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<mods:title id="1E4FA203B761694EE83A25093622D92D">On the identity of the species described in the genus Lithobius Leach, 1814 by L. J. Dobroruka from the former Czechoslovakia (Czech and Slovak Republics) (Chilopoda: Lithobiomorpha)</mods:title>
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<mods:namePart id="39E5B58D2842CE5785E3D323281CA6BC">Tuf, Ivan Hadrián</mods:namePart>
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<mods:namePart id="B1A680B92D849D99B9C7927A30F00A73">Wytwer, Jolanta</mods:namePart>
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<mods:namePart id="88282F3EDED4A9AF4ECD17DB1CCDC76D">Tajovský, Karel</mods:namePart>
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<treatment id="0392AA42FFC9FFEC34C4F98250B3817A" ID-DOI="http://doi.org/10.5281/zenodo.6229887" ID-GBIF-Taxon="119369025" ID-Zenodo-Dep="6229887" LSID="urn:lsid:plazi:treatment:0392AA42FFC9FFEC34C4F98250B3817A" httpUri="http://treatment.plazi.org/id/0392AA42FFC9FFEC34C4F98250B3817A" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42">
<subSubSection id="C32148DFFFC9FFEF34C4F9825090850F" box="[151,659,1618,1644]" pageId="5" pageNumber="42" type="nomenclature">
<paragraph id="8B841B54FFC9FFEF34C4F9825090850F" blockId="5.[151,659,1618,1644]" box="[151,659,1618,1644]" pageId="5" pageNumber="42">
<heading id="D0CCAC38FFC9FFEF34C4F9825090850F" bold="true" box="[151,659,1618,1644]" fontSize="11" level="1" pageId="5" pageNumber="42" reason="1">
<taxonomicName id="4C3B60D7FFC9FFEF34C4F9825090850F" ID-CoL="7JXRX" authority="Dobroruka, 1959" authorityName="Dobroruka" authorityYear="1959" box="[151,659,1618,1644]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="pusillus" subSpecies="johni">
<emphasis id="B94FC746FFC9FFEF34C4F9825090850F" bold="true" box="[151,659,1618,1644]" pageId="5" pageNumber="42">
<emphasis id="B94FC746FFC9FFEF34C4F98253B4850F" bold="true" box="[151,439,1618,1644]" italics="true" pageId="5" pageNumber="42">Lithobius pusillus johni</emphasis>
<bibRefCitation id="EFAA66A5FFC9FFEF35EDF9825090850F" author="Dobroruka" box="[446,659,1618,1644]" pageId="5" pageNumber="45" refString="Dobroruka, L. J. (1959) Chilopoda statni prirodni reservace Mohelno. [Chilopoden des staatlichen Naturschutzgebeites Mohelno.] Ochrana p r irody, 14, 104 - 106. (in Czech, with German summary)" type="journal article" year="1959">Dobroruka, 1959</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="8B841B54FFC9FFEF34C4F94D504A85D7" blockId="5.[151,701,1693,1751]" box="[151,585,1693,1716]" pageId="5" pageNumber="42">
<treatmentCitationGroup id="AB2B3C7AFFC9FFEF34C4F94D504A85D7" box="[151,585,1693,1716]" pageId="5" pageNumber="42">
<taxonomicName id="4C3B60D7FFC9FFEF34C4F94D504785D7" ID-CoL="72KVJ" authority="Dobroruka 1956: 271" authorityName="Dobroruka" authorityPageNumber="271" authorityYear="1956" box="[151,580,1693,1716]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B94FC746FFC9FFEF34C4F94D535685D7" box="[151,341,1693,1716]" italics="true" pageId="5" pageNumber="42">Lithobius pusillus</emphasis>
<treatmentCitation id="0A9A3D45FFC9FFEF350FF94D504785D7" author="Dobroruka" box="[348,580,1693,1716]" page="271" pageId="5" pageNumber="42" year="1956">
<bibRefCitation id="EFAA66A5FFC9FFEF350FF94D500C85D7" author="Dobroruka" box="[348,527,1693,1716]" pageId="5" pageNumber="45" refString="Dobroruka, L. J. (1956) O nekterych zajimavych Lithobiidech (Chilopoda) z Mohelenske stepi. [Interessante Lithobius - Arten aus der Mohelno-Steppe.] Ochrana p r irody, 11, 271 - 272. (in Czech, with Russian and German summaries)" type="journal article" year="1956">Dobroruka 1956</bibRefCitation>
: 271
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8B841B54FFC9FFEF34C4F96F50BE85B5" blockId="5.[151,701,1693,1751]" box="[151,701,1725,1751]" pageId="5" pageNumber="42">
<treatmentCitationGroup id="AB2B3C7AFFC9FFEF34C4F96F50BE85B5" box="[151,701,1725,1751]" pageId="5" pageNumber="42">
<taxonomicName id="4C3B60D7FFC9FFEF34C4F96F507C85B5" ID-CoL="7JXRX" authority="Dobroruka 1959: 105" authorityName="Dobroruka" authorityPageNumber="105" authorityYear="1959" box="[151,639,1727,1750]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="pusillus" subSpecies="johni">
<emphasis id="B94FC746FFC9FFEF34C4F96F539085B5" box="[151,403,1727,1750]" italics="true" pageId="5" pageNumber="42">Lithobius pusillus johni</emphasis>
<treatmentCitation id="0A9A3D45FFC9FFEF35CAF96F507C85B5" author="Dobroruka" box="[409,639,1727,1750]" page="105" pageId="5" pageNumber="42" year="1959">
<bibRefCitation id="EFAA66A5FFC9FFEF35CAF96F504E85B5" author="Dobroruka" box="[409,589,1727,1750]" pageId="5" pageNumber="45" refString="Dobroruka, L. J. (1959) Chilopoda statni prirodni reservace Mohelno. [Chilopoden des staatlichen Naturschutzgebeites Mohelno.] Ochrana p r irody, 14, 104 - 106. (in Czech, with German summary)" type="journal article" year="1959">Dobroruka 1959</bibRefCitation>
: 105
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106.
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<subSubSection id="C32148DFFFC9FFEF34C4F8D750C58411" pageId="5" pageNumber="42" type="materials_examined">
<paragraph id="8B841B54FFC9FFEF34C4F8D757928441" blockId="5.[151,1436,1799,2026]" box="[151,1425,1799,1826]" pageId="5" pageNumber="42">
<emphasis id="B94FC746FFC9FFEF34C4F8D7533E8442" bold="true" box="[151,317,1799,1825]" pageId="5" pageNumber="42">
<typeStatus id="5480A5F6FFC9FFEF34C4F8D752D78442" box="[151,212,1799,1825]" pageId="5" pageNumber="42">Type</typeStatus>
locality.
</emphasis>
Mohelenská hadcová step NNR (= Serpentine Steppe of Mohelno), Moravia,
<collectingCountry id="F32C5BC4FFC9FFEF309CF8D8578D8441" box="[1231,1422,1800,1826]" name="Czech Republic" pageId="5" pageNumber="42">Czech Republic</collectingCountry>
.
</paragraph>
<paragraph id="8B841B54FFC9FFEF3495F8FF56D68429" blockId="5.[151,1436,1799,2026]" box="[198,1237,1839,1866]" pageId="5" pageNumber="42">
<emphasis id="B94FC746FFC9FFEF3495F8FF5392842A" bold="true" box="[198,401,1839,1865]" pageId="5" pageNumber="42">
<typeStatus id="5480A5F6FFC9FFEF3495F8FF5300842A" box="[198,259,1839,1865]" pageId="5" pageNumber="42">Type</typeStatus>
specimens.
</emphasis>
The description of
<taxonomicName id="4C3B60D7FFC9FFEF3625F8E05137842A" box="[630,820,1840,1865]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="pusillus" subSpecies="johni">
<emphasis id="B94FC746FFC9FFEF3625F8E05137842A" box="[630,820,1840,1865]" italics="true" pageId="5" pageNumber="42">L. pusillus johni</emphasis>
</taxonomicName>
was based on 3 ɗ, coll. Dobroruka.
</paragraph>
<paragraph id="8B841B54FFC9FFEF3495F88750C58411" blockId="5.[151,1436,1799,2026]" box="[198,710,1879,1906]" pageId="5" pageNumber="42">
<emphasis id="B94FC746FFC9FFEF3495F88753B98412" bold="true" box="[198,442,1879,1905]" pageId="5" pageNumber="42">Material examined.</emphasis>
No material available.
</paragraph>
</subSubSection>
<subSubSection id="C32148DFFFC9FFEC3495F8AF50B3817A" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42" type="discussion">
<paragraph id="8B841B54FFC9FFEC3495F8AF504D82A9" blockId="5.[151,1436,1799,2026]" lastBlockId="6.[151,1436,152,538]" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42">
<emphasis id="B94FC746FFC9FFEF3495F8AF534384FA" bold="true" box="[198,320,1919,1945]" pageId="5" pageNumber="42">Remarks.</emphasis>
<taxonomicName id="4C3B60D7FFC9FFEF351AF85050BA84F9" authority="Latzel, 1880" authorityName="Latzel" authorityYear="1880" box="[329,697,1920,1946]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B94FC746FFC9FFEF351AF850501F84FA" box="[329,540,1920,1945]" italics="true" pageId="5" pageNumber="42">Lithobius pusillus</emphasis>
<bibRefCitation id="EFAA66A5FFC9FFEF3677F85050BA84F9" author="Latzel" box="[548,697,1920,1946]" pageId="5" pageNumber="46" refString="Latzel, R. (1880) Die Myriapoden der Osterreichisch-Ungarischen Monarchie. I.: Die Chilopoden. A. Holder, Wien, 228 pp." type="book" year="1880">Latzel, 1880</bibRefCitation>
</taxonomicName>
was synonymized with
<taxonomicName id="4C3B60D7FFC9FFEF378FF850572684F9" authority="Meinert, 1872" authorityName="Meinert" authorityYear="1872" box="[988,1317,1920,1946]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="lapidicola">
<emphasis id="B94FC746FFC9FFEF378FF850567784FA" box="[988,1140,1920,1945]" italics="true" pageId="5" pageNumber="42">L. lapidicola</emphasis>
<bibRefCitation id="EFAA66A5FFC9FFEF302EF850572684F9" author="Meinert" box="[1149,1317,1920,1946]" pageId="5" pageNumber="46" refString="Meinert, F. (1872) Myriapoda Musaei Hauniensis. Bidrag til Myriapodernes Morphologi og Systematik. II. Lithobiini. Naturhistorisk Tidskrift, 3, Raekke, 8, 281 - 344." type="journal article" year="1872">Meinert, 1872</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EFAA66A5FFC9FFEF3107F85052EA84A1" author="Eason" pageId="5" pageNumber="45" refString="Eason, E. H. (1982) A review of the north-west European species of Lithobiomorpha with a revised key to their identification. Zoological Journal of the Linnean Society, 74, 9 - 33." type="journal article" year="1982">Eason (1982)</bibRefCitation>
. Dobroruka characterized his taxon in two papers. At first he considered 2 ɗ as atypical specimens of
<taxonomicName id="4C3B60D7FFC9FFEF34C4F80053F18489" authority="Dobroruka 1956" authorityName="Dobroruka" authorityYear="1956" box="[151,498,2000,2026]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B94FC746FFC9FFEF34C4F8005311848A" box="[151,274,2000,2025]" italics="true" pageId="5" pageNumber="42">L. pusillus</emphasis>
(
<bibRefCitation id="EFAA66A5FFC9FFEF3572F80053EA8489" author="Dobroruka" box="[289,489,2000,2026]" pageId="5" pageNumber="45" refString="Dobroruka, L. J. (1956) O nekterych zajimavych Lithobiidech (Chilopoda) z Mohelenske stepi. [Interessante Lithobius - Arten aus der Mohelno-Steppe.] Ochrana p r irody, 11, 271 - 272. (in Czech, with Russian and German summaries)" type="journal article" year="1956">Dobroruka 1956</bibRefCitation>
)
</taxonomicName>
, but later, after a study of the third ɗ,
<bibRefCitation id="EFAA66A5FFC9FFEF37E2F80056858489" author="Dobroruka" box="[945,1158,2000,2026]" pageId="5" pageNumber="45" refString="Dobroruka, L. J. (1959) Chilopoda statni prirodni reservace Mohelno. [Chilopoden des staatlichen Naturschutzgebeites Mohelno.] Ochrana p r irody, 14, 104 - 106. (in Czech, with German summary)" type="journal article" year="1959">Dobroruka (1959)</bibRefCitation>
described them all as a new subspecies. The main distinguishing characters mentioned by Dobroruka were the spinulation of the 15th prefemur 1/3 (whole leg D: 0,0,1,0,0, V: 0,1,3,1,0, respectively) and the antennae with 25 (2 ɗ) or 30 articles (1 ɗ). The number of antennomeres in the original description of
<taxonomicName id="4C3B60D7FFCAFFEC37CBFF38562D8262" box="[920,1070,232,257]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="lapidicola">
<emphasis id="B94FC746FFCAFFEC37CBFF38562D8262" box="[920,1070,232,257]" italics="true" pageId="6" pageNumber="43">L. lapidicola</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFAA66A5FFCAFFEC3008FF38570E8261" author="Meinert" box="[1115,1293,232,258]" pageId="6" pageNumber="46" refString="Meinert, F. (1872) Myriapoda Musaei Hauniensis. Bidrag til Myriapodernes Morphologi og Systematik. II. Lithobiini. Naturhistorisk Tidskrift, 3, Raekke, 8, 281 - 344." type="journal article" year="1872">Meinert (1872)</bibRefCitation>
varies from 26 to 40, from 21 to 33, according to
<bibRefCitation id="EFAA66A5FFCAFFEC3607FEC050F58249" author="Latzel" box="[596,758,272,298]" pageId="6" pageNumber="46" refString="Latzel, R. (1880) Die Myriapoden der Osterreichisch-Ungarischen Monarchie. I.: Die Chilopoden. A. Holder, Wien, 228 pp." type="book" year="1880">Latzel (1880)</bibRefCitation>
. The spinulation of the last legs in the course of development grows from D: 0,0,0,0,0 to D: 1,0,2,0,0 (
<bibRefCitation id="EFAA66A5FFCAFFEC369AFEE8515E8231" author="Latzel" box="[713,861,312,338]" pageId="6" pageNumber="46" refString="Latzel, R. (1880) Die Myriapoden der Osterreichisch-Ungarischen Monarchie. I.: Die Chilopoden. A. Holder, Wien, 228 pp." type="book" year="1880">Latzel 1880</bibRefCitation>
). However, Dobroruka found the same 15th leg spinulation formula as he described in his new subspecies shared also with a specimen of
<taxonomicName id="4C3B60D7FFCAFFEC3097FEB05758821A" box="[1220,1371,352,377]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="lapidicola">
<emphasis id="B94FC746FFCAFFEC3097FEB05758821A" box="[1220,1371,352,377]" italics="true" pageId="6" pageNumber="43">L. lapidicola</emphasis>
</taxonomicName>
from the same locality (
<bibRefCitation id="EFAA66A5FFCAFFEC3523FE58503B82C1" author="Dobroruka" box="[368,568,392,418]" pageId="6" pageNumber="45" refString="Dobroruka, L. J. (1959) Chilopoda statni prirodni reservace Mohelno. [Chilopoden des staatlichen Naturschutzgebeites Mohelno.] Ochrana p r irody, 14, 104 - 106. (in Czech, with German summary)" type="journal article" year="1959">Dobroruka 1959</bibRefCitation>
). Anyway, the absence of prefemoral spines on the 15th legs seems to be insufficient to distinguish subspecies.
</paragraph>
<paragraph id="8B841B54FFCAFFEC3495FE0750B3817A" blockId="6.[151,1436,152,538]" pageId="6" pageNumber="43">
<emphasis id="B94FC746FFCAFFEC3495FE0753968292" bold="true" box="[198,405,471,497]" pageId="6" pageNumber="43">Taxonomic note.</emphasis>
We consider
<taxonomicName id="4C3B60D7FFCAFFEC366AFE0856208291" authority="Dobroruka, 1959" authorityName="Dobroruka" authorityYear="1959" box="[569,1059,472,498]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="pusillus" subSpecies="johni">
<emphasis id="B94FC746FFCAFFEC366AFE0851538292" box="[569,848,472,497]" italics="true" pageId="6" pageNumber="43">Lithobius pusillus johni</emphasis>
<bibRefCitation id="EFAA66A5FFCAFFEC370AFE0856208291" author="Dobroruka" box="[857,1059,472,498]" pageId="6" pageNumber="45" refString="Dobroruka, L. J. (1959) Chilopoda statni prirodni reservace Mohelno. [Chilopoden des staatlichen Naturschutzgebeites Mohelno.] Ochrana p r irody, 14, 104 - 106. (in Czech, with German summary)" type="journal article" year="1959">Dobroruka, 1959</bibRefCitation>
</taxonomicName>
as a junior subjective synonym of
<taxonomicName id="4C3B60D7FFCAFFEC34E5FDD050508179" authority="Meinert, 1872" authorityName="Meinert" authorityYear="1872" box="[182,595,512,538]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="lapidicola">
<emphasis id="B94FC746FFCAFFEC34E5FDD053A7817A" box="[182,420,512,537]" italics="true" pageId="6" pageNumber="43">Lithobius lapidicola</emphasis>
<bibRefCitation id="EFAA66A5FFCAFFEC35F8FDD050508179" author="Meinert" box="[427,595,512,538]" pageId="6" pageNumber="46" refString="Meinert, F. (1872) Myriapoda Musaei Hauniensis. Bidrag til Myriapodernes Morphologi og Systematik. II. Lithobiini. Naturhistorisk Tidskrift, 3, Raekke, 8, 281 - 344." type="journal article" year="1872">Meinert, 1872</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A27C7A3DFFCAFFEC360CFE2F50B3817A" box="[607,688,511,537]" pageId="6" pageNumber="43">
<emphasis id="B94FC746FFCAFFEC360CFE2F50B3817A" bold="true" box="[607,688,511,537]" pageId="6" pageNumber="43">syn. n.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>