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<document id="10512C90A604DF8C52580EDEE5B15EDB" ID-CLB-Dataset="8556" ID-DOI="10.11646/zootaxa.4843.1.1" ID-GBIF-Dataset="a2aef684-44f1-429d-9dd5-dd394539fa36" ID-ISSN="1175-5326" ID-Zenodo-Dep="4405729" ID-ZooBank="57044928-1950-4525-850E-E381A52599A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609419131752" checkinUser="plazi" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="03938025FF95A90BA7D8B212FAFBFA60" docLanguage="en" docName="zootaxa.4843.1.1.pdf" docOrigin="Zootaxa 4843 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole flammea Salata &amp; Fisher 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="30" masterDocId="FFAAF85DFF8EA916A74FB608FFC6FFFD" masterDocTitle="The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group" masterLastPageNumber="64" masterPageNumber="1" pageNumber="28" updateTime="1698850390174" updateUser="ExternalLinkService">
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<mods:title id="127CA69EBC3E627E2A4280D5DEBA4919">The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group</mods:title>
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<mods:namePart id="2F60054A4018CEC0F34E333B8F89A0FF">Salata, Sebastian</mods:namePart>
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<mods:namePart id="7DAB2B1192F47DA50FBE63816420981D">Fisher, Brian L.</mods:namePart>
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<mods:nameIdentifier id="3E79C80AAD0E3AFE9F389CE15C1C43D2" type="email">bfisher@calacademy.org</mods:nameIdentifier>
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<mods:title id="F2CC33877701DC858F0685416C959886">Zootaxa</mods:title>
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<mods:part id="0B67389CDED0EF85DB6BA792A8B65ED6">
<mods:date id="82D505DEFA3D802956E2AA1614267124">2020</mods:date>
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<mods:number id="CBEED9A65CD80F040EFDC3E30B7B3A6C">2020-08-31</mods:number>
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<mods:number id="7CC345CD213010CF80519F368589580F">4843</mods:number>
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<mods:identifier id="D2FA3F91AACFF88708B229FC88BD9C92" type="DOI">10.11646/zootaxa.4843.1.1</mods:identifier>
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<treatment id="03938025FF95A90BA7D8B212FAFBFA60" ID-DOI="http://doi.org/10.5281/zenodo.4488385" ID-GBIF-Taxon="177202125" ID-Zenodo-Dep="4488385" LSID="urn:lsid:plazi:treatment:03938025FF95A90BA7D8B212FAFBFA60" httpUri="http://treatment.plazi.org/id/03938025FF95A90BA7D8B212FAFBFA60" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
<subSubSection id="C32062B8FF95A90DA7D8B212FE11FBC8" box="[151,471,1050,1077]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8B853133FF95A90DA7D8B212FE11FBC8" blockId="27.[151,471,1050,1113]" box="[151,471,1050,1077]" pageId="27" pageNumber="28">
<heading id="D0CD865FFF95A90DA7D8B212FE11FBC8" bold="true" box="[151,471,1050,1077]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
<emphasis id="B94EED21FF95A90DA7D8B212FE11FBC8" bold="true" box="[151,471,1050,1077]" pageId="27" pageNumber="28">
<taxonomicName id="4C3A4AB0FF95A90DA7D8B212FEB4FBC8" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,370,1050,1077]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="flammea" status="sp. nov.">
<emphasis id="B94EED21FF95A90DA7D8B212FEB4FBC8" bold="true" box="[151,370,1050,1077]" italics="true" pageId="27" pageNumber="28">Pheidole flammea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D505AFF95A90DA635B213FE11FBC8" box="[378,471,1051,1077]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF95A90DA7D8B237FE4DFBA4" box="[151,395,1087,1113]" pageId="27" pageNumber="28" type="description">
<paragraph id="8B853133FF95A90DA7D8B237FE4DFBA4" blockId="27.[151,471,1050,1113]" box="[151,395,1087,1113]" pageId="27" pageNumber="28">
<figureCitation id="13012DB6FF95A90DA7D8B237FF2EFBA4" box="[151,232,1087,1113]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="27" pageNumber="28">Figs 14</figureCitation>
AF, 27F, 28F
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF95A90DA7D8B28FFCB4FAF0" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8B853133FF95A90DA7D8B28FFBA9FB14" blockId="27.[151,1437,1158,1941]" pageId="27" pageNumber="28">
<materialsCitation id="3B523B6EFF95A90DA7D8B28FFB7EFB38" ID-GBIF-Occurrence="3027954625" accessionNumber="BLF30615" collectingDate="2013-02-17" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="520" latitude="-13.58487" location="Mont Galoko" longLatPrecision="1" longitude="48.71818" municipality="Galoko" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<typeStatus id="54818F91FF95A90DA7D8B28FFEE9FB5C" box="[151,303,1159,1185]" pageId="27" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
: 1s.,
<collectingCountry id="F32D71A3FF95A90DA621B28FFE31FB5C" box="[366,503,1159,1185]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
,
<collectingRegion id="49FEFFD1FF95A90DA54BB28FFD48FB5C" box="[516,654,1159,1185]" country="Madagascar" name="Antsiranana" pageId="27" pageNumber="28">Antsiranana</collectingRegion>
,
<collectingMunicipality id="6BE1AB49FF95A90DA5D3B28FFD28FB5C" box="[668,750,1159,1185]" pageId="27" pageNumber="28">Galoko</collectingMunicipality>
chain,
<location id="8EE567E8FF95A90DA408B28FFC25FB5C" LSID="urn:lsid:plazi:treatment:03938025FF95A90BA7D8B212FAFBFA60:8EE567E8FF95A90DA408B28FFC25FB5C" box="[839,995,1159,1185]" country="Madagascar" latitude="-13.58487" longLatPrecision="1" longitude="48.71818" municipality="Galoko" name="Mont Galoko" pageId="27" pageNumber="28" stateProvince="Antsiranana">Mont Galoko</location>
,
<geoCoordinate id="EE0E57F4FF95A90DA4BDB28EFBA2FB5C" box="[1010,1124,1158,1185]" degrees="13.58487" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="-13.58487">-13.58487</geoCoordinate>
<geoCoordinate id="EE0E57F4FF95A90DA321B28FFB1CFB5C" box="[1134,1242,1159,1185]" degrees="48.71818" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="1" value="48.71818">48.71818</geoCoordinate>
,
<quantity id="4CC29CD6FF95A90DA3A7B28EFAF3FB5C" box="[1256,1333,1158,1185]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="27" pageNumber="28" unit="m" value="520.0">
<elevation id="0017D600FF95A90DA3A7B28EFAF3FB5C" box="[1256,1333,1158,1185]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="27" pageNumber="28" unit="m" value="520.0">520 m</elevation>
</quantity>
,
<date id="FF8417F3FF95A90DA20BB28FFF14FB38" pageId="27" pageNumber="28" value="2013-02-17">
<collectingDate id="EFC0EE1BFF95A90DA20BB28FFF14FB38" pageId="27" pageNumber="28" value="2013-02-17">17-Feb- 2013</collectingDate>
</date>
, rainforest, ex rotten log,
<collectorName id="26CF54E5FF95A90DA6B8B2A3FDB9FB38" box="[503,639,1195,1221]" pageId="27" pageNumber="28">B. L. Fisher</collectorName>
<emphasis id="B94EED21FF95A90DA5C8B2A3FD7FFB38" box="[647,697,1195,1221]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. leg. BLF30615, CASENT0923251 (
<collectionCode id="ED2BA9F6FF95A90DA32EB2A3FB76FB38" box="[1121,1200,1195,1221]" pageId="27" pageNumber="28">CASC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B523B6EFF95A90DA38CB2A3FBACFB14" ID-GBIF-Occurrence="3027954520" accessionNumber="BLF30615" collectingDate="2013-02-17" collectionCode="CASC, MHNG" collectorName="B. L. Fisher" country="Madagascar" elevation="520" latitude="-13.58487" location="Mont Galoko" longLatPrecision="1" longitude="48.71818" municipality="Galoko" pageId="27" pageNumber="28" specimenCount="2" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus id="54818F91FF95A90DA38CB2A3FAA4FB38" box="[1219,1378,1195,1221]" pageId="27" pageNumber="28" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D3CFABAFF95A90DA221B2A3FA51FB38" box="[1390,1431,1195,1221]" pageId="27" pageNumber="28" type="generic">1w.</specimenCount>
,
<specimenCount id="9D3CFABAFF95A90DA7D8B2C7FF7AFB14" box="[151,188,1231,1257]" pageId="27" pageNumber="28" type="generic">1q.</specimenCount>
, the same data as holotype, CASENT0298332, CASENT0298333 (
<collectionCode id="ED2BA9F6FF95A90DA4FEB2C7FC38FB14" box="[945,1022,1231,1257]" pageId="27" pageNumber="28">CASC</collectionCode>
,
<collectionCode id="ED2BA9F6FF95A90DA346B2C7FB99FB14" box="[1033,1119,1231,1257]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="27" pageNumber="28" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B853133FF95A90DA788B2FAFCB4FAF0" blockId="27.[151,1437,1158,1941]" box="[199,882,1266,1293]" pageId="27" pageNumber="28">
<emphasis id="B94EED21FF95A90DA788B2FAFE60FAF0" bold="true" box="[199,422,1266,1293]" pageId="27" pageNumber="28">Geographic range.</emphasis>
<collectingCountry id="F32D71A3FF95A90DA6E2B2FBFDF0FAF0" box="[429,566,1267,1293]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
,
<collectingRegion id="49FEFFD1FF95A90DA570B2FBFD0FFAF0" box="[575,713,1267,1293]" country="Madagascar" name="Antsiranana" pageId="27" pageNumber="28">Antsiranana</collectingRegion>
, Mont Galoko.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF95A90DA788B31EFEB4F8D4" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8B853133FF95A90DA788B31EFEB4F8D4" blockId="27.[151,1437,1158,1941]" pageId="27" pageNumber="28">
<emphasis id="B94EED21FF95A90DA788B31EFDC5FACC" bold="true" box="[199,515,1302,1329]" pageId="27" pageNumber="28">Diagnosis. Major workers.</emphasis>
Head in full-face view sub-rectangular, not widening posteriorly, with lateral margins slightly convex; side of head with moderately dense, long, suberect to erect pilosity; medial frons with sparse costulae; interspaces between costulae smooth; frons laterally with thick and dense rugae; interspaces between rugae smooth to indistinctly rugulate; occipital lobes with thinner rugae; interspaces between rugae smooth; scape, when laid back, exceeding the midlength of head by two-fifths of its length; inner hypostomal tooth small, bulge-like, and arching posteroventrally; outer hypostomal tooth lobe-like, high, and wide, with top arching posterolaterally; median tooth absent; promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct and tubercle-like; mesosoma microreticulate; promesonotal dorsum with reduced microreticulae and additional transverse rugae; lateral pronotum with smooth notch; katepisternum and lateral sides of propodeum with additional rugae; gaster indistinctly shagreened; body brown.
<emphasis id="B94EED21FF95A90DA343B052FB0EF988" bold="true" box="[1036,1224,1626,1653]" pageId="27" pageNumber="28">Minor workers.</emphasis>
Occiput elongated forming short and moderately narrow neck; nuchal collar distinct; head sculpture smooth; scape, when laid back, exceeding the posterior head margin by three-fifths of its length; promesonotum low, long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines moderately large and thin; pronotum, mesonotum, and dorsal propodeum smooth; anepisternum, katepisternum, and lateral sides of propodeum with sparse puncta and rugae; body brown.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF95A90BA788B13AFC89FBA4" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="8B853133FF95A90DA788B13AFF16F868" blockId="27.[151,1437,1158,1941]" pageId="27" pageNumber="28">
<emphasis id="B94EED21FF95A90DA788B13AFDDBF8B0" bold="true" box="[199,541,1842,1869]" pageId="27" pageNumber="28">Description. Major workers.</emphasis>
Measurements (n=1): HL: 1.65; HW: 1.5; SL: 1.19; EL: 0.18; WL: 1.45; PSL: 0.3; MTL: 1.21; PNW: 0.56; PTW: 0.18; PPW: 0.46; CI: 110.7; SI: 79.7; PSLI: 17.9; PPI: 38.8; PNI: 37.6; MTI: 81.1.
</paragraph>
<caption id="DF4561BBFF92A90AA7D8B0A0FB61F919" ID-DOI="http://doi.org/10.5281/zenodo.4405767" ID-Zenodo-Dep="4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1704,1728]" targetBox="[151,1436,181,1680]" targetPageId="28">
<paragraph id="8B853133FF92A90AA7D8B0A0FB61F919" blockId="28.[151,1436,1704,1764]" pageId="28" pageNumber="29">
<emphasis id="B94EED21FF92A90AA7D8B0A0FEDBF93D" bold="true" box="[151,285,1704,1728]" pageId="28" pageNumber="29">FIGURE 14.</emphasis>
<taxonomicName id="4C3A4AB0FF92A90AA66EB0A0FE13F93D" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[289,469,1704,1728]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FF92A90AA66EB0A0FE13F93D" box="[289,469,1704,1728]" italics="true" pageId="28" pageNumber="29">Pheidole flammea</emphasis>
</taxonomicName>
, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251).
</paragraph>
</caption>
<paragraph id="8B853133FF92A90BA788B11BFDCFFD9C" blockId="28.[151,1437,1810,2017]" lastBlockId="29.[151,1437,151,609]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="B94EED21FF92A90AA788B11BFEC8F8D0" bold="true" box="[199,270,1811,1837]" pageId="28" pageNumber="29">Head.</emphasis>
In full-face view sub-rectangular, not widening posteriorly with lateral sides slightly convex (
<figureCitation id="13012DB6FF92A90AA266B11AFA49F8D0" box="[1321,1423,1810,1837]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="28" pageNumber="29">Fig. 14B</figureCitation>
). In lateral view sub-oval, not depressed posteriorly; ventral and dorsal margins convex; inner hypostomal tooth visible. Side of head with moderately dense, long, suberect to erect pilosity; head dorsum with relatively dense, long, decumbent to erect pilosity. Medial frons with sparse costulae; interspaces between costulae smooth. Frons laterally with thick and dense rugae; interspaces between rugae smooth to indistinctly rugulate. Occipital lobes with thinner rugae; interspaces between rugae smooth. Gena with sparse and moderately thick costulae; interspaces between costulae smooth. Sides posterolateral from eyes with dense but very thin network of rugoreticulae; sculpture weakening posteriorly. Center of clypeus shiny and smooth, lateral sides with distinct rugulae; median notch present, moderately wide and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, exceeding the midlength of head by two-fifths of its length; pilosity subdecumbent to erect (
<figureCitation id="13012DB6FF93A90BA247B70BFA56FEE0" box="[1288,1424,259,285]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="29" pageNumber="30">Fig. 14B, D</figureCitation>
). Inner hypostomal tooth small, bulge-like, and arching posteroventrally; outer hypostomal tooth lobe-like, high, and wide, with top arching posterolaterally; median tooth absent (
<figureCitation id="13012DB6FF93A90BA40AB743FC6FFE98" box="[837,937,331,357]" captionStart="FIGURE 27" captionStartId="60.[151,250,1604,1628]" captionTargetBox="[151,1436,1137,1581]" captionTargetId="figure-432@60.[151,1436,1137,1581]" captionTargetPageId="60" captionText="FIGURE 27. Major worker, hypostomal tooth. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole decollata (E). Pheidole flammea (F). Pheidole flavodepressa (G). Pheidole grallatrix (H). Pheidole madecassa (I). Pheidole mantadioflava (J). Pheidole maro (K). Pheidole oswaldi (L). Pheidole ovalinoda (M). Pheidole similis (N). Pheidole tenebrovulgaris (O). Pheidole uranus (P). Pheidole voreios (Q). Pheidole zirafy (R)." figureDoi="http://doi.org/10.5281/zenodo.4405803" httpUri="https://zenodo.org/record/4405803/files/figure.png" pageId="29" pageNumber="30">Fig. 27F</figureCitation>
).
<emphasis id="B94EED21FF93A90BA4F3B743FB87FE98" bold="true" box="[956,1089,331,357]" pageId="29" pageNumber="30">Mesosoma.</emphasis>
In lateral view, promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct and tuberclelike; promesonotal groove absent; metanotal groove absent; propodeal spines long with narrow base and acute top; humeral tubercle weakly produced (
<figureCitation id="13012DB6FF93A90BA52CB7BFFD0BFE2C" box="[611,717,439,465]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="29" pageNumber="30">Fig. 14D</figureCitation>
). Surface shiny and microreticulate; promesonotal dorsum with reduced microreticulae and additional transverse rugae; lateral pronotum with smooth notch; katepisternum and lateral sides of propodeum with additional rugae. Pilosity dense, long, and erect (
<figureCitation id="13012DB6FF93A90BA37FB7F7FB7EFDE4" box="[1072,1208,511,537]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="29" pageNumber="30">Fig. 14D, F</figureCitation>
).
<emphasis id="B94EED21FF93A90BA382B7F6FAE4FDE5" bold="true" box="[1229,1314,510,536]" pageId="29" pageNumber="30">Gaster.</emphasis>
Shiny and indistinctly shagreened; pilosity moderately sparse, long, and erect (Fig. D, F).
<emphasis id="B94EED21FF93A90BA357B42AFBA7FDC1" bold="true" box="[1048,1121,546,572]" pageId="29" pageNumber="30">Color.</emphasis>
Brown, antenna, mandibles and legs yellowish (
<figureCitation id="13012DB6FF93A90BA638B44FFE3BFD9C" box="[375,509,583,609]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="29" pageNumber="30">Fig. 14D, F</figureCitation>
).
</paragraph>
<paragraph id="8B853133FF93A90BA788B486FF18FD0C" blockId="29.[151,1437,654,1437]" pageId="29" pageNumber="30">
<emphasis id="B94EED21FF93A90BA788B486FDDDFD54" bold="true" box="[199,539,654,681]" pageId="29" pageNumber="30">Description. Minor workers.</emphasis>
Measurements (n=1): HL: 0.79; HW: 0.49; SL: 1.17; EL: 0.13; WL: 1.03; PSL: 0.14; MTL: 1.03; PNW: 0.36; PTW: 0.08; PPW: 0.13; CI: 161.7; SI: 238.3; PSLI: 17.1; PPI: 60.3; PNI: 73.7; MTI: 208.8.
</paragraph>
<paragraph id="8B853133FF93A90BA788B4F3FA5AFC58" blockId="29.[151,1437,654,1437]" pageId="29" pageNumber="30">
<emphasis id="B94EED21FF93A90BA788B4F3FEC8FCE8" bold="true" box="[199,270,763,789]" pageId="29" pageNumber="30">Head.</emphasis>
In full-face view oval, posterior region elongated forming short and moderately narrow neck; nuchal collar distinct (
<figureCitation id="13012DB6FF93A90BA60EB517FE6FFCC4" box="[321,425,799,825]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="29" pageNumber="30">Fig. 14A</figureCitation>
). Pilosity relatively sparse, long, and subdecumbent to erect. Sculpture shiny and smooth; antennal sockets with few indistinct rugulae that are curved posterolaterally; interspaces between rugulae smooth. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by three-fifths of its length; pilosity dense, subdecumbent to erect (
<figureCitation id="13012DB6FF93A90BA248B583FA49FC58" box="[1287,1423,907,933]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="29" pageNumber="30">Fig. 14A, C</figureCitation>
).
</paragraph>
<paragraph id="8B853133FF93A90BA7D8B5A7FC89FBA4" blockId="29.[151,1437,654,1437]" pageId="29" pageNumber="30">
<emphasis id="B94EED21FF93A90BA7D8B5A7FEDAFC34" bold="true" box="[151,284,943,969]" pageId="29" pageNumber="30">Mesosoma.</emphasis>
In lateral view, promesonotum low, long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines moderately large, thin, and triangular (
<figureCitation id="13012DB6FF93A90BA4C2B5DBFC3EFC10" box="[909,1016,979,1005]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="29" pageNumber="30">Fig. 14C</figureCitation>
). Pronotum, mesonotum, and dorsal propodeum smooth; anepisternum, katepisternum, and lateral sides of propodeum with sparse puncta and rugae. Pilosity moderately sparse, long, and erect (
<figureCitation id="13012DB6FF93A90BA536B213FD3BFBC8" box="[633,765,1051,1077]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="29" pageNumber="30">Fig. 14C, E</figureCitation>
).
<emphasis id="B94EED21FF93A90BA440B212FCA2FBC9" bold="true" box="[783,868,1050,1076]" pageId="29" pageNumber="30">Gaster.</emphasis>
With sparse and erect pilosity (
<figureCitation id="13012DB6FF93A90BA3F1B213FA87FBC8" box="[1214,1345,1051,1077]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="29" pageNumber="30">Fig. 14C, E</figureCitation>
).
<emphasis id="B94EED21FF93A90BA21CB212FA5AFBC9" bold="true" box="[1363,1436,1050,1076]" pageId="29" pageNumber="30">Color.</emphasis>
Brown, legs, antennae and mandibles yellowish (
<figureCitation id="13012DB6FF93A90BA5F2B237FC85FBA4" box="[701,835,1087,1113]" captionStart="FIGURE14" captionStartId="28.[151,250,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-35@28.[151,1436,181,1680]" captionTargetPageId="28" captionText="FIGURE14. Pheidoleflammea, full-faceview (A), profile(C), anddorsal view (E)of paratype minorworker(CASENT0298333) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0923251)." figureDoi="http://doi.org/10.5281/zenodo.4405767" httpUri="https://zenodo.org/record/4405767/files/figure.png" pageId="29" pageNumber="30">Fig. 14C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF93A90BA788B26AFA83FB80" box="[199,1349,1122,1149]" pageId="29" pageNumber="30" type="biology_ecology">
<paragraph id="8B853133FF93A90BA788B26AFA83FB80" blockId="29.[151,1437,654,1437]" box="[199,1349,1122,1149]" pageId="29" pageNumber="30">
<emphasis id="B94EED21FF93A90BA788B26AFEE1FB81" bold="true" box="[199,295,1122,1148]" pageId="29" pageNumber="30">Biology.</emphasis>
The species was collected at
<quantity id="4CC29CD6FF93A90BA53DB26AFD7FFB80" box="[626,697,1122,1149]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="29" pageNumber="30" unit="m" value="520.0">520 m</quantity>
elevation, in rainforest. A nest was located in a rotten log.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF93A90BA788B28EFB96FA84" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8B853133FF93A90BA788B28EFB96FA84" blockId="29.[151,1437,654,1437]" pageId="29" pageNumber="30">
<emphasis id="B94EED21FF93A90BA788B28EFE89FB5D" bold="true" box="[199,335,1158,1184]" pageId="29" pageNumber="30">Comments.</emphasis>
<taxonomicName id="4C3A4AB0FF93A90BA619B28FFDD9FB5C" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[342,543,1159,1185]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FF93A90BA619B28FFDD9FB5C" box="[342,543,1159,1185]" italics="true" pageId="29" pageNumber="30">Pheidole flammea</emphasis>
</taxonomicName>
is known only from Galoko Mountain. Its majors are most similar to
<taxonomicName id="4C3A4AB0FF93A90BA26BB28FFF75FB38" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FF93A90BA26BB28FFF75FB38" italics="true" pageId="29" pageNumber="30">P. ovalinoda</emphasis>
</taxonomicName>
and differ in moderately dense, long and suberect to erect pilosity on the side of the head, large outer hypostomal tooth, and mostly smooth interspaces between rugae on head. Majors of
<taxonomicName id="4C3A4AB0FF93A90BA4F0B2C7FB8DFB14" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[959,1099,1231,1257]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FF93A90BA4F0B2C7FB8DFB14" box="[959,1099,1231,1257]" italics="true" pageId="29" pageNumber="30">P. ovalinoda</emphasis>
</taxonomicName>
have dense, short and decumbent to suberect setae on sides of head, strongly reduced outer hypostomal tooth, and more distinctly sculptured interspaces between rugae on head. Minors of
<taxonomicName id="4C3A4AB0FF93A90BA5EBB31FFCE6FACC" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[676,800,1303,1329]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FF93A90BA5EBB31FFCE6FACC" box="[676,800,1303,1329]" italics="true" pageId="29" pageNumber="30">P. flammea</emphasis>
</taxonomicName>
are most similar to parapatric
<taxonomicName id="4C3A4AB0FF93A90BA335B31FFACFFACC" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1146,1289,1303,1329]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="antsahabe">
<emphasis id="B94EED21FF93A90BA335B31FFACFFACC" box="[1146,1289,1303,1329]" italics="true" pageId="29" pageNumber="30">P. antsahabe</emphasis>
</taxonomicName>
and differ in brown body coloration, smooth promesonotum, and larger propodeal spines. In contrast, minors of
<taxonomicName id="4C3A4AB0FF93A90BA244B333FA5AFAA8" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1291,1436,1339,1365]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="antsahabe">
<emphasis id="B94EED21FF93A90BA244B333FA5AFAA8" box="[1291,1436,1339,1365]" italics="true" pageId="29" pageNumber="30">P. antsahabe</emphasis>
</taxonomicName>
have black body coloration, sculptured promesonotum, and shorter propodeal spines.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF93A90BA788B38BFAFBFA60" box="[199,1341,1411,1437]" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8B853133FF93A90BA788B38BFAFBFA60" blockId="29.[151,1437,654,1437]" box="[199,1341,1411,1437]" pageId="29" pageNumber="30">
<emphasis id="B94EED21FF93A90BA788B38BFE88FA60" bold="true" box="[199,334,1411,1437]" pageId="29" pageNumber="30">Etymology.</emphasis>
Latin for flames, in reference to the shape and density of setae on head of minor workers.
</paragraph>
</subSubSection>
</treatment>
</document>