502 lines
85 KiB
XML
502 lines
85 KiB
XML
<document id="90A153B3669366E15BA1977241F3451F" ID-CLB-Dataset="55193" ID-DOI="10.11646/zootaxa.5284.2.8" ID-GBIF-Dataset="b4bc4bdf-927c-43d9-ac2b-c39d33137048" ID-ISSN="1175-5326" ID-Zenodo-Dep="7923530" ID-ZooBank="770F88D2-FA38-499B-AC81-61B8BE61992F" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683787118133" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Tri, Ngo Van" docDate="2023" docId="039387F1FFBD3A08FF2E5E07FCDBF94C" docLanguage="en" docName="zootaxa.5284.2.8.pdf" docOrigin="Zootaxa 5284 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5284.2.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tortomon distinctum Ng & Tri 2023, n. sp." docType="treatment" docVersion="6" lastPageNumber="369" masterDocId="FFAAFF89FFBC3A0DFFB95804FFC4FFBF" masterDocTitle="Two new species of semiaquatic freshwater crabs of the genera Tortomon and Aiyunamon from karst forests in northern Vietnam (Crustacea: Brachyura: Potamidae)" masterLastPageNumber="376" masterPageNumber="364" pageNumber="365" updateTime="1699515580356" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="04085855B09F13991878BAFD77BA44A6">Two new species of semiaquatic freshwater crabs of the genera Tortomon and Aiyunamon from karst forests in northern Vietnam (Crustacea: Brachyura: Potamidae)</mods:title>
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<mods:name id="6FF1A752D6B88CA6CE854809FFD16914" type="personal">
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<mods:roleTerm id="8225AE85D3A6102A17D6CFEF0CAF25ED">Author</mods:roleTerm>
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<mods:namePart id="7C6CA694E90C56950E82B01C293ED236">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="A580278CFC99B0F9F7DDC3700225C120">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore.</mods:affiliation>
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<mods:roleTerm id="983FE232442C310955B44B715F0679B1">Author</mods:roleTerm>
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<mods:namePart id="FCBEAB6CE07042706CA957DECDE4B057">Tri, Ngo Van</mods:namePart>
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<mods:affiliation id="5E47A61629B46D40ED55891C538AD8C4">National Key Laboratory, Institute of Tropical Biology, Vietnamese Academy of Sciences and Technology, 9 / 621 Hanoi Highway, Thu Duc City, Ho Chi Minh City, Vietnam.</mods:affiliation>
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<mods:typeOfResource id="56FF81A9256402D50BCC37CABF0D80CE">text</mods:typeOfResource>
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<mods:title id="033C9976E37DB698C2F7C3F2A5CD85F0">Zootaxa</mods:title>
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<mods:part id="9BD2D8B3EA995BFE1B7B6C07BDB0836E">
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<mods:date id="4FC3CC7A262BE0D45FF4F5C1C917D636">2023</mods:date>
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<mods:number id="4CA2BC11E4B90A95F5376FF7EBB04420">2023-05-11</mods:number>
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<treatment id="039387F1FFBD3A08FF2E5E07FCDBF94C" ID-DOI="http://doi.org/10.5281/zenodo.7923524" ID-GBIF-Taxon="209616448" ID-Zenodo-Dep="7923524" LSID="urn:lsid:plazi:treatment:039387F1FFBD3A08FF2E5E07FCDBF94C" httpUri="http://treatment.plazi.org/id/039387F1FFBD3A08FF2E5E07FCDBF94C" lastPageId="5" lastPageNumber="369" pageId="1" pageNumber="365">
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<heading id="D0CD818BFFBD3A0CFF2E5E07FE19F9A2" bold="true" box="[151,477,1539,1566]" fontSize="11" level="1" pageId="1" pageNumber="365" reason="1">
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<emphasis id="B94EEAF5FFBD3A0CFF2E5E07FE19F9A2" bold="true" box="[151,477,1539,1566]" pageId="1" pageNumber="365">
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<taxonomicName id="4C3A4D64FFBD3A0CFF2E5E07FE57F9A1" ID-CoL="fafe31b9-511b-4880-8aa6-2bbeff1804b6" authority="Ng & Tri, 2023" authorityName="Ng & Tri" authorityYear="2023" box="[151,403,1539,1566]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="distinctum" status="sp. nov.">
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<emphasis id="B94EEAF5FFBD3A0CFF2E5E07FE57F9A1" bold="true" box="[151,403,1539,1566]" italics="true" pageId="1" pageNumber="365">Tortomon distinctum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27D578EFFBD3A0CFE225E07FE19F9A2" box="[411,477,1539,1565]" pageId="1" pageNumber="365" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C320656CFFBD3A0CFF2E5E23FED6F9FD" box="[151,274,1575,1602]" pageId="1" pageNumber="365" type="description">
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<paragraph id="8B8536E7FFBD3A0CFF2E5E23FED6F9FD" blockId="1.[151,477,1539,1602]" box="[151,274,1575,1602]" pageId="1" pageNumber="365">
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(
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<figureCitation id="13012A62FFBD3A0CFF265E23FECDF9FD" box="[159,265,1575,1602]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1181,1205]" captionStartId-1="3.[151,250,1973,1997]" captionStartId-2="4.[151,250,1900,1924]" captionStartId-3="5.[151,250,1070,1094]" captionStartId-4="6.[151,250,1960,1984]" captionTargetBox-0="[168,1420,345,1158]" captionTargetBox-1="[183,1405,597,1950]" captionTargetBox-2="[167,1421,181,1876]" captionTargetBox-3="[153,1434,183,1044]" captionTargetBox-4="[166,1421,921,1937]" captionTargetId-0="figure-104@2.[168,1420,345,1158]" captionTargetId-1="figure-277@3.[183,1405,597,1950]" captionTargetId-2="figure-19@4.[167,1421,181,1876]" captionTargetId-3="figure-32@5.[151,1436,181,1046]" captionTargetId-4="figure-406@6.[166,1421,921,1937]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Tortomon distinctum n. sp., colour in life. Paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam. A, overall dorsal habitus; B, frontal view; C, major chela; D, ventral surface and gonopods." captionText-1="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." captionText-2="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." captionText-3="FIGURE 4. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsomesial view); D, distal part of left G1 (dorsal view); E, left G2 (ventral view); F, distal part of left G2 (ventral view); G, distal part of left G2 (dorsal view). Scales: A, E = 0.5 mm; B–D, F, G = 0.2 mm." captionText-4="FIGURE 5. Tortomon distinctum n. sp., paratype female (18.0 × 12.3 mm) (ZRC 2022.798), Vietnam.A, overall dorsal habitus; B, buccal cavity, anterior thoracic sternum and pleon; C, sternopleonal cavity showing vulvae." figureDoi-0="http://doi.org/10.5281/zenodo.7923532" figureDoi-1="http://doi.org/10.5281/zenodo.7923534" figureDoi-2="http://doi.org/10.5281/zenodo.7923536" figureDoi-3="http://doi.org/10.5281/zenodo.7923538" figureDoi-4="http://doi.org/10.5281/zenodo.7923540" httpUri-0="https://zenodo.org/record/7923532/files/figure.png" httpUri-1="https://zenodo.org/record/7923534/files/figure.png" httpUri-2="https://zenodo.org/record/7923536/files/figure.png" httpUri-3="https://zenodo.org/record/7923538/files/figure.png" httpUri-4="https://zenodo.org/record/7923540/files/figure.png" pageId="1" pageNumber="365">Figs. 1–5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C320656CFFBD3A0CFF2E5E6BFEDBF8A6" pageId="1" pageNumber="365" type="materials_examined">
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<emphasis id="B94EEAF5FFBD3A0CFF2E5E6BFE87F936" bold="true" box="[151,323,1647,1673]" pageId="1" pageNumber="365">Type material</emphasis>
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.
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: male (19.2 × 13.0 mm) (
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<specimenCode id="DB9C9E9CFFBD3A0CFC905E6BFC18F936" box="[809,988,1647,1673]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="365">ZRC 2022.797</specimenCode>
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),
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<location id="8EE5603CFFBD3A0CFC4D5E6BFB95F935" LSID="urn:lsid:plazi:treatment:039387F1FFBD3A08FF2E5E07FCDBF94C:8EE5603CFFBD3A0CFC4D5E6BFB95F935" box="[1012,1105,1647,1674]" country="Vietnam" county="Tam Dy" latitude="22.2" longLatPrecision="1260" longitude="103.6" municipality="Khun Ha" name="Phu Tra" pageId="1" pageNumber="365" stateProvince="Lai Chau">Phu Tra</location>
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limestome forest,
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commune,
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ờng district,
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,
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<collectingCountry id="F32D7677FFBD3A0CFD505E97FC8FF912" box="[745,843,1683,1709]" name="Vietnam" pageId="1" pageNumber="365">Vietnam</collectingCountry>
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,
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,
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,
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<quantity id="4CC29B02FFBD3A0CFBFC5E97FB58F911" box="[1093,1180,1683,1710]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="365" unit="m" value="1800.0">1800 m</quantity>
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asl
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</elevation>
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, coll.
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<collectionCode id="ED2BAE22FFBD3A0CFAB75E90FAE6F911" box="[1294,1314,1684,1710]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="365" type="Museum">V</collectionCode>
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.
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<collectionCode id="ED2BAE22FFBD3A0CFA975E90FA84F911" box="[1326,1344,1684,1710]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="365">T</collectionCode>
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. Ngo,
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||
<date id="FF841027FFBD3A0CFA375E97FECDF96E" pageId="1" pageNumber="365" value="2022-07-05">
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<collectingDate id="EFC0E9CFFFBD3A0CFA375E97FECDF96E" pageId="1" pageNumber="365" value="2022-07-05">5 July 2022</collectingDate>
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</date>
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</materialsCitation>
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.
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<typeStatus id="54818845FFBD3A0CFEAB5EB3FE75F96E" box="[274,433,1719,1745]" pageId="1" pageNumber="365" type="paratype">PARATYPES</typeStatus>
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:
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||
<specimenCount id="9D3CFD6EFFBD3A0CFE025EB3FDD5F96E" box="[443,529,1719,1745]" pageId="1" pageNumber="365" type="male">6 males</specimenCount>
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(14.8 ×
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<quantity id="4CC29B02FFBD3A0CFDD65EB3FD11F96D" box="[623,725,1719,1746]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="1" pageNumber="365" unit="mm" value="10.2">10.2 mm</quantity>
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||
, 17.6 ×
|
||
<quantity id="4CC29B02FFBD3A0CFC975EB3FC50F96D" box="[814,916,1719,1746]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="1" pageNumber="365" unit="mm" value="12.2">12.2 mm</quantity>
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||
, 18.2 ×
|
||
<quantity id="4CC29B02FFBD3A0CFC545EB3FB97F96D" box="[1005,1107,1719,1746]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="365" unit="mm" value="12.5">12.5 mm</quantity>
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||
, 18.3 ×
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||
<quantity id="4CC29B02FFBD3A0CFB155EB3FAD6F96D" box="[1196,1298,1719,1746]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="1" pageNumber="365" unit="mm" value="12.4">12.4 mm</quantity>
|
||
, 19.5 ×
|
||
<quantity id="4CC29B02FFBD3A0CFAD25EB3FF01F949" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="1" pageNumber="365" unit="mm" value="12.8">12.8 mm</quantity>
|
||
, 20.4 ×
|
||
<quantity id="4CC29B02FFBD3A0CFE985EDFFE4EF94A" box="[289,394,1755,1781]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="1" pageNumber="365" unit="mm" value="13.6">13.6 mm</quantity>
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),
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||
<materialsCitation id="3B523CBAFFBD3A0CFE275EDFFED8F8A6" ID-GBIF-Occurrence="4104291304" collectingDate="2022-07-05" collectionCode="ZRC" country="Vietnam" county="Tam Dy" elevation="1800" latitude="22.2" location="Phu Tra" longLatPrecision="1260" longitude="103.6" municipality="Khun Ha" pageId="1" pageNumber="365" specimenCode="ZRC 2022.798" specimenCount="3" specimenCount-female="3" stateProvince="Lai Chau" typeStatus="paratype">
|
||
<specimenCount id="9D3CFD6EFFBD3A0CFE275EDFFDCEF94A" box="[414,522,1755,1781]" pageId="1" pageNumber="365" type="female">3 females</specimenCount>
|
||
(16.1 ×
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<quantity id="4CC29B02FFBD3A0CFDD25EDFFD15F949" box="[619,721,1755,1782]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="1" pageNumber="365" unit="mm" value="11.2">11.2 mm</quantity>
|
||
, 16.9 ×
|
||
<quantity id="4CC29B02FFBD3A0CFC945EDFFC57F949" box="[813,915,1755,1782]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="1" pageNumber="365" unit="mm" value="11.6">11.6 mm</quantity>
|
||
, 18.0 ×
|
||
<quantity id="4CC29B02FFBD3A0CFC565EDFFB90F94A" box="[1007,1108,1755,1781]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="1" pageNumber="365" unit="mm" value="12.3">12.3 mm</quantity>
|
||
) (
|
||
<specimenCode id="DB9C9E9CFFBD3A0CFBD25EDFFADDF94A" box="[1131,1305,1755,1781]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="365">ZRC 2022.798</specimenCode>
|
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), same data as holotype
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||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C320656CFFBD3A0FFF7E5F27FCD3F912" lastPageId="2" lastPageNumber="366" pageId="1" pageNumber="365" type="diagnosis">
|
||
<paragraph id="8B8536E7FFBD3A0FFF7E5F27FBA0FEFE" blockId="1.[151,1437,1647,2034]" lastBlockId="2.[151,1436,151,321]" lastPageId="2" lastPageNumber="366" pageId="1" pageNumber="365">
|
||
<emphasis id="B94EEAF5FFBD3A0CFF7E5F27FEFEF882" bold="true" box="[199,314,1827,1853]" pageId="1" pageNumber="365">Diagnosis</emphasis>
|
||
. Carapace transversely ovate, proportionately very wide carapace, width 1.44–1.52 times length (
|
||
<figureCitation id="13012A62FFBD3A0CFF265F43FEDCF8DE" box="[159,280,1863,1889]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="1" pageNumber="365">Fig. 2A, B</figureCitation>
|
||
); epigastric cristae very low, evident only as flattened rugae, postfrontal cristae not visible (
|
||
<figureCitation id="13012A62FFBD3A0CFAAC5F43FA49F8DE" box="[1301,1421,1863,1889]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="1" pageNumber="365">Fig. 2A, B</figureCitation>
|
||
); anterolateral margin with very low crista, surface appears rounded in frontal view (
|
||
<figureCitation id="13012A62FFBD3A0CFBFE5F6FFB6EF83A" box="[1095,1194,1899,1925]" captionStart="FIGURE 1" captionStartId="2.[151,250,1181,1205]" captionTargetBox="[168,1420,345,1158]" captionTargetId="figure-104@2.[168,1420,345,1158]" captionTargetPageId="2" captionText="FIGURE 1. Tortomon distinctum n. sp., colour in life. Paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam. A, overall dorsal habitus; B, frontal view; C, major chela; D, ventral surface and gonopods." figureDoi="http://doi.org/10.5281/zenodo.7923532" httpUri="https://zenodo.org/record/7923532/files/figure.png" pageId="1" pageNumber="365">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBD3A0CFB015F6FFB18F83A" box="[1208,1244,1899,1925]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="1" pageNumber="365">2C</figureCitation>
|
||
); median lobe of posterior margin of epistome relatively acutely triangular (
|
||
<figureCitation id="13012A62FFBD3A0CFCA75F8BFC53F816" box="[798,919,1935,1961]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="1" pageNumber="365">Fig. 2C, D</figureCitation>
|
||
). Third maxilliped distinctly elongate, ischium almost twice as long as wide (
|
||
<figureCitation id="13012A62FFBD3A0CFE565FB7FD83F872" box="[495,583,1971,1997]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="1" pageNumber="365">Fig. 2E</figureCitation>
|
||
); inner angle of carpus of cheliped with low tubercle (
|
||
<figureCitation id="13012A62FFBD3A0CFB085FB7FAD2F872" box="[1201,1302,1971,1997]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="1" pageNumber="365">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBD3A0CFA9A5FB7FA8DF871" box="[1315,1353,1971,1998]" captionStart="FIGURE 5" captionStartId="6.[151,250,1960,1984]" captionTargetBox="[166,1421,921,1937]" captionTargetId="figure-406@6.[166,1421,921,1937]" captionTargetPageId="6" captionText="FIGURE 5. Tortomon distinctum n. sp., paratype female (18.0 × 12.3 mm) (ZRC 2022.798), Vietnam.A, overall dorsal habitus; B, buccal cavity, anterior thoracic sternum and pleon; C, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.7923540" httpUri="https://zenodo.org/record/7923540/files/figure.png" pageId="1" pageNumber="365">5A</figureCitation>
|
||
). Adult major male chela forming distinct basal gape between closed fingers (
|
||
<figureCitation id="13012A62FFBD3A0CFC0E5FD3FBDEF84E" box="[951,1050,2007,2033]" captionStart="FIGURE 1" captionStartId="2.[151,250,1181,1205]" captionTargetBox="[168,1420,345,1158]" captionTargetId="figure-104@2.[168,1420,345,1158]" captionTargetPageId="2" captionText="FIGURE 1. Tortomon distinctum n. sp., colour in life. Paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam. A, overall dorsal habitus; B, frontal view; C, major chela; D, ventral surface and gonopods." figureDoi="http://doi.org/10.5281/zenodo.7923532" httpUri="https://zenodo.org/record/7923532/files/figure.png" pageId="1" pageNumber="365">Figs. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBD3A0CFB905FD3FB8EF84E" box="[1065,1098,2007,2033]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="1" pageNumber="365">2F</figureCitation>
|
||
). P2–5 meri slender, elongate (
|
||
<figureCitation id="13012A62FFBE3A0FFF265893FEC2FF0E" box="[159,262,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="2" pageNumber="366">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBE3A0FFEAF5893FEFEFF0E" box="[278,314,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="2" pageNumber="366">3C</figureCitation>
|
||
). Sternal tubercle of male pleonal locking mechanism relatively small, submedian in position on sternite 5 (
|
||
<figureCitation id="13012A62FFBE3A0FFEB458BFFEA0FF6A" box="[269,356,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="2" pageNumber="366">Fig. 3B</figureCitation>
|
||
). Male pleon distinctly wide, triangular, proximal margin almost twice length of somite 6; lateral margin of telson gently concave (
|
||
<figureCitation id="13012A62FFBE3A0FFDA658DBFD67FF46" box="[543,675,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="2" pageNumber="366">Fig. 3A, D</figureCitation>
|
||
). G1 distinctly bent, terminal segment relatively slender, tapering distally (
|
||
<figureCitation id="13012A62FFBE3A0FFF435907FE45FEA2" box="[250,385,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="2" pageNumber="366">Figs. 3G, H</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBE3A0FFE345907FE12FEA2" box="[397,470,259,285]" captionStart="FIGURE 4" captionStartId="5.[151,250,1070,1094]" captionTargetBox="[153,1434,183,1044]" captionTargetId="figure-32@5.[151,1436,181,1046]" captionTargetPageId="5" captionText="FIGURE 4. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsomesial view); D, distal part of left G1 (dorsal view); E, left G2 (ventral view); F, distal part of left G2 (ventral view); G, distal part of left G2 (dorsal view). Scales: A, E = 0.5 mm; B–D, F, G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7923538" httpUri="https://zenodo.org/record/7923538/files/figure.png" pageId="2" pageNumber="366">4A–D</figureCitation>
|
||
). G2 with long flagellum, ca. two-thirds length of basal segment (
|
||
<figureCitation id="13012A62FFBE3A0FFB045907FAD2FEA2" box="[1213,1302,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="2" pageNumber="366">Figs. 3I</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBE3A0FFA9A5907FA82FEA2" box="[1315,1350,259,285]" captionStart="FIGURE 4" captionStartId="5.[151,250,1070,1094]" captionTargetBox="[153,1434,183,1044]" captionTargetId="figure-32@5.[151,1436,181,1046]" captionTargetPageId="5" captionText="FIGURE 4. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsomesial view); D, distal part of left G1 (dorsal view); E, left G2 (ventral view); F, distal part of left G2 (ventral view); G, distal part of left G2 (dorsal view). Scales: A, E = 0.5 mm; B–D, F, G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7923538" httpUri="https://zenodo.org/record/7923538/files/figure.png" pageId="2" pageNumber="366">4E</figureCitation>
|
||
). Vulva ovate, large, on proximal half of sternite 6, pressing on suture with sternite 5 (
|
||
<figureCitation id="13012A62FFBE3A0FFC465923FB9CFEFE" box="[1023,1112,294,321]" captionStart="FIGURE 5" captionStartId="6.[151,250,1960,1984]" captionTargetBox="[166,1421,921,1937]" captionTargetId="figure-406@6.[166,1421,921,1937]" captionTargetPageId="6" captionText="FIGURE 5. Tortomon distinctum n. sp., paratype female (18.0 × 12.3 mm) (ZRC 2022.798), Vietnam.A, overall dorsal habitus; B, buccal cavity, anterior thoracic sternum and pleon; C, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.7923540" httpUri="https://zenodo.org/record/7923540/files/figure.png" pageId="2" pageNumber="366">Fig. 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF45666FFFBE3A0FFF2E5C99FC0EFB66" ID-DOI="http://doi.org/10.5281/zenodo.7923532" ID-Zenodo-Dep="7923532" httpUri="https://zenodo.org/record/7923532/files/figure.png" pageId="2" pageNumber="366" startId="2.[151,250,1181,1205]" targetBox="[168,1420,345,1158]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8B8536E7FFBE3A0FFF2E5C99FC0EFB66" blockId="2.[151,1436,1181,1241]" pageId="2" pageNumber="366">
|
||
<emphasis id="B94EEAF5FFBE3A0FFF2E5C99FED0FB0A" bold="true" box="[151,276,1181,1205]" pageId="2" pageNumber="366">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C3A4D64FFBE3A0FFEA25C9AFE29FB0A" authority="Ng & Tri, 2023" authorityName="Ng & Tri" authorityYear="2023" box="[283,493,1181,1205]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="distinctum" status="sp. nov.">
|
||
<emphasis id="B94EEAF5FFBE3A0FFEA25C9AFE29FB0A" box="[283,493,1181,1205]" italics="true" pageId="2" pageNumber="366">Tortomon distinctum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94EEAF5FFBE3A0FFE4D5C99FDEAFB0A" bold="true" box="[500,558,1181,1205]" pageId="2" pageNumber="366">
|
||
<taxonomicNameLabel id="A27D578EFFBE3A0FFE4D5C99FDEAFB0A" box="[500,558,1181,1205]" pageId="2" pageNumber="366" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, colour in life. Paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam. A, overall dorsal habitus; B, frontal view; C, major chela; D, ventral surface and gonopods.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8536E7FFBE3A0FFF7E5D03FE54F9D9" blockId="2.[151,1437,1287,2034]" pageId="2" pageNumber="366">
|
||
<emphasis id="B94EEAF5FFBE3A0FFF7E5D03FD8FFA9E" bold="true" box="[199,587,1287,1313]" pageId="2" pageNumber="366">Female characters and variation</emphasis>
|
||
. The prominence of the large teeth on the cutting edges of the fingers of the major male chela is variable: in the
|
||
<typeStatus id="54818845FFBE3A0FFD895D2FFD56FAFA" box="[560,658,1323,1349]" pageId="2" pageNumber="366" type="holotype">holotype</typeStatus>
|
||
male, the dactylus has two or three larger teeth, the largest being on the proximal one-third, while there is one large tooth on the basal part of the pollex (
|
||
<figureCitation id="13012A62FFBE3A0FFBE15D4BFB6BFAD6" box="[1112,1199,1359,1385]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="2" pageNumber="366">Fig. 2F</figureCitation>
|
||
); in a
|
||
<typeStatus id="54818845FFBE3A0FFB4F5D54FA92FAD5" box="[1270,1366,1360,1386]" pageId="2" pageNumber="366" type="paratype">paratype</typeStatus>
|
||
male, there are two very large teeth on each of the proximal halves of the dactylus and pollex (
|
||
<figureCitation id="13012A62FFBE3A0FFBC55D77FB10FA32" box="[1148,1236,1395,1421]" captionStart="FIGURE 1" captionStartId="2.[151,250,1181,1205]" captionTargetBox="[168,1420,345,1158]" captionTargetId="figure-104@2.[168,1420,345,1158]" captionTargetPageId="2" captionText="FIGURE 1. Tortomon distinctum n. sp., colour in life. Paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam. A, overall dorsal habitus; B, frontal view; C, major chela; D, ventral surface and gonopods." figureDoi="http://doi.org/10.5281/zenodo.7923532" httpUri="https://zenodo.org/record/7923532/files/figure.png" pageId="2" pageNumber="366">Fig. 1C</figureCitation>
|
||
). Both specimens are of similar sizes and there are no other major differences. Females of
|
||
<taxonomicName id="4C3A4D64FFBE3A0FFC785D9CFB96FA0E" authority="Ng & Tri, 2023" authorityName="Ng & Tri" authorityYear="2023" box="[961,1106,1431,1457]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="distinctum" status="sp. nov.">
|
||
<emphasis id="B94EEAF5FFBE3A0FFC785D9CFB96FA0E" box="[961,1106,1431,1457]" italics="true" pageId="2" pageNumber="366">T. distinctum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94EEAF5FFBE3A0FFBE05D93FB53FA0E" bold="true" box="[1113,1175,1431,1457]" pageId="2" pageNumber="366">
|
||
<taxonomicNameLabel id="A27D578EFFBE3A0FFBE05D93FB53FA0E" box="[1113,1175,1431,1457]" pageId="2" pageNumber="366" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
have distinctly smaller and symmetrical chelae (
|
||
<figureCitation id="13012A62FFBE3A0FFE175DBFFDC3FA69" box="[430,519,1467,1494]" captionStart="FIGURE 5" captionStartId="6.[151,250,1960,1984]" captionTargetBox="[166,1421,921,1937]" captionTargetId="figure-406@6.[166,1421,921,1937]" captionTargetPageId="6" captionText="FIGURE 5. Tortomon distinctum n. sp., paratype female (18.0 × 12.3 mm) (ZRC 2022.798), Vietnam.A, overall dorsal habitus; B, buccal cavity, anterior thoracic sternum and pleon; C, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.7923540" httpUri="https://zenodo.org/record/7923540/files/figure.png" pageId="2" pageNumber="366">Fig. 5A</figureCitation>
|
||
). The length/width proportions of the male pleon varies and are not taxonomically reliable. In the
|
||
<typeStatus id="54818845FFBE3A0FFEFA5DDBFE61FA46" box="[323,421,1503,1529]" pageId="2" pageNumber="366" type="holotype">holotype</typeStatus>
|
||
male, somite 6 is relatively wider, with the proximal margin almost twice the length (
|
||
<figureCitation id="13012A62FFBE3A0FFAD65DDBFF78F9A2" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="2" pageNumber="366">Fig. 3D</figureCitation>
|
||
), but in two similar sized-paratype males, somite 6 is distinctly longer (
|
||
<figureCitation id="13012A62FFBE3A0FFC525E07FBA6F9A2" box="[1003,1122,1539,1565]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="2" pageNumber="366">Fig. 3E, F</figureCitation>
|
||
). Similar variations in male pleonal proportions have been reported in the Myanmarese potamid
|
||
<taxonomicName id="4C3A4D64FFBE3A0FFC2B5E23FA84F9FE" authority="Ng, Win Mar & Shih, 2020" authorityName="Ng, Win Mar & Shih" authorityYear="2020" box="[914,1344,1575,1602]" class="Malacostraca" family="Potamidae" genus="Indawmon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="366" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94EEAF5FFBE3A0FFC2B5E23FBCCF9FE" box="[914,1032,1575,1601]" italics="true" pageId="2" pageNumber="366">Indawmon</emphasis>
|
||
Ng, Win Mar & Shih, 2020
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFAB4B16FFBE3A0FFAC35E2CFE8EF9D9" author="Ng, P. K. L. & Mar, W. & Shih, H. - T." pageId="2" pageNumber="366" pagination="529 - 538" refId="ref5796" refString="Ng, P. K. L., Mar, W. & Shih, H. - T. (2020) Indawmon thanayei, a new genus and new species of freshwater crab (Crustacea: Brachyura: Potamidae) from Indawgyi Lake, Kachin State, Myanmar. Raffles Bulletin of Zoology, 68, 529 - 538." type="journal article" year="2020">
|
||
Ng
|
||
<emphasis id="B94EEAF5FFBE3A0FFF2E5E48FF0BF9DA" box="[151,207,1611,1637]" italics="true" pageId="2" pageNumber="366">et al.</emphasis>
|
||
2020: 535
|
||
</bibRefCitation>
|
||
, 537).
|
||
</paragraph>
|
||
<paragraph id="8B8536E7FFBE3A0FFF7E5E6BFCD3F912" blockId="2.[151,1437,1287,2034]" pageId="2" pageNumber="366">
|
||
<emphasis id="B94EEAF5FFBE3A0FFF7E5E6BFEDFF936" bold="true" box="[199,283,1647,1673]" pageId="2" pageNumber="366">Colour</emphasis>
|
||
. Carapace, chelipeds and ambulatory legs are dark purple; with the ventral surfaces somewhat paler and the distal half of the fingers of the chelae orange (
|
||
<figureCitation id="13012A62FFBE3A0FFD7D5E97FCC2F912" box="[708,774,1683,1709]" captionStart="FIGURE 1" captionStartId="2.[151,250,1181,1205]" captionTargetBox="[168,1420,345,1158]" captionTargetId="figure-104@2.[168,1420,345,1158]" captionTargetPageId="2" captionText="FIGURE 1. Tortomon distinctum n. sp., colour in life. Paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam. A, overall dorsal habitus; B, frontal view; C, major chela; D, ventral surface and gonopods." figureDoi="http://doi.org/10.5281/zenodo.7923532" httpUri="https://zenodo.org/record/7923532/files/figure.png" pageId="2" pageNumber="366">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C320656CFFBE3A0FFF7E5EB3FED4F949" pageId="2" pageNumber="366" type="etymology">
|
||
<paragraph id="8B8536E7FFBE3A0FFF7E5EB3FED4F949" blockId="2.[151,1437,1287,2034]" pageId="2" pageNumber="366">
|
||
<emphasis id="B94EEAF5FFBE3A0FFF7E5EB3FE8CF96E" bold="true" box="[199,328,1719,1745]" pageId="2" pageNumber="366">Etymology</emphasis>
|
||
. The species is named for its many distinctive features, easily distinguishing it from its Chinese congeners.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C320656CFFBE3A08FF7E5EFBFCC1F980" lastPageId="5" lastPageNumber="369" pageId="2" pageNumber="366" type="discussion">
|
||
<paragraph id="8B8536E7FFBE3A0EFF7E5EFBFAC4FD82" blockId="2.[151,1437,1287,2034]" lastBlockId="3.[151,1437,151,573]" lastPageId="3" lastPageNumber="367" pageId="2" pageNumber="366">
|
||
<emphasis id="B94EEAF5FFBE3A0FFF7E5EFBFEF0F8A6" bold="true" box="[199,308,1791,1817]" pageId="2" pageNumber="366">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C3A4D64FFBE3A0FFEF15F04FDF2F8A6" authority="Ng & Tri, 2023" authorityName="Ng & Tri" authorityYear="2023" box="[328,566,1791,1817]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="distinctum" status="sp. nov.">
|
||
<emphasis id="B94EEAF5FFBE3A0FFEF15F04FDF2F8A6" box="[328,566,1791,1817]" italics="true" pageId="2" pageNumber="366">Tortomon distinctum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94EEAF5FFBE3A0FFDFB5EFBFD43F8A6" bold="true" box="[578,647,1791,1817]" pageId="2" pageNumber="366">
|
||
<taxonomicNameLabel id="A27D578EFFBE3A0FFDFB5EFBFD43F8A6" box="[578,647,1791,1817]" pageId="2" pageNumber="366" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can easily be distinguished from
|
||
<taxonomicName id="4C3A4D64FFBE3A0FFB9A5F04FBBCF8A6" authorityName="Huang, Wang & Shih" authorityYear="2020" box="[1059,1144,1792,1817]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="puer">
|
||
<emphasis id="B94EEAF5FFBE3A0FFB9A5F04FBBCF8A6" box="[1059,1144,1792,1817]" italics="true" pageId="2" pageNumber="366">T. puer</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A4D64FFBE3A0FFB025F04FAEEF8A6" authorityName="C Huang, J Wang & HT Shih" authorityYear="2020" box="[1211,1322,1792,1817]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="gejiuin">
|
||
<emphasis id="B94EEAF5FFBE3A0FFB025F04FAEEF8A6" box="[1211,1322,1792,1817]" italics="true" pageId="2" pageNumber="366">T. gejiuin</emphasis>
|
||
</taxonomicName>
|
||
having a proportionately wider carapace with the width 1.44–1.52 times length (
|
||
<figureCitation id="13012A62FFBE3A0FFC765F27FB89F882" box="[975,1101,1827,1853]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="2" pageNumber="366">Fig. 2A, B</figureCitation>
|
||
) (versus proportionately less wide, width 1.28–1.45 times length;
|
||
<bibRefCitation id="EFAB4B16FFBE3A0FFD8E5F4CFCCCF8DE" author="Huang, C. & Wang, J. & Shih, H. - T." box="[567,776,1863,1890]" pageId="2" pageNumber="366" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBE3A0FFD335F4CFD00F8DE" box="[650,708,1863,1889]" italics="true" pageId="2" pageNumber="366">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 1, 4); the median lobe of the posterior margin of the epistome is more acutely triangular (
|
||
<figureCitation id="13012A62FFBE3A0FFD925F6FFD64F83A" box="[555,672,1899,1925]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="2" pageNumber="366">Fig. 2C, D</figureCitation>
|
||
) (vs. median lobe proportionately wider;
|
||
<bibRefCitation id="EFAB4B16FFBE3A0FFBD35F68FAF7F83A" author="Huang, C. & Wang, J. & Shih, H. - T." box="[1130,1331,1899,1926]" pageId="2" pageNumber="366" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBE3A0FFB025F68FB36F83A" box="[1211,1266,1899,1925]" italics="true" pageId="2" pageNumber="366">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 2A, 5A); the third maxilliped is distinctly elongate, with the ischium almost twice as long as wide (
|
||
<figureCitation id="13012A62FFBE3A0FFB785F8BFAD2F816" box="[1217,1302,1935,1961]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="2" pageNumber="366">Fig. 2E</figureCitation>
|
||
) (vs. length of ischium of third maxilliped ca. 1.5 times width;
|
||
<bibRefCitation id="EFAB4B16FFBE3A0FFD745FB0FC5CF872" author="Huang, C. & Wang, J. & Shih, H. - T." box="[717,920,1971,1998]" pageId="2" pageNumber="366" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBE3A0FFCA75FB0FC92F872" box="[798,854,1971,1997]" italics="true" pageId="2" pageNumber="366">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 3A, 6A); the inner angle of the carpus of the cheliped only has a low tubercle (
|
||
<figureCitation id="13012A62FFBE3A0FFD825FD3FD5BF84E" box="[571,671,2007,2033]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="2" pageNumber="366">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBE3A0FFD125FD3FD0AF84D" box="[683,718,2007,2034]" captionStart="FIGURE 5" captionStartId="6.[151,250,1960,1984]" captionTargetBox="[166,1421,921,1937]" captionTargetId="figure-406@6.[166,1421,921,1937]" captionTargetPageId="6" captionText="FIGURE 5. Tortomon distinctum n. sp., paratype female (18.0 × 12.3 mm) (ZRC 2022.798), Vietnam.A, overall dorsal habitus; B, buccal cavity, anterior thoracic sternum and pleon; C, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.7923540" httpUri="https://zenodo.org/record/7923540/files/figure.png" pageId="2" pageNumber="366">5A</figureCitation>
|
||
) (vs. with a sharp tooth;
|
||
<bibRefCitation id="EFAB4B16FFBE3A0FFC5E5FDCFB71F84E" author="Huang, C. & Wang, J. & Shih, H. - T." box="[999,1205,2007,2034]" pageId="2" pageNumber="366" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBE3A0FFB805FDCFBB6F84E" box="[1081,1138,2007,2033]" italics="true" pageId="2" pageNumber="366">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 1, 4); the adult major male chela forms a distinct basal gape when the fingers are closed (
|
||
<figureCitation id="13012A62FFBF3A0EFC635893FBF8FF0E" box="[986,1084,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1181,1205]" captionTargetBox="[168,1420,345,1158]" captionTargetId="figure-104@2.[168,1420,345,1158]" captionTargetPageId="2" captionText="FIGURE 1. Tortomon distinctum n. sp., colour in life. Paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam. A, overall dorsal habitus; B, frontal view; C, major chela; D, ventral surface and gonopods." figureDoi="http://doi.org/10.5281/zenodo.7923532" httpUri="https://zenodo.org/record/7923532/files/figure.png" pageId="3" pageNumber="367">Figs. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBF3A0EFBF05893FBACFF0E" box="[1097,1128,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="3" pageNumber="367">2F</figureCitation>
|
||
) (vs. without distinct gape;
|
||
<bibRefCitation id="EFAB4B16FFBF3A0EFF2E58BFFEA1FF6A" author="Huang, C. & Wang, J. & Shih, H. - T." box="[151,357,187,213]" pageId="3" pageNumber="367" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBF3A0EFF5058BFFEE6FF6A" box="[233,290,187,213]" italics="true" pageId="3" pageNumber="367">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 3F, 6F); the P2–5 meri are proportionately more slender and longer (
|
||
<figureCitation id="13012A62FFBF3A0EFB1458BFFAD5FF6A" box="[1197,1297,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1997]" captionTargetBox="[183,1405,597,1950]" captionTargetId="figure-277@3.[183,1405,597,1950]" captionTargetPageId="3" captionText="FIGURE 2. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae." figureDoi="http://doi.org/10.5281/zenodo.7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="3" pageNumber="367">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBF3A0EFAA458BFFAFAFF6A" box="[1309,1342,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="3" pageNumber="367">3C</figureCitation>
|
||
) than in
|
||
<taxonomicName id="4C3A4D64FFBF3A0EFF2E58DBFF22FF47" authorityName="Huang, Wang & Shih" authorityYear="2020" box="[151,230,223,248]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="367" phylum="Arthropoda" rank="species" species="puer">
|
||
<emphasis id="B94EEAF5FFBF3A0EFF2E58DBFF22FF47" box="[151,230,223,248]" italics="true" pageId="3" pageNumber="367">T. puer</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A4D64FFBF3A0EFEA458DBFE43FF47" authorityName="C Huang, J Wang & HT Shih" authorityYear="2020" box="[285,391,223,248]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="367" phylum="Arthropoda" rank="species" species="gejiuin">
|
||
<emphasis id="B94EEAF5FFBF3A0EFEA458DBFE43FF47" box="[285,391,223,248]" italics="true" pageId="3" pageNumber="367">T. gejiuin</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFAB4B16FFBF3A0EFE2F58DBFDA1FF46" author="Huang, C. & Wang, J. & Shih, H. - T." box="[406,613,223,249]" pageId="3" pageNumber="367" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBF3A0EFE5058DBFDE6FF46" box="[489,546,223,249]" italics="true" pageId="3" pageNumber="367">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 1, 4); the sternal tubercle of the male pleonal locking mechanism is distinctly smaller and less prominent (
|
||
<figureCitation id="13012A62FFBF3A0EFDFD5907FD5EFEA2" box="[580,666,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="3" pageNumber="367">Fig. 3B</figureCitation>
|
||
) than in
|
||
<taxonomicName id="4C3A4D64FFBF3A0EFD475907FC89FEA3" authorityName="Huang, Wang & Shih" authorityYear="2020" box="[766,845,259,284]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="367" phylum="Arthropoda" rank="species" species="puer">
|
||
<emphasis id="B94EEAF5FFBF3A0EFD475907FC89FEA3" box="[766,845,259,284]" italics="true" pageId="3" pageNumber="367">T. puer</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A4D64FFBF3A0EFC3D5907FC2AFEA3" authorityName="C Huang, J Wang & HT Shih" authorityYear="2020" box="[900,1006,259,284]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="367" phylum="Arthropoda" rank="species" species="gejiuin">
|
||
<emphasis id="B94EEAF5FFBF3A0EFC3D5907FC2AFEA3" box="[900,1006,259,284]" italics="true" pageId="3" pageNumber="367">T. gejiuin</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFAB4B16FFBF3A0EFC445907FB08FEA2" author="Huang, C. & Wang, J. & Shih, H. - T." box="[1021,1228,259,285]" pageId="3" pageNumber="367" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBF3A0EFBE95907FB4DFEA2" box="[1104,1161,259,285]" italics="true" pageId="3" pageNumber="367">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 2D, 5D); the male pleon is distinctly broader and the lateral margin of the telson is gently concave (
|
||
<figureCitation id="13012A62FFBF3A0EFBDC5923FB1BFEFE" box="[1125,1247,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="3" pageNumber="367">Fig. 3A, D</figureCitation>
|
||
) (vs. male pleon proportionately narrower and lateral margin of telson strongly concave;
|
||
<bibRefCitation id="EFAB4B16FFBF3A0EFC79594FFB49FEDA" author="Huang, C. & Wang, J. & Shih, H. - T." box="[960,1165,331,357]" pageId="3" pageNumber="367" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBF3A0EFBAB594FFB8EFEDA" box="[1042,1098,331,357]" italics="true" pageId="3" pageNumber="367">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 2B, C, 5B, C); the G1 is more distinctly bent, with the terminal segment relatively slender, tapering distally (
|
||
<figureCitation id="13012A62FFBF3A0EFB32596BFAD6FE36" box="[1163,1298,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="3" pageNumber="367">Figs. 3G, H</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBF3A0EFAA7596BFAA0FE36" box="[1310,1380,367,393]" captionStart="FIGURE 4" captionStartId="5.[151,250,1070,1094]" captionTargetBox="[153,1434,183,1044]" captionTargetId="figure-32@5.[151,1436,181,1046]" captionTargetPageId="5" captionText="FIGURE 4. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsomesial view); D, distal part of left G1 (dorsal view); E, left G2 (ventral view); F, distal part of left G2 (ventral view); G, distal part of left G2 (dorsal view). Scales: A, E = 0.5 mm; B–D, F, G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7923538" httpUri="https://zenodo.org/record/7923538/files/figure.png" pageId="3" pageNumber="367">4A–D</figureCitation>
|
||
) (vs. G1 gently curved to almost straight, the terminal segment being relatively stouter, shorter, conical;
|
||
<bibRefCitation id="EFAB4B16FFBF3A0EFAB75997FF17FE6E" author="Huang, C. & Wang, J. & Shih, H. - T." pageId="3" pageNumber="367" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBF3A0EFADB5997FA59FE12" box="[1378,1437,403,429]" italics="true" pageId="3" pageNumber="367">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 3C–E, 6C–E); the G2 is distinctly longer than the G1 (
|
||
<figureCitation id="13012A62FFBF3A0EFCC559B3FC3DFE6E" box="[892,1017,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="3" pageNumber="367">Figs. 3G–I</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBF3A0EFBBD59B3FB83FE6E" box="[1028,1095,439,465]" captionStart="FIGURE 4" captionStartId="5.[151,250,1070,1094]" captionTargetBox="[153,1434,183,1044]" captionTargetId="figure-32@5.[151,1436,181,1046]" captionTargetPageId="5" captionText="FIGURE 4. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsomesial view); D, distal part of left G1 (dorsal view); E, left G2 (ventral view); F, distal part of left G2 (ventral view); G, distal part of left G2 (dorsal view). Scales: A, E = 0.5 mm; B–D, F, G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7923538" httpUri="https://zenodo.org/record/7923538/files/figure.png" pageId="3" pageNumber="367">4A, E</figureCitation>
|
||
) (vs. G2 subequal in length to G1;
|
||
<bibRefCitation id="EFAB4B16FFBF3A0EFF7E59DFFE54FE4A" author="Huang, C. & Wang, J. & Shih, H. - T." box="[199,400,475,501]" pageId="3" pageNumber="367" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBF3A0EFEAE59DFFE8AFE4A" box="[279,334,475,501]" italics="true" pageId="3" pageNumber="367">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 3B–E, 6B–E); the G2 has a distinctly longer flagellum which is ca. two-thirds the length of the basal segment (
|
||
<figureCitation id="13012A62FFBF3A0EFE3159FBFE1BFDA6" box="[392,479,511,537]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="3" pageNumber="367">Figs. 3I</figureCitation>
|
||
,
|
||
<figureCitation id="13012A62FFBF3A0EFE5359FBFDCFFDA6" box="[490,523,511,537]" captionStart="FIGURE 4" captionStartId="5.[151,250,1070,1094]" captionTargetBox="[153,1434,183,1044]" captionTargetId="figure-32@5.[151,1436,181,1046]" captionTargetPageId="5" captionText="FIGURE 4. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsomesial view); D, distal part of left G1 (dorsal view); E, left G2 (ventral view); F, distal part of left G2 (ventral view); G, distal part of left G2 (dorsal view). Scales: A, E = 0.5 mm; B–D, F, G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7923538" httpUri="https://zenodo.org/record/7923538/files/figure.png" pageId="3" pageNumber="367">4E</figureCitation>
|
||
) (vs. G2 flagellum shorter, ca. half the length of basal segment;
|
||
<bibRefCitation id="EFAB4B16FFBF3A0EFB7659FBFA5CFDA6" author="Huang, C. & Wang, J. & Shih, H. - T." box="[1231,1432,511,537]" pageId="3" pageNumber="367" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBF3A0EFAA659FBFA93FDA6" box="[1311,1367,511,537]" italics="true" pageId="3" pageNumber="367">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 3B, 6B); and the vulva is smaller and less prominent (
|
||
<figureCitation id="13012A62FFBF3A0EFC935A27FC44FD82" box="[810,896,546,573]" captionStart="FIGURE 5" captionStartId="6.[151,250,1960,1984]" captionTargetBox="[166,1421,921,1937]" captionTargetId="figure-406@6.[166,1421,921,1937]" captionTargetPageId="6" captionText="FIGURE 5. Tortomon distinctum n. sp., paratype female (18.0 × 12.3 mm) (ZRC 2022.798), Vietnam.A, overall dorsal habitus; B, buccal cavity, anterior thoracic sternum and pleon; C, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.7923540" httpUri="https://zenodo.org/record/7923540/files/figure.png" pageId="3" pageNumber="367">Fig. 5C</figureCitation>
|
||
) (
|
||
<bibRefCitation id="EFAB4B16FFBF3A0EFC2E5A27FBA2FD82" author="Huang, C. & Wang, J. & Shih, H. - T." box="[919,1126,547,573]" pageId="3" pageNumber="367" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. & Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
|
||
Huang
|
||
<emphasis id="B94EEAF5FFBF3A0EFC525A27FBE7FD82" box="[1003,1059,547,573]" italics="true" pageId="3" pageNumber="367">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: figs. 2F, 5F).
|
||
</paragraph>
|
||
<caption id="DF45666FFFBF3A0EFF2E5FB1FA55F84E" ID-DOI="http://doi.org/10.5281/zenodo.7923534" ID-Zenodo-Dep="7923534" httpUri="https://zenodo.org/record/7923534/files/figure.png" pageId="3" pageNumber="367" startId="3.[151,250,1973,1997]" targetBox="[183,1405,597,1950]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8B8536E7FFBF3A0EFF2E5FB1FA55F84E" blockId="3.[151,1436,1973,2033]" pageId="3" pageNumber="367">
|
||
<emphasis id="B94EEAF5FFBF3A0EFF2E5FB1FED7F872" bold="true" box="[151,275,1973,1997]" pageId="3" pageNumber="367">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C3A4D64FFBF3A0EFEA35FB2FE28F872" authority="Ng & Tri, 2023" authorityName="Ng & Tri" authorityYear="2023" box="[282,492,1973,1997]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="367" phylum="Arthropoda" rank="species" species="distinctum" status="sp. nov.">
|
||
<emphasis id="B94EEAF5FFBF3A0EFEA35FB2FE28F872" box="[282,492,1973,1997]" italics="true" pageId="3" pageNumber="367">Tortomon distinctum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94EEAF5FFBF3A0EFE4B5FB1FDEFF872" bold="true" box="[498,555,1973,1997]" pageId="3" pageNumber="367">
|
||
<taxonomicNameLabel id="A27D578EFFBF3A0EFE4B5FB1FDEFF872" box="[498,555,1973,1997]" pageId="3" pageNumber="367" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, overall dorsal habitus; B, dorsal view of carapace, C, frontal view of cephalothorax; D, epistome and antennules; E, right third maxilliped; F, chelae.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF45666FFFB83A09FF2E5F68FC46F84F" ID-DOI="http://doi.org/10.5281/zenodo.7923536" ID-Zenodo-Dep="7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="4" pageNumber="368" startId="4.[151,250,1900,1924]" targetBox="[167,1421,181,1876]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B8536E7FFB83A09FF2E5F68FC46F84F" blockId="4.[151,1437,1900,2032]" pageId="4" pageNumber="368">
|
||
<emphasis id="B94EEAF5FFB83A09FF2E5F68FED6F83B" bold="true" box="[151,274,1900,1924]" pageId="4" pageNumber="368">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C3A4D64FFB83A09FEAE5F69FE2CF83B" authority="Ng & Tri, 2023" authorityName="Ng & Tri" authorityYear="2023" box="[279,488,1900,1924]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="368" phylum="Arthropoda" rank="species" species="distinctum" status="sp. nov.">
|
||
<emphasis id="B94EEAF5FFB83A09FEAE5F69FE2CF83B" box="[279,488,1900,1924]" italics="true" pageId="4" pageNumber="368">Tortomon distinctum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94EEAF5FFB83A09FE545F68FDE1F83B" bold="true" box="[493,549,1900,1924]" pageId="4" pageNumber="368">
|
||
<taxonomicNameLabel id="A27D578EFFB83A09FE545F68FDE1F83B" box="[493,549,1900,1924]" pageId="4" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF45666FFFB93A08FF2E5C2AFEF6FB0D" ID-DOI="http://doi.org/10.5281/zenodo.7923538" ID-Zenodo-Dep="7923538" httpUri="https://zenodo.org/record/7923538/files/figure.png" pageId="5" pageNumber="369" startId="5.[151,250,1070,1094]" targetBox="[153,1434,183,1044]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B8536E7FFB93A08FF2E5C2AFEF6FB0D" blockId="5.[151,1436,1070,1203]" pageId="5" pageNumber="369">
|
||
<emphasis id="B94EEAF5FFB93A08FF2E5C2AFED7FBF9" bold="true" box="[151,275,1070,1094]" pageId="5" pageNumber="369">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C3A4D64FFB93A08FEA05C2BFE2FFBF9" authority="Ng & Tri, 2023" authorityName="Ng & Tri" authorityYear="2023" box="[281,491,1070,1094]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="369" phylum="Arthropoda" rank="species" species="distinctum" status="sp. nov.">
|
||
<emphasis id="B94EEAF5FFB93A08FEA05C2BFE2FFBF9" box="[281,491,1070,1094]" italics="true" pageId="5" pageNumber="369">Tortomon distinctum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94EEAF5FFB93A08FE485C2BFDEEFBF8" bold="true" box="[497,554,1071,1095]" pageId="5" pageNumber="369">
|
||
<taxonomicNameLabel id="A27D578EFFB93A08FE485C2BFDEEFBF8" box="[497,554,1071,1095]" pageId="5" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsomesial view); D, distal part of left G1 (dorsal view); E, left G2 (ventral view); F, distal part of left G2 (ventral view); G, distal part of left G2 (dorsal view). Scales: A, E = 0.5 mm; B–D, F, G = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8536E7FFB93A08FF7E5CE5FC1FFAD8" blockId="5.[151,1437,1249,1779]" pageId="5" pageNumber="369">
|
||
The characters of
|
||
<taxonomicName id="4C3A4D64FFB93A08FE205CE5FDE8FB44" authority="Ng & Tri, 2023" authorityName="Ng & Tri" authorityYear="2023" box="[409,556,1249,1275]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="369" phylum="Arthropoda" rank="species" species="distinctum" status="sp. nov.">
|
||
<emphasis id="B94EEAF5FFB93A08FE205CE5FDE8FB44" box="[409,556,1249,1275]" italics="true" pageId="5" pageNumber="369">T. distinctum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94EEAF5FFB93A08FD8F5CE5FDB3FB44" bold="true" box="[566,631,1249,1275]" pageId="5" pageNumber="369">
|
||
<taxonomicNameLabel id="A27D578EFFB93A08FD8F5CE5FDB3FB44" box="[566,631,1249,1275]" pageId="5" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, notably in the longer third maxilliped, male pleonal shape and more elongate G2, may argue for placing the species in a separate genus from
|
||
<taxonomicName id="4C3A4D64FFB93A08FC705D01FBF1FAA1" authorityName="Huang, Wang & Shih" authorityYear="2020" box="[969,1077,1285,1310]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="369" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94EEAF5FFB93A08FC705D01FBF1FAA1" box="[969,1077,1285,1310]" italics="true" pageId="5" pageNumber="369">Tortomon</emphasis>
|
||
</taxonomicName>
|
||
. That being said, the distinctive structure of the G1 and twisted G2 flagellum suggest treating all three species as congeneric for now. To this effect, we have amended the diagnosis for the genus to reflect these observations.
|
||
</paragraph>
|
||
<paragraph id="8B8536E7FFB93A08FF7E5D75FC30FA48" blockId="5.[151,1437,1249,1779]" pageId="5" pageNumber="369">
|
||
The smooth ovate carapace, entire anterolateral margins and the absence of a flagellum on the exopod of the third maxilliped of
|
||
<taxonomicName id="4C3A4D64FFB93A08FECC5D91FDC3FA10" authority="Ng & Tri, 2023" authorityName="Ng & Tri" authorityYear="2023" box="[373,519,1429,1455]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="369" phylum="Arthropoda" rank="species" species="distinctum" status="sp. nov.">
|
||
<emphasis id="B94EEAF5FFB93A08FECC5D91FDC3FA10" box="[373,519,1429,1455]" italics="true" pageId="5" pageNumber="369">T. distinctum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94EEAF5FFB93A08FDB65D91FD8BFA10" bold="true" box="[527,591,1429,1455]" pageId="5" pageNumber="369">
|
||
<taxonomicNameLabel id="A27D578EFFB93A08FDB65D91FD8BFA10" box="[527,591,1429,1455]" pageId="5" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also allies it with
|
||
<taxonomicName id="4C3A4D64FFB93A08FC9D5D91FB84FA10" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[804,1088,1429,1455]" class="Malacostraca" family="Potamidae" genus="Xestomon" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="369" phylum="Arthropoda" rank="species" species="tacu">
|
||
<emphasis id="B94EEAF5FFB93A08FC9D5D91FC0EFA11" box="[804,970,1429,1454]" italics="true" pageId="5" pageNumber="369">Xestomon tacu</emphasis>
|
||
<bibRefCitation id="EFAB4B16FFB93A08FC6A5D91FB84FA10" author="Ng, P. K. L." box="[979,1088,1429,1455]" pageId="5" pageNumber="369" pagination="65 - 73" refId="ref5630" refString="Ng, P. K. L. (2021) Description of a new genus and new species of freshwater crab (Brachyura: Potamoidea) from southern Vietnam. Crustacean Research, 50, 65 - 73. https: // doi. org / 10.18353 / crustacea. 50.0 _ 65" type="journal article" year="2021">Ng, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, described from a mountain in southern
|
||
<collectingCountry id="F32D7677FFB93A08FF445DBDFE98FA6C" box="[253,348,1465,1491]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFAB4B16FFB93A08FED05DBDFE0AFA6C" author="Ng, P. K. L." box="[361,462,1465,1491]" pageId="5" pageNumber="369" pagination="65 - 73" refId="ref5630" refString="Ng, P. K. L. (2021) Description of a new genus and new species of freshwater crab (Brachyura: Potamoidea) from southern Vietnam. Crustacean Research, 50, 65 - 73. https: // doi. org / 10.18353 / crustacea. 50.0 _ 65" type="journal article" year="2021">Ng 2021</bibRefCitation>
|
||
). The form of the posterior margin of the epistome, male anterior thoracic sternum, male pleon, G1, and most significantly, the G2, however, are completely different.
|
||
</paragraph>
|
||
<paragraph id="8B8536E7FFB93A08FF7E5E05FCC1F980" blockId="5.[151,1437,1249,1779]" pageId="5" pageNumber="369">
|
||
The colour of
|
||
<taxonomicName id="4C3A4D64FFB93A08FED05E05FE7CF9A5" authorityName="Huang, Wang & Shih" authorityYear="2020" box="[361,440,1537,1562]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="369" phylum="Arthropoda" rank="species" species="puer">
|
||
<emphasis id="B94EEAF5FFB93A08FED05E05FE7CF9A5" box="[361,440,1537,1562]" italics="true" pageId="5" pageNumber="369">T. puer</emphasis>
|
||
</taxonomicName>
|
||
is brownish purple and most similar to the distinct purple colour of
|
||
<taxonomicName id="4C3A4D64FFB93A08FB045E05FA8AF9A4" authority="Ng & Tri, 2023" authorityName="Ng & Tri" authorityYear="2023" box="[1213,1358,1537,1563]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="369" phylum="Arthropoda" rank="species" species="distinctum" status="sp. nov.">
|
||
<emphasis id="B94EEAF5FFB93A08FB045E05FA8AF9A4" box="[1213,1358,1537,1563]" italics="true" pageId="5" pageNumber="369">T. distinctum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94EEAF5FFB93A08FAEC5E05FA51F9A4" bold="true" box="[1365,1429,1537,1563]" pageId="5" pageNumber="369">
|
||
<taxonomicNameLabel id="A27D578EFFB93A08FAEC5E05FA51F9A4" box="[1365,1429,1537,1563]" pageId="5" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, being distinct from the greyish-white colour of
|
||
<taxonomicName id="4C3A4D64FFB93A08FD135E21FD3AF981" authorityName="Huang, Wang & Shih" authorityYear="2020" box="[682,766,1573,1598]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="369" phylum="Arthropoda" rank="species" species="gejiu">
|
||
<emphasis id="B94EEAF5FFB93A08FD135E21FD3AF981" box="[682,766,1573,1598]" italics="true" pageId="5" pageNumber="369">T. gejiu</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C320656CFFB93A08FF7E5E4DFCDBF94C" pageId="5" pageNumber="369" type="biology_ecology">
|
||
<paragraph id="8B8536E7FFB93A08FF7E5E4DFB53F938" blockId="5.[151,1437,1249,1779]" pageId="5" pageNumber="369">
|
||
<emphasis id="B94EEAF5FFB93A08FF7E5E4DFEE8F9DC" bold="true" box="[199,300,1609,1635]" pageId="5" pageNumber="369">Ecology.</emphasis>
|
||
This is a semi-aquatic freshwater crab, inhabiting the banks of a small stream in a small karst outcrop in a black cardamon plantation (
|
||
<emphasis id="B94EEAF5FFB93A08FDB85E69FDA1F938" box="[513,613,1645,1671]" italics="true" pageId="5" pageNumber="369">Amomon</emphasis>
|
||
sp.,
|
||
<taxonomicName id="4C3A4D64FFB93A08FD205E69FCFEF938" authorityName="Martinov" authorityYear="1820" box="[665,826,1645,1671]" class="Liliopsida" family="Zingiberaceae" kingdom="Plantae" order="Zingiberales" pageId="5" pageNumber="369" phylum="Tracheophyta" rank="family">Zingiberaceae</taxonomicName>
|
||
) at high altitudes (
|
||
<quantity id="4CC29B02FFB93A08FBB35E69FBA4F938" box="[1034,1120,1645,1671]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="369" unit="m" value="1800.0">1800 m</quantity>
|
||
asl).
|
||
</paragraph>
|
||
<paragraph id="8B8536E7FFB93A08FF7E5E94FCDBF94C" blockId="5.[151,1437,1249,1779]" pageId="5" pageNumber="369">
|
||
<emphasis id="B94EEAF5FFB93A08FF7E5E94FEABF915" bold="true" box="[199,367,1680,1706]" pageId="5" pageNumber="369">Conservation.</emphasis>
|
||
This is a small species and does not appear to be under immediate threat. The area it occurs in, however, is not protected, and until its distribution is better understood, it should be regarded as a vulnerable species with a very restricted range (cf.
|
||
<bibRefCitation id="EFAB4B16FFB93A08FE475EDDFCD7F94C" author="Cumberlidge, N. & Ng, P. K. L. & Yeo, D. C. J. & Magalhaes, C. & Campos, M. R. & Alvarez, F. & Naruse, T. & Daniels, S. R. & Esser, L. J. & Attipoe, F. Y. K. & Clotilde-Ba, F. - L. & Darwall, W. & Mcivor, A. & Collen, B. & Ram, M." box="[510,787,1753,1779]" pageId="5" pageNumber="369" pagination="1665 - 1673" refId="ref5090" refString="Cumberlidge, N., Ng, P. K. L., Yeo, D. C. J., Magalhaes, C., Campos, M. R., Alvarez, F., Naruse, T., Daniels, S. R., Esser, L. J., Attipoe, F. Y. K. Clotilde-Ba, F. - L., Darwall, W., Mcivor, A., Collen, B. & Ram, M. (2009) Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation, 142, 1665 - 1673. https: // doi. org / 10.1016 / j. biocon. 2009.02.038" type="journal article" year="2009">
|
||
Cumberlidge
|
||
<emphasis id="B94EEAF5FFB93A08FD2E5EDDFD0BF94C" box="[663,719,1753,1779]" italics="true" pageId="5" pageNumber="369">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |