="4CC2979F621330416606E743FF02F862" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="65" unit="mm" value="0.27">0.27 mm</quantity>
; claval commissure 
<quantity id="4CC2979F6213304162D9E76EFDE5F862" box="[440,559,1899,1925]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" pageId="2" pageNumber="65" unit="mm" value="0.525">0.525 mm</quantity>
long; costal fracture 
<quantity id="4CC2979F62133041604BE76EFC59F862" box="[810,915,1899,1925]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="65" unit="mm" value="0.25">0.25 mm</quantity>
long; hind femora 
<quantity id="4CC2979F621330416714E76EFB14F862" box="[1141,1246,1899,1925]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="2" pageNumber="65" unit="mm" value="0.87">0.87 mm</quantity>
long, (
<quantity id="4CC2979F621330416653E76EFA56F862" box="[1330,1436,1899,1925]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="65" unit="mm" value="1.05">1.05 mm</quantity>
for PA 6524), 
<quantity id="4CC2979F621330416221E794FE63F84C" box="[320,425,1937,1963]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="65" unit="mm" value="0.25">0.25 mm</quantity>
wide; hind tibia 
<quantity id="4CC2979F621330416112E794FD16F84C" box="[627,732,1937,1963]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="65" unit="mm" value="0.85">0.85 mm</quantity>
long, width mm, tarsal joints I 
<quantity id="4CC2979F621330416732E794FB04F84C" box="[1107,1230,1937,1963]" metricMagnitude="-5" metricUnit="m" metricValue="8.3" pageId="2" pageNumber="65" unit="mm" value="0.083">0.083 mm</quantity>
, II+III 
<quantity id="4CC2979F621330416645E794FA56F84C" box="[1316,1436,1937,1963]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="2" pageNumber="65" unit="mm" value="0.18">0.180 mm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="C32069F16213304063A4E7DEFAF5FE8B" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65" type="discussion">
<paragraph id="8B853A7A6213304063A4E7DEFAF5FE8B" blockId="2.[151,1438,1675,2037]" lastBlockId="3.[151,1437,152,551]" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65">
<emphasis id="B94EE6686213304163A4E7DEFE9DF812" bold="true" box="[197,343,2011,2037]" pageId="2" pageNumber="65">Discussion</emphasis>
. After 
<bibRefCitation id="EFAB478B6213304162D6E7DEFD34F812" author="Herczek" box="[439,766,2011,2037]" pageId="2" pageNumber="68" refString="Herczek, A. &amp; Popov, Yu. A. (1992) A remarkable psallopinous bug from Baltic amber (Miridae). Mitteilungen der Geologische-Palaontologische Institute der Universitat Hamburg, 73, 235 - 239." type="journal article" year="1992">Herczek &amp; Popov (1992)</bibRefCitation>
and Popov &amp; 
<bibRefCitation id="EFAB478B6213304160DCE7DEFB46F812" box="[957,1164,2011,2037]" pageId="2" pageNumber="68" refString="Popov, Yu. A. &amp; Herczek, A. (2006) Cylapopsallops kerhneri gen. et sp. n. - a new peculiar mirid from Baltic amber (Heteroptera: Miridae: Psallopinae). Russian Entomological Journal, 15 (2), 187 - 188." type="journal article">Herczek (2006)</bibRefCitation>
, the mirid subfamily Psallopinae is characterized by the following features: large eyes covering greater part of head, very deeply inserted antennae, absence of ocelli, and presence of subapical tooth on claws. These authors placed the two genera 
<taxonomicName id="4C3A41F962123040638DE0E7FE60FF1C" box="[236,426,226,251]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE66862123040638DE0E7FE60FF1C" box="[236,426,226,251]" italics="true" pageId="3" pageNumber="66">Isometopsallops</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3A41F9621230406283E0E7FD52FF1C" box="[482,664,226,251]" class="Insecta" family="Miridae" genus="Cylapopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE668621230406283E0E7FD52FF1C" box="[482,664,226,251]" italics="true" pageId="3" pageNumber="66">Cylapopsallops</emphasis>
</taxonomicName>
in this subfamily on the basis of these characters. 
<collectingCountry id="F32D7AEA621230406780E0E7FAC8FF1B" box="[1249,1282,226,252]" name="American Samoa" pageId="3" pageNumber="66">As</collectingCountry>
they are also shared by 
<taxonomicName id="4C3A41F9621230406275E10DFDF1FEC6" box="[276,571,264,289]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="prokopi" status="sp. nov.">
<emphasis id="B94EE668621230406275E10DFDF1FEC6" box="[276,571,264,289]" italics="true" pageId="3" pageNumber="66">Isometopsallops prokopi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D5B13621230406122E10DFD69FEC5" box="[579,675,264,290]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
, we consider that it also belongs to this subfamily. One further character that potentially supports an attribution of 
<taxonomicName id="4C3A41F9621230406073E128FC40FEA1" box="[786,906,301,327]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="prokopi">
<emphasis id="B94EE668621230406073E128FCD6FEA1" box="[786,796,301,326]" italics="true" pageId="3" pageNumber="66">I</emphasis>
. 
<emphasis id="B94EE66862123040604DE128FC40FEA1" box="[812,906,301,326]" italics="true" pageId="3" pageNumber="66">prokopi</emphasis>
</taxonomicName>
to the Psallopinae is the presence of seven metafemoral trichobothria, a putative apomorphy also present in the 
<taxonomicName id="4C3A41F96212304060DAE157FBEFFE8B" box="[955,1061,338,364]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
, after 
<bibRefCitation id="EFAB478B62123040670CE157FAF1FE8B" author="Schuh" box="[1133,1339,338,364]" pageId="3" pageNumber="68" refString="Schuh, R. T. (1976) Pretarsal structure in the Miridae (Hemiptera) with a cladistic analysis of relationships within the family. American Museum Novitates, 2601, 1 - 26." type="journal article" year="1976">Schuh (1976: 13)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32069F16212304063A4E17DFCF0F93C" pageId="3" pageNumber="66" type="reference_group">
<paragraph id="8B853A7A6212304063A4E17DFDA6FDC0" blockId="3.[151,1437,152,551]" pageId="3" pageNumber="66">
The 
<taxonomicName id="4C3A41F962123040639CE17DFE6CFE75" box="[253,422,376,402]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
, 
<taxonomicName id="4C3A41F96212304062D2E17DFDF9FE75" box="[435,563,376,402]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Cylapinae">Cylapinae</taxonomicName>
, and Psallopinae share also plesiomorphic characters, i.e., parempodia setiform (
<bibRefCitation id="EFAB478B621230406269E198FDFEFE50" author="Schuh" box="[264,564,413,439]" pageId="3" pageNumber="68" refString="Schuh, R. T. &amp; Schwartz, M. D. (1984) Carvalhoma (Hemiptera: Miridae): revised subfamily placement. Journal of the New York Entomological Society, 92, 48 - 52." type="journal article" year="1984">Schuh &amp; Schwartz, 1984</bibRefCitation>
; 
<bibRefCitation id="EFAB478B621230406120E198FCF8FE50" author="Schuh" box="[577,818,413,439]" pageId="3" pageNumber="68" refString="Schuh, R. T. &amp; Stys, P. 1991. Phylogenetic analysis of cimicomorphan family relationships (Heteroptera). Journal of the New York Entomological Society, 99, 298 - 350." type="journal article" year="1991">Schuh &amp; Štys, 1991</bibRefCitation>
); and absence of pulvilli (
<bibRefCitation id="EFAB478B621230406705E198FACEFE50" author="Schuh" box="[1124,1284,413,439]" pageId="3" pageNumber="68" refString="Schuh, R. T. (1976) Pretarsal structure in the Miridae (Hemiptera) with a cladistic analysis of relationships within the family. American Museum Novitates, 2601, 1 - 26." type="journal article" year="1976">Schuh [1976]</bibRefCitation>
observed no pulvilli in 
<taxonomicName id="4C3A41F9621230406271E1C7FE7BFE3B" box="[272,433,450,476]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
, Psallopinae, and 
<taxonomicName id="4C3A41F96212304061E2E1C7FD3FFE3B" box="[643,757,450,476]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phasmida" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Phyllinae">Phyllinae</taxonomicName>
; for 
<taxonomicName id="4C3A41F9621230406048E1C7FC69FE3B" box="[809,931,450,476]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Cylapinae">Cylapinae</taxonomicName>
, pulvilli are usually present and attached to ventral surface of claw which is an apomorphic character); antennae in inner side of eyes (
<bibRefCitation id="EFAB478B621230406676E1EDFE9BFDC0" author="Schuh" pageId="3" pageNumber="68" refString="Schuh, R. T. &amp; Schwartz, M. D. (1984) Carvalhoma (Hemiptera: Miridae): revised subfamily placement. Journal of the New York Entomological Society, 92, 48 - 52." type="journal article" year="1984">Schuh and Schwartz, 1984</bibRefCitation>
; 
<bibRefCitation id="EFAB478B62123040623CE208FD95FDC0" author="Schuh" box="[349,607,525,551]" pageId="3" pageNumber="68" refString="Schuh, R. T. &amp; Stys, P. 1991. Phylogenetic analysis of cimicomorphan family relationships (Heteroptera). Journal of the New York Entomological Society, 99, 298 - 350." type="journal article" year="1991">Schuh and Štys, 1991</bibRefCitation>
).
</paragraph>
<caption id="DF456AF26212304063F6E4BEFB0EFB34" box="[151,1220,1211,1235]" httpUri="https://zenodo.org/record/195789/files/figure.png" pageId="3" pageNumber="66" targetBox="[226,1360,607,1186]" targetPageId="3">
<paragraph id="8B853A7A6212304063F6E4BEFB0EFB34" blockId="3.[151,1220,1211,1235]" box="[151,1220,1211,1235]" pageId="3" pageNumber="66">
<emphasis id="B94EE6686212304063F6E4BEFED3FB34" bold="true" box="[151,281,1211,1235]" pageId="3" pageNumber="66">FIGURE 3.</emphasis>
<taxonomicName id="4C3A41F9621230406241E4B9FDECFB34" ID-CoL="6MZ5C" box="[288,550,1212,1235]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="prokopi" status="sp. nov.">
<emphasis id="B94EE668621230406241E4B9FDECFB34" box="[288,550,1212,1235]" italics="true" pageId="3" pageNumber="66">Isometopsallops prokopi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D5B1362123040614DE4BEFD48FB34" box="[556,642,1211,1235]" pageId="3" pageNumber="66" rank="species">
<emphasis id="B94EE66862123040614DE4BEFDB1FB34" bold="true" box="[556,635,1211,1235]" pageId="3" pageNumber="66">sp. nov</emphasis>
.
</taxonomicNameLabel>
paratype MNHN A30854, photograph of dorsal view.
</paragraph>
</caption>
<paragraph id="8B853A7A6212304063A4E504FCF0F93C" blockId="3.[151,1437,1281,2016]" pageId="3" pageNumber="66">
The Psallopinae is currently considered to be related to the two subfamilies, 
<taxonomicName id="4C3A41F962123040672CE504FB0EFAFC" box="[1101,1220,1281,1307]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Cylapinae">Cylapinae</taxonomicName>
and 
<taxonomicName id="4C3A41F962123040679FE504FA56FAFC" box="[1278,1436,1281,1307]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
(
<bibRefCitation id="EFAB478B6212304063FEE523FEA2FAA7" author="Gorczyca" box="[159,360,1318,1344]" pageId="3" pageNumber="68" refString="Gorczyca, J. (2000) A systematic study on Cylapinae with a revision of the Afrotropical region (Heteroptera, Miridae). Wydawnictwo Uniwersystetu Slaskiego, Katowice, 176 pp." type="book" year="2000">Gorczyca, 2000</bibRefCitation>
; 
<bibRefCitation id="EFAB478B621230406214E523FD91FAA7" author="Lin" box="[373,603,1318,1344]" pageId="3" pageNumber="68" refString="Lin, C. - S. &amp; Yang, C. - T. (2005) External male genitalia of the Miridae (Hemiptera: Heteroptera). National Museum of Natural Science Special Publication, 9, 1 - 174." type="journal article" year="2005">Lin &amp; Yang, 2005</bibRefCitation>
). 
<bibRefCitation id="EFAB478B621230406111E523FC8AFAA7" author="Gorczyca" box="[624,832,1318,1344]" pageId="3" pageNumber="68" refString="Gorczyca, J. (2000) A systematic study on Cylapinae with a revision of the Afrotropical region (Heteroptera, Miridae). Wydawnictwo Uniwersystetu Slaskiego, Katowice, 176 pp." type="book" year="2000">Gorczyca (2000)</bibRefCitation>
supported the clade (
<taxonomicName id="4C3A41F962123040672BE523FB02FAA7" box="[1098,1224,1318,1344]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Cylapinae">Cylapinae</taxonomicName>
+ (Psallopinae + 
<taxonomicName id="4C3A41F96212304063F6E54EFEF3FA82" box="[151,313,1355,1381]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
)) on the basis of two putative synapomorphies: presence of claws with a subapical tooth, and tarsi two-segmented. The presence of three-segmented tarsi in 
<taxonomicName id="4C3A41F9621230406013E574FC2EFA6D" ID-CoL="6MZ5C" box="[882,996,1393,1419]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="prokopi">
<emphasis id="B94EE668621230406013E574FCB6FA6D" box="[882,892,1393,1418]" italics="true" pageId="3" pageNumber="66">I</emphasis>
. 
<emphasis id="B94EE6686212304060EBE574FC2EFA6D" box="[906,996,1393,1418]" italics="true" pageId="3" pageNumber="66">prokopi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3A41F9621230406093E574FB9CFA6D" ID-CoL="6MYTB" box="[1010,1110,1393,1419]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B94EE668621230406093E574FC36FA6D" box="[1010,1020,1393,1418]" italics="true" pageId="3" pageNumber="66">I</emphasis>
. 
<emphasis id="B94EE66862123040676BE574FB9CFA6D" box="[1034,1110,1393,1418]" italics="true" pageId="3" pageNumber="66">schuhi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3A41F96212304067F4E574FA81FA6D" ID-CoL="62Q2J" box="[1173,1355,1393,1418]" class="Insecta" family="Miridae" genus="Cylapopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE6686212304067F4E574FA81FA6D" box="[1173,1355,1393,1418]" italics="true" pageId="3" pageNumber="66">Cylapopsallops</emphasis>
</taxonomicName>
would then be problematic for their attribution to the Psallopinae. 
<bibRefCitation id="EFAB478B621230406017E593FB7CFA57" author="Schuh" box="[886,1206,1430,1456]" pageId="3" pageNumber="68" refString="Schuh, R. T. &amp; Schwartz, M. D. (1984) Carvalhoma (Hemiptera: Miridae): revised subfamily placement. Journal of the New York Entomological Society, 92, 48 - 52." type="journal article" year="1984">Schuh &amp; Schwartz (1984)</bibRefCitation>
and 
<bibRefCitation id="EFAB478B621230406792E593FF23FA32" author="Schuh" pageId="3" pageNumber="68" refString="Schuh, R. T. &amp; Stys, P. 1991. Phylogenetic analysis of cimicomorphan family relationships (Heteroptera). Journal of the New York Entomological Society, 99, 298 - 350." type="journal article" year="1991">Schuh &amp; Štys (1991)</bibRefCitation>
considered three-segmented tarsi as plesiomorphic. This character state is present in other mirid subfamilies. However, there are some cylapine genera as 
<taxonomicName id="4C3A41F9621230406058E5E4FC14FA1D" ID-CoL="89R7" box="[825,990,1505,1530]" class="Insecta" family="Miridae" genus="Yamatofulvius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE668621230406058E5E4FC14FA1D" box="[825,990,1505,1530]" italics="true" pageId="3" pageNumber="66">Yamatofulvius</emphasis>
</taxonomicName>
where all the species have apparently functional three-segmented tarsi (
<bibRefCitation id="EFAB478B621230406140E603FD29F9C7" author="Yasunaga" box="[545,739,1542,1568]" pageId="3" pageNumber="68" refString="Yasunaga, T. (2000) The mirid subfamily Cylapinae (Heteroptera: Miridae) or fungal inhabiting plant bugs in Japan. Tijdschrift voor Entomologie, 143, 183 - 209." type="journal article" year="2000">Yasunaga, 2000</bibRefCitation>
). 
<bibRefCitation id="EFAB478B621230406196E603FC5FF9C7" author="Schuh" box="[759,917,1542,1568]" pageId="3" pageNumber="68" refString="Schuh, R. T. (1976) Pretarsal structure in the Miridae (Hemiptera) with a cladistic analysis of relationships within the family. American Museum Novitates, 2601, 1 - 26." type="journal article" year="1976">Schuh (1976)</bibRefCitation>
described also 
<taxonomicName id="4C3A41F962123040672CE603FB0EF9C7" box="[1101,1220,1542,1568]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Cylapinae">Cylapinae</taxonomicName>
and 
<taxonomicName id="4C3A41F962123040679FE603FA56F9C7" box="[1278,1436,1542,1568]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
with tarsi two- or three-segmented (for 
<taxonomicName id="4C3A41F9621230406106E62EFCC2F9A2" box="[615,776,1579,1605]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
, see also 
<bibRefCitation id="EFAB478B621230406016E62EFBF9F9A2" author="Herczek" box="[887,1075,1579,1605]" pageId="3" pageNumber="68" refString="Herczek, A. (1993) Systematic position of the Isometopinae Fieb. (Miridae, Heteroptera) and their interrelationships. Prace Naukowe Uniwersytetu S l q skiego, 1357, 1 - 86." type="journal article" year="1993">Herczek [1993]</bibRefCitation>
). Thus this character is at least homoplasic. Lastly, 
<bibRefCitation id="EFAB478B6212304062EBE654FDB8F98C" author="Lin" box="[394,626,1617,1643]" pageId="3" pageNumber="68" refString="Lin, C. - S. &amp; Yang, C. - T. (2005) External male genitalia of the Miridae (Hemiptera: Heteroptera). National Museum of Natural Science Special Publication, 9, 1 - 174." type="journal article" year="2005">Lin &amp; Yang (2005)</bibRefCitation>
indicated: &quot;according to Schuh &amp; Slater (1996: 172) tarsi are twosegmented in the 
<taxonomicName id="4C3A41F9621230406217E673FDD4F977" box="[374,542,1654,1680]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
and Psallopinae; two- or three-segmented in the 
<taxonomicName id="4C3A41F96212304067ECE673FADAF977" box="[1165,1296,1654,1680]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Cylapinae">Cylapinae</taxonomicName>
; and threesegmented in the 
<taxonomicName id="4C3A41F962123040620EE69EFDC6F952" box="[367,524,1691,1717]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Orthotylinae">Orthotylinae</taxonomicName>
, 
<taxonomicName id="4C3A41F9621230406179E69EFD4CF952" box="[536,646,1691,1717]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
, 
<taxonomicName id="4C3A41F96212304061F2E69EFCE6F952" box="[659,812,1691,1717]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Bryocorinae">Bryocorinae</taxonomicName>
, 
<taxonomicName id="4C3A41F962123040605BE69EFC24F952" box="[826,1006,1691,1717]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Deraeocorinae">Deraeocorinae</taxonomicName>
, and 
<taxonomicName id="4C3A41F962123040674EE69EFB5CF952" box="[1071,1174,1691,1717]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Mirinae">Mirinae</taxonomicName>
.&quot; The two-segmented tarsus is judged to be plesiomorphic by these last authors.
</paragraph>
</subSubSection>
<subSubSection id="C32069F16212304063A4E6E3FABDF807" pageId="3" pageNumber="66" type="description">
<paragraph id="8B853A7A6212304063A4E6E3FCEBF8AC" blockId="3.[151,1437,1281,2016]" pageId="3" pageNumber="66">
The presence of large eyes (also present in the three fossils 
<taxonomicName id="4C3A41F96212304060C1E6E2FBD2F8E7" box="[928,1048,1766,1792]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="prokopi">
<emphasis id="B94EE6686212304060C1E6E2FC60F8E7" box="[928,938,1767,1792]" italics="true" pageId="3" pageNumber="66">I</emphasis>
. 
<emphasis id="B94EE6686212304060DAE6E2FBD2F8E7" box="[955,1048,1767,1792]" italics="true" pageId="3" pageNumber="66">prokopi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3A41F9621230406749E6E2FB5BF8E7" box="[1064,1169,1766,1792]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B94EE668621230406749E6E2FBF8F8E7" box="[1064,1074,1767,1792]" italics="true" pageId="3" pageNumber="66">I</emphasis>
. 
<emphasis id="B94EE668621230406723E6E2FB5BF8E7" box="[1090,1169,1767,1792]" italics="true" pageId="3" pageNumber="66">schuhi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3A41F96212304067B7E6E2FA58F8E7" box="[1238,1426,1767,1792]" class="Insecta" family="Miridae" genus="Cylapopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE6686212304067B7E6E2FA58F8E7" box="[1238,1426,1767,1792]" italics="true" pageId="3" pageNumber="66">Cylapopsallops</emphasis>
</taxonomicName>
) would be the main apomorphic characters supporting the clade (
<taxonomicName id="4C3A41F96212304060D9E70EFBABF8C2" box="[952,1121,1803,1829]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
+ Psallopinae) (
<bibRefCitation id="EFAB478B62123040664CE70EFE9BF8AC" author="Schuh" pageId="3" pageNumber="68" refString="Schuh, R. T. &amp; Schwartz, M. D. (1984) Carvalhoma (Hemiptera: Miridae): revised subfamily placement. Journal of the New York Entomological Society, 92, 48 - 52." type="journal article" year="1984">Schuh &amp; Schwartz, 1984</bibRefCitation>
; 
<bibRefCitation id="EFAB478B62123040623DE734FD80F8AC" author="Schuh" box="[348,586,1841,1867]" pageId="3" pageNumber="68" refString="Schuh, R. T. &amp; Stys, P. 1991. Phylogenetic analysis of cimicomorphan family relationships (Heteroptera). Journal of the New York Entomological Society, 99, 298 - 350." type="journal article" year="1991">Schuh &amp; Štys, 1991</bibRefCitation>
; 
<bibRefCitation id="EFAB478B621230406137E734FCDFF8AC" author="Gorczyca" box="[598,789,1841,1867]" pageId="3" pageNumber="68" refString="Gorczyca, J. (2000) A systematic study on Cylapinae with a revision of the Afrotropical region (Heteroptera, Miridae). Wydawnictwo Uniwersystetu Slaskiego, Katowice, 176 pp." type="book" year="2000">Gorczyca, 2000</bibRefCitation>
).
</paragraph>
<paragraph id="8B853A7A6212304063A4E753FABDF807" blockId="3.[151,1437,1281,2016]" pageId="3" pageNumber="66">
Within the clade (
<taxonomicName id="4C3A41F96212304062F9E753FDF3F897" box="[408,569,1878,1904]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
+ Psallopinae), the absence of ocelli characterises the Psallopinae and is the unique apomorphy visible on the three fossils 
<taxonomicName id="4C3A41F962123040619EE779FCBDF872" box="[767,887,1915,1941]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="prokopi">
<emphasis id="B94EE66862123040619EE779FCC3F872" box="[767,777,1916,1941]" italics="true" pageId="3" pageNumber="66">I</emphasis>
. 
<emphasis id="B94EE668621230406078E779FCBDF872" box="[793,887,1916,1941]" italics="true" pageId="3" pageNumber="66">prokopi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3A41F96212304060E6E779FC3FF872" box="[903,1013,1915,1941]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B94EE6686212304060E6E779FC5BF872" box="[903,913,1916,1941]" italics="true" pageId="3" pageNumber="66">I</emphasis>
. 
<emphasis id="B94EE6686212304060C3E779FC3FF872" box="[930,1013,1916,1941]" italics="true" pageId="3" pageNumber="66">schuhi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3A41F9621230406757E779FB39F872" box="[1078,1267,1916,1941]" class="Insecta" family="Miridae" genus="Cylapopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE668621230406757E779FB39F872" box="[1078,1267,1916,1941]" italics="true" pageId="3" pageNumber="66">Cylapopsallops</emphasis>
</taxonomicName>
. For 
<bibRefCitation id="EFAB478B621230406656E77EFF22F85C" author="Herczek" pageId="3" pageNumber="68" refString="Herczek, A. (1993) Systematic position of the Isometopinae Fieb. (Miridae, Heteroptera) and their interrelationships. Prace Naukowe Uniwersytetu S l q skiego, 1357, 1 - 86." type="journal article" year="1993">Herczek (1993)</bibRefCitation>
, it seems obvious that the occurrence of ocelli is plesiomorphic. 
<bibRefCitation id="EFAB478B621230406091E7A4FAECF85C" author="Schuh" box="[1008,1318,1953,1979]" pageId="3" pageNumber="68" refString="Schuh, R. T. &amp; Schwartz, M. D. (1984) Carvalhoma (Hemiptera: Miridae): revised subfamily placement. Journal of the New York Entomological Society, 92, 48 - 52." type="journal article" year="1984">Schuh &amp; Schwartz (1984)</bibRefCitation>
preferred the interpretation of character distributions as a convergent lose of the ocelli in Psallopinae and 
<taxonomicName id="4C3A41F9621230406799E7C3FAB9F807" box="[1272,1395,1990,2016]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Cylapinae">Cylapinae</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF456AF26215304763F6E431FB50FBAC" box="[151,1178,1076,1100]" httpUri="https://zenodo.org/record/195790/files/figure.png" pageId="4" pageNumber="67" targetBox="[151,1434,194,1053]" targetPageId="4">
<paragraph id="8B853A7A6215304763F6E431FB50FBAC" blockId="4.[151,1178,1076,1100]" box="[151,1178,1076,1100]" pageId="4" pageNumber="67">
<emphasis id="B94EE6686215304763F6E431FED3FBAB" bold="true" box="[151,281,1076,1100]" pageId="4" pageNumber="67">FIGURE 4.</emphasis>
<taxonomicName id="4C3A41F9621530476241E430FDECFBAB" box="[288,550,1077,1100]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="prokopi" status="sp. nov.">
<emphasis id="B94EE668621530476241E430FDECFBAB" box="[288,550,1077,1100]" italics="true" pageId="4" pageNumber="67">Isometopsallops prokopi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D5B1362153047614DE431FD48FBAC" box="[556,642,1076,1100]" pageId="4" pageNumber="67" rank="species">
<emphasis id="B94EE66862153047614DE431FDB1FBAB" bold="true" box="[556,635,1076,1100]" pageId="4" pageNumber="67">sp. nov</emphasis>
.
</taxonomicNameLabel>
allotype MNHN A30852, photograph of hind leg.
</paragraph>
</caption>
<subSubSection id="C32069F16215304763A4E47FFC59FAA9" pageId="4" pageNumber="67" type="reference_group">
<paragraph id="8B853A7A6215304763A4E47FFB8BFB39" blockId="4.[151,1437,1146,2030]" pageId="4" pageNumber="67">
<taxonomicName id="4C3A41F96215304763A4E47FFE88FB74" ID-CoL="6MZ5C" box="[197,322,1146,1172]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="prokopi">
<emphasis id="B94EE6686215304763A4E47FFF05FB74" box="[197,207,1146,1171]" italics="true" pageId="4" pageNumber="67">I</emphasis>
. 
<emphasis id="B94EE668621530476380E47FFE88FB74" box="[225,322,1146,1171]" italics="true" pageId="4" pageNumber="67">prokopi</emphasis>
</taxonomicName>
differs from the recent 
<taxonomicName id="4C3A41F9621530476113E47FFD14FB74" ID-CoL="6Y8Z" box="[626,734,1146,1171]" class="Insecta" family="Miridae" genus="Psallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE668621530476113E47FFD14FB74" box="[626,734,1146,1171]" italics="true" pageId="4" pageNumber="67">Psallops</emphasis>
</taxonomicName>
in that the right genital style is larger than the left one (
<bibRefCitation id="EFAB478B6215304763FEE49AFEABFB5E" author="Yasunaga" box="[159,353,1183,1209]" pageId="4" pageNumber="68" refString="Yasunaga, T. (1999) First record of the plant bug subfamily Psallopinae (Heteroptera: Miridae) from Japan, with descriptions of three new species of the genus Psallops Usinger. Proceedings of the Entomological Society of Washington, 101, 737 - 741." type="journal article" year="1999">Yasunaga, 1999</bibRefCitation>
; 
<bibRefCitation id="EFAB478B62153047620FE49AFE2CFB5E" author="Lin" box="[366,486,1183,1209]" pageId="4" pageNumber="68" refString="Lin, C. - S. (2004) First record of the plant bug subfamily Psallopinae (Heteroptera: Miridae) from Taiwan and China, with descriptions of three new species of the genus Psallops Usinger. Formosan Entomology, 24, 273 - 279." type="journal article" year="2004">Lin, 2004</bibRefCitation>
; 
<bibRefCitation id="EFAB478B621530476292E49AFD1BFB5E" author="Lin" box="[499,721,1183,1209]" pageId="4" pageNumber="68" refString="Lin, C. - S. &amp; Yang, C. - T. (2005) External male genitalia of the Miridae (Hemiptera: Heteroptera). National Museum of Natural Science Special Publication, 9, 1 - 174." type="journal article" year="2005">Lin &amp; Yang, 2005</bibRefCitation>
). In recent 
<taxonomicName id="4C3A41F9621530476036E49AFC19FB5E" box="[855,979,1183,1209]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="subFamily" subFamily="Cylapinae">Cylapinae</taxonomicName>
, the genital styles are asymmetrical in shape and size, one either left or right larger, depending on the species or genus.
</paragraph>
<paragraph id="8B853A7A6215304763A4E4EFFC59FAA9" blockId="4.[151,1437,1146,2030]" pageId="4" pageNumber="67">
Thus nearly all the characters currently used to support the Psallopinae, 
<taxonomicName id="4C3A41F962153047675BE4EFFB72FAE3" box="[1082,1208,1258,1284]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="subFamily" subFamily="Cylapinae">Cylapinae</taxonomicName>
, and 
<taxonomicName id="4C3A41F9621530476798E4EFFA56FAE3" box="[1273,1436,1258,1284]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
relationships are homoplastic. And so the positions of the three fossils (
<taxonomicName id="4C3A41F9621530476735E515FB1CFACE" ID-CoL="6MZ5C" box="[1108,1238,1295,1321]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="prokopi">
<emphasis id="B94EE668621530476735E515FB94FACE" box="[1108,1118,1296,1321]" italics="true" pageId="4" pageNumber="67">I</emphasis>
. 
<emphasis id="B94EE668621530476713E515FB1CFACE" box="[1138,1238,1296,1321]" italics="true" pageId="4" pageNumber="67">prokopi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3A41F9621530476788E515FA96FACE" ID-CoL="6MYTB" box="[1257,1372,1295,1321]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B94EE668621530476788E515FB39FACE" box="[1257,1267,1296,1321]" italics="true" pageId="4" pageNumber="67">I</emphasis>
. 
<emphasis id="B94EE668621530476666E515FA96FACE" box="[1287,1372,1296,1321]" italics="true" pageId="4" pageNumber="67">schuhi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3A41F96215304763F6E530FE87FAA9" ID-CoL="62Q2J" box="[151,333,1333,1358]" class="Insecta" family="Miridae" genus="Cylapopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE6686215304763F6E530FE87FAA9" box="[151,333,1333,1358]" italics="true" pageId="4" pageNumber="67">Cylapopsallops</emphasis>
</taxonomicName>
) may perhaps be subject to changes in the future.
</paragraph>
</subSubSection>
<subSubSection id="C32069F16215304663A4E55FFEDCFDC0" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67" type="description">
<paragraph id="8B853A7A6215304763A4E55FFE1AFA59" blockId="4.[151,1437,1146,2030]" pageId="4" pageNumber="67">
<taxonomicName id="4C3A41F96215304763A4E55FFEF2FA94" box="[197,312,1370,1396]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="prokopi">
<emphasis id="B94EE6686215304763A4E55FFF05FA94" box="[197,207,1370,1395]" italics="true" pageId="4" pageNumber="67">I</emphasis>
. 
<emphasis id="B94EE6686215304763BFE55FFEF2FA94" box="[222,312,1370,1395]" italics="true" pageId="4" pageNumber="67">prokopi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C3A41F96215304762B7E55FFD47FA94" box="[470,653,1370,1395]" class="Insecta" family="Miridae" genus="Cylapopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE6686215304762B7E55FFD47FA94" box="[470,653,1370,1395]" italics="true" pageId="4" pageNumber="67">Cylapopsallops</emphasis>
</taxonomicName>
because its body is less elongate (only two times as long as wide, not three times as long as wide); broad femora, not flattened in 
<taxonomicName id="4C3A41F9621530476018E585FBE5FA7E" box="[889,1071,1408,1433]" class="Insecta" family="Miridae" genus="Cylapopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE668621530476018E585FBE5FA7E" box="[889,1071,1408,1433]" italics="true" pageId="4" pageNumber="67">Cylapopsallops</emphasis>
</taxonomicName>
; collar absent, instead of being distinct in 
<taxonomicName id="4C3A41F9621530476272E5A0FE03FA59" box="[275,457,1445,1470]" class="Insecta" family="Miridae" genus="Cylapopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE668621530476272E5A0FE03FA59" box="[275,457,1445,1470]" italics="true" pageId="4" pageNumber="67">Cylapopsallops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B853A7A6215304763A4E5CFFE71F923" blockId="4.[151,1437,1146,2030]" pageId="4" pageNumber="67">
The habitus of 
<taxonomicName id="4C3A41F962153047621DE5CFFE3BFA04" box="[380,497,1482,1508]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="prokopi">
<emphasis id="B94EE66862153047621DE5CFFE4CFA04" box="[380,390,1482,1507]" italics="true" pageId="4" pageNumber="67">I</emphasis>
. 
<emphasis id="B94EE6686215304762F4E5CFFE3BFA04" box="[405,497,1482,1507]" italics="true" pageId="4" pageNumber="67">prokopi</emphasis>
</taxonomicName>
is very similar to that of 
<taxonomicName id="4C3A41F9621530476044E5CFFC47FA04" box="[805,909,1482,1508]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B94EE668621530476044E5CFFCE5FA04" box="[805,815,1482,1507]" italics="true" pageId="4" pageNumber="67">I</emphasis>
. 
<emphasis id="B94EE66862153047605EE5CFFC47FA04" box="[831,909,1482,1507]" italics="true" pageId="4" pageNumber="67">schuhi</emphasis>
</taxonomicName>
. They share the same characters mentioned above: tarsi three-segmented (apparently pseudo-trisegmentation); subapical tooth on each claw; trapezoidal pronotum; and two-celled membrane of hemelytra. The ratio length/width of the pronotum = 
<quantity id="4CC2979F6215304767BFE611FAE8F9C9" box="[1246,1314,1556,1582]" metricMagnitude="-2" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="67" unit="in" value="0.4">0.4 in</quantity>
<taxonomicName id="4C3A41F9621530476648E610FA56F9C9" box="[1321,1436,1556,1582]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="prokopi">
<emphasis id="B94EE668621530476648E610FAF9F9C9" box="[1321,1331,1557,1582]" italics="true" pageId="4" pageNumber="67">I</emphasis>
. 
<emphasis id="B94EE668621530476623E610FA56F9C9" box="[1346,1436,1557,1582]" italics="true" pageId="4" pageNumber="67">prokopi</emphasis>
</taxonomicName>
but 
<quantity id="4CC2979F6215304763A5E63FFEDCF9B3" box="[196,278,1594,1620]" metricMagnitude="-2" metricUnit="m" metricValue="1.6764" pageId="4" pageNumber="67" unit="in" value="0.66">0.66 in</quantity>
<taxonomicName id="4C3A41F962153047627FE63FFDFBF9B4" box="[286,561,1594,1619]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B94EE66862153047627FE63FFDFBF9B4" box="[286,561,1594,1619]" italics="true" pageId="4" pageNumber="67">Isometopsallops schuhi</emphasis>
</taxonomicName>
. We lack information on the number of femoral trichobothria in the latter, which renders their comparison for now a little less positive. With the present state of knowledge we prefer to include our fossil in a new species of the same genus 
<taxonomicName id="4C3A41F9621530476075E680FC19F979" box="[788,979,1669,1694]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE668621530476075E680FC19F979" box="[788,979,1669,1694]" italics="true" pageId="4" pageNumber="67">Isometopsallops</emphasis>
</taxonomicName>
. Unfortunately the male genitalia of 
<taxonomicName id="4C3A41F96215304766EAE680FF2EF924" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B94EE6686215304766EAE680FA5FF979" box="[1419,1429,1669,1694]" italics="true" pageId="4" pageNumber="67">I</emphasis>
. 
<emphasis id="B94EE6686215304763F6E6AFFF2EF924" box="[151,228,1706,1731]" italics="true" pageId="4" pageNumber="67">schuhi</emphasis>
</taxonomicName>
are not described.
</paragraph>
<paragraph id="8B853A7A6215304763A4E6CAFEDAF8D3" blockId="4.[151,1437,1146,2030]" pageId="4" pageNumber="67">
The presence of a new species belonging to the Baltic amber genus 
<taxonomicName id="4C3A41F962153047609DE6D5FB75F90E" box="[1020,1215,1744,1769]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE66862153047609DE6D5FB75F90E" box="[1020,1215,1744,1769]" italics="true" pageId="4" pageNumber="67">Isometopsallops</emphasis>
</taxonomicName>
in the Lowermost Eocene French amber is not surprising because similar fossils occur in other orders (Psocoptera, etc.) (
<bibRefCitation id="EFAB478B621530476632E6F1FECEF8D3" author="Nel" pageId="4" pageNumber="68" refString="Nel, A., Prokop, J., De Ploeg, G. &amp; Millet, J. (2005) New Psocoptera (Insecta) from the Lowermost Eocene amber of Oise, France. Journal of Systematic Palaeontology, 3, 371 - 391." type="journal article" year="2005">
Nel 
<emphasis id="B94EE6686215304766E6E6F0FF64F8D4" italics="true" pageId="4" pageNumber="67">et al</emphasis>
., 2005
</bibRefCitation>
).
</paragraph>
<paragraph id="8B853A7A6215304763A4E73AFA50F809" blockId="4.[151,1437,1146,2030]" pageId="4" pageNumber="67">
The genus 
<taxonomicName id="4C3A41F9621530476228E745FE65F8BE" box="[329,431,1856,1881]" class="Insecta" family="Miridae" genus="Psallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE668621530476228E745FE65F8BE" box="[329,431,1856,1881]" italics="true" pageId="4" pageNumber="67">Psallops</emphasis>
</taxonomicName>
comprises an undescribed Eocene species from Baltic amber, plus 10 known recent species in the world (
<bibRefCitation id="EFAB478B6215304762FCE761FDF1F899" author="Schuh" box="[413,571,1892,1918]" pageId="4" pageNumber="68" refString="Schuh, R. T. (1995) Plant bugs of the World (Insecta: Heteroptera: Miridae): systematic catalog, distributions, host list, and bibliography. The New York Entomological Society, New York, 1329 pp." type="book" year="1995">Schuh, 1995</bibRefCitation>
; 
<bibRefCitation id="EFAB478B621530476129E761FCC7F899" author="Yasunaga" box="[584,781,1892,1918]" pageId="4" pageNumber="68" refString="Yasunaga, T. (1999) First record of the plant bug subfamily Psallopinae (Heteroptera: Miridae) from Japan, with descriptions of three new species of the genus Psallops Usinger. Proceedings of the Entomological Society of Washington, 101, 737 - 741." type="journal article" year="1999">Yasunaga, 1999</bibRefCitation>
; 
<bibRefCitation id="EFAB478B62153047607AE761FC5CF899" author="Lin" box="[795,918,1892,1918]" pageId="4" pageNumber="68" refString="Lin, C. - S. (2004) First record of the plant bug subfamily Psallopinae (Heteroptera: Miridae) from Taiwan and China, with descriptions of three new species of the genus Psallops Usinger. Formosan Entomology, 24, 273 - 279." type="journal article" year="2004">Lin, 2004</bibRefCitation>
; Popov &amp; 
<bibRefCitation id="EFAB478B621530476779E761FB1BF899" author="Herczek" box="[1048,1233,1892,1918]" pageId="4" pageNumber="68" refString="Popov, Yu. A. &amp; Herczek, A. (2006) Cylapopsallops kerhneri gen. et sp. n. - a new peculiar mirid from Baltic amber (Heteroptera: Miridae: Psallopinae). Russian Entomological Journal, 15 (2), 187 - 188." type="journal article" year="2006">Herczek, 2006</bibRefCitation>
): 
<taxonomicName id="4C3A41F9621530476787E760FE21F843" authority="Linnavuori &amp; Alamy, 1982" authorityName="Linnavuori &amp; Alamy" authorityYear="1982" class="Insecta" family="Miridae" genus="Psallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="grandoculus">
<emphasis id="B94EE668621530476787E760FA56F899" box="[1254,1436,1893,1918]" italics="true" pageId="4" pageNumber="67">P. grandoculus</emphasis>
Linnavuori &amp; Alamy, 1982
</taxonomicName>
from 
<collectingCountry id="F32D7AEA621530476159E78FFD2BF843" box="[568,737,1930,1956]" name="Saudi Arabia" pageId="4" pageNumber="67">Saudi Arabia</collectingCountry>
; 
<taxonomicName id="4C3A41F962153047618FE78FFBFAF843" authority="Usinger, 1946" authorityName="Usinger" authorityYear="1946" box="[750,1072,1930,1956]" class="Insecta" family="Miridae" genus="Psallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="B94EE66862153047618FE78FFCB2F844" box="[750,888,1930,1955]" italics="true" pageId="4" pageNumber="67">P. oculatus</emphasis>
Usinger, 1946
</taxonomicName>
from 
<collectingCountry id="F32D7AEA62153047671CE78FFB02F843" box="[1149,1224,1930,1956]" name="Guam" pageId="4" pageNumber="67">Guam</collectingCountry>
and the 
<collectingCountry id="F32D7AEA621530476654E78FFF24F82E" name="Northern Mariana Islands" pageId="4" pageNumber="67">Mariana Islands</collectingCountry>
, Saipan; 
<taxonomicName id="4C3A41F962153047623DE7B5FD74F82E" authority="Carvalho, 1956" authorityName="Carvalho" authorityYear="1956" box="[348,702,1967,1993]" class="Insecta" family="Miridae" genus="Psallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="ponapensis">
<emphasis id="B94EE66862153047623DE7B5FE35F82E" box="[348,511,1968,1993]" italics="true" pageId="4" pageNumber="67">P. ponapensis</emphasis>
Carvalho, 1956
</taxonomicName>
and 
<taxonomicName id="4C3A41F962153047619BE7B5FBF7F82E" authority="Carvalho, 1956" authorityName="Carvalho" authorityYear="1956" box="[762,1085,1967,1993]" class="Insecta" family="Miridae" genus="Psallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="yapensis">
<emphasis id="B94EE66862153047619BE7B5FCB7F82E" box="[762,893,1968,1993]" italics="true" pageId="4" pageNumber="67">P. yapensis</emphasis>
Carvalho, 1956
</taxonomicName>
from the 
<collectingCountry id="F32D7AEA6215304767D5E7AAFAB0F82E" box="[1204,1402,1967,1993]" name="Micronesia" pageId="4" pageNumber="67">Caroline Islands</collectingCountry>
; 
<taxonomicName id="4C3A41F96215304766E7E7B5FE76F809" authority="Yasunaga" authorityName="Yasunaga" class="Insecta" family="Miridae" genus="Psallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="myiocephalus">
<emphasis id="B94EE6686215304766E7E7B5FEF7F809" italics="true" pageId="4" pageNumber="67">P. myiocephalus</emphasis>
Yasunaga
</taxonomicName>
, 
<taxonomicName id="4C3A41F96215304762ABE7D0FCECF809" authority="Yasunaga, 1999" authorityName="Yasunaga" authorityYear="1999" box="[458,806,2004,2030]" class="Insecta" family="Miridae" genus="Psallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="nakatanii">
<emphasis id="B94EE6686215304762ABE7D0FD91F809" box="[458,603,2005,2030]" italics="true" pageId="4" pageNumber="67">P. nakatanii</emphasis>
<bibRefCitation id="EFAB478B621530476102E7D1FCECF809" author="Yasunaga" box="[611,806,2004,2030]" pageId="4" pageNumber="68" refString="Yasunaga, T. (1999) First record of the plant bug subfamily Psallopinae (Heteroptera: Miridae) from Japan, with descriptions of three new species of the genus Psallops Usinger. Proceedings of the Entomological Society of Washington, 101, 737 - 741." type="journal article" year="1999">Yasunaga, 1999</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C3A41F9621530476009E7D0FB2EF809" authority="Yasunaga, 1999" authorityName="Yasunaga" authorityYear="1999" box="[872,1252,2004,2030]" class="Insecta" family="Miridae" genus="Psallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="yaeyamanus">
<emphasis id="B94EE668621530476009E7D0FBD6F809" box="[872,1052,2005,2030]" italics="true" pageId="4" pageNumber="67">P. yaeyamanus</emphasis>
<bibRefCitation id="EFAB478B621530476745E7D1FB2EF809" author="Yasunaga" box="[1060,1252,2004,2030]" pageId="4" pageNumber="68" refString="Yasunaga, T. (1999) First record of the plant bug subfamily Psallopinae (Heteroptera: Miridae) from Japan, with descriptions of three new species of the genus Psallops Usinger. Proceedings of the Entomological Society of Washington, 101, 737 - 741." type="journal article" year="1999">Yasunaga, 1999</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F32D7AEA621530476651E7D1FAB3F809" box="[1328,1401,2004,2030]" name="Japan" pageId="4" pageNumber="67">Japan</collectingCountry>
; 
<emphasis id="B94EE6686215304766E7E7D0FA50F809" box="[1414,1434,2005,2030]" italics="true" pageId="4" pageNumber="67">P.</emphasis>
</paragraph>
<paragraph id="8B853A7A6214304663F6E09DFCBEFF30" blockId="5.[151,1437,152,551]" pageId="5" pageNumber="68">
<emphasis id="B94EE6686214304663F6E09DFEECFF56" box="[151,294,152,177]" italics="true" pageId="5" pageNumber="68">formosanus</emphasis>
<bibRefCitation id="EFAB478B62143046624EE09DFE6DFF55" author="Lin" box="[303,423,152,178]" pageId="5" pageNumber="68" refString="Lin, C. - S. (2004) First record of the plant bug subfamily Psallopinae (Heteroptera: Miridae) from Taiwan and China, with descriptions of three new species of the genus Psallops Usinger. Formosan Entomology, 24, 273 - 279." type="journal article" year="2004">Lin, 2004</bibRefCitation>
and 
<emphasis id="B94EE668621430466285E09DFD8BFF56" box="[484,577,152,177]" italics="true" pageId="5" pageNumber="68">P. l e e a e</emphasis>
<bibRefCitation id="EFAB478B62143046612CE09DFD0CFF55" author="Lin" box="[589,710,152,178]" pageId="5" pageNumber="68" refString="Lin, C. - S. (2004) First record of the plant bug subfamily Psallopinae (Heteroptera: Miridae) from Taiwan and China, with descriptions of three new species of the genus Psallops Usinger. Formosan Entomology, 24, 273 - 279." type="journal article" year="2004">Lin, 2004</bibRefCitation>
from 
<collectingCountry id="F32D7AEA621430466072E09DFCBBFF55" box="[787,881,152,178]" name="Taiwan" pageId="5" pageNumber="68">Taiwan</collectingCountry>
; and 
<emphasis id="B94EE6686214304660D2E09DFB88FF56" box="[947,1090,152,177]" italics="true" pageId="5" pageNumber="68">P. c h i n e n s i s</emphasis>
<bibRefCitation id="EFAB478B62143046672FE09DFB0CFF55" author="Lin" box="[1102,1222,152,178]" pageId="5" pageNumber="68" refString="Lin, C. - S. (2004) First record of the plant bug subfamily Psallopinae (Heteroptera: Miridae) from Taiwan and China, with descriptions of three new species of the genus Psallops Usinger. Formosan Entomology, 24, 273 - 279." type="journal article" year="2004">Lin, 2004</bibRefCitation>
from 
<collectingCountry id="F32D7AEA621430466672E09DFAABFF55" box="[1299,1377,152,178]" name="China" pageId="5" pageNumber="68">China</collectingCountry>
. The second recent psallopsine genus 
<taxonomicName id="4C3A41F9621430466174E0B8FD67FF31" box="[533,685,189,214]" class="Insecta" family="Miridae" genus="Isometocoris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE668621430466174E0B8FD67FF31" box="[533,685,189,214]" italics="true" pageId="5" pageNumber="68">Isometocoris</emphasis>
</taxonomicName>
is from 
<collectingCountry id="F32D7AEA62143046606EE0B8FCBAFF30" box="[783,880,189,215]" name="Panama" pageId="5" pageNumber="68">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B853A7A6214304663A4E0E7FEDCFDC0" blockId="5.[151,1437,152,551]" pageId="5" pageNumber="68">
Thus the recent Psallopsinae live under very different climates even if all are warm. One would infer a warm palaeoclimate for the three fossils 
<taxonomicName id="4C3A41F96214304661E1E10DFD3CFEC6" box="[640,758,264,290]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="prokopi">
<emphasis id="B94EE6686214304661E1E10DFD40FEC6" box="[640,650,264,289]" italics="true" pageId="5" pageNumber="68">I</emphasis>
. 
<emphasis id="B94EE6686214304661FBE10DFD3CFEC6" box="[666,758,264,289]" italics="true" pageId="5" pageNumber="68">prokopi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3A41F9621430466064E10DFCA1FEC6" box="[773,875,264,290]" class="Insecta" family="Miridae" genus="Isometopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B94EE668621430466064E10DFCC5FEC6" box="[773,783,264,289]" italics="true" pageId="5" pageNumber="68">I</emphasis>
. 
<emphasis id="B94EE66862143046607FE10DFCA1FEC6" box="[798,875,264,289]" italics="true" pageId="5" pageNumber="68">schuhi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3A41F96214304660CEE10DFBA2FEC6" box="[943,1128,264,289]" class="Insecta" family="Miridae" genus="Cylapopsallops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B94EE6686214304660CEE10DFBA2FEC6" box="[943,1128,264,289]" italics="true" pageId="5" pageNumber="68">Cylapopsallops</emphasis>
</taxonomicName>
. But as we know nothing about the exact phylogenetic relationships between them and the recent Psallopsinae, and also because the recent representatives of the 
<taxonomicName id="4C3A41F9621430466295E157FD5CFE8B" box="[500,662,338,364]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="subFamily" subFamily="Isometopinae">Isometopinae</taxonomicName>
(putative psallopsine sister group) can be found in very diverse environments and climate, in Europe, North 
<collectingCountry id="F32D7AEA6214304661FFE17DFCCCFE75" box="[670,774,376,402]" name="United States of America" pageId="5" pageNumber="68">America</collectingCountry>
, Africa, etc., it is in fact not possible to infer anything on the biology of these fossil taxa after their taxonomic position (
<bibRefCitation id="EFAB478B6214304660E7E198FBC8FE50" author="Nel" box="[902,1026,413,439]" pageId="5" pageNumber="68" refString="Nel, A. (1997) The probabilistic inference of unknown data in phylogenetic analysis. In: Grancolas, P. (ed.), The origin of biodiversity in insects: phylogenetic tests of evolutionary scenarios. Memoires du Museum National d'Histoire Naturelle, Paris, 173, 305 - 327." type="journal article" year="1997">Nel, 1997</bibRefCitation>
). Nevertheless, recent Psallopinae seem to live on tree trunks and bark (
<bibRefCitation id="EFAB478B621430466135E1C7FCDDFE3B" author="Yasunaga" box="[596,791,450,476]" pageId="5" pageNumber="68" refString="Yasunaga, T. (1999) First record of the plant bug subfamily Psallopinae (Heteroptera: Miridae) from Japan, with descriptions of three new species of the genus Psallops Usinger. Proceedings of the Entomological Society of Washington, 101, 737 - 741." type="journal article" year="1999">Yasunaga, 1999</bibRefCitation>
). If the corticolous habits may also be inferred for the Eocene Psallopinae, it would not be a surprise to have an overrepresentation of this subfamily in amber inclusions.
</paragraph>
</subSubSection>
</treatment>
</document>