259 lines
32 KiB
XML
259 lines
32 KiB
XML
<document id="AAE9D249FF5E68204A89637D86AAEC43" ID-DOI="10.11646/zootaxa.5336.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8272417" ID-ZooBank="7BD6357E-BB67-44E9-B028-4CA7A2769660" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1692697841468" checkinUser="plazi" docAuthor="Uiblein, Franz & Nielsen, Jørgen G." docDate="2023" docId="039287FEFC33FF9476BAEDF7FD5F97E4" docLanguage="en" docName="Zootaxa.5336.2.2.pdf" docOrigin="Zootaxa 5336 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5336.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neobythites monocellatus Nielsen 1999" docType="treatment" docVersion="2" lastPageNumber="202" masterDocId="FFABFF86FC25FF83762DEE25FFB09649" masterDocTitle="Five new ocellus-bearing species of the cusk-eel genus Neobythites (Ophidiidae, Ophidiiformes) from the West Pacific, with establishment of three new species groups" masterLastPageNumber="205" masterPageNumber="179" pageNumber="201" updateTime="1692698371970" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DD6153BAE280DBB669D632CB96ABFB8F">Five new ocellus-bearing species of the cusk-eel genus Neobythites (Ophidiidae, Ophidiiformes) from the West Pacific, with establishment of three new species groups</mods:title>
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<mods:namePart id="4DC89AF34E8F083A2AAF182C0DE4B0D2">Uiblein, Franz</mods:namePart>
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<mods:affiliation id="B109DC52C4DE8D2C53F0E0DDF0375B00">Institute of Marine Research, P. O. Box 1870 Nordnes, N- 5817 Bergen, Norway. Corresponding author; & National Research Foundation - South African Institute for Aquatic Biodiversity, Makhanda / Grahamstown, South Africa & Vietnam National Museum of Nature, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="5B2F62F4797B17FA2C83E6CCA60E00B5">Nielsen, Jørgen G.</mods:namePart>
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<mods:affiliation id="F618D9270F11C440D18B6EFF41017076">Natural History Museum of Denmark, University of Copenhagen, Denmark</mods:affiliation>
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<mods:typeOfResource id="521AA446190C855EB13BAF30D8BB32A6">text</mods:typeOfResource>
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<mods:title id="9C0D0B107468382E9D5E67E34A098798">Zootaxa</mods:title>
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<mods:part id="714AC3E3AC78798CA9EF6D07109C7705">
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<mods:date id="CFA0AFC5056DE4F3E79905A6484B8A3D">2023</mods:date>
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<mods:number id="1E1F4F93B6ADD067A72AC71AF18A10AE">2023-08-22</mods:number>
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<subSubSection id="C3216563FC33FF9576BAEDF7FDCA95A5" box="[151,634,978,1005]" pageId="22" pageNumber="201" type="nomenclature">
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<paragraph id="8B8436E8FC33FF9576BAEDF7FDCA95A5" blockId="22.[151,1427,978,1074]" box="[151,634,978,1005]" pageId="22" pageNumber="201">
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<heading id="D0CC8184FC33FF9576BAEDF7FDCA95A5" bold="true" box="[151,634,978,1005]" fontSize="11" level="1" pageId="22" pageNumber="201" reason="1">
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<taxonomicName id="4C3B4D6BFC33FF9576BAEDF7FDCA95A5" ID-CoL="46C3W" authority="Nielsen, 1999" authorityName="Nielsen" authorityYear="1999" box="[151,634,978,1005]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="monocellatus">
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<emphasis id="B94FEAFAFC33FF9576BAEDF7FDCA95A5" bold="true" box="[151,634,978,1005]" pageId="22" pageNumber="201">
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<emphasis id="B94FEAFAFC33FF9576BAEDF7FE7D95A4" bold="true" box="[151,461,978,1005]" italics="true" pageId="22" pageNumber="201">Neobythites monocellatus</emphasis>
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<bibRefCitation id="EFAA4B19FC33FF9577F8EDF7FDCA95A5" author="Nielsen, J. G." box="[469,634,978,1004]" pageId="22" pageNumber="201" pagination="335 - 372" refId="ref21162" refString="Nielsen, J. G. (1999) A review of the genus Neobythites (Pisces, Ophidiidae) in the Atlantic, with three new species. Bulletin of Marine Science, 64 (2), 335 - 372." type="journal article" year="1999">Nielsen, 1999</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3216563FC33FF9576BAEDD2FECD9258" box="[151,381,1014,1041]" pageId="22" pageNumber="201" type="description">
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<paragraph id="8B8436E8FC33FF9576BAEDD2FECD9258" blockId="22.[151,1427,978,1074]" box="[151,381,1014,1041]" pageId="22" pageNumber="201">
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<figureCitation id="13002A6DFC33FF9576BAEDD2FF479258" box="[151,247,1015,1041]" captionStart="FIGURE 6" captionStartId="18.[151,250,1829,1853]" captionTargetBox="[152,1436,382,1805]" captionTargetId="figure-50@18.[151,1436,381,1805]" captionTargetPageId="18" captionText="FIGURE 6. Four species of the steatiticus group. (A) Neobythites pako n. sp., HT, MNHN 2006-0337, SL 165 mm, Solomon Sea (side-reversed; M. Krag), (B) N. malayanus ZMUC P77742, 138 mm SL, off Lombok (listed in Uiblein & Nielsen 2018; M. Krag); (C) N. stefanovi, ZMUC P77841, PT, 198 mm SL, Gulf of Aden (listed in Uiblein & Nielsen 2018; M. Krag); (D) N. monocellatus, MOVI 39139, 97 mm SL, off E Brazil (listed in Nielsen et al. 2009; G. Brovad); with 10 mm scale bars." figureDoi="http://doi.org/10.5281/zenodo.8272439" httpUri="https://zenodo.org/record/8272439/files/figure.png" pageId="22" pageNumber="201">Figure 6</figureCitation>
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;
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<tableCitation id="C6B90353FC33FF957729EDD2FED19258" box="[260,353,1015,1041]" captionStart="TABLE 1" captionStartId="2.[151,239,476,500]" captionText="TABLE 1. Ocellus-bearing Neobythites species, with number and category of ocelli, preopercular spine number, and" pageId="22" pageNumber="201">Tables 1</tableCitation>
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,
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<tableCitation id="C6B90353FC33FF957742EDD3FECD9258" box="[367,381,1014,1041]" captionStart="TABLE 5" captionStartId="20.[151,239,151,175]" captionText="TABLE 5. Meristic, morphometric, quantitative colour, and otolith characters in Neobythites pako n. sp. and the eight" pageId="22" pageNumber="201">5</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3216563FC33FF9576BAEA32FA239266" box="[151,1427,1047,1074]" pageId="22" pageNumber="201" type="reference_group">
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<paragraph id="8B8436E8FC33FF9576BAEA32FA239266" blockId="22.[151,1427,978,1074]" box="[151,1427,1047,1074]" pageId="22" pageNumber="201">
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<treatmentCitation id="0A9A10F9FC33FF9576BAEA32FDD59266" author="Nielsen, J. G." box="[151,613,1047,1071]" page="351" pageId="22" pageNumber="201" year="1999">
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<taxonomicName id="4C3B4D6BFC33FF9576BAEA32FDD59266" ID-CoL="46C3W" authority="Nielsen, 1999: 351" authorityName="Nielsen" authorityPageNumber="351" authorityYear="1999" box="[151,613,1047,1071]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="monocellatus">
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<emphasis id="B94FEAFAFC33FF9576BAEA32FE2C9266" box="[151,412,1047,1071]" italics="true" pageId="22" pageNumber="201">Neobythites monocellatus</emphasis>
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<materialsCitation id="3B533CB5FC33FF95778EEA32FDD59266" box="[419,613,1047,1071]" collectionCode="USNM" country="Venezuela" elevation="229" latitude="9.883333" location="Venezuela" longLatPrecision="1298" longitude="-59.883335" pageId="22" pageNumber="201" specimenCode="USNM 309234" specimenCount="1" typeStatus="holotype">
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<bibRefCitation id="EFAA4B19FC33FF95778EEA32FDD59266" author="Nielsen, J. G." box="[419,613,1047,1071]" pageId="22" pageNumber="201" pagination="335 - 372" refId="ref21162" refString="Nielsen, J. G. (1999) A review of the genus Neobythites (Pisces, Ophidiidae) in the Atlantic, with three new species. Bulletin of Marine Science, 64 (2), 335 - 372." type="journal article" year="1999">Nielsen, 1999: 351</bibRefCitation>
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, figs. 8–9 (off
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<collectingCountry id="F32C7678FC33FF9574D7EA32FCD59266" box="[762,869,1047,1071]" name="Venezuela" pageId="22" pageNumber="201">Venezuela</collectingCountry>
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,
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<geoCoordinate id="EE0F502FFC33FF957543EA3CFC75927B" box="[878,965,1049,1074]" degrees="09" direction="north" minutes="53" orientation="latitude" pageId="22" pageNumber="201" precision="925" value="9.883333">09°53′N</geoCoordinate>
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,
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<geoCoordinate id="EE0F502FFC33FF9575FDEA3CFB9A927B" box="[976,1066,1049,1074]" degrees="59" direction="west" minutes="53" orientation="longitude" pageId="22" pageNumber="201" precision="925" value="-59.883335">59°53′W</geoCoordinate>
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,
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<quantity id="4CC39B0DFC33FF957219EA32FBC89266" box="[1076,1144,1047,1071]" metricMagnitude="2" metricUnit="m" metricValue="2.29" pageId="22" pageNumber="201" unit="m" value="229.0">
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<elevation id="0016D1DBFC33FF957219EA32FBC89266" box="[1076,1144,1047,1071]" metricMagnitude="2" metricUnit="m" metricValue="2.29" pageId="22" pageNumber="201" unit="m" value="229.0">229 m</elevation>
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</quantity>
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;
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<typeStatus id="5480884AFC33FF9572AFEA32FB6F9266" box="[1154,1247,1047,1071]" pageId="22" pageNumber="201">holotype</typeStatus>
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:
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<specimenCode id="DB9D9E93FC33FF9572C5EA32FA389266" box="[1256,1416,1047,1071]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="201" type="Museum">USNM 309234</specimenCode>
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).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3216563FC33FF9576BAEA78FE88933A" pageId="22" pageNumber="201" type="diagnosis">
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<paragraph id="8B8436E8FC33FF9576BAEA78FE88933A" blockId="22.[151,1437,1117,2008]" pageId="22" pageNumber="201">
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<emphasis id="B94FEAFAFC33FF9576BAEA78FEA2923E" bold="true" box="[151,274,1117,1143]" pageId="22" pageNumber="201">Diagnosis.</emphasis>
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Spine on hind margin of preopercle absent or thin and flat; dorsal-fin rays 93–99; anal-fin rays 78–83; pectoral-fin rays 24–27; precaudal vertebrae 12–13; total vertebrae 54–58; pseudobranchial filaments 4–6; long rakers on anterior gill arch 13–15; head length 21–24% SL; pelvic-fin length 17–23% SL, fins not extending beyond anus; orbit length 4.9–7.0% SL and 22–30% HL; longest gill filament 1.4–2.0% SL and 6.2–9.0% HL; dorsal fin with central ocellus with black spot placed well behind vertical line through anus, the ocellus spot distance being 41–51% SL and the spot covering 6–7 dorsal-fin rays, not extending ventrally to body; no vertical bars on body; otolith length 5.0–5.8% SL, sulcus length 4.0–4.7% SL, and ostium height 18–22% sulcus length and 28–34% ostium length.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3216563FC33FF9476EAEB58FC039708" lastPageId="23" lastPageNumber="202" pageId="22" pageNumber="201" type="distribution">
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<paragraph id="8B8436E8FC33FF9576EAEB58FE2693A9" blockId="22.[151,1437,1117,2008]" pageId="22" pageNumber="201">
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<emphasis id="B94FEAFAFC33FF9576EAEB58FE7893DE" bold="true" box="[199,456,1405,1431]" pageId="22" pageNumber="201">Distribution and size.</emphasis>
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Found in the tropical W and SW Atlantic from off
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<collectingCountry id="F32C7678FC33FF957221EB58FBC993DE" box="[1036,1145,1405,1431]" name="Honduras" pageId="22" pageNumber="201">Honduras</collectingCountry>
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and along the north coast of South America to
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<collectingCountry id="F32C7678FC33FF9577ACEB84FD9793F2" box="[385,551,1441,1467]" name="French Guiana" pageId="22" pageNumber="201">French Guiana</collectingCountry>
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and off
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<collectingRegion id="49FFF80AFC33FF9574A9EB84FD7893F2" box="[644,712,1441,1467]" country="Brazil" name="Bahia" pageId="22" pageNumber="201">Bahia</collectingRegion>
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, E
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<collectingCountry id="F32C7678FC33FF9574C1EB84FC8393F2" box="[748,819,1441,1467]" name="Brazil" pageId="22" pageNumber="201">Brazil</collectingCountry>
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||
, on the continental shelf and upper slope (
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<quantity id="4CC39B0DFC33FF957321EB84FA2093F2" box="[1292,1424,1441,1467]" metricMagnitude="2" metricUnit="m" metricValue="2.78" metricValueMax="4.39" metricValueMin="1.17" pageId="22" pageNumber="201" unit="m" value="278.0" valueMax="439.0" valueMin="117.0">117–439 m</quantity>
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||
). Max. size
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||
<quantity id="4CC39B0DFC33FF957726EBE0FED893A9" box="[267,360,1477,1504]" metricMagnitude="-1" metricUnit="m" metricValue="1.54" pageId="22" pageNumber="201" unit="mm" value="154.0">154 mm</quantity>
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SL.
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</paragraph>
|
||
<paragraph id="8B8436E8FC33FF9576EAEBCCFA389125" blockId="22.[151,1437,1117,2008]" pageId="22" pageNumber="201">
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<emphasis id="B94FEAFAFC33FF9576EAEBCCFEDD904A" bold="true" box="[199,365,1513,1539]" pageId="22" pageNumber="201">Comparisons.</emphasis>
|
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<taxonomicName id="4C3B4D6BFC33FF957758EBCCFD87904A" authority="Uiblein & Nielsen, 2023" authorityName="Uiblein & Nielsen" authorityYear="2023" box="[373,567,1513,1539]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="pako" status="n. sp.">
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<emphasis id="B94FEAFAFC33FF957758EBCCFD87904A" box="[373,567,1513,1539]" italics="true" pageId="22" pageNumber="201">Neobythites pako</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B94FEAFAFC33FF95746DEBCCFD30904A" bold="true" box="[576,640,1513,1539]" pageId="22" pageNumber="201">
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||
<taxonomicNameLabel id="A27C5781FC33FF95746DEBCCFD30904A" box="[576,640,1513,1539]" pageId="22" pageNumber="201" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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differs clearly from all other
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<taxonomicName id="4C3B4D6BFC33FF9575FCEBCFFBF3904A" authorityName="Alcock" authorityYear="1894" box="[977,1091,1514,1539]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="steatiticus">
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<emphasis id="B94FEAFAFC33FF9575FCEBCFFBF3904A" box="[977,1091,1514,1539]" italics="true" pageId="22" pageNumber="201">steatiticus</emphasis>
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</taxonomicName>
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species in having a very short but distinct single preopercular spine vs. none or only a very weakly formed, flat, or indistinct spine, and in the combination of several colour, meristic, morphometric, and otolith characters (
|
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<tableCitation id="C6B90353FC33FF95720CE814FBCB9005" box="[1057,1147,1585,1612]" captionStart="TABLE 5" captionStartId="20.[151,239,151,175]" captionText="TABLE 5. Meristic, morphometric, quantitative colour, and otolith characters in Neobythites pako n. sp. and the eight" pageId="22" pageNumber="201">Table 5</tableCitation>
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||
). It is most similar to
|
||
<taxonomicName id="4C3B4D6BFC33FF9573AEE817FEA09026" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="malayanus">
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||
<emphasis id="B94FEAFAFC33FF9573AEE817FEA09026" italics="true" pageId="22" pageNumber="201">N. malayanus</emphasis>
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||
</taxonomicName>
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||
and
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||
<taxonomicName id="4C3B4D6BFC33FF957769E873FE789026" box="[324,456,1621,1647]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="stefanovi">
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<emphasis id="B94FEAFAFC33FF957769E873FE789026" box="[324,456,1621,1647]" italics="true" pageId="22" pageNumber="201">N. stefanovi</emphasis>
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||
</taxonomicName>
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||
. From
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||
<taxonomicName id="4C3B4D6BFC33FF95743AE873FD1F9026" box="[535,687,1621,1647]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="malayanus">
|
||
<emphasis id="B94FEAFAFC33FF95743AE873FD1F9026" box="[535,687,1621,1647]" italics="true" pageId="22" pageNumber="201">N. malayanus</emphasis>
|
||
</taxonomicName>
|
||
, it differs in longer upper jaw relative to head length in combination with slightly deeper body, shorter pelvic fins and postorbital distance (
|
||
<figureCitation id="13002A6DFC33FF9575F9E85CFBAD90DA" box="[980,1053,1657,1683]" captionStart="FIGURE 7" captionStartId="19.[151,250,1331,1355]" captionTargetBox="[171,1419,189,1305]" captionTargetId="figure-23@19.[163,1424,181,1307]" captionTargetPageId="19" captionText="FIGURE 7. Plots of four morphometric characters and SL for visual comparisons between N. pako n. sp. and N. malayanus (the latter separated into 4 populations)." figureDoi="http://doi.org/10.5281/zenodo.8272443" httpUri="https://zenodo.org/record/8272443/files/figure.png" pageId="22" pageNumber="201">Fig. 7</figureCitation>
|
||
), and it differs in several otolith characters including shorter and shallower otolith, and shallower ostium relative to SL, sulcus, and ostium length (
|
||
<tableCitation id="C6B90353FC33FF9576B2E8E4FF459095" box="[159,245,1729,1756]" captionStart="TABLE 5" captionStartId="20.[151,239,151,175]" captionText="TABLE 5. Meristic, morphometric, quantitative colour, and otolith characters in Neobythites pako n. sp. and the eight" pageId="22" pageNumber="201">Table 5</tableCitation>
|
||
;
|
||
<figureCitation id="13002A6DFC33FF95772CE8E4FEC49092" box="[257,372,1729,1755]" captionStart="FIGURE 3" captionStartId="7.[151,250,1861,1885]" captionTargetBox="[151,1436,633,1838]" captionTargetId="figure-205@7.[151,1436,633,1838]" captionTargetPageId="7" captionText="FIGURE 3. Neobythites otoliths. (A) N.superocellatus n. sp., HT, MNHN 2021-0457, Solomon Sea (M. Krag). (B) N. japonicus n. sp., HT, KAUM -I. 37071, off Okinawa (from left side of head, shown side-reversed; M. Krag); (C) N. jonathan n. sp., HT, MNHN 2002-3883, Solomon Sea (C. Høegh-Guldberg); (D) N. zora n. sp., HT, ZMUC P771324, Fiji (C. Høegh-Guldberg); (E) N. pako n. sp., HT, MNHN 2006-0337, otolith number MNHN-ICOT-001969, Solomon Sea (M. Krag); (F) N. malayanus, ZMUC P77742, Lombok (C. Høegh-Guldberg); all otoliths from right side of head except for (B); with 1 mm scale bars." figureDoi="http://doi.org/10.5281/zenodo.8272427" httpUri="https://zenodo.org/record/8272427/files/figure.png" pageId="22" pageNumber="201">Fig. 3E, F</figureCitation>
|
||
). It differs from the Indonesian population (
|
||
<typeStatus id="5480884AFC33FF957576E8E4FC3D9092" box="[859,909,1729,1755]" pageId="22" pageNumber="201">type</typeStatus>
|
||
locality) of
|
||
<taxonomicName id="4C3B4D6BFC33FF957238E8E7FB1E9092" box="[1045,1198,1729,1755]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="malayanus">
|
||
<emphasis id="B94FEAFAFC33FF957238E8E7FB1E9092" box="[1045,1198,1729,1755]" italics="true" pageId="22" pageNumber="201">N. malayanus</emphasis>
|
||
</taxonomicName>
|
||
in having more anal-fin rays, fewer pseudobranchial filaments, and shorter preanal distance, from the
|
||
<collectingCountry id="F32C7678FC33FF95721DE8C3FB3C9149" box="[1072,1164,1766,1792]" name="Vanuatu" pageId="22" pageNumber="201">Vanuatu</collectingCountry>
|
||
population in having in longer gill-filaments and from the Philippine population (which is most similar) in having fewer pseudobranchial filaments (
|
||
<tableCitation id="C6B90353FC33FF957720E908FED3910E" box="[269,355,1837,1863]" captionStart="TABLE 6" captionStartId="21.[151,239,151,175]" captionText="TABLE 6. Meristic, morphometric, quantitative colour, and otolith characters in Neobythites pako n. sp. and N. malayanus." pageId="22" pageNumber="201">Table 6</tableCitation>
|
||
). It differs from
|
||
<taxonomicName id="4C3B4D6BFC33FF957436E90BFD10910E" box="[539,672,1837,1863]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="stefanovi">
|
||
<emphasis id="B94FEAFAFC33FF957436E90BFD10910E" box="[539,672,1837,1863]" italics="true" pageId="22" pageNumber="201">N. stefanovi</emphasis>
|
||
</taxonomicName>
|
||
in its absence of dark dorsal-fin margin, higher number of precaudal vertebrae, vertebrae number below dorsal-fin origin, smaller ocellus spot, and shallower otolith ostium (
|
||
<tableCitation id="C6B90353FC33FF957309E974FACA9125" box="[1316,1402,1873,1900]" captionStart="TABLE 5" captionStartId="20.[151,239,151,175]" captionText="TABLE 5. Meristic, morphometric, quantitative colour, and otolith characters in Neobythites pako n. sp. and the eight" pageId="22" pageNumber="201">Table 5</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8436E8FC33FF9476EAE953FC039708" blockId="22.[151,1437,1117,2008]" lastBlockId="23.[151,1437,151,429]" lastPageId="23" lastPageNumber="202" pageId="22" pageNumber="201">
|
||
For further comparisons among all other
|
||
<taxonomicName id="4C3B4D6BFC33FF9574BFE953FCB491C6" authorityName="Alcock" authorityYear="1894" box="[658,772,1910,1935]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="steatiticus">
|
||
<emphasis id="B94FEAFAFC33FF9574BFE953FCB491C6" box="[658,772,1910,1935]" italics="true" pageId="22" pageNumber="201">steatiticus</emphasis>
|
||
</taxonomicName>
|
||
group members except for the W Atlantic
|
||
<taxonomicName id="4C3B4D6BFC33FF9572F2E953FA2591C6" authorityName="Nielsen" authorityYear="1999" box="[1247,1429,1909,1935]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="monocellatus">
|
||
<emphasis id="B94FEAFAFC33FF9572F2E953FA2591C6" box="[1247,1429,1909,1935]" italics="true" pageId="22" pageNumber="201">N. monocellatus</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="EFAA4B19FC33FF9576E9E9BCFE5D91FA" author="Uiblein F. & Nielsen J. G." box="[196,493,1945,1971]" pageId="22" pageNumber="201" pagination="157 - 173" refId="ref21961" refString="Uiblein F. & Nielsen J. G. (2018) Review of the steatiticus species group of the cuskeel genus Neobythites (Ophidiidae) from the Indo-Pacific, with description of two new species. Zootaxa, 4387 (1), 157 - 173. https: // doi. org / 10.11646 / zootaxa. 4387.1.7" type="journal article" year="2018">Uiblein & Nielsen (2018)</bibRefCitation>
|
||
.
|
||
<taxonomicName id="4C3B4D6BFC33FF9577D7E9BCFCAC91FA" authorityName="Nielsen" authorityYear="1999" box="[506,796,1945,1971]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="monocellatus">
|
||
<emphasis id="B94FEAFAFC33FF9577D7E9BCFCAC91FA" box="[506,796,1945,1971]" italics="true" pageId="22" pageNumber="201">Neobythites monocellatus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3B4D6BFC33FF95759AE9BFFBA091FA" authority="Uiblein & Nielsen, 2023" authorityName="Uiblein & Nielsen" authorityYear="2023" box="[951,1040,1945,1971]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="201" phylum="Chordata" rank="species" species="pako" status="n. sp.">
|
||
<emphasis id="B94FEAFAFC33FF95759AE9BFFBA091FA" box="[951,1040,1945,1971]" italics="true" pageId="22" pageNumber="201">N. pako</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94FEAFAFC33FF957237E9BCFBEB91FA" bold="true" box="[1050,1115,1945,1971]" pageId="22" pageNumber="201">
|
||
<taxonomicNameLabel id="A27C5781FC33FF957237E9BCFBEB91FA" box="[1050,1115,1945,1971]" pageId="22" pageNumber="201" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in lack of dark stripe along anal fin, more dorsal-, anal- and pectoral fin rays, more pseudobranchial filaments, more developed gill rakers, and longer pelvic fins;
|
||
<taxonomicName id="4C3B4D6BFC32FF94775DEEB2FD9896F8" authorityName="Nielsen" authorityYear="1999" box="[368,552,151,177]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="monocellatus">
|
||
<emphasis id="B94FEAFAFC32FF94775DEEB2FD9896F8" box="[368,552,151,177]" italics="true" pageId="23" pageNumber="202">N. monocellatus</emphasis>
|
||
</taxonomicName>
|
||
is most similar to six
|
||
<emphasis id="B94FEAFAFC32FF947501EEB2FC3E96F9" box="[812,910,151,176]" italics="true" pageId="23" pageNumber="202">steaticus</emphasis>
|
||
group species,
|
||
<taxonomicName id="4C3B4D6BFC32FF94726FEEB2FB0596F8" authorityName="Uiblein & Nielsen" authorityYear="2018" box="[1090,1205,151,177]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="gloriae">
|
||
<emphasis id="B94FEAFAFC32FF94726FEEB2FB0596F8" box="[1090,1205,151,177]" italics="true" pageId="23" pageNumber="202">N. gloriae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D6BFC32FF9472E8EEB2FAC296F8" authorityName="Uiblein & Nielsen" authorityYear="2018" box="[1221,1394,151,177]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="lombokensis">
|
||
<emphasis id="B94FEAFAFC32FF9472E8EEB2FAC296F8" box="[1221,1394,151,177]" italics="true" pageId="23" pageNumber="202">N. lombokensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D6BFC32FF9473AEEEB2FEA0969C" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="malayanus">
|
||
<emphasis id="B94FEAFAFC32FF9473AEEEB2FEA0969C" italics="true" pageId="23" pageNumber="202">N. malayanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D6BFC32FF947732EE9EFE27969D" authorityName="Nielsen" authorityYear="1995" box="[287,407,187,212]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="meteori">
|
||
<emphasis id="B94FEAFAFC32FF947732EE9EFE27969D" box="[287,407,187,212]" italics="true" pageId="23" pageNumber="202">N. meteori</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D6BFC32FF947788EE9EFD89969D" authorityName="Alcock" authorityYear="1894" box="[421,569,187,212]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="steatiticus">
|
||
<emphasis id="B94FEAFAFC32FF947788EE9EFD89969D" box="[421,569,187,212]" italics="true" pageId="23" pageNumber="202">N. steatiticus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3B4D6BFC32FF94745CEE9EFD47969C" box="[625,759,187,213]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="stefanovi">
|
||
<emphasis id="B94FEAFAFC32FF94745CEE9EFD47969C" box="[625,759,187,213]" italics="true" pageId="23" pageNumber="202">N. stefanovi</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs in having a combination of relatively high number of dorsal- and anal-fin rays and gill rakers, relatively longer pelvic fins, shorter postorbital distance, and smaller-sized ocellus spot; from the remaining species,
|
||
<taxonomicName id="4C3B4D6BFC32FF947500EF26FC7B9754" authorityName="Nielsen" authorityYear="1995" box="[813,971,259,285]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="malhaensis">
|
||
<emphasis id="B94FEAFAFC32FF947500EF26FC7B9754" box="[813,971,259,285]" italics="true" pageId="23" pageNumber="202">N. malhaensis</emphasis>
|
||
</taxonomicName>
|
||
, it differs in fewer pectoral-fin rays, more pseudobranchial filaments, longer pelvic fins, and smaller ocellus spot.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3216563FC32FF9476EAEF6EFD5F97E4" pageId="23" pageNumber="202" type="discussion">
|
||
<paragraph id="8B8436E8FC32FF9476EAEF6EFD5F97E4" blockId="23.[151,1437,151,429]" pageId="23" pageNumber="202">
|
||
<emphasis id="B94FEAFAFC32FF9476EAEF6EFE8B972C" bold="true" box="[199,315,331,357]" pageId="23" pageNumber="202">Remarks.</emphasis>
|
||
One
|
||
<taxonomicName id="4C3B4D6BFC32FF947750EF6EFD35972C" box="[381,645,331,357]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="malayanus">
|
||
<emphasis id="B94FEAFAFC32FF947750EF6EFD35972C" box="[381,645,331,357]" italics="true" pageId="23" pageNumber="202">Neobythites malayanus</emphasis>
|
||
</taxonomicName>
|
||
specimen (length 171+ mm) from off Lombok could not be studied in detail due to missing its tail (
|
||
<tableCitation id="C6B90353FC32FF947424EF4AFDD397C0" box="[521,611,367,393]" captionStart="TABLE 6" captionStartId="21.[151,239,151,175]" captionText="TABLE 6. Meristic, morphometric, quantitative colour, and otolith characters in Neobythites pako n. sp. and N. malayanus." pageId="23" pageNumber="202">Table 6</tableCitation>
|
||
). Like
|
||
<taxonomicName id="4C3B4D6BFC32FF947495EF4AFCC297C0" authorityName="Smith & Radcliffe" authorityYear="1913" box="[696,882,367,393]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="unimaculatus">
|
||
<emphasis id="B94FEAFAFC32FF947495EF4AFCC297C0" box="[696,882,367,393]" italics="true" pageId="23" pageNumber="202">N. unimaculatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D6BFC32FF9475AFEF4AFBAF97C0" box="[898,1055,367,393]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="malayanus">
|
||
<emphasis id="B94FEAFAFC32FF9475AFEF4AFBAF97C0" box="[898,1055,367,393]" italics="true" pageId="23" pageNumber="202">N. malayanus</emphasis>
|
||
</taxonomicName>
|
||
should be studied in more detail regarding geographic differentiation (
|
||
<tableCitation id="C6B90353FC32FF947416EFB6FD2297E4" box="[571,658,403,429]" captionStart="TABLE 6" captionStartId="21.[151,239,151,175]" captionText="TABLE 6. Meristic, morphometric, quantitative colour, and otolith characters in Neobythites pako n. sp. and N. malayanus." pageId="23" pageNumber="202">Table 6</tableCitation>
|
||
,
|
||
<figureCitation id="13002A6DFC32FF9474B3EFB6FD5197E4" box="[670,737,403,429]" captionStart="FIGURE 7" captionStartId="19.[151,250,1331,1355]" captionTargetBox="[171,1419,189,1305]" captionTargetId="figure-23@19.[163,1424,181,1307]" captionTargetPageId="19" captionText="FIGURE 7. Plots of four morphometric characters and SL for visual comparisons between N. pako n. sp. and N. malayanus (the latter separated into 4 populations)." figureDoi="http://doi.org/10.5281/zenodo.8272443" httpUri="https://zenodo.org/record/8272443/files/figure.png" pageId="23" pageNumber="202">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |