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<document id="999F6DDAC027AC34DD484E91F439E7FB" ID-DOI="10.5281/zenodo.178383" ID-GBIF-Dataset="ff588456-e455-4555-bfe9-bb1f4cd80fbf" ID-ISSN="1175-5326" ID-Zenodo-Dep="178383" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459892246009" checkinUser="plazi" docAuthor="Anker, Arthur &amp; Ahyong, Shane T." docDate="2007" docId="03929D142A53761AFF3A821D2D4FFDF1" docLanguage="en" docName="zt01563p030.pdf" docOrigin="Zootaxa 1563" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Athanas ivoiriensis Anker &amp; Ahyong, 2007, n. sp." docType="treatment" docVersion="10" lastPageNumber="28" masterDocId="FFABE56C2A557611FFAD803A2E26FFEB" masterDocTitle="Description of two species in the alpheid shrimp genus Athanas Leach, 1814, with remarks on A. amazone Holthuis, 1951 (Decapoda, Caridea)" masterLastPageNumber="30" masterPageNumber="17" pageNumber="23" updateTime="1698228108777" updateUser="plazi">
<mods:mods id="FE3508C6031DF1F4E6678A355AC6CF36" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="96A3EA8A30EBF7515DCD77CF228B3D69">
<mods:title id="66672521135EEF3374E8146FB0CAD436">Description of two species in the alpheid shrimp genus Athanas Leach, 1814, with remarks on A. amazone Holthuis, 1951 (Decapoda, Caridea)</mods:title>
</mods:titleInfo>
<mods:name id="04DA6C08B483989FC70DA99E0ED0D0CA" type="personal">
<mods:role id="1A597634C2F2469F873E2D99B0D5AF1A">
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</mods:role>
<mods:namePart id="7DB6B2F2AC6E051D5D9AB7F7E8D08566">Anker, Arthur</mods:namePart>
</mods:name>
<mods:name id="DF06A89D8DA34EF117EC4BA0B86AECED" type="personal">
<mods:role id="B16AC775F4618746A715E7EF76BD1A0C">
<mods:roleTerm id="90E30F3722987F391263D871AB585FF1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="51BE03D28CE425CE0F117DDBAE56B3FC">Ahyong, Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B348F596B9A66561613E04EE4FE046D7">text</mods:typeOfResource>
<mods:relatedItem id="3E50F238C92724E61DA7575F99D511DD" type="host">
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<mods:title id="88DD2D5025EA676F582E413F8AFDFBEB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0C60243CE26B9B49ADD14FDE1009208A">
<mods:date id="E7DACFE16BF3A71ECED732C3314D44F4">2007</mods:date>
<mods:detail id="9566DE29AF7060EFA320C1B27AEDDFFA" type="volume">
<mods:number id="49E756BB916F327D84595B93EA684B7F">1563</mods:number>
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<mods:extent id="91BCC537E1395485169FBFF754048319" unit="page">
<mods:start id="67140DDFFF4F71D15CB2AD1F41B74C13">17</mods:start>
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<mods:classification id="6CF961F61F93B3E87FFDC887D43D15A2">journal article</mods:classification>
<mods:identifier id="B996599CCF6A6DD73C8EF224C884B72C" type="DOI">10.5281/zenodo.178383</mods:identifier>
<mods:identifier id="22A5AEFD598B03C65E5F7A0C0D95F119" type="GBIF-Dataset">ff588456-e455-4555-bfe9-bb1f4cd80fbf</mods:identifier>
<mods:identifier id="ADF2C7CD4BAF507B7D107864F62307A4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="215B6548E1100839C2012AFC7337CD17" type="Zenodo-Dep">178383</mods:identifier>
</mods:mods>
<treatment id="03929D142A53761AFF3A821D2D4FFDF1" ID-DOI="http://doi.org/10.5281/zenodo.5685976" ID-GBIF-Taxon="119372136" ID-Zenodo-Dep="5685976" LSID="urn:lsid:plazi:treatment:03929D142A53761AFF3A821D2D4FFDF1" httpUri="http://treatment.plazi.org/id/03929D142A53761AFF3A821D2D4FFDF1" lastPageId="11" lastPageNumber="28" pageId="6" pageNumber="23">
<subSubSection id="C3217F892A537617FF3A821D2F3EFD8F" pageId="6" pageNumber="23" type="nomenclature">
<paragraph id="8B842C022A537617FF3A821D2FEEFDAA" blockId="6.[151,456,551,612]" box="[151,456,551,577]" pageId="6" pageNumber="23">
<heading id="D0CC9B6E2A537617FF3A821D2FEEFDAA" bold="true" box="[151,456,551,577]" fontSize="11" level="1" pageId="6" pageNumber="23" reason="1">
<emphasis id="B94FF0102A537617FF3A821D2FEEFDAA" bold="true" box="[151,456,551,577]" pageId="6" pageNumber="23">
<taxonomicName id="4C3B57812A537617FF3A821D2F59FDAA" ID-CoL="85MC6" box="[151,383,551,577]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="ivoiriensis" status="sp. nov.">
<emphasis id="B94FF0102A537617FF3A821D2F59FDAA" bold="true" box="[151,383,551,577]" italics="true" pageId="6" pageNumber="23">Athanas ivoiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A537617FE2B821D2FEEFDAA" box="[390,456,551,577]" pageId="6" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B842C022A537617FF3A82702F3EFD8F" blockId="6.[151,456,551,612]" box="[151,280,586,612]" pageId="6" pageNumber="23">
(
<figureCitation id="130030872A537617FF3282702ED4FD8F" box="[159,242,586,612]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="6" pageNumber="23">Figs. 3</figureCitation>
, 4)
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A537617FF3A82A02DBBFBAF" pageId="6" pageNumber="23" type="materials_examined">
<paragraph id="8B842C022A537617FF3A82A02A1DFD37" blockId="6.[151,1437,666,2012]" pageId="6" pageNumber="23">
<emphasis id="B94FF0102A537617FF3A82A02F62FD5F" bold="true" box="[151,324,666,692]" pageId="6" pageNumber="23">
<typeStatus id="548092A02A537617FF3A82A02EF4FD5F" box="[151,210,666,692]" pageId="6" pageNumber="23">Type</typeStatus>
material
</emphasis>
:
<typeStatus id="548092A02A537617FEFB82A02FEEFD5F" box="[342,456,666,692]" pageId="6" pageNumber="23" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D3DE78B2A537617FE7982A02C01FD5F" box="[468,551,666,692]" pageId="6" pageNumber="23" type="male">1 male</specimenCount>
, CL 4.0 mm, TL
<quantity id="4CC381E72A537617FD5A82A02D42FD5F" box="[759,868,666,692]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="6" pageNumber="23" unit="mm" value="10.4">10.4 mm</quantity>
, MNHN-Na 11278,
<collectingCountry id="F32C6C922A537617FBF882A02AC9FD5F" box="[1109,1263,666,692]" name="Ivory Coast" pageId="6" pageNumber="23">Côte d'Ivoire</collectingCountry>
(
<collectingCountry id="F32C6C922A537617FB5282A02BA9FD5F" box="[1279,1423,666,692]" name="Ivory Coast" pageId="6" pageNumber="23">Ivory Coast</collectingCountry>
), Grand Bassam, depth
<quantity id="4CC381E72A537617FE3782F82FF1FD37" box="[410,471,706,732]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="23" unit="m" value="20.0">20 m</quantity>
, &quot;Benne Aberdeen&quot;, coll. P. LeLoeuff,
<date id="FF850AC22A537617FC0C82F82A11FD37" box="[929,1079,706,732]" pageId="6" pageNumber="23" value="1970-02-20">20 Feb 1970</date>
.
</paragraph>
<paragraph id="8B842C022A537617FF6B82D02DBBFBAF" blockId="6.[151,1437,666,2012]" pageId="6" pageNumber="23">
<emphasis id="B94FF0102A537617FF6B82D02FFBFCEF" bold="true" box="[198,477,746,772]" pageId="6" pageNumber="23">Comparative material</emphasis>
:
<taxonomicName id="4C3B57812A537617FE4082D02DB2FCEF" authority="Holthuis, 1951: 1" authorityName="Holthuis" authorityPageNumber="1" authorityYear="1951" box="[493,916,746,772]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A537617FE4082D02C98FCE8" box="[493,702,746,771]" italics="true" pageId="6" pageNumber="23">Athanas amazone</emphasis>
<bibRefCitation id="EFAA51F32A537617FD6B82D02D5CFCEF" author="Holthuis" box="[710,890,746,772]" pageId="6" pageNumber="29" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis, 1951</bibRefCitation>
:
<specimenCount id="9D3DE78B2A537617FC2B82D02DB2FCEF" box="[902,916,746,772]" pageId="6" pageNumber="23" type="male">1</specimenCount>
</taxonomicName>
male, CL
<quantity id="4CC381E72A537617FBBE82D02A57FCEF" box="[1043,1137,746,772]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="23" unit="mm" value="3.2">3.2 mm</quantity>
, TL ~
<quantity id="4CC381E72A537617FB6F82D02B2CFCEF" box="[1218,1290,746,772]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="23" unit="mm" value="9.0">9 mm</quantity>
, MNHN-Na 8806, off Pointe Noire,
<collectingCountry id="F32C6C922A537617FE0283282C26FCC7" box="[431,512,786,812]" name="Republic of the Congo" pageId="6" pageNumber="23">Congo</collectingCountry>
, Ombango Sta. 411, between
<geoCoordinate id="EE0F4AC52A537617FCF283282D8BFCC7" box="[863,941,786,812]" direction="south" orientation="latitude" pageId="6" pageNumber="23" precision="925" value="-4.8166666">4°49'S</geoCoordinate>
and
<geoCoordinate id="EE0F4AC52A537617FC4A83282A10FCC7" box="[999,1078,786,812]" direction="south" orientation="latitude" pageId="6" pageNumber="23" precision="925" value="-4.9666667">4°58'S</geoCoordinate>
, dredge,
<quantity id="4CC381E72A537617FB0F83282AC9FCC7" box="[1186,1263,786,812]" metricMagnitude="2" metricUnit="m" metricValue="1.04" pageId="6" pageNumber="23" unit="m" value="104.0">104 m</quantity>
,
<date id="FF850AC22A537617FB5183282BA7FCC7" box="[1276,1409,786,812]" pageId="6" pageNumber="23" value="1969-06-06">6 Jun 1969</date>
;
<specimenCount id="9D3DE78B2A537617FA2083282EF5FCBF" pageId="6" pageNumber="23" type="male">1 male</specimenCount>
, CL
<quantity id="4CC381E72A537617FEA383002F4BFCBF" box="[270,365,826,852]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="6" pageNumber="23" unit="mm" value="5.4">5.4 mm</quantity>
, TL
<quantity id="4CC381E72A537617FE0B83002FDBFCBF" box="[422,509,826,852]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="23" unit="mm" value="15.0">15 mm</quantity>
, MNHN-Na 13708, Banyuls-sur-Mer (
<collectionCode id="ED2AB4C72A537617FC6683002A28FCBF" box="[971,1038,826,852]" httpUri="http://grbio.org/cool/c14k-izbs" name="Berliner Staatisches Museum" pageId="6" pageNumber="23">BSM</collectionCode>
), southern
<collectingCountry id="F32C6C922A537617FB3C83002AC2FCBF" box="[1169,1252,826,852]" name="France" pageId="6" pageNumber="23">France</collectingCountry>
,
<geoCoordinate id="EE0F4AC52A537617FB5C83002B64FCBF" box="[1265,1346,826,852]" direction="east" orientation="longitude" pageId="6" pageNumber="23" precision="925" value="3.2166667">3°13'E</geoCoordinate>
,
<geoCoordinate id="EE0F4AC52A537617FAFD83002E89FC97" direction="north" orientation="latitude" pageId="6" pageNumber="23" precision="925" value="42.483334">42°29' N</geoCoordinate>
, dredge,
<quantity id="4CC381E72A537617FEB483582F70FC97" box="[281,342,866,892]" metricMagnitude="1" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="23" unit="m" value="85.0">85 m</quantity>
, sand-mud, coll. P. Noël,
<date id="FF850AC22A537617FD2F83582D34FC97" box="[642,786,866,892]" pageId="6" pageNumber="23" value="1976-05-06">6 May 1976</date>
, det. P. Noël,
<date id="FF850AC22A537617FC1E83582A74FC97" box="[947,1106,866,892]" pageId="6" pageNumber="23" value="1976-05-11">11 May 1976</date>
;
<specimenCount id="9D3DE78B2A537617FBF083582A89FC97" box="[1117,1199,866,892]" pageId="6" pageNumber="23" type="male">1 male</specimenCount>
, CL
<quantity id="4CC381E72A537617FB4583582B62FC97" box="[1256,1348,866,892]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="6" pageNumber="23" unit="mm" value="4.9">4.9 mm</quantity>
, TL ca.
<quantity id="4CC381E72A537617FF3A83B02ECAFC4F" box="[151,236,906,932]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="23" unit="mm" value="14.0">14 mm</quantity>
, MNHN-Na 13748, Baie de Banyuls off
<collectionCode id="ED2AB4C72A537617FD6783B02D2DFC4F" box="[714,779,906,932]" httpUri="http://grbio.org/cool/c14k-izbs" name="Berliner Staatisches Museum" pageId="6" pageNumber="23">BSM</collectionCode>
, southern
<collectingCountry id="F32C6C922A537617FC2F83B02DF3FC4F" box="[898,981,906,932]" name="France" pageId="6" pageNumber="23">France</collectingCountry>
, dredge,
<quantity id="4CC381E72A537617FB9383B02A5AFC4F" box="[1086,1148,906,932]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="23" unit="m" value="30.0">30 m</quantity>
, mud, coll. Labat, det. P. Noël,
<date id="FF850AC22A537617FF4F83882FACFC27" box="[226,394,946,972]" pageId="6" pageNumber="23" value="1976-05-19">19 May 1976</date>
;
<specimenCount id="9D3DE78B2A537617FE3783882C22FC27" box="[410,516,946,972]" pageId="6" pageNumber="23" type="female">1 female</specimenCount>
(CL and TL not measured), MNHN-Na 13752, off
<collectionCode id="ED2AB4C72A537617FB2983882AE3FC27" box="[1156,1221,946,972]" httpUri="http://grbio.org/cool/c14k-izbs" name="Berliner Staatisches Museum" pageId="6" pageNumber="23">BSM</collectionCode>
, southern
<collectingCountry id="F32C6C922A537617FAE883882BBEFC27" box="[1349,1432,946,972]" name="France" pageId="6" pageNumber="23">France</collectingCountry>
, dredge,
<quantity id="4CC381E72A537617FF5883E02F15FC1F" box="[245,307,986,1012]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="23" unit="m" value="35.0">35 m</quantity>
, coll. Labat, det. P. Noël,
<date id="FF850AC22A537617FDCC83E02D2EFC1F" box="[609,776,986,1012]" pageId="6" pageNumber="23" value="1977-05-26">26. May 1977</date>
; 1 ovigerous female (CL and TL not measured, carrying around
<specimenCount id="9D3DE78B2A537617FF5C84382F76FBF7" box="[241,336,1026,1052]" pageId="6" pageNumber="23" type="egg">20 eggs</specimenCount>
),
<specimenCount id="9D3DE78B2A537617FECA84382F9CFBF7" box="[359,442,1026,1052]" pageId="6" pageNumber="23" type="male">1 male</specimenCount>
, CL 4.0 mm, MNHN-Na 1745, dredge,
<quantity id="4CC381E72A537617FC3484382A20FBF7" box="[921,1030,1026,1052]" metricMagnitude="1" metricUnit="m" metricValue="5.75" metricValueMax="7.2" metricValueMin="4.3" pageId="6" pageNumber="23" unit="m" value="57.5" valueMax="72.0" valueMin="43.0">43–72 m</quantity>
, off Cap l'Abeille,
<collectionCode id="ED2AB4C72A537617FB4484382B0CFBF7" box="[1257,1322,1026,1052]" httpUri="http://grbio.org/cool/c14k-izbs" name="Berliner Staatisches Museum" pageId="6" pageNumber="23">BSM</collectionCode>
, southern
<collectingCountry id="F32C6C922A537617FF3A84102ECCFBAF" box="[151,234,1066,1092]" name="France" pageId="6" pageNumber="23">France</collectingCountry>
, coll. Guille, collection data unknown, det. J. Forest, 1967.
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A537619FF6B84682C68FAD3" lastPageId="8" lastPageNumber="25" pageId="6" pageNumber="23" type="description">
<paragraph id="8B842C022A537617FF6B84682B24FADF" blockId="6.[151,1437,666,2012]" pageId="6" pageNumber="23">
<emphasis id="B94FF0102A537617FF6B84682F71FB87" bold="true" box="[198,343,1106,1132]" pageId="6" pageNumber="23">Description</emphasis>
: Body elongate, slender. Carapace glabrous, not setose. Rostrum laterally compressed, straight, with acute tip, reaching slightly beyond mid-length of second segment of antennular peduncle (
<figureCitation id="130030872A537617FAC084402E83FB57" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="6" pageNumber="23">Fig. 3</figureCitation>
A), proximally broadened; rostral carina very feebly marked, posteriorly not reaching beyond rostral base. Extra-corneal spines acute, triangular, almost reaching anterior margin of cornea (
<figureCitation id="130030872A537617FBC484F02A96FB0F" box="[1129,1200,1226,1252]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="6" pageNumber="23">Fig. 3</figureCitation>
A, C); infra-corneal and supra-corneal spines absent. Eyes exposed in dorsal and lateral views; cornea well pigmented, anteromesial margin with small process (
<figureCitation id="130030872A537617FDA785202C76FADF" box="[522,592,1306,1332]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="6" pageNumber="23">Fig. 3</figureCitation>
C). Pterygostomial angle rounded, without spine (
<figureCitation id="130030872A537617FB3485202AF8FADF" box="[1177,1246,1306,1332]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="6" pageNumber="23">Fig. 3</figureCitation>
B).
</paragraph>
<paragraph id="8B842C022A537617FF6A85782F2FF9A7" blockId="6.[151,1437,666,2012]" pageId="6" pageNumber="23">
Antennular peduncle with second segment much shorter than dorsally visible portion of first segment, and shorter than third segment (
<figureCitation id="130030872A537617FE4E85502C0CFA6F" box="[483,554,1386,1412]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="6" pageNumber="23">Fig. 3</figureCitation>
A); stylocerite acute, reaching or almost reaching distal margin of second segment (
<figureCitation id="130030872A537617FEA485A82F69FA47" box="[265,335,1426,1452]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="6" pageNumber="23">Fig. 3</figureCitation>
A, B); ventromesial carina with well developed, acute spine; outer flagellum biramous, fused portion composed of four segments (
<figureCitation id="130030872A537617FDF285802C8CFA3F" box="[607,682,1466,1492]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="6" pageNumber="23">Fig. 3</figureCitation>
B). Antenna with basicerite bearing acute ventrolateral spine; scaphocerite exceeding antennular peduncle (
<figureCitation id="130030872A537617FD1E85D82CDFFA17" box="[691,761,1506,1532]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="6" pageNumber="23">Fig. 3</figureCitation>
A), broadly oval, lateral margin straight, anterior margin broadly convex, reaching slightly beyond distolateral spine; carpocerite slightly shorter than scaphocerite (
<figureCitation id="130030872A537617FF3286082EC3F9A7" box="[159,229,1586,1612]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="6" pageNumber="23">Fig. 3</figureCitation>
B).
</paragraph>
<paragraph id="8B842C022A537617FF6A86602ECCF907" blockId="6.[151,1437,666,2012]" pageId="6" pageNumber="23">Mouthparts without specific features in external view. Third maxilliped slender (Fig. 4J); ultimate segment about 1.3 times as long as penultimate segment and 0.6 times as long as antepenultimate segment; ultimate segment distally unarmed except for stiff setae; lateral plate of coxa as illustrated (Fig. 4K); arthrobranch absent.</paragraph>
<paragraph id="8B842C022A537616FF6A86C02FF8FEE9" blockId="6.[151,1437,666,2012]" lastBlockId="7.[151,1437,152,858]" lastPageId="7" lastPageNumber="24" pageId="6" pageNumber="23">Major cheliped slender (Fig. 4A) with basis bearing small exopod dorsally (Fig. 4D) and rounded process with row of curved spiniform setae ventromesially (Fig. 4E); ischium elongate, slightly depressed, slightly shorter than merus, ventromesially depressed; ventral margin irregularly toothed (Fig. 4A), dorsal margin and lateral margins proximally with slender spines (Fig. 4A, B); merus elongate, not expanded, ventromesially depressed, ventrolateral margin irregularly toothed (Fig. 4A); carpus about 3/4 length of merus, distally widening (Fig. 4B), ventral margin with two tubercles (Fig. 4A); chela subcylindrical, slightly compressed, ventral margin of palm with row of blunt tubercles (Fig. 4C), concave proximal to base of pollex; fingers shorter than palm, not gaping (Fig. 4C), tips barely crossing, cutting edges unarmed except for broad, blunt, proximal process on pollex (Fig. 4C).</paragraph>
<paragraph id="8B842C022A527616FF6A812A2CAEFE19" blockId="7.[151,1437,152,858]" pageId="7" pageNumber="24">Second pereiopod slender, relatively short (Fig. 4F); ischium shorter than merus; carpus five-segmented, first article as long as four others combined; ratio of carpal segments (from proximal to distal) approximately: 5: 1: 1: 1: 2.5; chela simple, longer than distal carpal article, fingers longer than palm. Third pereiopod slen- der, relatively short (Fig. 4G); ischium armed with one ventrolateral spinform seta; merus and carpus unarmed; propodus unarmed except for slender distal spiniform seta (Fig. 4H); dactylus about 0.7 length of propodus, simple, slender, slightly curved.</paragraph>
<paragraph id="8B842C022A527616FF6A823A2FCCFCD9" blockId="7.[151,1437,152,858]" pageId="7" pageNumber="24">
Pleura of first to fourth abdominal somites with rounded posteroventral angles; pleuron of fifth somite with angular posteroventral angle; sixth somite with articulated triangular flap posteroventrally (
<figureCitation id="130030872A527616FA8182122B53FDA9" box="[1324,1397,552,578]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="7" pageNumber="24">Fig. 3</figureCitation>
D). Second pleopod with appendix masculina exceeding appendix interna, tip with at least five spiniform setae (Fig.
<date id="FF850AC22A527616FF7582422ED2FD79" box="[216,244,632,658]" pageId="7" pageNumber="24">4I</date>
). Uropod with protopod ending distally in one larger acute and one smaller subacute lobe (
<figureCitation id="130030872A527616FA9C82422B51FD79" box="[1329,1399,632,658]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="7" pageNumber="24">Fig. 3</figureCitation>
E); diaeresis straight, lateral spiniform seta small (
<figureCitation id="130030872A527616FD6B829A2D28FD51" box="[710,782,672,698]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="7" pageNumber="24">Fig. 3</figureCitation>
E). Telson oval-rectangular, distally tapering, with two pairs of dorsal spiniform setae (
<figureCitation id="130030872A527616FDA482F22C6BFD09" box="[521,589,712,738]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="7" pageNumber="24">Fig. 3</figureCitation>
E); posterior margin medially convex, with two pairs of slender posterolateral spiniform setae, mesial almost three times as long as lateral (
<figureCitation id="130030872A527616FC6982CA2A2CFCE1" box="[964,1034,752,778]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="7" pageNumber="24">Fig. 3</figureCitation>
E). Gill formula typical for genus (see under previous species).
</paragraph>
<paragraph id="8B842C022A527616FF6A837A2DA4FCB1" blockId="7.[151,1437,152,858]" box="[199,898,832,858]" pageId="7" pageNumber="24">
Colour in life unknown. Size: CL 4.0 mm, TL ca.
<quantity id="4CC381E72A527616FCBF837A2D59FCB1" box="[786,895,832,858]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="7" pageNumber="24" unit="mm" value="10.4">10.4 mm</quantity>
.
</paragraph>
<caption id="DF447C8A2A527616FF3A87582D9FF855" httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="7" pageNumber="24" targetBox="[153,1416,902,1873]" targetPageId="7">
<paragraph id="8B842C022A527616FF3A87582D9FF855" blockId="7.[151,1436,1890,1982]" pageId="7" pageNumber="24">
<emphasis id="B94FF0102A527616FF3A87582F34F891" bold="true" box="[151,274,1890,1914]" pageId="7" pageNumber="24">FIGURE 3</emphasis>
.
<taxonomicName id="4C3B57812A527616FE8D87582FCBF892" box="[288,493,1890,1913]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="ivoiriensis" status="sp. nov.">
<emphasis id="B94FF0102A527616FE8D87582FCBF892" box="[288,493,1890,1913]" italics="true" pageId="7" pageNumber="24">Athanas ivoiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A527616FE5987582C09F892" box="[500,559,1890,1913]" pageId="7" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
, male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8B842C022A5D7619FF3A84A92C68FAD3" blockId="8.[151,1436,1171,1336]" pageId="8" pageNumber="25">
<emphasis id="B94FF0102A5D7619FF3A84A92F34FB40" bold="true" box="[151,274,1171,1195]" pageId="8" pageNumber="25">FIGURE 4</emphasis>
.
<taxonomicName id="4C3B57812A5D7619FE8D84AE2FCBFB40" box="[288,493,1172,1195]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="25" phylum="Arthropoda" rank="species" species="ivoiriensis" status="sp. nov.">
<emphasis id="B94FF0102A5D7619FE8D84AE2FCBFB40" box="[288,493,1172,1195]" italics="true" pageId="8" pageNumber="25">Athanas ivoiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A5D7619FE5984AE2C09FB40" box="[500,559,1172,1195]" pageId="8" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
, male,
<typeStatus id="548092A02A5D7619FDD184AE2CFAFB40" box="[636,732,1172,1195]" pageId="8" pageNumber="25" type="holotype">holotype</typeStatus>
, CL 4.0 mm, MNHN-Na 11278, Grand Bassam,
<collectingCountry id="F32C6C922A5D7619FB4484AE2B4CFB40" box="[1257,1386,1172,1195]" name="Ivory Coast" pageId="8" pageNumber="25">Ivory Coast</collectingCountry>
. A – left cheliped, dorsomesial view; B – same, lateral view; C – same, detail of carpus and chela; D – same, detail of coxa and basis, lateral view; E – same, ventromesial view; F – second pereiopod, lateral view; G – third pereiopod, lateral view; H – same, dactylus; I – endopod of second pleopod, mesial view; J – third maxilliped, lateral view; K – same, detail of lateral plate. Scale bars =
<quantity id="4CC381E72A5D7619FDA4851B2C6CFAD3" box="[521,586,1313,1336]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="25" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A5D7619FF6B855C2B70FA43" pageId="8" pageNumber="25" type="etymology">
<paragraph id="8B842C022A5D7619FF6B855C2B70FA43" blockId="8.[151,1437,1382,1968]" pageId="8" pageNumber="25">
<emphasis id="B94FF0102A5D7619FF6B855C2F6BFA6B" bold="true" box="[198,333,1382,1408]" pageId="8" pageNumber="25">Etymology</emphasis>
: The new species’ name is derived from the French name of the West African country of
<collectingCountry id="F32C6C922A5D7619FAF1855C2EFCFA43" name="Ivory Coast" pageId="8" pageNumber="25">Ivory Coast</collectingCountry>
(
<collectingCountry id="F32C6C922A5D7619FF4485B42C18FA43" box="[233,574,1422,1448]" name="Ivory Coast" pageId="8" pageNumber="25">République de Côte d’Ivoire</collectingCountry>
), where the
<typeStatus id="548092A02A5D7619FD6685B42CD8FA43" box="[715,766,1422,1448]" pageId="8" pageNumber="25">type</typeStatus>
locality of this species (Grand Bassam) is situated.
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A5D7619FF6B858C2AE4F9CB" pageId="8" pageNumber="25" type="materials_examined">
<paragraph id="8B842C022A5D7619FF6B858C2ECCFA13" blockId="8.[151,1437,1382,1968]" pageId="8" pageNumber="25">
<emphasis id="B94FF0102A5D7619FF6B858C2F4EFA3B" bold="true" box="[198,360,1462,1488]" pageId="8" pageNumber="25">
<typeStatus id="548092A02A5D7619FF6B858C2F25FA3B" box="[198,259,1462,1488]" pageId="8" pageNumber="25">Type</typeStatus>
locality
</emphasis>
: Grand Bassam, situated about
<quantity id="4CC381E72A5D7619FD44858C2D13FA3B" box="[745,821,1462,1488]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="25" unit="km" value="40.0">40 km</quantity>
east of Abidjan,
<collectingCountry id="F32C6C922A5D7619FBA5858C2A81FA3B" box="[1032,1191,1462,1488]" name="Ivory Coast" pageId="8" pageNumber="25">Côte d’Ivoire</collectingCountry>
(
<collectingCountry id="F32C6C922A5D7619FB17858C2B6AFA3B" box="[1210,1356,1462,1488]" name="Ivory Coast" pageId="8" pageNumber="25">Ivory Coast</collectingCountry>
), West Africa.
</paragraph>
<paragraph id="8B842C022A5D7619FF6B863C2AE4F9CB" blockId="8.[151,1437,1382,1968]" box="[198,1218,1542,1568]" pageId="8" pageNumber="25">
<emphasis id="B94FF0102A5D7619FF6B863C2F00F9CB" bold="true" box="[198,294,1542,1568]" pageId="8" pageNumber="25">Habitat</emphasis>
: The single specimen was collected from
<quantity id="4CC381E72A5D7619FCB9863C2D74F9CB" box="[788,850,1542,1568]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="25" unit="m" value="20.0">20 m</quantity>
, possibly on mud-sand bottom.
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A5D7619FF6B86142AE9F9A3" box="[198,1231,1582,1608]" pageId="8" pageNumber="25" type="distribution">
<paragraph id="8B842C022A5D7619FF6B86142AE9F9A3" blockId="8.[151,1437,1382,1968]" box="[198,1231,1582,1608]" pageId="8" pageNumber="25">
<emphasis id="B94FF0102A5D7619FF6B86142F79F9A3" bold="true" box="[198,351,1582,1608]" pageId="8" pageNumber="25">Distribution</emphasis>
: Presently known only from the
<typeStatus id="548092A02A5D7619FD4C86142D32F9A3" box="[737,788,1582,1608]" pageId="8" pageNumber="25">type</typeStatus>
locality off
<collectingCountry id="F32C6C922A5D7619FC0F86142A09F9A3" box="[930,1071,1582,1608]" name="Ivory Coast" pageId="8" pageNumber="25">Ivory Coast</collectingCountry>
, West Africa.
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A5D7619FF6B866C2CBEF85B" pageId="8" pageNumber="25" type="discussion">
<paragraph id="8B842C022A5D7619FF6B866C2CBEF85B" blockId="8.[151,1437,1382,1968]" pageId="8" pageNumber="25">
<emphasis id="B94FF0102A5D7619FF6B866C2F1EF99B" bold="true" box="[198,312,1622,1648]" pageId="8" pageNumber="25">Remarks</emphasis>
: With
<quantity id="4CC381E72A5D7619FE2A866C2FC9F99B" box="[391,495,1622,1648]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="8" pageNumber="25" unit="mm" value="10.4">10.4 mm</quantity>
TL the
<typeStatus id="548092A02A5D7619FDE7866C2C96F99B" box="[586,688,1622,1648]" pageId="8" pageNumber="25" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3B57812A5D7619FD7B866D2D56F99B" box="[726,880,1623,1648]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="25" phylum="Arthropoda" rank="species" species="ivoiriensis" status="sp. nov.">
<emphasis id="B94FF0102A5D7619FD7B866D2D56F99B" box="[726,880,1623,1648]" italics="true" pageId="8" pageNumber="25">A. ivoiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A5D7619FCDA866C2D90F99B" box="[887,950,1622,1648]" pageId="8" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
, is possibly a relatively young individual. Furthermore, it lacks the right cheliped, although the corresponding coxa is as robust as that of the detached left cheliped, indicating that the difference in size between the two chelipeds may not be so strong.
<taxonomicName id="4C3B57812A5D7619FF3A86F52F5FF903" box="[151,377,1743,1768]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="25" phylum="Arthropoda" rank="species" species="ivoiriensis" status="sp. nov.">
<emphasis id="B94FF0102A5D7619FF3A86F52F5FF903" box="[151,377,1743,1768]" italics="true" pageId="8" pageNumber="25">Athanas ivoiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A5D7619FE2F86F42FE5F903" box="[386,451,1742,1768]" pageId="8" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
, is closely related to
<taxonomicName id="4C3B57812A5D7619FD6D86F52D6CF903" box="[704,842,1743,1768]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="25" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5D7619FD6D86F52D6CF903" box="[704,842,1743,1768]" italics="true" pageId="8" pageNumber="25">A. amazone</emphasis>
</taxonomicName>
, but can be separated from
<taxonomicName id="4C3B57812A5D7619FB3886F52B39F903" box="[1173,1311,1743,1768]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="25" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5D7619FB3886F52B39F903" box="[1173,1311,1743,1768]" italics="true" pageId="8" pageNumber="25">A. amazone</emphasis>
</taxonomicName>
by (1) the absence of an infra-corneal spines or angles; (2) the much stouter antennular peduncles; (3) the distinctly longer stylocerite, reaching the distal margin of the second segment of the antennular peduncle; (4) the much shorter carpocerite, not reaching the distal margin of the scaphocerite blade; and (5) the different proportions and shape of the (possibly major) cheliped, which shows more similarities to the chelipeds of
<taxonomicName id="4C3B57812A5D7619FB4587552BBAF863" box="[1256,1436,1903,1928]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="25" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A5D7619FB4587552BBAF863" box="[1256,1436,1903,1928]" italics="true" pageId="8" pageNumber="25">A. phyllocheles</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3B57812A5D7619FF6587AD2F55F85B" box="[200,371,1943,1968]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="25" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A5D7619FF6587AD2F55F85B" box="[200,371,1943,1968]" italics="true" pageId="8" pageNumber="25">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A5D7619FED687AC2F9FF85B" box="[379,441,1942,1968]" pageId="8" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
(cf.
<figureCitation id="130030872A5D7619FE5D87AC2C16F85B" box="[496,560,1942,1968]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="8" pageNumber="25">Fig. 1</figureCitation>
D, E, 2).
</paragraph>
</subSubSection>
<caption id="DF447C8A2A5C7618FF3A866E2A19F8F2" httpUri="https://zenodo.org/record/178387/files/figure.png" pageId="9" pageNumber="26" targetBox="[169,1427,213,1585]" targetPageId="9">
<paragraph id="8B842C022A5C7618FF3A866E2A19F8F2" blockId="9.[151,1437,1620,1817]" pageId="9" pageNumber="26">
<emphasis id="B94FF0102A5C7618FF3A866E2F34F987" bold="true" box="[151,274,1620,1644]" pageId="9" pageNumber="26">FIGURE 5</emphasis>
.
<taxonomicName id="4C3B57812A5C7618FE8D866F2CA3F987" authority="Holthuis, 1951" authorityName="Holthuis" authorityYear="1951" box="[288,645,1621,1644]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5C7618FE8D866F2FFBF987" box="[288,477,1621,1644]" italics="true" pageId="9" pageNumber="26">Athanas amazone</emphasis>
Holthuis, 1951
</taxonomicName>
: male, CL 4.9 mm, MNHN-Na 13748, Banyuls-sur-Mer, southern France: A – habitus; B – major cheliped, chela and distal carpus, mesial view; male, CL 3.2 mm, MNHN-Na 8806, Pointe Noire, Congo: C – major cheliped, chela and distal carpus, ventral view; D – same, ventromesial view; E – same, dorsomesial view; F – same, basis and ischium, ventromesial view; G – same as B, fingers in slightly different angle (entire cheliped illustrated by Crosnier &amp; Forest 1973, fig. 48); ovigerous female, CL not measured, MNHN-Na 1745, Cap l'Abeille, Banyuls-sur-Mer, southern France, H - minor cheliped, lateral view. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="C3217F892A5C761AFF6A87772D4FFDF1" lastPageId="11" lastPageNumber="28" pageId="9" pageNumber="26" type="reference_group">
<paragraph id="8B842C022A5C761BFF6A87772DF6FD79" blockId="9.[151,1436,1869,2015]" lastBlockId="10.[151,1437,152,658]" lastPageId="10" lastPageNumber="27" pageId="9" pageNumber="26">
<taxonomicName id="4C3B57812A5C7618FF6A87772FB0F88D" ID-CoL="85MBL" box="[199,406,1869,1894]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5C7618FF6A87772FB0F88D" box="[199,406,1869,1894]" italics="true" pageId="9" pageNumber="26">Athanas amazone</emphasis>
</taxonomicName>
(
<figureCitation id="130030872A5C7618FE0B87772FCAF88C" box="[422,492,1869,1895]" captionStart="FIGURE 5" captionStartId="9.[151,255,1620,1644]" captionTargetBox="[169,1427,213,1585]" captionTargetId="figure@9.[151,1436,194,1597]" captionTargetPageId="9" captionText="FIGURE 5. Athanas amazone Holthuis, 1951: male, CL 4.9 mm, MNHN-Na 13748, Banyuls-sur-Mer, southern France: A – habitus; B – major cheliped, chela and distal carpus, mesial view; male, CL 3.2 mm, MNHN-Na 8806, Pointe Noire, Congo: C – major cheliped, chela and distal carpus, ventral view; D – same, ventromesial view; E – same, dorsomesial view; F – same, basis and ischium, ventromesial view; G – same as B, fingers in slightly different angle (entire cheliped illustrated by Crosnier &amp; Forest 1973, fig. 48); ovigerous female, CL not measured, MNHN-Na 1745, Cap l'Abeille, Banyuls-sur-Mer, southern France, H - minor cheliped, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178387/files/figure.png" pageId="9" pageNumber="26">Fig. 5</figureCitation>
) was reported from depths ranging between 40 and
<quantity id="4CC381E72A5C7618FBE387772ABDF88C" box="[1102,1179,1869,1895]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="9" pageNumber="26" unit="m" value="155.0">155 m</quantity>
, from
<collectingCountry id="F32C6C922A5C7618FB4B87772B18F88C" box="[1254,1342,1869,1895]" name="Nigeria" pageId="9" pageNumber="26">Nigeria</collectingCountry>
(
<bibRefCitation id="EFAA51F32A5C7618FAE387772ED9F864" author="Holthuis" pageId="9" pageNumber="29" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
),
<collectingCountry id="F32C6C922A5C7618FEB8874F2F72F864" box="[277,340,1909,1935]" name="Togo" pageId="9" pageNumber="26">Togo</collectingCountry>
,
<collectingCountry id="F32C6C922A5C7618FECD874F2F88F864" box="[352,430,1909,1935]" name="Republic of the Congo" pageId="9" pageNumber="26">Congo</collectingCountry>
(
<bibRefCitation id="EFAA51F32A5C7618FE13874F2CFDF864" author="Crosnier" box="[446,731,1909,1935]" pageId="9" pageNumber="29" refString="Crosnier, A. &amp; Forest, J. 1973. Les crevettes profondes de l'Atlantique Oriental Tropical. Faune Tropicale, 9. O. R. S. T. O. M, Paris. 409 pp." type="book" year="1973">Crosnier &amp; Forest 1973</bibRefCitation>
),
<collectingCountry id="F32C6C922A5C7618FD5C874F2D5FF864" box="[753,889,1909,1935]" name="Cape Verde" pageId="9" pageNumber="26">Cape Verde</collectingCountry>
(
<bibRefCitation id="EFAA51F32A5C7618FC27874F2A01F864" author="Turkay" box="[906,1063,1909,1935]" pageId="9" pageNumber="30" refString="Turkay, M. (1982) Marine Crustacea Decapoda von den Kapverdischen Inseln mit Bemerkungen zur Zoogeographie des Gebietes. Courier des Forschungs - Instituts Senckenberg, 52, 91 - 129." type="journal article" year="1982">Türkay 1982</bibRefCitation>
),
<collectingCountry id="F32C6C922A5C7618FB91874F2A80F864" box="[1084,1190,1909,1935]" name="Morocco" pageId="9" pageNumber="26">Morocco</collectingCountry>
(
<bibRefCitation id="EFAA51F32A5C7618FB1B874F2BA8F864" author="Garcia" box="[1206,1422,1909,1935]" pageId="9" pageNumber="29" refString="Garcia Raso, J. E. (1996) Crustacea Decapoda (excl. Sergestidae) from Ibero - Moroccan waters. Results of Balgim- 84 expedition. Bulletin of Marine Science, 58, 730 - 752." type="journal article" year="1996">Garcia Raso 1996</bibRefCitation>
), and throughout the Mediterranean Sea:
<collectingCountry id="F32C6C922A5C7618FDDD87A72C90F85C" box="[624,694,1949,1975]" name="Spain" pageId="9" pageNumber="26">Spain</collectingCountry>
,
<collectingCountry id="F32C6C922A5C7618FD6E87A72D30F85C" box="[707,790,1949,1975]" name="France" pageId="9" pageNumber="26">France</collectingCountry>
,
<collectingCountry id="F32C6C922A5C7618FC8E87A72D7CF85C" box="[803,858,1949,1975]" name="Italy" pageId="9" pageNumber="26">Italy</collectingCountry>
,
<collectingCountry id="F32C6C922A5C7618FCCA87A72D9BF85C" box="[871,957,1949,1975]" name="Greece" pageId="9" pageNumber="26">Greece</collectingCountry>
,
<collectingCountry id="F32C6C922A5C7618FC6787A72A2DF85C" box="[970,1035,1949,1975]" name="Israel" pageId="9" pageNumber="26">Israel</collectingCountry>
and
<collectingCountry id="F32C6C922A5C7618FBE587A72ABDF85C" box="[1096,1179,1949,1975]" name="Turkey" pageId="9" pageNumber="26">Turkey</collectingCountry>
(
<bibRefCitation id="EFAA51F32A5C7618FB0187A72EF3F834" author="Holthuis" lastPageId="9" lastPageNumber="30" pageId="9" pageNumber="29" refString="Holthuis, L. B. &amp; Gottlieb, E. (1958) An annotated list of the decapod Crustacea of the Mediterranean coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis &amp; Gottlieb 1958</bibRefCitation>
;
<bibRefCitation id="EFAA51F32A5C7618FF4E87FF2C27F834" author="Crosnier" box="[227,513,1989,2015]" pageId="9" pageNumber="29" refString="Crosnier, A. &amp; Forest, J. 1973. Les crevettes profondes de l'Atlantique Oriental Tropical. Faune Tropicale, 9. O. R. S. T. O. M, Paris. 409 pp." type="book" year="1973">Crosnier &amp; Forest 1973</bibRefCitation>
;
<bibRefCitation id="EFAA51F32A5C7618FDA287FF2C92F834" author="Kocatas" box="[527,692,1989,2015]" pageId="9" pageNumber="30" refString="Kocatas, A. (1981) Liste preliminaires et repartition des crustaces decapodes des eaux turques. Rapports de la Commission Internationale de la Mer Mediterranee, 27, 161 - 162." type="journal article" year="1981">Kocataş 1981</bibRefCitation>
;
<bibRefCitation id="EFAA51F32A5C7618FD6F87FF2DF6F834" author="Koukouras" box="[706,976,1989,2015]" pageId="9" pageNumber="30" refString="Koukouras, A., Dounas, C., Turkay, M. &amp; Voultsiadou-Koukoura, E. (1992) Decapod Crustacean fauna of the Aegean Sea: new information, checklist, affinities. Senckenbergiana maritima, 22, 217 - 244." type="journal article" year="1992">
Koukouras
<emphasis id="B94FF0102A5C7618FCE187FF2DAFF835" box="[844,905,1989,2014]" italics="true" pageId="9" pageNumber="26">et al.</emphasis>
1992
</bibRefCitation>
;
<bibRefCitation id="EFAA51F32A5C7618FC7387FF2ADAF834" author="Froglia" box="[990,1276,1989,2015]" pageId="9" pageNumber="29" refString="Froglia, C &amp; Argenti, L. (1993) Athanas amazone Holthuis, 1951, first record for the Italian seas (Crustacea, Decapoda, Alpheidae). Bios (Macedonia, Greece), Scientific Annals of the School of Biology, 1, 201 - 205." type="journal article" year="1993">Froglia &amp; Argenti 1993</bibRefCitation>
;
<bibRefCitation id="EFAA51F32A5C761BFAA787FF2EF3FF59" author="Garcia" lastPageId="10" lastPageNumber="27" pageId="9" pageNumber="29" refString="Garcia Raso, J. E. (1996) Crustacea Decapoda (excl. Sergestidae) from Ibero - Moroccan waters. Results of Balgim- 84 expedition. Bulletin of Marine Science, 58, 730 - 752." type="journal article" year="1996">García Raso 1996</bibRefCitation>
;
<bibRefCitation id="EFAA51F32A5F761BFF4C80A22FF3FF59" author="Atkinson" box="[225,469,152,178]" pageId="10" pageNumber="29" refString="Atkinson, R. J. A., Froglia, C., Arneri, E., &amp; Antolini, B. (1997) Observations on the burrows and burrowing behaviour of Squilla mantis (L.) (Crustacea: Stomatopoda). Pubblicazioni della Stazione Zoologica di Napoli: Marine Ecology, 18, 337 - 359." type="journal article" year="1997">
Atkinson
<emphasis id="B94FF0102A5F761BFEF980A22FA9FF5A" box="[340,399,152,177]" italics="true" pageId="10" pageNumber="27">et al.</emphasis>
1997
</bibRefCitation>
;
<bibRefCitation id="EFAA51F32A5F761BFE4C80A22D2AFF59" author="Froglia" box="[481,780,152,178]" pageId="10" pageNumber="29" refString="Froglia, C. &amp; Atkinson, R. J. A. (1998) Association between Athanas amazone (Decapoda: Alpheidae) and Squilla mantis (Stomatopoda: Squillidae). Journal of Crustacean Biology, 18, 529 - 532." type="journal article" year="1998">Froglia &amp; Atkinson 1998</bibRefCitation>
; see also d’Udekem
<bibRefCitation id="EFAA51F32A5F761BFBAD80A22ABBFF59" author="d'Acoz" box="[1024,1181,152,178]" pageId="10" pageNumber="30" refString="Udekem d'Acoz, C. d'. (1999) Inventaire et distribution des crustaces decapodes de l'Atlantique nord - oriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25 ° N. Institut d'Ecologie et de Gestion de la Biodiversite, S. P. N., Museum national d'Histoire naturelle, Paris. 383 pp." type="book" year="1999">d’Acoz 1999</bibRefCitation>
).
<bibRefCitation id="EFAA51F32A5F761BFB1C80A22EC3FF31" author="Holthuis" lastPageId="10" lastPageNumber="30" pageId="10" pageNumber="29" refString="Holthuis, L. B. &amp; Gottlieb, E. (1958) An annotated list of the decapod Crustacea of the Mediterranean coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis &amp; Gottlieb (1958)</bibRefCitation>
and
<bibRefCitation id="EFAA51F32A5F761BFEB380FA2C6EFF31" author="Crosnier" box="[286,584,192,218]" pageId="10" pageNumber="29" refString="Crosnier, A. &amp; Forest, J. 1973. Les crevettes profondes de l'Atlantique Oriental Tropical. Faune Tropicale, 9. O. R. S. T. O. M, Paris. 409 pp." type="book" year="1973">Crosnier &amp; Forest (1973)</bibRefCitation>
noted some variation in the development of the infra-corneal spines; the relative length of the stylocerite (compared to the distal margin of the first segment of the antennular peduncle); the length of the carpocerite (compared to the scaphocerite); and the shape of the chelipeds.
<bibRefCitation id="EFAA51F32A5F761BFA8A812A2F62FEB9" author="Banner" pageId="10" pageNumber="29" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner (1983)</bibRefCitation>
were &quot;not convinced that the specimens of
<taxonomicName id="4C3B57812A5F761BFCF481022DC2FEBA" ID-CoL="85MBL" box="[857,996,312,337]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5F761BFCF481022DC2FEBA" box="[857,996,312,337]" italics="true" pageId="10" pageNumber="27">A. amazone</emphasis>
</taxonomicName>
described and depicted [referring to
<bibRefCitation id="EFAA51F32A5F761BFF3A815A2F6FFE91" author="Holthuis" box="[151,329,352,378]" pageId="10" pageNumber="29" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis, 1951</bibRefCitation>
;
<bibRefCitation id="EFAA51F32A5F761BFEFB815A2CB6FE91" author="Holthuis" box="[342,656,352,378]" lastPageId="10" lastPageNumber="30" pageId="10" pageNumber="29" refString="Holthuis, L. B. &amp; Gottlieb, E. (1958) An annotated list of the decapod Crustacea of the Mediterranean coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis &amp; Gottlieb, 1958</bibRefCitation>
;
<bibRefCitation id="EFAA51F32A5F761BFD30815A2D99FE91" author="Crosnier" box="[669,959,352,378]" pageId="10" pageNumber="29" refString="Crosnier, A. &amp; Forest, J. 1973. Les crevettes profondes de l'Atlantique Oriental Tropical. Faune Tropicale, 9. O. R. S. T. O. M, Paris. 409 pp." type="book" year="1973">Crosnier &amp; Forest, 1973</bibRefCitation>
] are as mature as our holo- and allotype [of
<taxonomicName id="4C3B57812A5F761BFF6C81B22F55FE4A" ID-CoL="85MCS" box="[193,371,392,417]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A5F761BFF6C81B22F55FE4A" box="[193,371,392,417]" italics="true" pageId="10" pageNumber="27">A. phyllocheles</emphasis>
</taxonomicName>
], for even sexually mature males and females may not have yet developed the proportions and shape of ultimate condition of the chelipeds&quot;. Nevertheless, the
<typeStatus id="548092A02A5F761BFC69818A2A0CFE21" box="[964,1066,432,458]" pageId="10" pageNumber="27" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3B57812A5F761BFBF9818A2AF8FE22" ID-CoL="85MBL" box="[1108,1246,432,457]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5F761BFBF9818A2AF8FE22" box="[1108,1246,432,457]" italics="true" pageId="10" pageNumber="27">A. amazone</emphasis>
</taxonomicName>
is an ovigerous female, although its small size (TL
<quantity id="4CC381E72A5F761BFD9281E22CA0FE19" box="[575,646,472,498]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="27" unit="mm" value="7.0">7 mm</quantity>
) suggests that it must be a young female. The mostly fragmentary specimens reported from
<collectingCountry id="F32C6C922A5F761BFE6A823A2C2EFDF1" box="[455,520,512,538]" name="Israel" pageId="10" pageNumber="27">Israel</collectingCountry>
(
<bibRefCitation id="EFAA51F32A5F761BFDB4823A2D69FDF1" author="Holthuis" box="[537,847,512,538]" lastPageId="10" lastPageNumber="30" pageId="10" pageNumber="29" refString="Holthuis, L. B. &amp; Gottlieb, E. (1958) An annotated list of the decapod Crustacea of the Mediterranean coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis &amp; Gottlieb 1958</bibRefCitation>
) were
<quantity id="4CC381E72A5F761BFC0E823A2A33FDF1" box="[931,1045,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="10" pageNumber="27" unit="mm" value="7.0" valueMax="10.0" valueMin="4.0">4–10 mm</quantity>
in TL.
<bibRefCitation id="EFAA51F32A5F761BFBDF823A2BBBFDF1" author="Banner" box="[1138,1437,512,538]" pageId="10" pageNumber="29" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner (1983)</bibRefCitation>
noted: &quot;perhaps the Mediterranean specimens may be found to be a different species on the basis of their chelipeds, and the specimens described by Crosnier &amp; Forest different enough on the basis of their carpocerites that they may in the future be considered to be three separate species&quot;.
</paragraph>
<paragraph id="8B842C022A5F761BFF3A82FD2EDCFC3F" blockId="10.[151,1436,711,980]" pageId="10" pageNumber="27">
<emphasis id="B94FF0102A5F761BFF3A82FD2F2FFD34" bold="true" box="[151,265,711,735]" pageId="10" pageNumber="27">TABLE 2.</emphasis>
Variation among specimens of
<taxonomicName id="4C3B57812A5F761BFDF182F22CFEFD34" ID-CoL="85MBL" box="[604,728,712,735]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5F761BFDF182F22CFEFD34" box="[604,728,712,735]" italics="true" pageId="10" pageNumber="27">A. amazone</emphasis>
</taxonomicName>
(A–E), compared to
<taxonomicName id="4C3B57812A5F761BFC1682F22A6EFD34" ID-CoL="85MC6" box="[955,1096,712,735]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="ivoiriensis" status="sp. nov.">
<emphasis id="B94FF0102A5F761BFC1682F22A6EFD34" box="[955,1096,712,735]" italics="true" pageId="10" pageNumber="27">A. ivoiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A5F761BFBE282F22AAEFD34" box="[1103,1160,712,735]" pageId="10" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
(F).
<taxonomicName id="4C3B57812A5F761BFB1082F22B5CFD34" ID-CoL="85MBL" box="[1213,1402,712,735]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5F761BFB1082F22B5CFD34" box="[1213,1402,712,735]" italics="true" pageId="10" pageNumber="27">Athanas amazone</emphasis>
</taxonomicName>
: A – adult male (MNHN-Na 13748) from Banyuls-sur-Mer, southern
<collectingCountry id="F32C6C922A5F761BFCFA82D02DB9FCEA" box="[855,927,746,769]" name="France" pageId="10" pageNumber="27">France</collectingCountry>
(
<figureCitation id="130030872A5F761BFC0082D02DCDFCEA" box="[941,1003,746,769]" captionStart="FIGURE 5" captionStartId="9.[151,255,1620,1644]" captionTargetBox="[169,1427,213,1585]" captionTargetId="figure@9.[151,1436,194,1597]" captionTargetPageId="9" captionText="FIGURE 5. Athanas amazone Holthuis, 1951: male, CL 4.9 mm, MNHN-Na 13748, Banyuls-sur-Mer, southern France: A – habitus; B – major cheliped, chela and distal carpus, mesial view; male, CL 3.2 mm, MNHN-Na 8806, Pointe Noire, Congo: C – major cheliped, chela and distal carpus, ventral view; D – same, ventromesial view; E – same, dorsomesial view; F – same, basis and ischium, ventromesial view; G – same as B, fingers in slightly different angle (entire cheliped illustrated by Crosnier &amp; Forest 1973, fig. 48); ovigerous female, CL not measured, MNHN-Na 1745, Cap l'Abeille, Banyuls-sur-Mer, southern France, H - minor cheliped, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178387/files/figure.png" pageId="10" pageNumber="27">Fig. 5</figureCitation>
A, B); B – adult male (MNHN-Na 8806) from Pointe Noire,
<collectingCountry id="F32C6C922A5F761BFECA83372F88FCCF" box="[359,430,781,804]" name="Republic of the Congo" pageId="10" pageNumber="27">Congo</collectingCountry>
(cf.
<bibRefCitation id="EFAA51F32A5F761BFE4D83372CC6FCCF" author="Crosnier" box="[480,736,781,804]" pageId="10" pageNumber="29" refString="Crosnier, A. &amp; Forest, J. 1973. Les crevettes profondes de l'Atlantique Oriental Tropical. Faune Tropicale, 9. O. R. S. T. O. M, Paris. 409 pp." type="book" year="1973">Crosnier &amp; Forest 1973</bibRefCitation>
, fig. 48;
<figureCitation id="130030872A5F761BFC9283372D5BFCCF" box="[831,893,781,804]" captionStart="FIGURE 5" captionStartId="9.[151,255,1620,1644]" captionTargetBox="[169,1427,213,1585]" captionTargetId="figure@9.[151,1436,194,1597]" captionTargetPageId="9" captionText="FIGURE 5. Athanas amazone Holthuis, 1951: male, CL 4.9 mm, MNHN-Na 13748, Banyuls-sur-Mer, southern France: A – habitus; B – major cheliped, chela and distal carpus, mesial view; male, CL 3.2 mm, MNHN-Na 8806, Pointe Noire, Congo: C – major cheliped, chela and distal carpus, ventral view; D – same, ventromesial view; E – same, dorsomesial view; F – same, basis and ischium, ventromesial view; G – same as B, fingers in slightly different angle (entire cheliped illustrated by Crosnier &amp; Forest 1973, fig. 48); ovigerous female, CL not measured, MNHN-Na 1745, Cap l'Abeille, Banyuls-sur-Mer, southern France, H - minor cheliped, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178387/files/figure.png" pageId="10" pageNumber="27">Fig. 5</figureCitation>
C–H); C – possibly immature male from
<collectingCountry id="F32C6C922A5F761BFA9883372B56FCCF" box="[1333,1392,781,804]" name="Israel" pageId="10" pageNumber="27">Israel</collectingCountry>
(cf.
<bibRefCitation id="EFAA51F32A5F761BFF3A830A2F97FCAC" author="Holthuis" box="[151,433,816,839]" lastPageId="10" lastPageNumber="30" pageId="10" pageNumber="29" refString="Holthuis, L. B. &amp; Gottlieb, E. (1958) An annotated list of the decapod Crustacea of the Mediterranean coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis &amp; Gottlieb 1958</bibRefCitation>
, fig. 4); D – ovigerous
<typeStatus id="548092A02A5F761BFD19830A2D37FCAC" box="[692,785,816,839]" pageId="10" pageNumber="27" type="holotype">holotype</typeStatus>
female from
<collectingCountry id="F32C6C922A5F761BFC05830A2DDEFCAC" box="[936,1016,816,839]" name="Nigeria" pageId="10" pageNumber="27">Nigeria</collectingCountry>
(cf.
<bibRefCitation id="EFAA51F32A5F761BFB83830A2AECFCAC" author="Holthuis" box="[1070,1226,816,839]" pageId="10" pageNumber="29" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
, fig. 23); E – adult ovigerous female (MNHN-Na 1745) from Cap l'Abeille near Banyuls-sur-Mer, southern
<collectingCountry id="F32C6C922A5F761BFBE883682AA8FC82" box="[1093,1166,850,873]" name="France" pageId="10" pageNumber="27">France</collectingCountry>
(
<figureCitation id="130030872A5F761BFB3183682AFCFC82" box="[1180,1242,850,873]" captionStart="FIGURE 5" captionStartId="9.[151,255,1620,1644]" captionTargetBox="[169,1427,213,1585]" captionTargetId="figure@9.[151,1436,194,1597]" captionTargetPageId="9" captionText="FIGURE 5. Athanas amazone Holthuis, 1951: male, CL 4.9 mm, MNHN-Na 13748, Banyuls-sur-Mer, southern France: A – habitus; B – major cheliped, chela and distal carpus, mesial view; male, CL 3.2 mm, MNHN-Na 8806, Pointe Noire, Congo: C – major cheliped, chela and distal carpus, ventral view; D – same, ventromesial view; E – same, dorsomesial view; F – same, basis and ischium, ventromesial view; G – same as B, fingers in slightly different angle (entire cheliped illustrated by Crosnier &amp; Forest 1973, fig. 48); ovigerous female, CL not measured, MNHN-Na 1745, Cap l'Abeille, Banyuls-sur-Mer, southern France, H - minor cheliped, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178387/files/figure.png" pageId="10" pageNumber="27">Fig. 5</figureCitation>
H).
<taxonomicName id="4C3B57812A5F761BFAAE83682EF2FC67" ID-CoL="85MC6" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="ivoiriensis" status="sp. nov.">
<emphasis id="B94FF0102A5F761BFAAE83682EF2FC67" italics="true" pageId="10" pageNumber="27">Athanas ivoiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A5F761BFF71834F2F3DFC67" box="[220,283,885,908]" pageId="10" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
: F –
<typeStatus id="548092A02A5F761BFEF8834F2F94FC67" box="[341,434,885,908]" pageId="10" pageNumber="27" type="holotype">holotype</typeStatus>
male (MNHN-Na 11278) from
<collectingCountry id="F32C6C922A5F761BFCBE834F2DB4FC67" box="[787,914,885,908]" name="Ivory Coast" pageId="10" pageNumber="27">Ivory Coast</collectingCountry>
(
<figureCitation id="130030872A5F761BFC0E834F2DD6FC67" box="[931,1008,885,908]" captionStart="FIGURE 3" captionStartId="7.[151,255,1890,1914]" captionTargetBox="[153,1416,902,1873]" captionTargetId="figure@7.[151,1436,890,1884]" captionTargetPageId="7" captionText="FIGURE 3. Athanas ivoiriensis n. sp., male, holotype, CL 4.0 mm, MNHN-Na 11278, Grand Bassam, Ivory Coast. A – frontal region, dorsal view; B – same, lateral view; C – same, detail of rostrum and eyes; D – sixth abdominal segment and tail fan, right lateral view; E – tail fan, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178386/files/figure.png" pageId="10" pageNumber="27">Figs. 3</figureCitation>
, 4). Abbreviations: S1-2, S2-3 = limits between 1st and 2nd and 2nd and 3rd segments, respectively; Sc. = scaphocerite; Tb. = tubercle;&gt; = exceeding; &lt;= not reaching.
</paragraph>
<paragraph id="8B842C022A5F761AFF3283CE2DC9FE91" blockId="10.[159,1418,1012,1036]" lastBlockId="11.[151,1436,152,538]" lastPageId="11" lastPageNumber="28" pageId="10" pageNumber="27">
Species
<emphasis id="B94FF0102A5F761BFDB683CF2BACFBE7" box="[539,1418,1013,1036]" italics="true" pageId="10" pageNumber="27">
<taxonomicName id="4C3B57812A5F761BFDB683CF2CB1FBE7" ID-CoL="85MBL" box="[539,663,1013,1036]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="amazone">A. amazone</taxonomicName>
<taxonomicName id="4C3B57812A5F761BFB5083CF2BACFBE7" ID-CoL="85MC6" box="[1277,1418,1013,1036]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="ivoiriensis">A. ivoiriensis</taxonomicName>
</emphasis>
A series of
<taxonomicName id="4C3B57812A5E761AFEE180A22FF3FF5A" ID-CoL="85MBL" box="[332,469,152,177]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5E761AFEE180A22FF3FF5A" box="[332,469,152,177]" italics="true" pageId="11" pageNumber="28">A. amazone</emphasis>
</taxonomicName>
is deposited in the MNHN, Paris. The differences between the MNHN specimens, those reported by
<bibRefCitation id="EFAA51F32A5E761AFEC780FA2C00FF31" author="Holthuis" box="[362,550,192,218]" pageId="11" pageNumber="29" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis (1951)</bibRefCitation>
,
<bibRefCitation id="EFAA51F32A5E761AFD9E80FA2D55FF31" author="Holthuis" box="[563,883,192,218]" lastPageId="11" lastPageNumber="30" pageId="11" pageNumber="29" refString="Holthuis, L. B. &amp; Gottlieb, E. (1958) An annotated list of the decapod Crustacea of the Mediterranean coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis &amp; Gottlieb (1958)</bibRefCitation>
and
<bibRefCitation id="EFAA51F32A5E761AFC0180FA2AFEFF31" author="Crosnier" box="[940,1240,192,218]" pageId="11" pageNumber="29" refString="Crosnier, A. &amp; Forest, J. 1973. Les crevettes profondes de l'Atlantique Oriental Tropical. Faune Tropicale, 9. O. R. S. T. O. M, Paris. 409 pp." type="book" year="1973">Crosnier &amp; Forest (1973)</bibRefCitation>
, and
<taxonomicName id="4C3B57812A5E761AFABB80FA2E90FEEA" ID-CoL="85MC6" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="ivoiriensis" status="sp. nov.">
<emphasis id="B94FF0102A5E761AFABB80FA2E90FEEA" italics="true" pageId="11" pageNumber="28">A. ivoiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A5E761AFF1080D22ED8FEE9" box="[189,254,232,258]" pageId="11" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
, are summarized in Table 2. It shows that some features, e.g., the length of the carpocerite, the shape of the merus and the chela of the major cheliped, and the slenderness of the minor cheliped, may indeed be sexually dimorphic and/or age-dependent. However, more material with information on colour patterns and ecology is needed to conclude about the taxonomic status of
<taxonomicName id="4C3B57812A5E761AFCCD815A2DCEFE92" ID-CoL="85MBL" box="[864,1000,352,377]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5E761AFCCD815A2DCEFE92" box="[864,1000,352,377]" italics="true" pageId="11" pageNumber="28">A. amazone</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B842C022A5F761BFF3284272B52F874" pageId="10" pageNumber="27">
<table id="F93BDEA22A5F89EEFF3284272BB4F82A" box="[159,1426,1053,1985]" gridcols="7" gridrows="13" pageId="10" pageNumber="27">
<tr id="350B2E402A5F89EEFF3284272BB4FBDF" box="[159,1426,1053,1076]" gridrow="0" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF3284272C23FBDF" box="[159,517,1053,1076]" gridcol="0" gridrow="0" pageId="10" pageNumber="27">Features / Specimens</th>
<th id="76DA473C2A5F89EEFDB684272CA7FBDF" box="[539,641,1053,1076]" gridcol="1" gridrow="0" pageId="10" pageNumber="27">A (male)</th>
<th id="76DA473C2A5F89EEFD0184272D34FBDF" box="[684,786,1053,1076]" gridcol="2" gridrow="0" pageId="10" pageNumber="27">B (male)</th>
<th id="76DA473C2A5F89EEFC8384272DADFBDF" box="[814,907,1053,1076]" gridcol="3" gridrow="0" pageId="10" pageNumber="27">C (male)</th>
<th id="76DA473C2A5F89EEFC1D84272A0FFBDF" box="[944,1065,1053,1076]" gridcol="4" gridrow="0" pageId="10" pageNumber="27">D (female)</th>
<th id="76DA473C2A5F89EEFBE684272ACCFBDF" box="[1099,1258,1053,1076]" gridcol="5" gridrow="0" pageId="10" pageNumber="27">E (female)</th>
<th id="76DA473C2A5F89EEFB5084272BB4FBDF" box="[1277,1426,1053,1076]" gridcol="6" gridrow="0" pageId="10" pageNumber="27">F (male)</th>
</tr>
<tr id="350B2E402A5F89EEFF32847C2BB4FB6B" box="[159,1426,1094,1152]" gridrow="1" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF32847C2C23FB6B" box="[159,517,1094,1152]" gridcol="0" gridrow="1" pageId="10" pageNumber="27">1. Stylocerite in relation to anten- nular peduncle segments (S1-3)</th>
<td id="76DA473C2A5F89EEFDB6847C2CA7FB6B" box="[539,641,1094,1152]" gridcol="1" gridrow="1" pageId="10" pageNumber="27">slightly&gt; S1-2</td>
<td id="76DA473C2A5F89EEFD01847C2D34FB6B" box="[684,786,1094,1152]" gridcol="2" gridrow="1" pageId="10" pageNumber="27">slightly&gt; S1-2</td>
<td id="76DA473C2A5F89EEFC83847C2DADFB6B" box="[814,907,1094,1152]" gridcol="3" gridrow="1" pageId="10" pageNumber="27">&lt;S1-2</td>
<td id="76DA473C2A5F89EEFC1D847C2A0FFB6B" box="[944,1065,1094,1152]" gridcol="4" gridrow="1" pageId="10" pageNumber="27">distinctly&gt; S1-2</td>
<td id="76DA473C2A5F89EEFBE6847C2ACCFB6B" box="[1099,1258,1094,1152]" gridcol="5" gridrow="1" pageId="10" pageNumber="27">slightly&gt; S1-2</td>
<td id="76DA473C2A5F89EEFB50847C2BB4FB6B" box="[1277,1426,1094,1152]" gridcol="6" gridrow="1" pageId="10" pageNumber="27">reaching S2-3</td>
</tr>
<tr id="350B2E402A5F89EEFF3284A82BB4FB27" box="[159,1426,1170,1228]" gridrow="2" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF3284A82C23FB27" box="[159,517,1170,1228]" gridcol="0" gridrow="2" pageId="10" pageNumber="27">Second antennular segment, ratio length/width</th>
<td id="76DA473C2A5F89EEFDB684A82CA7FB27" box="[539,641,1170,1228]" gridcol="1" gridrow="2" pageId="10" pageNumber="27">~1.8</td>
<td id="76DA473C2A5F89EEFD0184A82D34FB27" box="[684,786,1170,1228]" gridcol="2" gridrow="2" pageId="10" pageNumber="27">~1.8</td>
<td id="76DA473C2A5F89EEFC8384A82DADFB27" box="[814,907,1170,1228]" gridcol="3" gridrow="2" pageId="10" pageNumber="27">~1.8</td>
<td id="76DA473C2A5F89EEFC1D84A82A0FFB27" box="[944,1065,1170,1228]" gridcol="4" gridrow="2" pageId="10" pageNumber="27">~1.4</td>
<td id="76DA473C2A5F89EEFBE684A82ACCFB27" box="[1099,1258,1170,1228]" gridcol="5" gridrow="2" pageId="10" pageNumber="27">? (not noted)</td>
<td id="76DA473C2A5F89EEFB5084A82BB4FB27" box="[1277,1426,1170,1228]" gridcol="6" gridrow="2" pageId="10" pageNumber="27">~0.7</td>
</tr>
<tr id="350B2E402A5F89EEFF3284E42BB4FAF3" box="[159,1426,1246,1304]" gridrow="3" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF3284E42C23FAF3" box="[159,517,1246,1304]" gridcol="0" gridrow="3" pageId="10" pageNumber="27">3. Antenna, carpocerite reaching beyond scaphocerite by</th>
<td id="76DA473C2A5F89EEFDB684E42CA7FAF3" box="[539,641,1246,1304]" gridcol="1" gridrow="3" pageId="10" pageNumber="27">full Sc. length</td>
<td id="76DA473C2A5F89EEFD0184E42D34FAF3" box="[684,786,1246,1304]" gridcol="2" gridrow="3" pageId="10" pageNumber="27">full Sc. length</td>
<td id="76DA473C2A5F89EEFC8384E42DADFAF3" box="[814,907,1246,1304]" gridcol="3" gridrow="3" pageId="10" pageNumber="27">4/5 Sc. length</td>
<td id="76DA473C2A5F89EEFC1D84E42A0FFAF3" box="[944,1065,1246,1304]" gridcol="4" gridrow="3" pageId="10" pageNumber="27">1/5 Sc. length</td>
<td id="76DA473C2A5F89EEFBE684E42ACCFAF3" box="[1099,1258,1246,1304]" gridcol="5" gridrow="3" pageId="10" pageNumber="27">2/5 Sc. length</td>
<td id="76DA473C2A5F89EEFB5084E42BB4FAF3" box="[1277,1426,1246,1304]" gridcol="6" gridrow="3" pageId="10" pageNumber="27">not reaching beyond Sc.</td>
</tr>
<tr id="350B2E402A5F89EEFF3285102BB4FA88" box="[159,1426,1322,1379]" gridrow="4" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF3285102C23FA88" box="[159,517,1322,1379]" gridcol="0" gridrow="4" pageId="10" pageNumber="27">4. Antenna, carpocerite with/ without small tubercles</th>
<td id="76DA473C2A5F89EEFDB685102CA7FA88" box="[539,641,1322,1379]" gridcol="1" gridrow="4" pageId="10" pageNumber="27">with</td>
<td id="76DA473C2A5F89EEFD0185102D34FA88" box="[684,786,1322,1379]" gridcol="2" gridrow="4" pageId="10" pageNumber="27">with</td>
<td id="76DA473C2A5F89EEFC8385102DADFA88" box="[814,907,1322,1379]" gridcol="3" gridrow="4" pageId="10" pageNumber="27">without</td>
<td id="76DA473C2A5F89EEFC1D85102A0FFA88" box="[944,1065,1322,1379]" gridcol="4" gridrow="4" pageId="10" pageNumber="27">without</td>
<td id="76DA473C2A5F89EEFBE685102ACCFA88" box="[1099,1258,1322,1379]" gridcol="5" gridrow="4" pageId="10" pageNumber="27">without</td>
<td id="76DA473C2A5F89EEFB5085102BB4FA88" box="[1277,1426,1322,1379]" gridcol="6" gridrow="4" pageId="10" pageNumber="27">without</td>
</tr>
<tr id="350B2E402A5F89EEFF32854F2BB4FA39" box="[159,1426,1397,1490]" gridrow="5" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF32854F2C23FA39" box="[159,517,1397,1490]" gridcol="0" gridrow="5" pageId="10" pageNumber="27">5. Anterolateral margin of cara- pace with/without conspicuous concavity</th>
<td id="76DA473C2A5F89EEFDB6854F2CA7FA39" box="[539,641,1397,1490]" gridcol="1" gridrow="5" pageId="10" pageNumber="27">with</td>
<td id="76DA473C2A5F89EEFD01854F2D34FA39" box="[684,786,1397,1490]" gridcol="2" gridrow="5" pageId="10" pageNumber="27">with</td>
<td id="76DA473C2A5F89EEFC83854F2DADFA39" box="[814,907,1397,1490]" gridcol="3" gridrow="5" pageId="10" pageNumber="27">without</td>
<td id="76DA473C2A5F89EEFC1D854F2A0FFA39" box="[944,1065,1397,1490]" gridcol="4" gridrow="5" pageId="10" pageNumber="27">without</td>
<td id="76DA473C2A5F89EEFBE6854F2ACCFA39" box="[1099,1258,1397,1490]" gridcol="5" gridrow="5" pageId="10" pageNumber="27">without</td>
<td id="76DA473C2A5F89EEFB50854F2BB4FA39" box="[1277,1426,1397,1490]" gridcol="6" gridrow="5" pageId="10" pageNumber="27">without</td>
</tr>
<tr id="350B2E402A5F89EEFF3285DE2BB4FA10" box="[159,1426,1508,1531]" gridrow="6" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF3285DE2C23FA10" box="[159,517,1508,1531]" gridcol="0" gridrow="6" pageId="10" pageNumber="27">6. Infra-corneal angle/spine</th>
<td id="76DA473C2A5F89EEFDB685DE2CA7FA10" box="[539,641,1508,1531]" gridcol="1" gridrow="6" pageId="10" pageNumber="27">acute</td>
<td id="76DA473C2A5F89EEFD0185DE2D34FA10" box="[684,786,1508,1531]" gridcol="2" gridrow="6" pageId="10" pageNumber="27">rounded</td>
<td id="76DA473C2A5F89EEFC8385DE2DADFA10" box="[814,907,1508,1531]" gridcol="3" gridrow="6" pageId="10" pageNumber="27">acute</td>
<td id="76DA473C2A5F89EEFC1D85DE2A0FFA10" box="[944,1065,1508,1531]" gridcol="4" gridrow="6" pageId="10" pageNumber="27">acute</td>
<td id="76DA473C2A5F89EEFBE685DE2ACCFA10" box="[1099,1258,1508,1531]" gridcol="5" gridrow="6" pageId="10" pageNumber="27">? (not noted)</td>
<td id="76DA473C2A5F89EEFB5085DE2BB4FA10" box="[1277,1426,1508,1531]" gridcol="6" gridrow="6" pageId="10" pageNumber="27">absent</td>
</tr>
<tr id="350B2E402A5F89EEFF3286372BB4F9AC" box="[159,1426,1549,1607]" gridrow="7" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF3286372C23F9AC" box="[159,517,1549,1607]" gridcol="0" gridrow="7" pageId="10" pageNumber="27">7. Major cheliped, ratio carpus/ merus</th>
<td id="76DA473C2A5F89EEFDB686372CA7F9AC" box="[539,641,1549,1607]" gridcol="1" gridrow="7" pageId="10" pageNumber="27">~0.7</td>
<td id="76DA473C2A5F89EEFD0186372D34F9AC" box="[684,786,1549,1607]" gridcol="2" gridrow="7" pageId="10" pageNumber="27">~0.8</td>
<td id="76DA473C2A5F89EEFC8386372DADF9AC" box="[814,907,1549,1607]" gridcol="3" gridrow="7" pageId="10" pageNumber="27">? (not noted)</td>
<td id="76DA473C2A5F89EEFC1D86372A0FF9AC" box="[944,1065,1549,1607]" gridcol="4" gridrow="7" pageId="10" pageNumber="27">~0.9</td>
<td id="76DA473C2A5F89EEFBE686372ACCF9AC" box="[1099,1258,1549,1607]" gridcol="5" gridrow="7" pageId="10" pageNumber="27">? (not noted)</td>
<td id="76DA473C2A5F89EEFB5086372BB4F9AC" box="[1277,1426,1549,1607]" gridcol="6" gridrow="7" pageId="10" pageNumber="27">~0.6</td>
</tr>
<tr id="350B2E402A5F89EEFF3286632BB4F979" box="[159,1426,1625,1682]" gridrow="8" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF3286632C23F979" box="[159,517,1625,1682]" gridcol="0" gridrow="8" pageId="10" pageNumber="27">8. Major cheliped, merus</th>
<td id="76DA473C2A5F89EEFDB686632CA7F979" box="[539,641,1625,1682]" gridcol="1" gridrow="8" pageId="10" pageNumber="27">expanded</td>
<td id="76DA473C2A5F89EEFD0186632D34F979" box="[684,786,1625,1682]" gridcol="2" gridrow="8" pageId="10" pageNumber="27">expanded</td>
<td id="76DA473C2A5F89EEFC8386632DADF979" box="[814,907,1625,1682]" gridcol="3" gridrow="8" pageId="10" pageNumber="27">? (not noted)</td>
<td id="76DA473C2A5F89EEFC1D86632A0FF979" box="[944,1065,1625,1682]" gridcol="4" gridrow="8" pageId="10" pageNumber="27">slender</td>
<td id="76DA473C2A5F89EEFBE686632ACCF979" box="[1099,1258,1625,1682]" gridcol="5" gridrow="8" pageId="10" pageNumber="27">slender</td>
<td id="76DA473C2A5F89EEFB5086632BB4F979" box="[1277,1426,1625,1682]" gridcol="6" gridrow="8" pageId="10" pageNumber="27">slender</td>
</tr>
<tr id="350B2E402A5F89EEFF32869E2BB4F935" box="[159,1426,1700,1758]" gridrow="9" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF32869E2C23F935" box="[159,517,1700,1758]" gridcol="0" gridrow="9" pageId="10" pageNumber="27">9. Major chela, ventral margin</th>
<td id="76DA473C2A5F89EEFDB6869E2CA7F935" box="[539,641,1700,1758]" gridcol="1" gridrow="9" pageId="10" pageNumber="27">straight</td>
<td id="76DA473C2A5F89EEFD01869E2D34F935" box="[684,786,1700,1758]" gridcol="2" gridrow="9" pageId="10" pageNumber="27">straight</td>
<td id="76DA473C2A5F89EEFC83869E2DADF935" box="[814,907,1700,1758]" gridcol="3" gridrow="9" pageId="10" pageNumber="27">? (not noted)</td>
<td id="76DA473C2A5F89EEFC1D869E2A0FF935" box="[944,1065,1700,1758]" gridcol="4" gridrow="9" pageId="10" pageNumber="27">concave</td>
<td id="76DA473C2A5F89EEFBE6869E2ACCF935" box="[1099,1258,1700,1758]" gridcol="5" gridrow="9" pageId="10" pageNumber="27">? (not noted)</td>
<td id="76DA473C2A5F89EEFB50869E2BB4F935" box="[1277,1426,1700,1758]" gridcol="6" gridrow="9" pageId="10" pageNumber="27">concave, Tb. row</td>
</tr>
<tr id="350B2E402A5F89EEFF3286CA2BB4F8C1" box="[159,1426,1776,1834]" gridrow="10" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF3286CA2C23F8C1" box="[159,517,1776,1834]" gridcol="0" gridrow="10" pageId="10" pageNumber="27">10. Major chela, ratio palm/fin- gers</th>
<td id="76DA473C2A5F89EEFDB686CA2CA7F8C1" box="[539,641,1776,1834]" gridcol="1" gridrow="10" pageId="10" pageNumber="27">~2.8</td>
<td id="76DA473C2A5F89EEFD0186CA2D34F8C1" box="[684,786,1776,1834]" gridcol="2" gridrow="10" pageId="10" pageNumber="27">~2.9</td>
<td id="76DA473C2A5F89EEFC8386CA2DADF8C1" box="[814,907,1776,1834]" gridcol="3" gridrow="10" pageId="10" pageNumber="27">? (not noted)</td>
<td id="76DA473C2A5F89EEFC1D86CA2A0FF8C1" box="[944,1065,1776,1834]" gridcol="4" gridrow="10" pageId="10" pageNumber="27">~1.6</td>
<td id="76DA473C2A5F89EEFBE686CA2ACCF8C1" box="[1099,1258,1776,1834]" gridcol="5" gridrow="10" pageId="10" pageNumber="27">? (not noted)</td>
<td id="76DA473C2A5F89EEFB5086CA2BB4F8C1" box="[1277,1426,1776,1834]" gridcol="6" gridrow="10" pageId="10" pageNumber="27">~1.6</td>
</tr>
<tr id="350B2E402A5F89EEFF3287062BB4F89E" box="[159,1426,1852,1909]" gridrow="11" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF3287062C23F89E" box="[159,517,1852,1909]" gridcol="0" gridrow="11" pageId="10" pageNumber="27">11. Minor cheliped, teeth on ven- tral margin of carpus</th>
<td id="76DA473C2A5F89EEFDB687062CA7F89E" box="[539,641,1852,1909]" gridcol="1" gridrow="11" pageId="10" pageNumber="27">absent</td>
<td id="76DA473C2A5F89EEFD0187062D34F89E" box="[684,786,1852,1909]" gridcol="2" gridrow="11" pageId="10" pageNumber="27">absent</td>
<td id="76DA473C2A5F89EEFC8387062DADF89E" box="[814,907,1852,1909]" gridcol="3" gridrow="11" pageId="10" pageNumber="27">absent</td>
<td id="76DA473C2A5F89EEFC1D87062A0FF89E" box="[944,1065,1852,1909]" gridcol="4" gridrow="11" pageId="10" pageNumber="27">present (&gt;15)</td>
<td id="76DA473C2A5F89EEFBE687062ACCF89E" box="[1099,1258,1852,1909]" gridcol="5" gridrow="11" pageId="10" pageNumber="27">absent</td>
<td id="76DA473C2A5F89EEFB5087062BB4F89E" box="[1277,1426,1852,1909]" gridcol="6" gridrow="11" pageId="10" pageNumber="27">? (missing)</td>
</tr>
<tr id="350B2E402A5F89EEFF3287B22BB4F82A" box="[159,1426,1928,1985]" gridrow="12" pageId="10" pageNumber="27">
<th id="76DA473C2A5F89EEFF3287B22C23F82A" box="[159,517,1928,1985]" gridcol="0" gridrow="12" pageId="10" pageNumber="27">12. Minor cheliped, ratio carpus length/width</th>
<td id="76DA473C2A5F89EEFDB687B22CA7F82A" box="[539,641,1928,1985]" gridcol="1" gridrow="12" pageId="10" pageNumber="27">18</td>
<td id="76DA473C2A5F89EEFD0187B22D34F82A" box="[684,786,1928,1985]" gridcol="2" gridrow="12" pageId="10" pageNumber="27">18</td>
<td id="76DA473C2A5F89EEFC8387B22DADF82A" box="[814,907,1928,1985]" gridcol="3" gridrow="12" pageId="10" pageNumber="27">20</td>
<td id="76DA473C2A5F89EEFC1D87B22A0FF82A" box="[944,1065,1928,1985]" gridcol="4" gridrow="12" pageId="10" pageNumber="27">14</td>
<td id="76DA473C2A5F89EEFBE687B22ACCF82A" box="[1099,1258,1928,1985]" gridcol="5" gridrow="12" pageId="10" pageNumber="27">17</td>
<td id="76DA473C2A5F89EEFB5087B22BB4F82A" box="[1277,1426,1928,1985]" gridcol="6" gridrow="12" pageId="10" pageNumber="27">? (missing)</td>
</tr>
</table>
</paragraph>
<paragraph id="8B842C022A5E761AFF6A81B22D4FFDF1" blockId="11.[151,1436,152,538]" pageId="11" pageNumber="28">
Interestingly, at least in
<taxonomicName id="4C3B57812A5E761AFE4981B22C48FE4A" ID-CoL="85MBL" box="[484,622,392,417]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A5E761AFE4981B22C48FE4A" box="[484,622,392,417]" italics="true" pageId="11" pageNumber="28">A. amazone</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3B57812A5E761AFD0481B22D63FE4A" ID-CoL="85MC6" box="[681,837,392,417]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="ivoiriensis" status="sp. nov.">
<emphasis id="B94FF0102A5E761AFD0481B22D63FE4A" box="[681,837,392,417]" italics="true" pageId="11" pageNumber="28">A. ivoiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A5E761AFCE381B22DA9FE49" box="[846,911,392,418]" pageId="11" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
, the ventromesial portion of the basis of the major and minor chelipeds bears a peculiar process furnished with a comb-like row of slender, curved spiniform setae (Fig. 4E, 5F). The process on the right cheliped is thus juxtaposed to that on the left cheliped. The function of these structures, if there is any, remains unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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